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<document id="AA28091239835C24246723A30BD010EC" ID-CLB-Dataset="55748" ID-DOI="10.5281/zenodo.5301289" ID-GBIF-Dataset="55b2b0bc-d79f-48e3-8be1-81424185dea7" ID-ISSN="0253-116X" ID-Zenodo-Dep="5301289" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630101010072" checkinUser="marcus" docAuthor="Assing, V." docDate="2013" docId="1974879EFFFDFFD0FF04E9D2FC39FC3A" docLanguage="en" docName="LinzerbiolBeitr.2013.45.2.1479-1520.pdf" docOrigin="Linzer biologische Beiträge 45 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Scymbalium nepalense Assing 2013, nov.sp." docType="treatment" docVersion="10" lastPageNumber="1488" masterDocId="E54DFFE6FFF5FFD9FFB6EA15FFE8FFA6" masterDocTitle="On the Palaearctic and Oriental species of Scymbalium and Micrillus (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="1520" masterPageNumber="1479" pageNumber="1487" updateTime="1699152599027" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A69571C56BC6172A568A342A4489FE0E">On the Palaearctic and Oriental species of Scymbalium and Micrillus (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<mods:namePart id="8196A3CD0EF98426CE2345E9E0888F27">Assing, V.</mods:namePart>
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<mods:title id="E38B52D849A54E5F0F1CABCC8ECA62C9">Linzer biologische Beiträge</mods:title>
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<mods:date id="45C544E4066EDD2E9DDAB6B0CBE97B5E">2013</mods:date>
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<mods:number id="2D093529DEF4A1819F768E767BCD7E05">2013-12-20</mods:number>
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<treatment id="1974879EFFFDFFD0FF04E9D2FC39FC3A" ID-DOI="http://doi.org/10.5281/zenodo.6512313" ID-GBIF-Taxon="194955306" ID-Zenodo-Dep="6512313" LSID="urn:lsid:plazi:treatment:1974879EFFFDFFD0FF04E9D2FC39FC3A" httpUri="http://treatment.plazi.org/id/1974879EFFFDFFD0FF04E9D2FC39FC3A" lastPageId="9" lastPageNumber="1488" pageId="8" pageNumber="1487">
<subSubSection id="D9C76503FFFDFFD1FF04E9D2FE1CFC7B" box="[178,500,967,989]" pageId="8" pageNumber="1487" type="nomenclature">
<paragraph id="91623688FFFDFFD1FF04E9D2FE1CFC7B" blockId="8.[178,679,967,989]" box="[178,500,967,989]" pageId="8" pageNumber="1487">
<heading id="CA2A81E4FFFDFFD1FF04E9D2FE1CFC7B" box="[178,500,967,989]" fontSize="8" level="2" pageId="8" pageNumber="1487" reason="2">
<emphasis id="A3A9EA9AFFFDFFD1FF04E9D2FE1CFC7B" bold="true" box="[178,500,967,989]" pageId="8" pageNumber="1487">
<taxonomicName id="56DD4D0BFFFDFFD1FF04E9D2FE4AFC7B" ID-CoL="9QRRW" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[178,418,967,989]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1487" phylum="Arthropoda" rank="species" species="nepalense" status="sp. nov.">
<emphasis id="A3A9EA9AFFFDFFD1FF04E9D2FE4AFC7B" bold="true" box="[178,418,967,989]" italics="true" pageId="8" pageNumber="1487">Scymbalium nepalense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89A57E1FFFDFFD1FE1EE9D2FE1CFC7B" box="[424,500,967,989]" pageId="8" pageNumber="1487" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<paragraph id="91623688FFFDFFD1FE4CE9D2FD4FFC7B" blockId="8.[178,679,967,989]" box="[506,679,967,989]" pageId="8" pageNumber="1487">
<heading id="CA2A81E4FFFDFFD1FE4CE9D2FD4FFC7B" box="[506,679,967,989]" fontSize="8" level="2" pageId="8" pageNumber="1487" reason="2">
(
<figureCitation id="09E62A0DFFFDFFD1FDB7E9D2FDB0FC7B" box="[513,600,967,989]" captionStart="Figs 2-8" captionStartId="7.[178,218,1503,1523]" captionTargetBox="[182,1080,219,1485]" captionTargetId="figure-59@7.[178,1085,219,1487]" captionTargetPageId="7" captionText="Figs 2-8: Scymbalium persimile CAMERON, holotype (2-4) and S. nepalense nov.sp., paratype (5- 8): (2, 5) habitus; (3, 6) forebody; (4) abdomen; (7) aedeagus in lateral view; (8) aedeagus in ventral view. Scale bars: 2-6: 1.0 mm; 7-8: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301293" httpUri="https://zenodo.org/record/5301293/files/figure.png" pageId="8" pageNumber="1487">Figs 5-8</figureCitation>
,
<figureCitation id="09E62A0DFFFDFFD1FDD5E9D2FD77FC7B" box="[611,671,967,989]" captionStart-0="Figs 56-62" captionStart-1="Figs 63-66" captionStartId-0="32.[178,218,1520,1540]" captionStartId-1="33.[178,218,834,854]" captionTargetBox-0="[190,1085,222,1489]" captionTargetBox-1="[191,1085,214,809]" captionTargetId-0="figure-68@32.[178,1085,212,1497]" captionTargetId-1="figure-296@33.[178,1085,212,809]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionText-0="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." captionText-1="Figs 63-66: Scymbalium nepalense nov.sp., holotype: (63) habitus; (64) male sternite VIII; (65-66) aedeagus in lateral and in ventral view. Scale bars: 63: 1.0 mm; 64: 0.5 mm; 65-66: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5301311" figureDoi-1="http://doi.org/10.5281/zenodo.5301317" httpUri-0="https://zenodo.org/record/5301311/files/figure.png" httpUri-1="https://zenodo.org/record/5301317/files/figure.png" pageId="8" pageNumber="1487">61-66</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFFDFFD1FF04E9FBFEA1FBE0" pageId="8" pageNumber="1487" type="materials_examined">
<paragraph id="91623688FFFDFFD1FF04E9FBFEA1FBE0" blockId="8.[178,1086,1006,1094]" pageId="8" pageNumber="1487">
Type material
<materialsCitation id="21B53CD5FFFDFFD1FE06E9FBFCA1FB89" ID-GBIF-Occurrence="3760604397" collectingDate="2009-07-02" collectionCode="NME" collectorName="M. Hartmann" country="Nepal" determinerName="V. Assing" elevation="210" latitude="28.970278" location="Mahendranagar" longitude="80.18361" municipality="Kanchanpur" pageId="8" pageNumber="1487" specimenCount="1" stateProvince="Mahakali" typeStatus="holotype">
<typeStatus id="4E66882AFFFDFFD1FE06E9FBFE14FBA4" box="[432,508,1006,1026]" pageId="8" pageNumber="1487" type="holotype">Holotype</typeStatus>
: &quot;
<collectingCountry id="E9CA7618FFFDFFD1FD8AE9FBFD9CFBA4" box="[572,628,1006,1026]" name="Nepal" pageId="8" pageNumber="1487">Nepal</collectingCountry>
P:
<collectingRegion id="5319F86AFFFDFFD1FD29E9FBFD1FFBA4" box="[671,759,1006,1026]" country="Nepal" name="Mahakali" pageId="8" pageNumber="1487">Mahakali</collectingRegion>
, D:
<collectingMunicipality id="7106ACF2FFFDFFD1FC9EE9FBFC71FBA4" box="[808,921,1006,1026]" pageId="8" pageNumber="1487">Kanchanpur</collectingMunicipality>
,
<location id="94026053FFFDFFD1FC1FE9FBFBD2FBA4" LSID="urn:lsid:plazi:treatment:1974879EFFFDFFD0FF04E9D2FC39FC3A:94026053FFFDFFD1FC1FE9FBFBD2FBA4" box="[937,1082,1006,1026]" country="Nepal" municipality="Kanchanpur" name="Mahendranagar" pageId="8" pageNumber="1487" stateProvince="Mahakali">Mahendranagar</location>
,
<location id="94026053FFFDFFD1FF04EE11FE81FBBE" LSID="urn:lsid:plazi:treatment:1974879EFFFDFFD0FF04E9D2FC39FC3A:94026053FFFDFFD1FF04EE11FE81FBBE" box="[178,361,1028,1048]" country="Nepal" municipality="Kanchanpur" name="Hotel Sweet Dream" pageId="8" pageNumber="1487" stateProvince="Mahakali">Hotel Sweet Dream</location>
/
<quantity id="56259B6DFFFDFFD1FEC9EE11FE55FBBE" box="[383,445,1028,1048]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="1487" unit="m" value="210.0">
<elevation id="1AF0D1BBFFFDFFD1FEC9EE11FE55FBBE" box="[383,445,1028,1048]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="1487" unit="m" value="210.0">210 m</elevation>
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,
<geoCoordinate id="F4E9504FFFFDFFD1FE7EEE11FDDCFBBE" box="[456,564,1028,1048]" degrees="28" direction="north" minutes="58" orientation="latitude" pageId="8" pageNumber="1487" precision="15" seconds="13" value="28.970278">N28°58'13''</geoCoordinate>
,
<geoCoordinate id="F4E9504FFFFDFFD1FD89EE11FD40FBBE" box="[575,680,1028,1048]" degrees="80" direction="east" minutes="11" orientation="longitude" pageId="8" pageNumber="1487" precision="15" seconds="01" value="80.18361">E80°11'01''</geoCoordinate>
,
<date id="E5631048FFFDFFD1FD02EE11FCCDFBBE" box="[692,805,1028,1048]" pageId="8" pageNumber="1487" value="2009-07-02">
<collectingDate id="F527E9A0FFFDFFD1FD02EE11FCCDFBBE" box="[692,805,1028,1048]" pageId="8" pageNumber="1487" value="2009-07-02">02.VII.2009</collectingDate>
</date>
, leg.
<collectorName id="3C28535EFFFDFFD1FCEFEE11FC30FBBE" box="[857,984,1028,1048]" pageId="8" pageNumber="1487">M. Hartmann</collectorName>
, LF 60 / Holotypus
<taxonomicName id="56DD4D0BFFFDFFD1FE86EE0EFDE9FB89" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[304,513,1051,1071]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1487" phylum="Arthropoda" rank="species" species="nepalense" status="sp. nov.">
<emphasis id="A3A9EA9AFFFDFFD1FE86EE0EFDE9FB89" box="[304,513,1051,1071]" italics="true" pageId="8" pageNumber="1487">Scymbalium nepalense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89A57E1FFFDFFD1FDB1EE0EFDDCFB89" box="[519,564,1051,1071]" pageId="8" pageNumber="1487" rank="species">sp.n.</taxonomicNameLabel>
, det.
<determinerName id="64BEDA4CFFFDFFD1FDD0EE0EFD29FB89" box="[614,705,1051,1071]" pageId="8" pageNumber="1487">V. Assing</determinerName>
2013&quot; (
<collectionCode id="F7CCAE4DFFFDFFD1FCBDEE0EFCABFB89" box="[779,835,1051,1071]" pageId="8" pageNumber="1487">NME</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="21B53CD5FFFDFFD1FCE5EE0EFEAEFBE0" ID-GBIF-Occurrence="3760604365" pageId="8" pageNumber="1487" typeStatus="paratype">
<typeStatus id="4E66882AFFFDFFD1FCE5EE0EFC4BFB89" box="[851,931,1051,1071]" pageId="8" pageNumber="1487" type="paratype">Paratype</typeStatus>
: same data as holotype (cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFFDFFD1FF04EE41FC5BFBCC" box="[178,947,1108,1130]" pageId="8" pageNumber="1487" type="etymology">
<paragraph id="91623688FFFDFFD1FF04EE41FC5BFBCC" blockId="8.[178,947,1108,1130]" box="[178,947,1108,1130]" pageId="8" pageNumber="1487">
Etymology: The specific epithet is an adjective derived from
<collectingCountry id="E9CA7618FFFDFFD1FCD9EE41FC47FBCC" box="[879,943,1108,1130]" name="Nepal" pageId="8" pageNumber="1487">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFFDFFD0FF04EE6CFEE1FE0E" lastPageId="9" lastPageNumber="1488" pageId="8" pageNumber="1487" type="description">
<paragraph id="91623688FFFDFFD1FF04EE6CFD5CFB6C" blockId="8.[178,1085,1145,1226]" pageId="8" pageNumber="1487">
Description: Size rather variable; ody length
<quantity id="56259B6DFFFDFFD1FD59EE6CFC83FB29" box="[751,875,1145,1167]" metricMagnitude="-3" metricUnit="m" metricValue="8.15" metricValueMax="8.5" metricValueMin="7.8" pageId="8" pageNumber="1487" unit="mm" value="8.15" valueMax="8.5" valueMin="7.8">
7.8
<emphasis id="A3A9EA9AFFFDFFD1FCB9EE6CFCFFFB29" bold="true" box="[783,791,1145,1167]" pageId="8" pageNumber="1487">-</emphasis>
8.5 mm
</quantity>
; length of forebody
<quantity id="56259B6DFFFDFFD1FF04EE82FED9FB0A" box="[178,305,1175,1197]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.7" metricValueMin="3.9" pageId="8" pageNumber="1487" unit="mm" value="4.3" valueMax="4.7" valueMin="3.9">
3.9
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4.7 mm
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. Habitus as in
<figureCitation id="09E62A0DFFFDFFD1FE69EE82FDCDFB0B" box="[479,549,1175,1197]" captionStart="Figs 2-8" captionStartId="7.[178,218,1503,1523]" captionTargetBox="[182,1080,219,1485]" captionTargetId="figure-59@7.[178,1085,219,1487]" captionTargetPageId="7" captionText="Figs 2-8: Scymbalium persimile CAMERON, holotype (2-4) and S. nepalense nov.sp., paratype (5- 8): (2, 5) habitus; (3, 6) forebody; (4) abdomen; (7) aedeagus in lateral view; (8) aedeagus in ventral view. Scale bars: 2-6: 1.0 mm; 7-8: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301293" httpUri="https://zenodo.org/record/5301293/files/figure.png" pageId="8" pageNumber="1487">Figs 5</figureCitation>
,
<figureCitation id="09E62A0DFFFDFFD1FD81EE82FDBBFB0B" box="[567,595,1175,1197]" captionStart="Figs 63-66" captionStartId="33.[178,218,834,854]" captionTargetBox="[191,1085,214,809]" captionTargetId="figure-296@33.[178,1085,212,809]" captionTargetPageId="33" captionText="Figs 63-66: Scymbalium nepalense nov.sp., holotype: (63) habitus; (64) male sternite VIII; (65-66) aedeagus in lateral and in ventral view. Scale bars: 63: 1.0 mm; 64: 0.5 mm; 65-66: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301317" httpUri="https://zenodo.org/record/5301317/files/figure.png" pageId="8" pageNumber="1487">63</figureCitation>
. Coloration: head reddish to blackish-brown; remainder of body, including appendages, reddish.
</paragraph>
<paragraph id="91623688FFFDFFD1FF04EECCFC4AFAC2" blockId="8.[178,1086,1241,1380]" pageId="8" pageNumber="1487">
Head (
<figureCitation id="09E62A0DFFFDFFD1FF40EECCFED0FB49" box="[246,312,1241,1263]" captionStart="Figs 2-8" captionStartId="7.[178,218,1503,1523]" captionTargetBox="[182,1080,219,1485]" captionTargetId="figure-59@7.[178,1085,219,1487]" captionTargetPageId="7" captionText="Figs 2-8: Scymbalium persimile CAMERON, holotype (2-4) and S. nepalense nov.sp., paratype (5- 8): (2, 5) habitus; (3, 6) forebody; (4) abdomen; (7) aedeagus in lateral view; (8) aedeagus in ventral view. Scale bars: 2-6: 1.0 mm; 7-8: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301293" httpUri="https://zenodo.org/record/5301293/files/figure.png" pageId="8" pageNumber="1487">Figs 6</figureCitation>
,
<figureCitation id="09E62A0DFFFDFFD1FEF3EECCFEB7FB49" box="[325,351,1241,1263]" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="8" pageNumber="1487">61</figureCitation>
) approximately as long as broad; lateral margins behind eyes subparallel; posterior angles marked; disc with sparse, barely noticeable micropunctation, anteriorly, laterally, and posteriorly with scattered coarser punctures; integument with very shallow and very fine transverse microsculpture visible only at higher magnification. Antenna
<quantity id="56259B6DFFFDFFD1FEB8EF5BFE6BFAC2" box="[270,387,1358,1380]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.3" metricValueMin="2.9" pageId="8" pageNumber="1487" unit="mm" value="3.0999999999999996" valueMax="3.3" valueMin="2.9">
2.9
<emphasis id="A3A9EA9AFFFDFFD1FE98EF5BFEDEFAC2" bold="true" box="[302,310,1358,1380]" pageId="8" pageNumber="1487">-</emphasis>
3.3 mm
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long and slender, all antennomeres distinctly oblong.
</paragraph>
<paragraph id="91623688FFFDFFD1FF04EF61FDF6FA47" blockId="8.[178,1086,1396,1505]" pageId="8" pageNumber="1487">
Pronotum (
<figureCitation id="09E62A0DFFFDFFD1FE93EF61FE8BFA2C" box="[293,355,1396,1418]" captionStart="Figs 2-8" captionStartId="7.[178,218,1503,1523]" captionTargetBox="[182,1080,219,1485]" captionTargetId="figure-59@7.[178,1085,219,1487]" captionTargetPageId="7" captionText="Figs 2-8: Scymbalium persimile CAMERON, holotype (2-4) and S. nepalense nov.sp., paratype (5- 8): (2, 5) habitus; (3, 6) forebody; (4) abdomen; (7) aedeagus in lateral view; (8) aedeagus in ventral view. Scale bars: 2-6: 1.0 mm; 7-8: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301293" httpUri="https://zenodo.org/record/5301293/files/figure.png" pageId="8" pageNumber="1487">Fig. 6</figureCitation>
) of rather variable shape, 1.08
<emphasis id="A3A9EA9AFFFDFFD1FD16EF61FD40FA2C" bold="true" box="[672,680,1396,1418]" pageId="8" pageNumber="1487">-</emphasis>
1.14 times as long and 0.9
<emphasis id="A3A9EA9AFFFDFFD1FC0FEF61FC29FA2C" bold="true" box="[953,961,1396,1418]" pageId="8" pageNumber="1487">-</emphasis>
1.0 times as broad and as broad as head, widest at anterior angles; lateral margins straight, weakly converging in dorsal view; posterior margin distinctly concave; punctation dense and very fine; microsculpture indistinct.
</paragraph>
<paragraph id="91623688FFFDFFD1FF04EFE4FBD9F9E7" blockId="8.[178,1085,1521,1601]" pageId="8" pageNumber="1487">
Elytra (
<figureCitation id="09E62A0DFFFDFFD1FF48EFE4FED3F9A1" box="[254,315,1521,1543]" captionStart="Figs 2-8" captionStartId="7.[178,218,1503,1523]" captionTargetBox="[182,1080,219,1485]" captionTargetId="figure-59@7.[178,1085,219,1487]" captionTargetPageId="7" captionText="Figs 2-8: Scymbalium persimile CAMERON, holotype (2-4) and S. nepalense nov.sp., paratype (5- 8): (2, 5) habitus; (3, 6) forebody; (4) abdomen; (7) aedeagus in lateral view; (8) aedeagus in ventral view. Scale bars: 2-6: 1.0 mm; 7-8: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301293" httpUri="https://zenodo.org/record/5301293/files/figure.png" pageId="8" pageNumber="1487">Fig. 6</figureCitation>
) approximately 1.05 times as long as pronotum; disc matt due to extremely dense punctation and microsculpture. Hind wings fully developed. Protarsomeres I-IV moderately dilated. Metatarsomere I approximately as long as combined length of II-IV.
</paragraph>
<paragraph id="91623688FFFCFFD0FF04EACFFEACFE8D" blockId="9.[178,1086,218,299]" pageId="9" pageNumber="1488">Abdomen narrower than elytra; punctation very dense and very fine; interstices with microsculpture; posterior margin of tergite VII with palisade fringe; tergite X and sternite IX very small.</paragraph>
<paragraph id="91623688FFFCFFD0FF71EB2EFEE1FE0E" blockId="9.[178,1085,314,424]" pageId="9" pageNumber="1488">
: posterior excision of sternite VIII weakly and broadly concave (
<figureCitation id="09E62A0DFFFCFFD0FC1DEB2FFBEAFEF6" box="[939,1026,314,336]" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="9" pageNumber="1488">Figs 62</figureCitation>
,
<figureCitation id="09E62A0DFFFCFFD0FBA2EB2FFBD9FEF6" box="[1044,1073,314,336]" captionStart="Figs 63-66" captionStartId="33.[178,218,834,854]" captionTargetBox="[191,1085,214,809]" captionTargetId="figure-296@33.[178,1085,212,809]" captionTargetPageId="33" captionText="Figs 63-66: Scymbalium nepalense nov.sp., holotype: (63) habitus; (64) male sternite VIII; (65-66) aedeagus in lateral and in ventral view. Scale bars: 63: 1.0 mm; 64: 0.5 mm; 65-66: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301317" httpUri="https://zenodo.org/record/5301317/files/figure.png" pageId="9" pageNumber="1488">64</figureCitation>
); aedeagus (
<figureCitation id="09E62A0DFFFCFFD0FE94EB4DFE97FEC8" box="[290,383,344,366]" captionStart="Figs 2-8" captionStartId="7.[178,218,1503,1523]" captionTargetBox="[182,1080,219,1485]" captionTargetId="figure-59@7.[178,1085,219,1487]" captionTargetPageId="7" captionText="Figs 2-8: Scymbalium persimile CAMERON, holotype (2-4) and S. nepalense nov.sp., paratype (5- 8): (2, 5) habitus; (3, 6) forebody; (4) abdomen; (7) aedeagus in lateral view; (8) aedeagus in ventral view. Scale bars: 2-6: 1.0 mm; 7-8: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301293" httpUri="https://zenodo.org/record/5301293/files/figure.png" pageId="9" pageNumber="1488">Figs 7-8</figureCitation>
,
<figureCitation id="09E62A0DFFFCFFD0FE26EB4DFE24FEC8" box="[400,460,344,366]" captionStart="Figs 63-66" captionStartId="33.[178,218,834,854]" captionTargetBox="[191,1085,214,809]" captionTargetId="figure-296@33.[178,1085,212,809]" captionTargetPageId="33" captionText="Figs 63-66: Scymbalium nepalense nov.sp., holotype: (63) habitus; (64) male sternite VIII; (65-66) aedeagus in lateral and in ventral view. Scale bars: 63: 1.0 mm; 64: 0.5 mm; 65-66: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301317" httpUri="https://zenodo.org/record/5301317/files/figure.png" pageId="9" pageNumber="1488">
65
<emphasis id="A3A9EA9AFFFCFFD0FE1FEB4DFE59FEC8" bold="true" box="[425,433,344,366]" pageId="9" pageNumber="1488">-</emphasis>
66
</figureCitation>
) of variable size,
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0.6
<emphasis id="A3A9EA9AFFFCFFD0FD0FEB4DFD29FEC8" bold="true" box="[697,705,344,366]" pageId="9" pageNumber="1488">-</emphasis>
0.7 mm
</quantity>
long, symmetric, and with weakly sclerotized, membranous internal structures; parameres nearly completely reduced.
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFFCFFD0FF04EBA2FC1BFCB9" pageId="9" pageNumber="1488" type="discussion">
<paragraph id="91623688FFFCFFD0FF04EBA2FC94FE4C" blockId="9.[178,1085,439,491]" pageId="9" pageNumber="1488">Comment: The short, in lateral view transverse plate above the ventral opening of the aedeagus is interpreted as a rudiment of fused ancestral parameres.</paragraph>
<paragraph id="91623688FFFCFFD0FF04EBEFFE7FFD04" blockId="9.[178,1086,506,674]" pageId="9" pageNumber="1488">
Intraspecific variation: The
<specimenCount id="87DBFD01FFFCFFD0FD39EBEEFD01FDB6" box="[655,745,507,528]" pageId="9" pageNumber="1488" type="generic" typeStatus="type">two type</specimenCount>
specimens differ considerably in body size, the size of the head (in relation to the pronotum), the shape of the pronotum, and the size of the aedeagus. However, the shapes of the aedeagus and of the male sternite VIII are practically identical. Also, both specimens were collected in the same sample, suggesting that the observed differences should be attributable to intra- rather than interspecific variation.
</paragraph>
<paragraph id="91623688FFFCFFD0FF04E8A7FC1BFCB9" blockId="9.[178,1086,690,799]" pageId="9" pageNumber="1488">
Comparative notes: From
<taxonomicName id="56DD4D0BFFFCFFD0FDE5E8A6FD55FD6E" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" box="[595,701,690,712]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1488" phylum="Arthropoda" rank="species" species="badium">
<emphasis id="A3A9EA9AFFFCFFD0FDE5E8A6FD55FD6E" box="[595,701,690,712]" italics="true" pageId="9" pageNumber="1488">L. badium</emphasis>
</taxonomicName>
, the only other geographically close described
<taxonomicName id="56DD4D0BFFFCFFD0FEADE8DAFE03FD43" box="[283,491,719,741]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1488" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="A3A9EA9AFFFCFFD0FEADE8DAFE7FFD43" box="[283,407,719,741]" italics="true" pageId="9" pageNumber="1488">Scymbalium</emphasis>
species
</taxonomicName>
, the new species is distinguished by the finer and sparser punctation of the head, the shallower posterior concavity of the male sternite VIII, and by the differently shaped apex of the ventral process of the aedagus (lateral view).
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFFCFFD0FF04E93AFC39FC3A" pageId="9" pageNumber="1488" type="distribution">
<paragraph id="91623688FFFCFFD0FF04E93AFC39FC3A" blockId="9.[178,1085,815,924]" pageId="9" pageNumber="1488">
Distribution and natural history: Thetypelocalityissituatedin
<collectingRegion id="5319F86AFFFCFFD0FF04E959FE99FCC4" box="[178,369,844,866]" country="Nepal" name="Mahakali" pageId="9" pageNumber="1488">Mahakali province</collectingRegion>
in the extreme southwest of
<collectingCountry id="E9CA7618FFFCFFD0FD2DE959FD32FCC4" box="[667,730,844,866]" name="Nepal" pageId="9" pageNumber="1488">Nepal</collectingCountry>
, close to the borders with Uttaranchal and Uttar Pradesh. The
<typeStatus id="4E66882AFFFCFFD0FE63E97CFDC6FCD9" box="[469,558,873,895]" pageId="9" pageNumber="1488" type="holotype">holotype</typeStatus>
was collected with a light trap at an altitude of
<quantity id="56259B6DFFFCFFD0FBA1E97CFF20FC3A" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="1488" unit="m" value="210.0">210 m</quantity>
, suggesting that
<taxonomicName id="56DD4D0BFFFCFFD0FEC7E992FE07FC3A" authorityName="Assing" authorityYear="2013" box="[369,495,902,924]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1488" phylum="Arthropoda" rank="species" species="nepalense">
<emphasis id="A3A9EA9AFFFCFFD0FEC7E992FE07FC3A" box="[369,495,902,924]" italics="true" pageId="9" pageNumber="1488">S. nepalense</emphasis>
</taxonomicName>
is widespread at least in the Himalayan region.
</paragraph>
</subSubSection>
</treatment>
</document>
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