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treatments-xml/data/19/74/87/1974879EFFECFFC5FF04E9AFFE30FD9C.xml
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<document id="89E252A163B2C35CC68187DF74E6AC4D" ID-CLB-Dataset="55748" ID-DOI="10.5281/zenodo.5301289" ID-GBIF-Dataset="55b2b0bc-d79f-48e3-8be1-81424185dea7" ID-ISSN="0253-116X" ID-Zenodo-Dep="5301289" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630101010072" checkinUser="marcus" docAuthor="Assing, V." docDate="2013" docId="1974879EFFECFFC5FF04E9AFFE30FD9C" docLanguage="en" docName="LinzerbiolBeitr.2013.45.2.1479-1520.pdf" docOrigin="Linzer biologische Beiträge 45 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Micrillus distortus Assing 2013, nov.sp." docType="treatment" docVersion="9" lastPageNumber="1507" masterDocId="E54DFFE6FFF5FFD9FFB6EA15FFE8FFA6" masterDocTitle="On the Palaearctic and Oriental species of Scymbalium and Micrillus (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="1520" masterPageNumber="1479" pageNumber="1504" updateTime="1699152599027" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="F730E4916F44DFE1F18E165ACEF234C8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3D53F856765D7D42492D273754030D15">On the Palaearctic and Oriental species of Scymbalium and Micrillus (Coleoptera: Staphylinidae: Paederinae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="F0F164841D15DA477D1715AF8F1C71DE">Assing, V.</mods:namePart>
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<mods:title id="531AD3D5E511DCF59112CAB9E6550E58">Linzer biologische Beiträge</mods:title>
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<mods:part id="B65F0700326511DA5BC908EC4775F1EF">
<mods:date id="F06A8BD2C578D4E5DFA12A36C0EAE569">2013</mods:date>
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<mods:number id="82E781793F32FD8C081E16B9B9C3C7B5">2013-12-20</mods:number>
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<mods:number id="DF8CE4734EE739DB52661019FDB346A0">45</mods:number>
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<mods:identifier id="D73C42BEA2A4009F3C6C6D7C221DDCD5" type="CLB-Dataset">55748</mods:identifier>
<mods:identifier id="6CC6D878608681B46222F55064A30D9C" type="DOI">10.5281/zenodo.5301289</mods:identifier>
<mods:identifier id="569DB8DDA247D09C8948B260387399F0" type="GBIF-Dataset">55b2b0bc-d79f-48e3-8be1-81424185dea7</mods:identifier>
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<treatment id="1974879EFFECFFC5FF04E9AFFE30FD9C" ID-DOI="http://doi.org/10.5281/zenodo.6512338" ID-GBIF-Taxon="194955321" ID-Zenodo-Dep="6512338" LSID="urn:lsid:plazi:treatment:1974879EFFECFFC5FF04E9AFFE30FD9C" httpUri="http://treatment.plazi.org/id/1974879EFFECFFC5FF04E9AFFE30FD9C" lastPageId="28" lastPageNumber="1507" pageId="25" pageNumber="1504">
<subSubSection id="D9C76503FFECFFC0FF04E9AFFE2CFC76" box="[178,452,954,976]" pageId="25" pageNumber="1504" type="nomenclature">
<paragraph id="91623688FFECFFC0FF04E9AFFE2CFC76" blockId="25.[178,663,954,976]" box="[178,452,954,976]" pageId="25" pageNumber="1504">
<heading id="CA2A81E4FFECFFC0FF04E9AFFE2CFC76" box="[178,452,954,976]" fontSize="8" level="2" pageId="25" pageNumber="1504" reason="2">
<emphasis id="A3A9EA9AFFECFFC0FF04E9AFFE2CFC76" bold="true" box="[178,452,954,976]" pageId="25" pageNumber="1504">
<taxonomicName id="56DD4D0BFFECFFC0FF04E9AFFE9AFC76" ID-CoL="9PLYY" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[178,370,954,976]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1504" phylum="Arthropoda" rank="species" species="distortus" status="sp. nov.">
<emphasis id="A3A9EA9AFFECFFC0FF04E9AFFE9AFC76" bold="true" box="[178,370,954,976]" italics="true" pageId="25" pageNumber="1504">Micrillus distortus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89A57E1FFECFFC0FECEE9AFFE2CFC76" box="[376,452,954,976]" pageId="25" pageNumber="1504" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFECFFC0FE7CE9AFFD7FFC76" box="[458,663,954,976]" pageId="25" pageNumber="1504" type="description">
<paragraph id="91623688FFECFFC0FE7CE9AFFD7FFC76" blockId="25.[178,663,954,976]" box="[458,663,954,976]" pageId="25" pageNumber="1504">
<heading id="CA2A81E4FFECFFC0FE7CE9AFFD7FFC76" box="[458,663,954,976]" fontSize="8" level="2" pageId="25" pageNumber="1504" reason="2">
(
<figureCitation id="09E62A0DFFECFFC0FE67E9AFFDAAFC76" box="[465,578,954,976]" captionStart="Figs 36-43" captionStartId="26.[178,218,1520,1540]" captionTargetBox="[186,1078,212,1484]" captionTargetId="figure-65@26.[178,1085,212,1497]" captionTargetPageId="26" captionText="Figs 36-43: Micrillus distortus nov.sp.: (36) habitus; (37) forebody; (38) male sternite VII; (39) male sternite VIII; (40-41) aedeagus in lateral and in ventral view; (42) ventral process of aedeagus in ventral view; (43) internal structure of aedeagus in dorsal view. Scale bars: 36-37: 1.0 mm; 38- 43: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301303" httpUri="https://zenodo.org/record/5301303/files/figure.png" pageId="25" pageNumber="1504">Figs 36-43</figureCitation>
,
<figureCitation id="09E62A0DFFECFFC0FDFAE9AEFD67FC76" box="[588,655,954,976]" captionStart="Map 4" captionStartId="27.[178,223,1091,1111]" captionTargetBox="[178,1085,212,1066]" captionTargetId="figure-174@27.[178,1085,212,1066]" captionTargetPageId="27" captionText="Map 4: Distributions of Micrillus distortus nov.sp. (triangles) and M. suturalis (CAMERON) (circles) in the Oriental region, based on examined records." figureDoi="http://doi.org/10.5281/zenodo.5301305" httpUri="https://zenodo.org/record/5301305/files/figure.png" pageId="25" pageNumber="1504">Map 4</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFECFFC0FF04E9F5FE68FB35" pageId="25" pageNumber="1504" type="materials_examined">
<paragraph id="91623688FFECFFC0FF04E9F5FE68FB35" blockId="25.[178,1085,992,1171]" pageId="25" pageNumber="1504">
Type material examined
<materialsCitation id="21B53CD5FFECFFC0FDDCE9F5FDECFB9E" ID-GBIF-Occurrence="3760604311" collectedFrom="Micrillus distortus sp. n." collectingDate="2008-06-17" collectingDateMax="2008-06-18" collectingDateMin="2008-06-17" collectionCode="ZMUC" collectorName="A. Solodovnikov &amp; J. Pedersen" country="Laos" determinerName="V. Assing" elevation="0" latitude="18.217117" location="Nahin" longLatPrecision="1" longitude="104.51466" municipality="Ban Khounkham" pageId="25" pageNumber="1504" specimenCount="1" stateProvince="Khammouane prov." typeStatus="holotype">
<typeStatus id="4E66882AFFECFFC0FDDCE9F5FD56FC52" box="[618,702,992,1012]" pageId="25" pageNumber="1504" type="holotype">Holotype</typeStatus>
: &quot;
<collectingCountry id="E9CA7618FFECFFC0FCB6E9F5FCD9FC52" box="[768,817,992,1012]" name="Laos" pageId="25" pageNumber="1504">Laos</collectingCountry>
:
<collectingRegion id="5319F86AFFECFFC0FCF2E9F5FBEEFC52" box="[836,1030,992,1012]" country="Laos" name="Khammouan" pageId="25" pageNumber="1504">Khammouane prov.</collectingRegion>
,
<collectingMunicipality id="7106ACF2FFECFFC0FBAEE9F5FECAFBAD" pageId="25" pageNumber="1504">Ban Khounkham</collectingMunicipality>
(
<location id="94026053FFECFFC0FE84E9E2FE86FBAD" LSID="urn:lsid:plazi:treatment:1974879EFFECFFC5FF04E9AFFE30FD9C:94026053FFECFFC0FE84E9E2FE86FBAD" box="[306,366,1015,1035]" country="Laos" latitude="18.217117" longLatPrecision="1" longitude="104.51466" municipality="Ban Khounkham" name="Nahin" pageId="25" pageNumber="1504" stateProvince="Khammouane prov.">Nahin</location>
),
<date id="E5631048FFECFFC0FE37E9E2FDEAFBAD" box="[385,514,1015,1035]" pageId="25" pageNumber="1504" value="2008-06-17" valueMax="2008-06-18" valueMin="2008-06-17">
<collectingDate id="F527E9A0FFECFFC0FE37E9E2FDEAFBAD" box="[385,514,1015,1035]" pageId="25" pageNumber="1504" value="2008-06-17" valueMax="2008-06-18" valueMin="2008-06-17">
17-18.vi.
<quantity id="56259B6DFFECFFC0FE78E9E2FDEAFBAD" box="[462,514,1015,1035]" metricMagnitude="6" metricUnit="m" metricValue="2.0082999999999998" pageId="25" pageNumber="1504" unit="m" value="2008300.0">2008</quantity>
</collectingDate>
</date>
, 300 m,
<geoCoordinate id="F4E9504FFFECFFC0FDEDE9E2FD26FBAD" box="[603,718,1015,1035]" degrees="18" direction="north" minutes="13.027" orientation="latitude" pageId="25" pageNumber="1504" precision="1" value="18.217117">18°13.027'N</geoCoordinate>
<geoCoordinate id="F4E9504FFFECFFC0FD61E9E2FCBAFBAD" box="[727,850,1015,1035]" degrees="104" direction="east" minutes="30.880" orientation="longitude" pageId="25" pageNumber="1504" precision="1" value="104.51466">104°30.880'E</geoCoordinate>
/
<collectorName id="3C28535EFFECFFC0FCDDE9E2FBE2FBAD" box="[875,1034,1015,1035]" pageId="25" pageNumber="1504">A. Solodovnikov</collectorName>
&amp;
<collectorName id="3C28535EFFECFFC0FB98E9E2FEECFB84" pageId="25" pageNumber="1504">J. Pedersen</collectorName>
leg. Disturbed primary rain forest; ZMUC collection / Holotypus
<collectedFrom id="22C17575FFECFFC0FC24EE1BFF37FB9E" pageId="25" pageNumber="1504">
<taxonomicName id="56DD4D0BFFECFFC0FC24EE1BFBD5FB84" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[914,1085,1038,1058]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1504" phylum="Arthropoda" rank="species" species="distortus" status="sp. nov.">
<emphasis id="A3A9EA9AFFECFFC0FC24EE1BFBD5FB84" box="[914,1085,1038,1058]" italics="true" pageId="25" pageNumber="1504">Micrillus distortus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89A57E1FFECFFC0FF04EE31FF37FB9E" box="[178,223,1060,1080]" pageId="25" pageNumber="1504" rank="species">sp.n.</taxonomicNameLabel>
</collectedFrom>
, det.
<determinerName id="64BEDA4CFFECFFC0FEA7EE31FE85FB9E" box="[273,365,1060,1080]" pageId="25" pageNumber="1504">V. Assing</determinerName>
2012&quot; (
<collectionCode id="F7CCAE4DFFECFFC0FE01EE31FE16FB9E" box="[439,510,1060,1080]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="25" pageNumber="1504">ZMUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="21B53CD5FFECFFC0FDB8EE31FBD2FB9E" ID-GBIF-Occurrence="3760604321" box="[526,1082,1060,1080]" collectingDate="2008-06-17" collectingDateMax="2008-06-18" collectingDateMin="2008-06-17" collectionCode="ZMUC" collectorName="A. Solodovnikov &amp; J. Pedersen" country="Laos" determinerName="V. Assing" latitude="18.217117" location="Nahin" longLatPrecision="1" longitude="104.51466" municipality="Ban Khounkham" pageId="25" pageNumber="1504" specimenCount="1" stateProvince="Khammouan" typeStatus="paratype">
<typeStatus id="4E66882AFFECFFC0FDB8EE31FD82FB9E" box="[526,618,1060,1080]" pageId="25" pageNumber="1504" type="paratype">Paratypes</typeStatus>
: 1, 5: same data as holotype (
<collectionCode id="F7CCAE4DFFECFFC0FC02EE31FC11FB9E" box="[948,1017,1060,1080]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="25" pageNumber="1504">ZMUC</collectionCode>
, cAss)
</materialsCitation>
;
<materialsCitation id="21B53CD5FFECFFC0FF04EE2EFE49FBC0" ID-GBIF-Occurrence="3760604405" collectingDate="2001-04-24" collectingDateMax="2001-04-29" collectingDateMin="2001-04-24" collectionCode="NHMB" collectorName="Pacholatko" country="Laos" elevation="200" latitude="18.116667" location="Ban Khoun Ngeun" longLatPrecision="1276" longitude="104.48333" municipality="Kham Mouan" pageId="25" pageNumber="1504" specimenCount="1" typeStatus="paratype">
<specimenCount id="87DBFD01FFECFFC0FF04EE2EFF55FBE9" box="[178,189,1083,1103]" pageId="25" pageNumber="1504">1</specimenCount>
: &quot;
<collectingCountry id="E9CA7618FFECFFC0FF53EE2EFEFBFBE9" box="[229,275,1083,1103]" name="Laos" pageId="25" pageNumber="1504">Laos</collectingCountry>
-C,
<collectingMunicipality id="7106ACF2FFECFFC0FE88EE2EFE55FBE9" box="[318,445,1083,1103]" pageId="25" pageNumber="1504">Kham Mouan</collectingMunicipality>
pr.,
<location id="94026053FFECFFC0FE5CEE2EFD72FBE9" LSID="urn:lsid:plazi:treatment:1974879EFFECFFC5FF04E9AFFE30FD9C:94026053FFECFFC0FE5CEE2EFD72FBE9" box="[490,666,1083,1103]" country="Laos" latitude="18.116667" longLatPrecision="1276" longitude="104.48333" municipality="Kham Mouan" name="Ban Khoun Ngeun" pageId="25" pageNumber="1504">Ban Khoun Ngeun</location>
, ç
<quantity id="56259B6DFFECFFC0FD18EE2EFD04FBE9" box="[686,748,1083,1103]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="1504" unit="m" value="200.0">
<elevation id="1AF0D1BBFFECFFC0FD18EE2EFD04FBE9" box="[686,748,1083,1103]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="1504" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<geoCoordinate id="F4E9504FFFECFFC0FD40EE2EFCA9FBE9" box="[758,833,1083,1103]" degrees="18" direction="north" minutes="07" orientation="latitude" pageId="25" pageNumber="1504" precision="925" value="18.116667">18°07'N</geoCoordinate>
<geoCoordinate id="F4E9504FFFECFFC0FCFEEE2EFC76FBE9" box="[840,926,1083,1103]" degrees="104" direction="east" minutes="29" orientation="longitude" pageId="25" pageNumber="1504" precision="925" value="104.48333">104°29'E</geoCoordinate>
,
<collectorName id="3C28535EFFECFFC0FC1FEE2EFBE6FBE9" box="[937,1038,1083,1103]" pageId="25" pageNumber="1504">Pacholátko</collectorName>
leg.,
<date id="E5631048FFECFFC0FF04EE47FED1FBC0" box="[178,313,1106,1126]" pageId="25" pageNumber="1504" value="2001-04-24" valueMax="2001-04-29" valueMin="2001-04-24">
<collectingDate id="F527E9A0FFECFFC0FF04EE47FED1FBC0" box="[178,313,1106,1126]" pageId="25" pageNumber="1504" value="2001-04-24" valueMax="2001-04-29" valueMin="2001-04-24">24.-29.iv.2001</collectingDate>
</date>
&quot; (
<collectionCode id="F7CCAE4DFFECFFC0FEE4EE47FE73FBC0" box="[338,411,1106,1126]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="25" pageNumber="1504">NHMB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="21B53CD5FFECFFC0FE06EE47FEE0FBDB" ID-GBIF-Occurrence="3760604325" collectionCode="BMNH" collectorName="M. Cameron. Bequest." country="Laos" location="Indo-China" municipality="Saigon" pageId="25" pageNumber="1504" specimenCount="1" typeStatus="paratype">
1: &quot;
<collectingMunicipality id="7106ACF2FFECFFC0FE51EE47FDC2FBC0" box="[487,554,1106,1126]" pageId="25" pageNumber="1504">Saigon</collectingMunicipality>
,
<location id="94026053FFECFFC0FD8EEE47FD48FBC0" LSID="urn:lsid:plazi:treatment:1974879EFFECFFC5FF04E9AFFE30FD9C:94026053FFECFFC0FD8EEE47FD48FBC0" box="[568,672,1106,1126]" country="Laos" municipality="Saigon" name="Indo-China" pageId="25" pageNumber="1504">Indo-China</location>
/
<collectorName id="3C28535EFFECFFC0FD0DEE47FC7CFBC0" box="[699,916,1106,1126]" pageId="25" pageNumber="1504">M. Cameron. Bequest.</collectorName>
B.M. 1955-147.&quot; (
<collectionCode id="F7CCAE4DFFECFFC0FF0EEE7CFEE9FBDB" box="[184,257,1129,1149]" pageId="25" pageNumber="1504">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="21B53CD5FFECFFC0FEA1EE7CFE94FB35" ID-GBIF-Occurrence="3760604343" collectingDate="1993-04-29" collectionCode="NHMW" country="Laos" pageId="25" pageNumber="1504" specimenCount="1" typeStatus="paratype">
1: &quot;T-Thailand, Chiang.M., NW Chiang Mai, Pai City,
<date id="E5631048FFECFFC0FCF9EE7CFC44FBDB" box="[847,940,1129,1149]" pageId="25" pageNumber="1504" value="1993-04-29">
<collectingDate id="F527E9A0FFECFFC0FCF9EE7CFC44FBDB" box="[847,940,1129,1149]" pageId="25" pageNumber="1504" value="1993-04-29">29.4.1993</collectingDate>
</date>
, Pacholatko &amp; Dembicky&quot; (
<collectionCode id="F7CCAE4DFFECFFC0FE90EE6AFE9EFB35" box="[294,374,1151,1171]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="1504" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFECFFC0FF04EEB7FE31FB73" pageId="25" pageNumber="1504" type="etymology">
<paragraph id="91623688FFECFFC0FF04EEB7FE31FB73" blockId="25.[178,1085,1186,1237]" pageId="25" pageNumber="1504">Etymology: The specific epithet (Latin, adjective: distorted) refers to the distinctly asymmetric aedeagus.</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFECFFC2FF04EEF1FCBDF980" lastPageId="27" lastPageNumber="1506" pageId="25" pageNumber="1504" type="description">
<paragraph id="91623688FFECFFC0FF04EEF1FD0DFA93" blockId="25.[178,1085,1252,1333]" pageId="25" pageNumber="1504">
Description: Small species; body length
<quantity id="56259B6DFFECFFC0FD0CEEF1FCDDFB5C" box="[698,821,1252,1274]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.7" metricValueMin="3.8" pageId="25" pageNumber="1504" unit="mm" value="4.25" valueMax="4.7" valueMin="3.8">3.8-4.7 mm</quantity>
; length of forebody: 2.0-
<quantity id="56259B6DFFECFFC0FF04EF14FEEAFAB1" box="[178,258,1281,1303]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="25" pageNumber="1504" unit="mm" value="2.3">2.3 mm</quantity>
. Habitus as in
<figureCitation id="09E62A0DFFECFFC0FE2EEF14FE0BFAB1" box="[408,483,1281,1303]" captionStart="Figs 36-43" captionStartId="26.[178,218,1520,1540]" captionTargetBox="[186,1078,212,1484]" captionTargetId="figure-65@26.[178,1085,212,1497]" captionTargetPageId="26" captionText="Figs 36-43: Micrillus distortus nov.sp.: (36) habitus; (37) forebody; (38) male sternite VII; (39) male sternite VIII; (40-41) aedeagus in lateral and in ventral view; (42) ventral process of aedeagus in ventral view; (43) internal structure of aedeagus in dorsal view. Scale bars: 36-37: 1.0 mm; 38- 43: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301303" httpUri="https://zenodo.org/record/5301303/files/figure.png" pageId="25" pageNumber="1504">Fig. 36</figureCitation>
. Coloration: head reddish-brown to dark-brown; remainder of body reddish; legs reddish-yellow; antennae reddish.
</paragraph>
<paragraph id="91623688FFECFFC0FF04EF51FC70FA69" blockId="25.[178,1085,1348,1487]" pageId="25" pageNumber="1504">
Head (
<figureCitation id="09E62A0DFFECFFC0FF4EEF51FEADFAFC" box="[248,325,1348,1370]" captionStart="Figs 36-43" captionStartId="26.[178,218,1520,1540]" captionTargetBox="[186,1078,212,1484]" captionTargetId="figure-65@26.[178,1085,212,1497]" captionTargetPageId="26" captionText="Figs 36-43: Micrillus distortus nov.sp.: (36) habitus; (37) forebody; (38) male sternite VII; (39) male sternite VIII; (40-41) aedeagus in lateral and in ventral view; (42) ventral process of aedeagus in ventral view; (43) internal structure of aedeagus in dorsal view. Scale bars: 36-37: 1.0 mm; 38- 43: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301303" httpUri="https://zenodo.org/record/5301303/files/figure.png" pageId="25" pageNumber="1504">Fig. 37</figureCitation>
) approximately as long as broad; punctation composed of sparse micropunctation and few interspersed macropunctures; interstices with very shallow microsculpture and glossy. Eyes 0.6-0.8 times as long as postocular region and weakly projecting from lateral contours of head in dorsal view. Apical joint of maxillary palpus of conical shape and distinctly less than half the length of preapical joint.
</paragraph>
<caption id="C5A26600FFEFFFC3FF04EFE5FEC9F9EE" ID-DOI="http://doi.org/10.5281/zenodo.5301303" ID-Zenodo-Dep="5301303" httpUri="https://zenodo.org/record/5301303/files/figure.png" pageId="26" pageNumber="1505" startId="26.[178,218,1520,1540]" targetBox="[186,1078,212,1484]" targetPageId="26">
<paragraph id="91623688FFEFFFC3FF04EFE5FEC9F9EE" blockId="26.[178,1086,1520,1608]" pageId="26" pageNumber="1505">
<emphasis id="A3A9EA9AFFEFFFC3FF04EFE5FEF0F9A2" bold="true" box="[178,280,1520,1540]" pageId="26" pageNumber="1505">Figs 36-43</emphasis>
:
<taxonomicName id="56DD4D0BFFEFFFC3FE91EFE5FE3AF9A2" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[295,466,1520,1540]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="1505" phylum="Arthropoda" rank="species" species="distortus" status="sp. nov.">
<emphasis id="A3A9EA9AFFEFFFC3FE91EFE5FE3AF9A2" box="[295,466,1520,1540]" italics="true" pageId="26" pageNumber="1505">Micrillus distortus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89A57E1FFEFFFC3FE6CEFE5FDC8F9A2" box="[474,544,1520,1540]" pageId="26" pageNumber="1505" rank="species">nov.sp.</taxonomicNameLabel>
: (
<emphasis id="A3A9EA9AFFEFFFC3FD85EFE5FDA2F9A2" bold="true" box="[563,586,1520,1540]" pageId="26" pageNumber="1505">36</emphasis>
) habitus; (
<emphasis id="A3A9EA9AFFEFFFC3FD04EFE5FD38F9A2" bold="true" box="[690,720,1520,1540]" pageId="26" pageNumber="1505">37)</emphasis>
forebody; (
<emphasis id="A3A9EA9AFFEFFFC3FCF7EFE5FCB0F9A2" bold="true" box="[833,856,1520,1540]" pageId="26" pageNumber="1505">38</emphasis>
) male sternite VII; (
<emphasis id="A3A9EA9AFFEFFFC3FBA9EFE5FBDEF9A2" bold="true" box="[1055,1078,1520,1540]" pageId="26" pageNumber="1505">39</emphasis>
) male sternite VIII; (
<emphasis id="A3A9EA9AFFEFFFC3FEDEEC12FE75F9BD" bold="true" box="[360,413,1543,1563]" pageId="26" pageNumber="1505">40-41</emphasis>
) aedeagus in lateral and in ventral view; (
<emphasis id="A3A9EA9AFFEFFFC3FCABEC12FCDCF9BD" bold="true" box="[797,820,1543,1563]" pageId="26" pageNumber="1505">42</emphasis>
) ventral process of aedeagus in ventral view; (
<emphasis id="A3A9EA9AFFEFFFC3FEE7EC08FE87F997" bold="true" box="[337,367,1565,1585]" pageId="26" pageNumber="1505">43)</emphasis>
internal structure of aedeagus in dorsal view. Scale bars: 36-37: 1.0 mm; 38- 43: 0.2 mm.
</paragraph>
</caption>
<caption id="C5A26600FFEEFFC2FF04EE56FD47FBC8" ID-DOI="http://doi.org/10.5281/zenodo.5301305" ID-Zenodo-Dep="5301305" httpUri="https://zenodo.org/record/5301305/files/figure.png" pageId="27" pageNumber="1506" startId="27.[178,223,1091,1111]" targetBox="[178,1085,212,1066]" targetPageId="27">
<paragraph id="91623688FFEEFFC2FF04EE56FD47FBC8" blockId="27.[178,1085,1091,1134]" pageId="27" pageNumber="1506">
<emphasis id="A3A9EA9AFFEEFFC2FF04EE56FF1BFBF1" bold="true" box="[178,243,1091,1111]" pageId="27" pageNumber="1506">Map 4</emphasis>
: Distributions of
<taxonomicName id="56DD4D0BFFEEFFC2FE2DEE56FDADFBF1" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[411,581,1091,1111]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="1506" phylum="Arthropoda" rank="species" species="distortus" status="sp. nov.">
<emphasis id="A3A9EA9AFFEEFFC2FE2DEE56FDADFBF1" box="[411,581,1091,1111]" italics="true" pageId="27" pageNumber="1506">Micrillus distortus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89A57E1FFEEFFC2FDF8EE56FD78FBF1" box="[590,656,1091,1111]" pageId="27" pageNumber="1506" rank="species">nov.sp.</taxonomicNameLabel>
(triangles) and
<taxonomicName id="56DD4D0BFFEEFFC2FC90EE56FBE6FBF1" authority="(CAMERON)" authorityName="Assing" authorityYear="2013" baseAuthorityName="Cameron" baseAuthorityYear="1930" box="[806,1038,1091,1111]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="1506" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="A3A9EA9AFFEEFFC2FC90EE56FC7EFBF1" box="[806,918,1091,1111]" italics="true" pageId="27" pageNumber="1506">M. suturalis</emphasis>
(CAMERON)
</taxonomicName>
(circles) in the Oriental region, based on examined records.
</paragraph>
</caption>
<paragraph id="91623688FFEEFFC2FF04EE68FEF4FB4F" blockId="27.[178,1085,1148,1257]" pageId="27" pageNumber="1506">
Pronotum (
<figureCitation id="09E62A0DFFEEFFC2FE91EE69FE9DFB34" box="[295,373,1148,1170]" captionStart="Figs 36-43" captionStartId="26.[178,218,1520,1540]" captionTargetBox="[186,1078,212,1484]" captionTargetId="figure-65@26.[178,1085,212,1497]" captionTargetPageId="26" captionText="Figs 36-43: Micrillus distortus nov.sp.: (36) habitus; (37) forebody; (38) male sternite VII; (39) male sternite VIII; (40-41) aedeagus in lateral and in ventral view; (42) ventral process of aedeagus in ventral view; (43) internal structure of aedeagus in dorsal view. Scale bars: 36-37: 1.0 mm; 38- 43: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301303" httpUri="https://zenodo.org/record/5301303/files/figure.png" pageId="27" pageNumber="1506">Fig. 37</figureCitation>
) approximately 1.1 times as long as broad and almost as broad as head, noticeably tapering posteriad; surface glossy and with extremely fine, barely noticeable micropunctation; on either side of midline with series of few (4-5) larger punctures.
</paragraph>
<paragraph id="91623688FFEEFFC2FF04EEECFEFDFAEF" blockId="27.[178,1086,1273,1354]" pageId="27" pageNumber="1506">
Elytra (
<figureCitation id="09E62A0DFFEEFFC2FF49EEECFEA2FAA9" box="[255,330,1273,1295]" captionStart="Figs 36-43" captionStartId="26.[178,218,1520,1540]" captionTargetBox="[186,1078,212,1484]" captionTargetId="figure-65@26.[178,1085,212,1497]" captionTargetPageId="26" captionText="Figs 36-43: Micrillus distortus nov.sp.: (36) habitus; (37) forebody; (38) male sternite VII; (39) male sternite VIII; (40-41) aedeagus in lateral and in ventral view; (42) ventral process of aedeagus in ventral view; (43) internal structure of aedeagus in dorsal view. Scale bars: 36-37: 1.0 mm; 38- 43: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301303" httpUri="https://zenodo.org/record/5301303/files/figure.png" pageId="27" pageNumber="1506">Fig. 37</figureCitation>
) approximately as long as pronotum; punctation dense and distinct. Hind wing fully developed. Metatarsomere I approximately as long as the combined length of II and III.
</paragraph>
<paragraph id="91623688FFEEFFC2FF04EF4CFD01FA0F" blockId="27.[178,1085,1369,1449]" pageId="27" pageNumber="1506">Abdomen slightly narrower than elytra; punctation rather dense, fine, but distinct; interstices with very shallow microsculpture; posterior margin of tergite VII with palisade fringe; posterior margin of tergite VIII strongly convex.</paragraph>
<paragraph id="91623688FFEEFFC2FF71EFACFCBDF980" blockId="27.[178,1085,1465,1574]" pageId="27" pageNumber="1506">
: sternite VII (
<figureCitation id="09E62A0DFFEEFFC2FEECEFACFE4DFA69" box="[346,421,1465,1487]" captionStart="Figs 36-43" captionStartId="26.[178,218,1520,1540]" captionTargetBox="[186,1078,212,1484]" captionTargetId="figure-65@26.[178,1085,212,1497]" captionTargetPageId="26" captionText="Figs 36-43: Micrillus distortus nov.sp.: (36) habitus; (37) forebody; (38) male sternite VII; (39) male sternite VIII; (40-41) aedeagus in lateral and in ventral view; (42) ventral process of aedeagus in ventral view; (43) internal structure of aedeagus in dorsal view. Scale bars: 36-37: 1.0 mm; 38- 43: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301303" httpUri="https://zenodo.org/record/5301303/files/figure.png" pageId="27" pageNumber="1506">Fig. 38</figureCitation>
) moderately transverse, posterior margin concave in the middle, pubescence unmodified; sternite VIII (
<figureCitation id="09E62A0DFFEEFFC2FD8DEFC3FD6DFA4A" box="[571,645,1494,1516]" captionStart="Figs 36-43" captionStartId="26.[178,218,1520,1540]" captionTargetBox="[186,1078,212,1484]" captionTargetId="figure-65@26.[178,1085,212,1497]" captionTargetPageId="26" captionText="Figs 36-43: Micrillus distortus nov.sp.: (36) habitus; (37) forebody; (38) male sternite VII; (39) male sternite VIII; (40-41) aedeagus in lateral and in ventral view; (42) ventral process of aedeagus in ventral view; (43) internal structure of aedeagus in dorsal view. Scale bars: 36-37: 1.0 mm; 38- 43: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301303" httpUri="https://zenodo.org/record/5301303/files/figure.png" pageId="27" pageNumber="1506">Fig. 39</figureCitation>
) oblong, with deep and very narrow posterior excision; aedeagus
<quantity id="56259B6DFFEEFFC2FE00EFE6FDA3F9AF" box="[438,587,1523,1545]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="27" pageNumber="1506" unit="mm" value="0.425" valueMax="0.45" valueMin="0.4">0.40-0.45 mm</quantity>
long, strongly asymmetric, and of distinctive shape; internal sac with long and asymmetric sclerotized structure.
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFE9FFC5FF04EACFFC42FE1B" pageId="28" pageNumber="1507" type="discussion">
<paragraph id="91623688FFE9FFC5FF04EACFFC42FE1B" blockId="28.[178,1086,218,445]" pageId="28" pageNumber="1507">
Comparative notes: This species is characterized by the distinctive morphology of the aedeagus and additionally distinguished from
<taxonomicName id="56DD4D0BFFE9FFC5FC80EAECFC4CFEA8" baseAuthorityName="Eppelsheim" baseAuthorityYear="1890" box="[822,932,248,270]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1507" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="A3A9EA9AFFE9FFC5FC80EAECFC4CFEA8" box="[822,932,248,270]" italics="true" pageId="28" pageNumber="1507">M. indicus</emphasis>
</taxonomicName>
,
<taxonomicName id="56DD4D0BFFE9FFC5FC02EAECFBDEFEA8" authorityName="Cameron" authorityYear="1931" box="[948,1078,249,270]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1507" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="A3A9EA9AFFE9FFC5FC02EAECFBDEFEA8" box="[948,1078,249,270]" italics="true" pageId="28" pageNumber="1507">M. nigriceps</emphasis>
</taxonomicName>
, and
<taxonomicName id="56DD4D0BFFE9FFC5FF56EB03FE86FE8D" authorityName="Motschulsky" authorityYear="1858" box="[224,366,277,299]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1507" phylum="Arthropoda" rank="species" species="flavescens">
<emphasis id="A3A9EA9AFFE9FFC5FF56EB03FE86FE8D" box="[224,366,277,299]" italics="true" pageId="28" pageNumber="1507">M. flavescens</emphasis>
</taxonomicName>
by different head shape (flattened, posteriorly not tapering, not oblong) and the almost obsolete microsculpture of the head, from
<taxonomicName id="56DD4D0BFFE9FFC5FCC4EB26FC0CFEEE" baseAuthorityName="Eppelsheim" baseAuthorityYear="1890" box="[882,996,306,328]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1507" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="A3A9EA9AFFE9FFC5FCC4EB26FC0CFEEE" box="[882,996,306,328]" italics="true" pageId="28" pageNumber="1507">M. indicus</emphasis>
</taxonomicName>
and
<taxonomicName id="56DD4D0BFFE9FFC5FB97EB26FEF8FEC3" authorityName="Cameron" authorityYear="1931" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1507" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="A3A9EA9AFFE9FFC5FB97EB26FEF8FEC3" italics="true" pageId="28" pageNumber="1507">M. nigriceps</emphasis>
</taxonomicName>
also by shorter antennae with less oblong preapical antennomeres, and from
<emphasis id="A3A9EA9AFFE9FFC5FB97EB44FEDEFE25" italics="true" pageId="28" pageNumber="1507">M. asperiventris</emphasis>
by the less convex, posteriorly distinctly tapering pronotum with much sparser and finer punctation, the longer and more slender antennae (preapical antennomeres not distinctly transverse), and the denser punctation of the abdomen.
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFE9FFC5FF04EBD8FE30FD9C" pageId="28" pageNumber="1507" type="distribution">
<paragraph id="91623688FFE9FFC5FF04EBD8FE30FD9C" blockId="28.[178,1086,461,570]" pageId="28" pageNumber="1507">
Distribution and natural history:
<taxonomicName id="56DD4D0BFFE9FFC5FCAFEBD8FC31FE45" authorityName="Assing" authorityYear="2013" box="[793,985,461,483]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1507" phylum="Arthropoda" rank="species" species="distortus">
<emphasis id="A3A9EA9AFFE9FFC5FCAFEBD8FC31FE45" box="[793,985,461,483]" italics="true" pageId="28" pageNumber="1507">Micrillus distortus</emphasis>
</taxonomicName>
was discovered in two localities in
<collectingRegion id="5319F86AFFE9FFC5FE66EBFFFD55FDA6" box="[464,701,490,512]" country="Laos" name="Khammouan" pageId="28" pageNumber="1507">Khammouane province</collectingRegion>
in central
<collectingCountry id="E9CA7618FFE9FFC5FC9BEBFEFC89FDA6" box="[813,865,491,512]" name="Laos" pageId="28" pageNumber="1507">Laos</collectingCountry>
, one in
<collectingCountry id="E9CA7618FFE9FFC5FC02EBFFFBE5FDA6" box="[948,1037,490,512]" name="Vietnam" pageId="28" pageNumber="1507">Vietnam</collectingCountry>
, and one in northern
<collectingCountry id="E9CA7618FFE9FFC5FEEFE812FE5BFDBB" box="[345,435,519,541]" name="Thailand" pageId="28" pageNumber="1507">Thailand</collectingCountry>
(
<figureCitation id="09E62A0DFFE9FFC5FE74E812FDE1FDBB" box="[450,521,519,541]" captionStart="Map 4" captionStartId="27.[178,223,1091,1111]" captionTargetBox="[178,1085,212,1066]" captionTargetId="figure-174@27.[178,1085,212,1066]" captionTargetPageId="27" captionText="Map 4: Distributions of Micrillus distortus nov.sp. (triangles) and M. suturalis (CAMERON) (circles) in the Oriental region, based on examined records." figureDoi="http://doi.org/10.5281/zenodo.5301305" httpUri="https://zenodo.org/record/5301305/files/figure.png" pageId="28" pageNumber="1507">Map 4</figureCitation>
). The specimens from the
<typeStatus id="4E66882AFFE9FFC5FC96E81DFCA4FDBB" box="[800,844,520,541]" pageId="28" pageNumber="1507">type</typeStatus>
locality were collected in a degraded primary forest.
</paragraph>
</subSubSection>
</treatment>
</document>
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