459 lines
53 KiB
XML
459 lines
53 KiB
XML
<document id="E44A6BCD550988C5952993DF1B110473" ID-CLB-Dataset="24056" ID-DOI="10.1080/00222933.2019.1709669" ID-GBIF-Dataset="175ef2e7-06f6-4404-a6be-224dac1a06ec" ID-ISSN="1464-5262" ID-Zenodo-Dep="3666538" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581606429779" checkinUser="valdenar" docAuthor="Freitag, Hendrik, Molls, Christian, Bouma, Aglaia M., Garces, Jhoana M., Rossato, Marzia, Cosentino, Emanuela & Delledonne, Massimo" docDate="2019" docId="052D87A15870130CFEBEAA71FB83FD6A" docLanguage="en" docName="J_NAT_HIST.53.41–42.2593–2620.pdf.imf" docOrigin="Journal of Natural History 53 (41)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Grouvellinus borneensis Freitag, Molls and Bouma 2019, sp. nov" docType="treatment" docVersion="12" lastPageNumber="2616" masterDocId="F914FFD958641314FF8EA961FFF3FFB2" masterDocTitle="Additional new species of Grouvellinus Champion 1923 (Insectaı Coleopteraı Elmidae) discovered by citizen scientists and DNA barcoded in the field applying a novel MinIONbased workflow" masterLastPageNumber="2620" masterPageNumber="2593" pageNumber="2612" updateTime="1698782781787" updateUser="ExternalLinkService">
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<mods:titleInfo id="6973678564F51AA08AD593CF73B10D19">
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<mods:title id="2E508DB8224CF6DDF98331F24801F209">Additional new species of Grouvellinus Champion 1923 (Insectaı Coleopteraı Elmidae) discovered by citizen scientists and DNA barcoded in the field applying a novel MinIONbased workflow</mods:title>
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<mods:namePart id="4809325FCB8110B175CEC9DBBEA40B55">Freitag, Hendrik</mods:namePart>
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<mods:namePart id="FC8F3283BF182A3F504784F0CCD7D574">Molls, Christian</mods:namePart>
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<mods:roleTerm id="2AF0AA80A3FFC998A38202F14F19190F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F6472A60BAD04344FEAC9FBE7474DDDF">Bouma, Aglaia M.</mods:namePart>
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<mods:roleTerm id="029B4CBB666B4AF554659C2B36917843">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="73FC585BB7D1E8DA05BA756D61BFD59C">Garces, Jhoana M.</mods:namePart>
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</mods:name>
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<mods:name id="686063EABD569920077D543892BF2165" type="personal">
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<mods:roleTerm id="66E347B49F5D14794FF45125064C7F11">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BBF5FB11245C7ED643EC6CF2D5B0CEBD">Rossato, Marzia</mods:namePart>
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</mods:role>
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<mods:namePart id="240F206C3A6FE234212E67457D3E62E5">Cosentino, Emanuela</mods:namePart>
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</mods:name>
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<mods:name id="1320C34E6B63EF0AB3C955E66B05AD50" type="personal">
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<mods:roleTerm id="1BB8A25B22B8A17153A693B32F8FA426">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2ED965B2F370E46F239D2E8CB5A942C6">Delledonne, Massimo</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="5C8106561912251F796D234CE9571509">text</mods:typeOfResource>
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<mods:title id="3DAE289A18DC1FE9693C6652BDC2AEA7">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="D10D0E5ECD2FFF8852BC9BD6254E2F8E">
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<mods:date id="97F563B4720A1832FD56FDB87FE70438">2019</mods:date>
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<mods:detail id="14E906E40F24707C24D99CFE0A1997A3" type="pubDate">
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<mods:number id="57CCA95CF1EAA54A86B27CB9543857E4">2020-01-24</mods:number>
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<mods:detail id="947F667F2EDB7134191823C84301633A" type="volume">
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<mods:number id="7A430DC0AA69146DCDAA681E277DB05D">53</mods:number>
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</mods:detail>
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<mods:detail id="C674AFB1F6AC340C299A2A5DA3EF94F1" type="issue">
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<mods:number id="86729DC30E3901E44B001CAA42DF2B72">41</mods:number>
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<mods:extent id="5DDF74D1D7CEC03DA7F75BE37C89400A" unit="page">
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<mods:start id="EF7FFACA69C2FF1CF18396C166321A24">2593</mods:start>
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<mods:end id="3858900D375B42FD098BBA7F6E3FAEAF">2620</mods:end>
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<mods:classification id="45F7B540AD7A1F14E842A10756FD855C">journal article</mods:classification>
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<mods:identifier id="31DDAEBF47472F88A2D908D71987E15A" type="CLB-Dataset">24056</mods:identifier>
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<mods:identifier id="B5849EECCEBFCA3A825C330679308C46" type="DOI">10.1080/00222933.2019.1709669</mods:identifier>
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<mods:identifier id="FB8B88F95055AF083F01E023D754F863" type="GBIF-Dataset">175ef2e7-06f6-4404-a6be-224dac1a06ec</mods:identifier>
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<mods:identifier id="8BE68DFC44187AA5015F08DE46924D4C" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="522A5E5991F02167C0479BB2F3B558AC" type="Zenodo-Dep">3666538</mods:identifier>
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<treatment id="052D87A15870130CFEBEAA71FB83FD6A" ID-DOI="http://doi.org/10.5281/zenodo.3671824" ID-GBIF-Taxon="162136041" ID-Zenodo-Dep="3671824" LSID="urn:lsid:plazi:treatment:052D87A15870130CFEBEAA71FB83FD6A" httpUri="http://treatment.plazi.org/id/052D87A15870130CFEBEAA71FB83FD6A" lastPageId="24" lastPageNumber="2616" pageId="20" pageNumber="2612">
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<subSubSection id="C59E653C58701300FEBEAA71FCC1FCF9" pageId="20" pageNumber="2612" type="nomenclature">
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<paragraph id="8D3B36B758701300FEBEAA71FC07FC9B" blockId="20.[304,1012,784,843]" box="[304,1012,784,809]" pageId="20" pageNumber="2612">
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<heading id="D67381DB58701300FEBEAA71FC03FC9B" box="[304,1008,784,809]" pageId="20" pageNumber="2612">
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||
<taxonomicName id="4A844D3458701300FEBEAA71FC7DFC9B" authority="Freitag, Molls and Bouma" authorityName="Freitag, Molls and Bouma" authorityYear="2019" box="[304,910,784,809]" class="Insecta" family="Elmidae" genus="Grouvellinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="2612" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
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||
<emphasis id="BFF0EAA558701300FEBEAA71FDA5FC9B" bold="true" box="[304,598,784,809]" italics="true" pageId="20" pageNumber="2612">Grouvellinus borneensis</emphasis>
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||
Freitag, Molls and Bouma
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</taxonomicName>
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,
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<taxonomicNameLabel id="A4C357DE58701300FC16AA71FC03FC9B" box="[920,1008,784,809]" pageId="20" pageNumber="2612" rank="species">sp. nov</taxonomicNameLabel>
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</heading>
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||
.
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||
</paragraph>
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||
<paragraph id="8D3B36B758701300FE7FAA53FCC1FCF9" blockId="20.[304,1012,784,843]" box="[497,818,818,843]" pageId="20" pageNumber="2612">
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<heading id="D67381DB58701300FE7FAA53FCC1FCF9" box="[497,818,818,843]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="2612" reason="5">
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(
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<figureCitation id="15BF2A3258701300FE77AA53FD96FCF9" box="[505,613,818,843]" captionStart="Figures 6–9" captionStartId="7.[160,235,1337,1360]" captionTargetBox="[163,1153,148,1289]" captionTargetId="figure@7.[163,1153,142,1306]" captionTargetPageId="7" captionText="Figures 6–9. Habitus of the Grouvellinus species collected from the Tawan Hills Park. 6: G. luciaensis, sp. nov. (paratype male from Lucia River); 7: G. borneensis, sp. nov. (paratype male from Tawau River environment); 8: G. sp.1 (female from Tawau River environment); 9: G. sp.2 (female from Lucia River). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3666544" httpUri="https://zenodo.org/record/3666544/files/figure.png" pageId="20" pageNumber="2612">Figures 7</figureCitation>
|
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,
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<figureCitation id="15BF2A3258701300FDFDAA53FD63FCF9" box="[627,656,818,843]" captionStart="Figure 16" captionStartId="21.[160,225,1038,1061]" captionTargetBox="[163,1153,142,1006]" captionTargetId="figure@21.[163,1153,142,1006]" captionTargetPageId="21" captionText="Figure 16. Grouvellinus borneensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum and elytra; (b) ventral aspect with prosternum and mesoventrite; (c) posteriodorsal aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666558" httpUri="https://zenodo.org/record/3666558/files/figure.png" pageId="20" pageNumber="2612">16</figureCitation>
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(a
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<emphasis id="BFF0EAA558701300FD2BAA53FD40FCF9" box="[677,691,818,843]" italics="true" pageId="20" pageNumber="2612">–</emphasis>
|
||
c), 17(a
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||
<emphasis id="BFF0EAA558701300FC89AA53FCE6FCF9" box="[775,789,818,843]" italics="true" pageId="20" pageNumber="2612">–</emphasis>
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||
c))
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C59E653C58701300FF2EAAE8FD4CFC77" pageId="20" pageNumber="2612" type="distribution">
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||
<paragraph id="8D3B36B758701300FF2EAAE8FD4CFC77" blockId="20.[160,1155,905,965]" pageId="20" pageNumber="2612">
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<emphasis id="BFF0EAA558701300FF2EAAE8FEB2FC10" bold="true" box="[160,321,905,930]" italics="true" pageId="20" pageNumber="2612">
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||
<typeStatus id="523F881558701300FF2EAAE8FF2BFC10" box="[160,216,905,930]" pageId="20" pageNumber="2612">Type</typeStatus>
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||
locality.
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||
</emphasis>
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||
<collectingCountry id="F593762758701300FEDFAAEBFE40FC10" box="[337,435,906,930]" name="Malaysia" pageId="20" pageNumber="2612">Malaysia</collectingCountry>
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||
,
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||
<collectingRegion id="4F40F85558701300FE4CAAEBFDF4FC10" box="[450,519,906,930]" country="Malaysia" name="Sabah" pageId="20" pageNumber="2612">Sabah</collectingRegion>
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||
(on Borneo Island), Tawau Hills Park, Tawau River,
|
||
<geoCoordinate id="E8B0507058701300FBE3AAEBFF08FC77" degrees="4" direction="north" minutes="24" orientation="latitude" pageId="20" pageNumber="2612" precision="15" seconds="14" value="4.403889">
|
||
4° 24
|
||
<emphasis id="BFF0EAA558701300FF35AACDFF32FC76" bold="true" box="[187,193,940,964]" italics="true" pageId="20" pageNumber="2612">ʹ</emphasis>
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||
14
|
||
<emphasis id="BFF0EAA558701300FF53AACDFF1AFC76" bold="true" box="[221,233,940,964]" italics="true" pageId="20" pageNumber="2612">ʹʹ</emphasis>
|
||
N
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||
</geoCoordinate>
|
||
,
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||
<geoCoordinate id="E8B0507058701300FE89AACCFE63FC77" box="[263,400,940,965]" degrees="117" direction="east" minutes="53" orientation="longitude" pageId="20" pageNumber="2612" precision="15" seconds="35" value="117.89305">
|
||
117°53
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||
<emphasis id="BFF0EAA558701300FEDAAACDFEA9FC76" bold="true" box="[340,346,940,964]" italics="true" pageId="20" pageNumber="2612">ʹ</emphasis>
|
||
35
|
||
<emphasis id="BFF0EAA558701300FEFBAACDFE72FC76" bold="true" box="[373,385,940,964]" italics="true" pageId="20" pageNumber="2612">ʹʹ</emphasis>
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||
E
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||
</geoCoordinate>
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||
,
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||
<quantity id="4A7C9B5258701300FE15AACCFE12FC77" box="[411,481,941,965]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="20" pageNumber="2612" unit="m" value="270.0">270 m</quantity>
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||
a.s.l. (
|
||
<figureCitation id="15BF2A3258701300FDA8AACCFD8FFC77" box="[550,636,941,965]" captionStart="Figure 1" captionStartId="3.[160,225,1413,1436]" captionTargetBox="[163,1153,142,1381]" captionTargetId="figure@3.[163,1153,142,1381]" captionTargetPageId="3" captionText="Figure 1. (a) Co-authors and assistants of the Taxon Expedition to Tawau Hills Park, Sabah, Malaysian Borneo, performing microhabitat-specific sampling; (b) Co-authors during beetle collection at a blacklight trap installed in the vicinity of the Tawau River; (c) The first author (second from the right) instructing Taxon Expedition participants on the identification of riffle beetles; (d) the transportable molecular field laboratory at the Tawau Park Station; (e) Type locality of Grouvellinus borneensis, sp. nov., G. cruxniger, sp. nov., G. nigerquadratus, sp. nov., and collection site of G. sp. 1; (f) Type locality of Grouvellinus luciaensis, sp. nov., and collection site of G. andrekuipersi, G. nigerquadratus, sp. nov. and G. sp. 2." figureDoi="http://doi.org/10.5281/zenodo.3666540" httpUri="https://zenodo.org/record/3666540/files/figure.png" pageId="20" pageNumber="2612">Figure 1</figureCitation>
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||
(a,e)).
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C59E653C58701300FF2EAA90FCE9FB05" pageId="20" pageNumber="2612" type="materials_examined">
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<paragraph id="8D3B36B758701300FF2EAA90FCE9FB05" blockId="20.[160,1156,1009,1207]" pageId="20" pageNumber="2612">
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<emphasis id="BFF0EAA558701300FF2EAA90FEBFFBB8" bold="true" box="[160,332,1009,1034]" italics="true" pageId="20" pageNumber="2612">Type material.</emphasis>
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<materialsCitation id="3DEC3CEA58701300FED2AA93FB73FBFD" ID-GBIF-Occurrence="2565372557" collectorName="Taxon Expedition" country="Malaysia" elevation="270" latitude="4.403889" location="Light trap near Park Headquarter" longLatPrecision="21" longitude="117.89305" municipality="Tawau River" pageId="20" pageNumber="2612" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
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||
<emphasis id="BFF0EAA558701300FED2AA93FE39FBB8" bold="true" box="[348,458,1010,1034]" pageId="20" pageNumber="2612">
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<typeStatus id="523F881558701300FED2AA93FE39FBB8" box="[348,458,1010,1034]" pageId="20" pageNumber="2612" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
3 [H7] (
|
||
<specimenCount id="9B82FD3E58701300FDA0AA93FDBCFBB8" box="[558,591,1010,1034]" pageId="20" pageNumber="2612" type="generic">SP</specimenCount>
|
||
):
|
||
<emphasis id="BFF0EAA558701300FDECAA93FD94FBB8" box="[610,615,1010,1034]" italics="true" pageId="20" pageNumber="2612">‘</emphasis>
|
||
<collectingCountry id="F593762758701300FDE9AA93FD2CFBB8" box="[615,735,1010,1034]" name="Malaysia" pageId="20" pageNumber="2612">MALAYSIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="4F40F85558701300FD65AA93FCC0FBB8" box="[747,819,1010,1034]" country="Malaysia" name="Sabah" pageId="20" pageNumber="2612">Sabah</collectingRegion>
|
||
:
|
||
<collectingMunicipality id="6D5FACCD58701300FCB1AA93FC38FBB8" box="[831,971,1010,1034]" pageId="20" pageNumber="2612">Tawau River</collectingMunicipality>
|
||
;
|
||
<location id="885B606C58701300FC56AA93FE9BFB9F" LSID="urn:lsid:plazi:treatment:052D87A15870130CFEBEAA71FB83FD6A:885B606C58701300FC56AA93FE9BFB9F" country="Malaysia" latitude="4.403889" longLatPrecision="21" longitude="117.89305" municipality="Tawau River" name="Light trap near Park Headquarter" pageId="20" pageNumber="2612" stateProvince="Sabah">Light trap near Park Headquarter</location>
|
||
; ca.
|
||
<geoCoordinate id="E8B0507058701300FE14AD74FDF8FB9F" box="[410,523,1044,1069]" degrees="4" direction="north" minutes="24" orientation="latitude" pageId="20" pageNumber="2612" precision="15" seconds="14" value="4.403889">
|
||
4°24
|
||
<emphasis id="BFF0EAA558701300FE45AD75FE22FB9E" bold="true" box="[459,465,1044,1068]" italics="true" pageId="20" pageNumber="2612">ʹ</emphasis>
|
||
14
|
||
<emphasis id="BFF0EAA558701300FE63AD75FE0AFB9E" bold="true" box="[493,505,1044,1068]" italics="true" pageId="20" pageNumber="2612">ʹʹ</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E8B0507058701300FD96AD74FD52FB9F" box="[536,673,1044,1069]" degrees="117" direction="east" minutes="53" orientation="longitude" pageId="20" pageNumber="2612" precision="15" seconds="35" value="117.89305">
|
||
117°53
|
||
<emphasis id="BFF0EAA558701300FDEAAD75FD99FB9E" bold="true" box="[612,618,1044,1068]" italics="true" pageId="20" pageNumber="2612">ʹ</emphasis>
|
||
35
|
||
<emphasis id="BFF0EAA558701300FD08AD75FD61FB9E" bold="true" box="[646,658,1044,1068]" italics="true" pageId="20" pageNumber="2612">ʹʹ</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
,
|
||
<quantity id="4A7C9B5258701300FD22AD74FD07FB9F" box="[684,756,1045,1069]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="20" pageNumber="2612" unit="m" value="270.0">
|
||
<elevation id="06A9D18458701300FD22AD74FD07FB9F" box="[684,756,1045,1069]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="20" pageNumber="2612" unit="m" value="270.0">270 m</elevation>
|
||
</quantity>
|
||
a.s.l.; leg.
|
||
<collectorName id="2071536158701300FCE7AD74FBDEFB9F" box="[873,1069,1045,1069]" pageId="20" pageNumber="2612">Taxon Expedition</collectorName>
|
||
participants 12.
|
||
<location id="885B606C58701300FE88AD56FEBDFBFD" LSID="urn:lsid:plazi:treatment:052D87A15870130CFEBEAA71FB83FD6A:885B606C58701300FE88AD56FEBDFBFD" box="[262,334,1079,1103]" country="Malaysia" latitude="4.403889" longLatPrecision="21" longitude="117.89305" municipality="Tawau River" name="March" pageId="20" pageNumber="2612" stateProvince="Sabah">March</location>
|
||
.2018 (1)L
|
||
<emphasis id="BFF0EAA558701300FE31AD56FE37FBFD" box="[447,452,1079,1103]" italics="true" pageId="20" pageNumber="2612">’</emphasis>
|
||
, terminal parts of abdomen incl. aedeagus glued separately
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3DEC3CEA58701300FF2EAD3BFCE9FB05" ID-GBIF-Occurrence="2565372558" pageId="20" pageNumber="2612" typeStatus="paratype">
|
||
<emphasis id="BFF0EAA558701300FF2EAD3BFEE4FBC0" bold="true" box="[160,279,1114,1138]" pageId="20" pageNumber="2612">
|
||
<typeStatus id="523F881558701300FF2EAD3BFEE4FBC0" box="[160,279,1114,1138]" pageId="20" pageNumber="2612" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
: 9 3,
|
||
<specimenCount id="9B82FD3E58701300FEECAD3BFE77FBC0" box="[354,388,1114,1138]" pageId="20" pageNumber="2612" type="female">4♀</specimenCount>
|
||
(SP) same data as
|
||
<typeStatus id="523F881558701300FDEAAD3BFD38FBC0" box="[612,715,1114,1138]" pageId="20" pageNumber="2612" type="holotype">holotype</typeStatus>
|
||
; 13,
|
||
<specimenCount id="9B82FD3E58701300FC87AD3BFCD8FBC0" box="[777,811,1114,1138]" pageId="20" pageNumber="2612" type="female">1♀</specimenCount>
|
||
[H49] (SP)
|
||
<emphasis id="BFF0EAA558701300FC21AD3BFC47FBC0" box="[943,948,1114,1138]" italics="true" pageId="20" pageNumber="2612">‘</emphasis>
|
||
<collectingCountry id="F593762758701300FC3AAD3BFBDFFBC0" box="[948,1068,1114,1138]" name="Malaysia" pageId="20" pageNumber="2612">MALAYSIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="4F40F85558701300FBB7AD3BFB72FBC0" box="[1081,1153,1114,1138]" country="Malaysia" name="Sabah" pageId="20" pageNumber="2612">Sabah</collectingRegion>
|
||
: Tawau River; bottom gravel, run; primary forest; ca.
|
||
<geoCoordinate id="E8B0507058701300FD63AD1CFCACFB27" box="[749,863,1148,1173]" degrees="4" direction="north" minutes="24" orientation="latitude" pageId="20" pageNumber="2612" precision="15" seconds="14" value="4.403889">
|
||
4°24
|
||
<emphasis id="BFF0EAA558701300FC91AD1DFCD6FB26" bold="true" box="[799,805,1148,1172]" italics="true" pageId="20" pageNumber="2612">ʹ</emphasis>
|
||
14
|
||
<emphasis id="BFF0EAA558701300FCCEAD1DFCBFFB26" bold="true" box="[832,844,1148,1172]" italics="true" pageId="20" pageNumber="2612">ʹʹ</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E8B0507058701300FCE5AD1CFC06FB27" box="[875,1013,1148,1173]" degrees="117" direction="east" minutes="53" orientation="longitude" pageId="20" pageNumber="2612" precision="15" seconds="35" value="117.89305">
|
||
117°53
|
||
<emphasis id="BFF0EAA558701300FC36AD1DFC4DFB26" bold="true" box="[952,958,1148,1172]" italics="true" pageId="20" pageNumber="2612">ʹ</emphasis>
|
||
35
|
||
<emphasis id="BFF0EAA558701300FC57AD1DFC16FB26" bold="true" box="[985,997,1148,1172]" italics="true" pageId="20" pageNumber="2612">ʹʹ</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
,
|
||
<quantity id="4A7C9B5258701300FC71AD1CFBB4FB27" box="[1023,1095,1149,1173]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="20" pageNumber="2612" unit="m" value="280.0">280 m</quantity>
|
||
a.s.l.; leg. Taxon Expedition participants 12.March.2018 (1c)M
|
||
<emphasis id="BFF0EAA558701300FC80ADFEFCE7FB05" box="[782,788,1183,1207]" italics="true" pageId="20" pageNumber="2612">’</emphasis>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C59E653C58701300FF2EAD85FBE7FAAD" box="[160,1044,1252,1311]" pageId="20" pageNumber="2612" type="etymology">
|
||
<paragraph id="8D3B36B758701300FF2EAD85FEEEFB4F" blockId="20.[160,1044,1252,1311]" box="[160,285,1252,1277]" pageId="20" pageNumber="2612">
|
||
<emphasis id="BFF0EAA558701300FF2EAD85FEEEFB4F" bold="true" box="[160,285,1252,1277]" italics="true" pageId="20" pageNumber="2612">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758701300FF2EAC66FBE7FAAD" blockId="20.[160,1044,1252,1311]" box="[160,1044,1287,1311]" pageId="20" pageNumber="2612">The species is named for the island of Borneo, where it is most likely endemic.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C59E653C5870130CFF2EAC2DFE26FDD8" lastPageId="24" lastPageNumber="2616" pageId="20" pageNumber="2612" type="description">
|
||
<paragraph id="8D3B36B758701300FF2EAC2DFED5FAD7" blockId="20.[160,1156,1356,1727]" box="[160,294,1356,1381]" pageId="20" pageNumber="2612">
|
||
<emphasis id="BFF0EAA558701300FF2EAC2DFED5FAD7" bold="true" box="[160,294,1356,1381]" italics="true" pageId="20" pageNumber="2612">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758701300FF2EAC0EFBBDFA18" blockId="20.[160,1156,1356,1727]" pageId="20" pageNumber="2612">
|
||
Body elongate obovate, males
|
||
<quantity id="4A7C9B5258701300FD84AC0EFD5DFA35" box="[522,686,1391,1415]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" metricValueMax="1.56" metricValueMin="1.5" pageId="20" pageNumber="2612" unit="mm" value="1.53" valueMax="1.56" valueMin="1.5">
|
||
1.50
|
||
<emphasis id="BFF0EAA558701300FDB6AC0EFDB6FA35" box="[568,581,1391,1415]" italics="true" pageId="20" pageNumber="2612">–</emphasis>
|
||
1.56 mm
|
||
</quantity>
|
||
, females
|
||
<quantity id="4A7C9B5258701300FC93AC0EFC4DFA35" box="[797,958,1391,1415]" metricMagnitude="-3" metricUnit="m" metricValue="1.635" metricValueMax="1.67" metricValueMin="1.6" pageId="20" pageNumber="2612" unit="mm" value="1.635" valueMax="1.67" valueMin="1.6">
|
||
1.60
|
||
<emphasis id="BFF0EAA558701300FCC5AC0EFCABFA35" box="[843,856,1391,1415]" italics="true" pageId="20" pageNumber="2612">–</emphasis>
|
||
1.67 mm
|
||
</quantity>
|
||
long (CL), males
|
||
<quantity id="4A7C9B5258701300FF2EACF3FEB2FA18" box="[160,321,1426,1450]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" metricValueMax="7.8" metricValueMin="7.4" pageId="20" pageNumber="2612" unit="mm" value="0.76" valueMax="0.78" valueMin="0.74">
|
||
0.74
|
||
<emphasis id="BFF0EAA558701300FF41ACF3FF2FFA18" box="[207,220,1426,1450]" italics="true" pageId="20" pageNumber="2612">–</emphasis>
|
||
0.78 mm
|
||
</quantity>
|
||
, females
|
||
<quantity id="4A7C9B5258701300FE27ACF3FDB4FA18" box="[425,583,1426,1450]" metricMagnitude="-4" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.299999999999999" metricValueMin="8.0" pageId="20" pageNumber="2612" unit="mm" value="0.815" valueMax="0.83" valueMin="0.8">
|
||
0.80
|
||
<emphasis id="BFF0EAA558701300FE56ACF3FE16FA18" box="[472,485,1426,1450]" italics="true" pageId="20" pageNumber="2612">–</emphasis>
|
||
0.83 mm
|
||
</quantity>
|
||
wide (EW), 2.0 times as long as wide (CL/EW).
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758701300FF4EACD4FE65F9A0" blockId="20.[160,1156,1356,1727]" pageId="20" pageNumber="2612">
|
||
Dorsal colouration (
|
||
<figureCitation id="15BF2A3258701300FE13ACD4FE09FA7F" box="[413,506,1461,1485]" captionStart="Figures 6–9" captionStartId="7.[160,235,1337,1360]" captionTargetBox="[163,1153,148,1289]" captionTargetId="figure@7.[163,1153,142,1306]" captionTargetPageId="7" captionText="Figures 6–9. Habitus of the Grouvellinus species collected from the Tawan Hills Park. 6: G. luciaensis, sp. nov. (paratype male from Lucia River); 7: G. borneensis, sp. nov. (paratype male from Tawau River environment); 8: G. sp.1 (female from Tawau River environment); 9: G. sp.2 (female from Lucia River). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3666544" httpUri="https://zenodo.org/record/3666544/files/figure.png" pageId="20" pageNumber="2612">Figure 7</figureCitation>
|
||
) almost entirely dark brown to black, only tarsi, antennae and maxillary palps golden brown; pubescence pale brown. Ventral side (
|
||
<figureCitation id="15BF2A3258701300FC6CACB6FBBEFA5D" box="[994,1101,1495,1519]" captionStart="Figure 16" captionStartId="21.[160,225,1038,1061]" captionTargetBox="[163,1153,142,1006]" captionTargetId="figure@21.[163,1153,142,1006]" captionTargetPageId="21" captionText="Figure 16. Grouvellinus borneensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum and elytra; (b) ventral aspect with prosternum and mesoventrite; (c) posteriodorsal aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666558" httpUri="https://zenodo.org/record/3666558/files/figure.png" pageId="20" pageNumber="2612">Figure 16</figureCitation>
|
||
(b,c)) brown to pale brown.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758701300FF4EAF7CFE41F90D" blockId="20.[160,1156,1356,1727]" pageId="20" pageNumber="2612">
|
||
Head (
|
||
<figureCitation id="15BF2A3258701300FE84AF7CFE85F987" box="[266,374,1565,1589]" captionStart="Figure 16" captionStartId="21.[160,225,1038,1061]" captionTargetBox="[163,1153,142,1006]" captionTargetId="figure@21.[163,1153,142,1006]" captionTargetPageId="21" captionText="Figure 16. Grouvellinus borneensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum and elytra; (b) ventral aspect with prosternum and mesoventrite; (c) posteriodorsal aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666558" httpUri="https://zenodo.org/record/3666558/files/figure.png" pageId="20" pageNumber="2612">Figure 16</figureCitation>
|
||
(a,b)) in males
|
||
<quantity id="4A7C9B5258701300FD94AF7CFD4FF987" box="[538,700,1564,1589]" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.5" metricValueMin="3.3" pageId="20" pageNumber="2612" unit="mm" value="0.33999999999999997" valueMax="0.35" valueMin="0.33">
|
||
0.33
|
||
<emphasis id="BFF0EAA558701300FDC7AF7DFDA5F986" box="[585,598,1564,1588]" italics="true" pageId="20" pageNumber="2612">–</emphasis>
|
||
0.35 mm
|
||
</quantity>
|
||
, in females
|
||
<quantity id="4A7C9B5258701300FCCCAF7CFC13F987" box="[834,992,1564,1589]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" metricValueMax="3.9" metricValueMin="3.7" pageId="20" pageNumber="2612" unit="mm" value="0.38" valueMax="0.39" valueMin="0.37">
|
||
0.37
|
||
<emphasis id="BFF0EAA558701300FCFEAF7DFC8EF986" box="[880,893,1564,1588]" italics="true" pageId="20" pageNumber="2612">–</emphasis>
|
||
0.39 mm
|
||
</quantity>
|
||
wide (HW); ID
|
||
<quantity id="4A7C9B5258701300FF2EAF5EFEB0F9E5" box="[160,323,1599,1623]" metricMagnitude="-4" metricUnit="m" metricValue="1.5500000000000003" metricValueMax="1.7000000000000002" metricValueMin="1.4000000000000001" pageId="20" pageNumber="2612" unit="mm" value="0.155" valueMax="0.17" valueMin="0.14">
|
||
0.14
|
||
<emphasis id="BFF0EAA558701300FF41AF5EFF2FF9E5" box="[207,220,1599,1623]" italics="true" pageId="20" pageNumber="2612">–</emphasis>
|
||
0.17 mm
|
||
</quantity>
|
||
; frons, clypeus, and labrum sparsely pubescent, slightly denser in lateral portions; punctures very small and scattered; intervals flat, glabrous. Frontoclypeal suture almost straight. Eyes large (eye diameter larger than ID), very slightly protruding. Antennae genus-typical.
|
||
</paragraph>
|
||
<caption id="D9FB663F58711301FF2EAD6FFE70FBED" ID-DOI="http://doi.org/10.5281/zenodo.3666558" ID-Zenodo-Dep="3666558" httpUri="https://zenodo.org/record/3666558/files/figure.png" pageId="21" pageNumber="2613" startId="21.[160,225,1038,1061]" targetBox="[163,1153,142,1006]" targetPageId="21">
|
||
<paragraph id="8D3B36B758711301FF2EAD6FFE70FBED" blockId="21.[160,1156,1037,1119]" pageId="21" pageNumber="2613">
|
||
<emphasis id="BFF0EAA558711301FF2EAD6FFEFBFB97" bold="true" box="[160,264,1038,1061]" pageId="21" pageNumber="2613">Figure 16.</emphasis>
|
||
<taxonomicName id="4A844D3458711301FE81AD6CFE03FB96" authority="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne, 2019" authorityName="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne" authorityYear="2019" box="[271,496,1037,1060]" class="Insecta" family="Elmidae" genus="Grouvellinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="2613" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="BFF0EAA558711301FE81AD6CFE03FB96" box="[271,496,1037,1060]" italics="true" pageId="21" pageNumber="2613">Grouvellinus borneensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A4C357DE58711301FE73AD6CFDBCFB97" box="[509,591,1037,1061]" pageId="21" pageNumber="2613" rank="species">
|
||
<emphasis id="BFF0EAA558711301FE73AD6CFDB9FB96" bold="true" box="[509,586,1037,1060]" pageId="21" pageNumber="2613">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(paratypes): (a) anteriodorsal aspect with pronotum and elytra; (b) ventral aspect with prosternum and mesoventrite; (c) posteriodorsal aspect with metaventrite and ventrites 1
|
||
<emphasis id="BFF0EAA558711301FEEAAD29FE82FBED" box="[356,369,1096,1119]" italics="true" pageId="21" pageNumber="2613">–</emphasis>
|
||
5.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8D3B36B758711301FF4EADFEFB87FA5D" blockId="21.[160,1156,1183,1727]" pageId="21" pageNumber="2613">
|
||
Pronotum (
|
||
<figureCitation id="15BF2A3258711301FEB3ADFEFE56FB05" box="[317,421,1183,1207]" captionStart="Figure 16" captionStartId="21.[160,225,1038,1061]" captionTargetBox="[163,1153,142,1006]" captionTargetId="figure@21.[163,1153,142,1006]" captionTargetPageId="21" captionText="Figure 16. Grouvellinus borneensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum and elytra; (b) ventral aspect with prosternum and mesoventrite; (c) posteriodorsal aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666558" httpUri="https://zenodo.org/record/3666558/files/figure.png" pageId="21" pageNumber="2613">Figure 16</figureCitation>
|
||
(a)) in males
|
||
<quantity id="4A7C9B5258711301FDB8ADFEFD20FB05" box="[566,723,1183,1207]" metricMagnitude="-4" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="21" pageNumber="2613" unit="mm" value="0.445" valueMax="0.45" valueMin="0.44">
|
||
0.44
|
||
<emphasis id="BFF0EAA558711301FDECADFEFD9CFB05" box="[610,623,1183,1207]" italics="true" pageId="21" pageNumber="2613">–</emphasis>
|
||
0.45 mm
|
||
</quantity>
|
||
, in females
|
||
<quantity id="4A7C9B5258711301FCD4ADFEFC06FB05" box="[858,1013,1183,1207]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="21" pageNumber="2613" unit="mm" value="0.48" valueMax="0.5" valueMin="0.46">
|
||
0.46
|
||
<emphasis id="BFF0EAA558711301FC08ADFEFC60FB05" box="[902,915,1183,1207]" italics="true" pageId="21" pageNumber="2613">–</emphasis>
|
||
0.50 mm
|
||
</quantity>
|
||
long (PL), in males
|
||
<quantity id="4A7C9B5258711301FF69ADA3FE70FB68" box="[231,387,1218,1242]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.6" metricValueMin="5.2" pageId="21" pageNumber="2613" unit="mm" value="0.54" valueMax="0.56" valueMin="0.52">
|
||
0.52
|
||
<emphasis id="BFF0EAA558711301FE9DADA3FED3FB68" box="[275,288,1218,1242]" italics="true" pageId="21" pageNumber="2613">–</emphasis>
|
||
0.56 mm
|
||
</quantity>
|
||
, in females
|
||
<quantity id="4A7C9B5258711301FD88ADA3FD53FB68" box="[518,672,1218,1242]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" metricValueMax="6.3" metricValueMin="5.8999999999999995" pageId="21" pageNumber="2613" unit="mm" value="0.61" valueMax="0.63" valueMin="0.59">
|
||
0.59
|
||
<emphasis id="BFF0EAA558711301FDBCADA3FDCCFB68" box="[562,575,1218,1242]" italics="true" pageId="21" pageNumber="2613">–</emphasis>
|
||
0.63 mm
|
||
</quantity>
|
||
wide (PW), wider than long (PL/PW), widest basally, narrower than elytra, anteriorly attenuate; anterior margin moderately convex; pronotal disc slightly vaulted; entire pronotum moderately densely punctate; punctures small and shallowly impressed; setae moderately short; median carina absent; pair of posteriormedian rugose patches distinct; sublateral carinae indistinct and short; laterobasal impression shallow, rugulose; oblique impressions shallow, partly rugulose, extending obliquely from anterior 0.2
|
||
<emphasis id="BFF0EAA558711301FEAEACF3FEDEFA18" box="[288,301,1426,1450]" italics="true" pageId="21" pageNumber="2613">–</emphasis>
|
||
0.6, then continuing aside sublateral carinae up to posterior margin; entire anterior portion and portion between sublateral carinae and posterior-median patches glabrous; disc anterior of these patches slightly rugose. Hypomeron very densely punctate.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758711301FF4EAC9BFDF9F9E5" blockId="21.[160,1156,1183,1727]" pageId="21" pageNumber="2613">
|
||
Prosternum (
|
||
<figureCitation id="15BF2A3258711301FEDAAC9BFE33F9A0" box="[340,448,1530,1554]" captionStart="Figure 16" captionStartId="21.[160,225,1038,1061]" captionTargetBox="[163,1153,142,1006]" captionTargetId="figure@21.[163,1153,142,1006]" captionTargetPageId="21" captionText="Figure 16. Grouvellinus borneensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum and elytra; (b) ventral aspect with prosternum and mesoventrite; (c) posteriodorsal aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666558" httpUri="https://zenodo.org/record/3666558/files/figure.png" pageId="21" pageNumber="2613">Figure 16</figureCitation>
|
||
(c)) short; lateral portions with inconspicuous plastron pubescence; median portion including process very shallowly impressed, slightly rugulose; prosternal process subquadrate.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758711303FF4EAF03FE93FEBC" blockId="21.[160,1156,1183,1727]" lastBlockId="23.[160,1156,142,1414]" lastPageId="23" lastPageNumber="2615" pageId="21" pageNumber="2613">
|
||
Scutellum (
|
||
<figureCitation id="15BF2A3258711301FECAAF03FE40F9C8" box="[324,435,1634,1658]" captionStart="Figure 16" captionStartId="21.[160,225,1038,1061]" captionTargetBox="[163,1153,142,1006]" captionTargetId="figure@21.[163,1153,142,1006]" captionTargetPageId="21" captionText="Figure 16. Grouvellinus borneensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum and elytra; (b) ventral aspect with prosternum and mesoventrite; (c) posteriodorsal aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666558" httpUri="https://zenodo.org/record/3666558/files/figure.png" pageId="21" pageNumber="2613">Figure 16</figureCitation>
|
||
(a)) subcordiform, flat, rugose, with few setiferous punctures. Elytra (
|
||
<figureCitation id="15BF2A3258711301FF62AFE4FEBBF92F" box="[236,328,1669,1693]" captionStart="Figures 6–9" captionStartId="7.[160,235,1337,1360]" captionTargetBox="[163,1153,148,1289]" captionTargetId="figure@7.[163,1153,142,1306]" captionTargetPageId="7" captionText="Figures 6–9. Habitus of the Grouvellinus species collected from the Tawan Hills Park. 6: G. luciaensis, sp. nov. (paratype male from Lucia River); 7: G. borneensis, sp. nov. (paratype male from Tawau River environment); 8: G. sp.1 (female from Tawau River environment); 9: G. sp.2 (female from Lucia River). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3666544" httpUri="https://zenodo.org/record/3666544/files/figure.png" pageId="21" pageNumber="2613">Figure 7</figureCitation>
|
||
) elongate, moderately convex dorsally, in males
|
||
<quantity id="4A7C9B5258711301FCE6AFE4FBFBF92F" box="[872,1032,1668,1693]" metricMagnitude="-3" metricUnit="m" metricValue="1.085" metricValueMax="1.1" metricValueMin="1.07" pageId="21" pageNumber="2613" unit="mm" value="1.085" valueMax="1.1" valueMin="1.07">
|
||
1.07
|
||
<emphasis id="BFF0EAA558711301FC18AFE5FC50F92E" box="[918,931,1668,1692]" italics="true" pageId="21" pageNumber="2613">–</emphasis>
|
||
1.10 mm
|
||
</quantity>
|
||
, in females
|
||
<quantity id="4A7C9B5258711301FF2EAFC6FECFF90D" box="[160,316,1703,1727]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" metricValueMax="1.24" metricValueMin="1.22" pageId="21" pageNumber="2613" unit="mm" value="1.23" valueMax="1.24" valueMin="1.22">
|
||
1.22
|
||
<emphasis id="BFF0EAA558711301FF41AFC6FF2FF90D" box="[207,220,1703,1727]" italics="true" pageId="21" pageNumber="2613">–</emphasis>
|
||
1.24 mm
|
||
</quantity>
|
||
long (EL), in males ca. 1.4 times, in females ca. 1.3 times as long as wide (EL/ EW), conspicuously widest at anterior 0.6, slightly attenuating anteriad, roundly attenuating posteriad; apices wide, almost conjointly rounded; with eight longitudinal, slightly impressed rows of primary punctures (striae); primary punctures large and moderately deeply impressed at median disc, slightly smaller in basal portion and increasingly smaller and more shallowly impressed apicad, regularly arranged in median rows, less regularly arranged in lateral rows; interstices and interval rugose; interval 8 with (genus-typical) serrate carina; interval 2 slightly carinate; all intervals with setiferous secondary punctures (inconspicuous due to rugose surface); pubescence yellowish, consisting of moderately large, erected setae and small adpressed setae; lateral elytral margin more or less distinctly serrate.
|
||
</paragraph>
|
||
<caption id="D9FB663F58721302FF2EAC3EFD92FA21" ID-DOI="http://doi.org/10.5281/zenodo.3666560" ID-Zenodo-Dep="3666560" httpUri="https://zenodo.org/record/3666560/files/figure.png" pageId="22" pageNumber="2614" startId="22.[160,225,1375,1398]" targetBox="[163,1153,142,1343]" targetPageId="22">
|
||
<paragraph id="8D3B36B758721302FF2EAC3EFD92FA21" blockId="22.[160,1155,1374,1427]" pageId="22" pageNumber="2614">
|
||
<emphasis id="BFF0EAA558721302FF2EAC3EFEF5FAC4" bold="true" box="[160,262,1375,1398]" pageId="22" pageNumber="2614">Figure 17.</emphasis>
|
||
<taxonomicName id="4A844D3458721302FE85AC3FFE18FAC7" authority="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne, 2019" authorityName="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne" authorityYear="2019" box="[267,491,1374,1397]" class="Insecta" family="Elmidae" genus="Grouvellinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="2614" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="BFF0EAA558721302FE85AC3FFE18FAC7" box="[267,491,1374,1397]" italics="true" pageId="22" pageNumber="2614">Grouvellinus borneensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A4C357DE58721302FE7BAC3EFDB5FAC4" box="[501,582,1375,1398]" pageId="22" pageNumber="2614" rank="species">
|
||
<emphasis id="BFF0EAA558721302FE7BAC3EFDB2FAC4" bold="true" box="[501,577,1375,1398]" pageId="22" pageNumber="2614">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(paratypes): (a) aedeagus, lateral; (b) aedeagus, ventral; (c) ovipositor (half), ventral. Scale bar = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8D3B36B758731303FF4EA878FDB0FE0E" blockId="23.[160,1156,142,1414]" pageId="23" pageNumber="2615">
|
||
Mesoventrite (
|
||
<figureCitation id="15BF2A3258731303FED3A878FE30FE83" box="[349,451,281,305]" captionStart="Figure 16" captionStartId="21.[160,225,1038,1061]" captionTargetBox="[163,1153,142,1006]" captionTargetId="figure@21.[163,1153,142,1006]" captionTargetPageId="21" captionText="Figure 16. Grouvellinus borneensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum and elytra; (b) ventral aspect with prosternum and mesoventrite; (c) posteriodorsal aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666558" httpUri="https://zenodo.org/record/3666558/files/figure.png" pageId="23" pageNumber="2615">Figure 16</figureCitation>
|
||
(c)) with 2 pairs of subtrapezoidal grooves, median pair shallowly impressed, lateral pair (behind procoxae) deeply impressed. Metaventrite (
|
||
<figureCitation id="15BF2A3258731303FC7DA85DFB93FEE6" box="[1011,1120,316,340]" captionStart="Figure 16" captionStartId="21.[160,225,1038,1061]" captionTargetBox="[163,1153,142,1006]" captionTargetId="figure@21.[163,1153,142,1006]" captionTargetPageId="21" captionText="Figure 16. Grouvellinus borneensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum and elytra; (b) ventral aspect with prosternum and mesoventrite; (c) posteriodorsal aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666558" httpUri="https://zenodo.org/record/3666558/files/figure.png" pageId="23" pageNumber="2615">Figure 16</figureCitation>
|
||
(d)) with distinct median suture over the entire length. Median disc glabrous, slightly dimorphic between sexes; lateral and marginal portions moderately densely covered with setiferous tubercles and inconspicuous plastron.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758731303FF4EA8A7FB9CFDF4" blockId="23.[160,1156,142,1414]" pageId="23" pageNumber="2615">
|
||
Ventrite 1 (
|
||
<figureCitation id="15BF2A3258731303FEB5A8A7FE57FE6C" box="[315,420,454,478]" captionStart="Figure 16" captionStartId="21.[160,225,1038,1061]" captionTargetBox="[163,1153,142,1006]" captionTargetId="figure@21.[163,1153,142,1006]" captionTargetPageId="21" captionText="Figure 16. Grouvellinus borneensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum and elytra; (b) ventral aspect with prosternum and mesoventrite; (c) posteriodorsal aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666558" httpUri="https://zenodo.org/record/3666558/files/figure.png" pageId="23" pageNumber="2615">Figure 16</figureCitation>
|
||
(d)) comparably short; carinae bordering glabrous disc oblique and rather inconspicuous; ventrites 1
|
||
<emphasis id="BFF0EAA558731303FD83A888FDE9FDB3" box="[525,538,489,513]" italics="true" pageId="23" pageNumber="2615">–</emphasis>
|
||
2 only glabrous in most anteriomedian portion; lateral portions of ventrites 1
|
||
<emphasis id="BFF0EAA558731303FE05AB6AFE6BFD91" box="[395,408,523,547]" italics="true" pageId="23" pageNumber="2615">–</emphasis>
|
||
2 and entire ventrites 4
|
||
<emphasis id="BFF0EAA558731303FD1EAB6AFD6EFD91" box="[656,669,523,547]" italics="true" pageId="23" pageNumber="2615">–</emphasis>
|
||
5 moderately densely covered with setiferous tubercles and inconspicuous plastron; apical margin of ventrite 5 fringed with pubescence.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758731303FF4EAB30FC5CFD46" blockId="23.[160,1156,142,1414]" pageId="23" pageNumber="2615">
|
||
Legs (
|
||
<figureCitation id="15BF2A3258731303FE8CAB30FEADFDDB" box="[258,350,593,617]" captionStart="Figures 6–9" captionStartId="7.[160,235,1337,1360]" captionTargetBox="[163,1153,148,1289]" captionTargetId="figure@7.[163,1153,142,1306]" captionTargetPageId="7" captionText="Figures 6–9. Habitus of the Grouvellinus species collected from the Tawan Hills Park. 6: G. luciaensis, sp. nov. (paratype male from Lucia River); 7: G. borneensis, sp. nov. (paratype male from Tawau River environment); 8: G. sp.1 (female from Tawau River environment); 9: G. sp.2 (female from Lucia River). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3666544" httpUri="https://zenodo.org/record/3666544/files/figure.png" pageId="23" pageNumber="2615">Figure 7</figureCitation>
|
||
) approximately of the same length as body; tibiae longer than tarsi and femora; trochanter, femura and tibiae almost entirely densely covered with setiferous tubercles and plastron setae except almost glabrous outer, dorsal face of tibiae and inner face of tibiae and femora. Mesofemur with loose fringe of short trichoid setae on the anterior
|
||
<emphasis id="BFF0EAA558731303FF76ABBAFEF6FD41" box="[248,261,731,755]" italics="true" pageId="23" pageNumber="2615">–</emphasis>
|
||
dorsal ridge. Legs not conspicuously varying between sexes.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758731303FF4EAB9FFF04FB6B" blockId="23.[160,1156,142,1414]" pageId="23" pageNumber="2615">
|
||
Aedeagus (
|
||
<figureCitation id="15BF2A3258731303FECAAB9FFE46FCA4" box="[324,437,766,790]" captionStart="Figure 17" captionStartId="22.[160,225,1375,1398]" captionTargetBox="[163,1153,142,1343]" captionTargetId="figure@22.[163,1153,142,1343]" captionTargetPageId="22" captionText="Figure 17. Grouvellinus borneensis, sp. nov. (paratypes): (a) aedeagus, lateral; (b) aedeagus, ventral; (c) ovipositor (half), ventral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3666560" httpUri="https://zenodo.org/record/3666560/files/figure.png" pageId="23" pageNumber="2615">Figure 17</figureCitation>
|
||
(a,b)) ca. 510 μm long, ca. 115 μm wide. Phallobase slightly asymmetrical basally, reaching ca. basal 0.31 of total aedeagus length. Median lobe moderately wide (ventral view), moderately overreaching parameres and ventral sac, very slightly conical in apical half, then abruptly tapered at the level of paramere tips towards slender apex; apex slightly bent ventrad (lateral view). Ventral sac internally densely stippled and with dense fringe of moderately short, thin and comparably delicate spines; few of those overreaching apical fringe. Parameres moderately shorter than median lobe and distinctly bent ventrad (lateral view), moderately broad and evenly conical from insertion to apex in lateral view, usually with more than 15 trichoid setae in apical half, most of them at the inner face; most apical setae of subequal length. Male sternite IX with median strut moderately long and almost rectangularly bent subdistally; posterior portion entirely fringed with a broad, distinctly sclerotised margin; posterior margin rounded; paraprocts subequal in length, not reaching apical margin.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758731303FF4EAD85FE2DFA34" blockId="23.[160,1156,142,1414]" pageId="23" pageNumber="2615">
|
||
Ovipositor (
|
||
<figureCitation id="15BF2A3258731303FECAAD85FE5CFB4E" box="[324,431,1252,1276]" captionStart="Figure 17" captionStartId="22.[160,225,1375,1398]" captionTargetBox="[163,1153,142,1343]" captionTargetId="figure@22.[163,1153,142,1343]" captionTargetPageId="22" captionText="Figure 17. Grouvellinus borneensis, sp. nov. (paratypes): (a) aedeagus, lateral; (b) aedeagus, ventral; (c) ovipositor (half), ventral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3666560" httpUri="https://zenodo.org/record/3666560/files/figure.png" pageId="23" pageNumber="2615">Figure 17</figureCitation>
|
||
(c)). Total length ca. 560 μm. Stylus ca. 15 μm long, very slightly bent outwards, with ca. four short sensilla. Coxite ca. 240 μm long, apically moderately broadened with scattered extremely short, acute setae and four apical sensilla. Valvifer ca. 325 μm long; caudal portion slightly sclerotised and with scattered, extremely short, acute setae; fibula almost straight.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758731303FF2EACA2FE77FA6E" blockId="23.[160,1156,1475,1673]" box="[160,388,1475,1500]" pageId="23" pageNumber="2615">
|
||
<emphasis id="BFF0EAA558731303FF2EACA2FE77FA6E" bold="true" box="[160,388,1475,1500]" italics="true" pageId="23" pageNumber="2615">Sexual dimorphism</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758731303FF2EAC87FDF9F93B" blockId="23.[160,1156,1475,1673]" pageId="23" pageNumber="2615">
|
||
Females with distinct rugulose elongate elytral patches extending from elytral basis approximately to apical 0.35 and between intervals 1 and 3 (comp.
|
||
<figureCitation id="15BF2A3258731303FC58AF68FBBBF993" box="[982,1096,1545,1569]" captionStart="Figure 16" captionStartId="21.[160,225,1038,1061]" captionTargetBox="[163,1153,142,1006]" captionTargetId="figure@21.[163,1153,142,1006]" captionTargetPageId="21" captionText="Figure 16. Grouvellinus borneensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum and elytra; (b) ventral aspect with prosternum and mesoventrite; (c) posteriodorsal aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666558" httpUri="https://zenodo.org/record/3666558/files/figure.png" pageId="23" pageNumber="2615">Figure 16</figureCitation>
|
||
(a,b)). Glabrous portions of metaventral disc more extended in males and entirely very broadly impressed along median suture, while rather narrow and only slightly impressed in posterior half and subanteriorly.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B7587C130CFF2EA9EFFE6EFF15" blockId="24.[160,1156,142,618]" box="[160,413,142,167]" pageId="24" pageNumber="2616">
|
||
<emphasis id="BFF0EAA5587C130CFF2EA9EFFE6EFF15" bold="true" box="[160,413,142,167]" italics="true" pageId="24" pageNumber="2616">Differential diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8D3B36B7587C130CFF2EA9D0FE19FD96" blockId="24.[160,1156,142,618]" pageId="24" pageNumber="2616">
|
||
<taxonomicName id="4A844D34587C130CFF2EA9D0FE55FF78" authority="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne, 2019" authorityName="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne" authorityYear="2019" box="[160,422,177,202]" class="Insecta" family="Elmidae" genus="Grouvellinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="2616" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="BFF0EAA5587C130CFF2EA9D0FE55FF78" box="[160,422,177,202]" italics="true" pageId="24" pageNumber="2616">Grouvellinus borneensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A4C357DE587C130CFE21A9D3FDFAFF78" box="[431,521,178,202]" pageId="24" pageNumber="2616" rank="species">sp. nov.</taxonomicNameLabel>
|
||
superficially resembles
|
||
<taxonomicName id="4A844D34587C130CFC90A9D0FC89FF78" box="[798,890,177,202]" class="Insecta" family="Elmidae" genus="Grouvellinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="2616" phylum="Arthropoda" rank="species" species="quest">
|
||
<emphasis id="BFF0EAA5587C130CFC90A9D0FC89FF78" box="[798,890,177,202]" italics="true" pageId="24" pageNumber="2616">G. quest</emphasis>
|
||
</taxonomicName>
|
||
in its size, proportions and entirely black colour, but can be distinguished from the latter by the rather smooth, moderately densely punctate pronotum (vs. very densely punctate), the basally smoother elytral surface with only intervals 2 and 8 partly carinate (vs. intervals 3, 5, 7 and 8 crenulate-carinate). From all known Bornean
|
||
<taxonomicName id="4A844D34587C130CFC3CA85AFBC9FEE6" authorityName="Champion" authorityYear="1923" box="[946,1082,315,340]" class="Insecta" family="Elmidae" genus="Grouvellinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="2616" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BFF0EAA5587C130CFC3CA85AFBC9FEE6" box="[946,1082,315,340]" italics="true" pageId="24" pageNumber="2616">Grouvellinus</emphasis>
|
||
</taxonomicName>
|
||
spp.,
|
||
<taxonomicName id="4A844D34587C130CFF2EA83FFEC7FEC5" authority="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne, 2019" authorityName="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne" authorityYear="2019" box="[160,308,350,375]" class="Insecta" family="Elmidae" genus="Grouvellinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="2616" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="BFF0EAA5587C130CFF2EA83FFEC7FEC5" box="[160,308,350,375]" italics="true" pageId="24" pageNumber="2616">G. borneensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A4C357DE587C130CFEB3A83EFE65FEC5" box="[317,406,351,375]" pageId="24" pageNumber="2616" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs by the large eyes (eye diameter larger than ID vs. smaller than ID), the sexual dimorphism in the mediobasal elytra (but presumably also present in the still unnamed species
|
||
<emphasis id="BFF0EAA5587C130CFD8BA8C2FDE5FE0E" box="[517,534,419,444]" italics="true" pageId="24" pageNumber="2616">G</emphasis>
|
||
. sp.1.). Furthermore, it is easily recognisable by its aedeagus with median lobe slightly conical in apical half, then abruptly tapered towards the slender apex, the short aedeagal base and the long parameres with about 15 equally long setae.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B7587C130CFF4EAB4EFE26FDD8" blockId="24.[160,1156,142,618]" pageId="24" pageNumber="2616">
|
||
The species varies by about 15
|
||
<emphasis id="BFF0EAA5587C130CFD95AB4EFDDBFDF5" box="[539,552,559,583]" italics="true" pageId="24" pageNumber="2616">–</emphasis>
|
||
18% genetic distance (654 bp CO1 barcode) from the known Bornean congeners.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C59E653C587C130CFF2EABFDFB83FD6A" box="[160,1136,668,728]" pageId="24" pageNumber="2616" type="distribution">
|
||
<paragraph id="8D3B36B7587C130CFF2EABFDFEDFFD07" blockId="24.[160,1136,668,728]" box="[160,300,668,693]" pageId="24" pageNumber="2616">
|
||
<emphasis id="BFF0EAA5587C130CFF2EABFDFEDFFD07" bold="true" box="[160,300,668,693]" italics="true" pageId="24" pageNumber="2616">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8D3B36B7587C130CFF2EABA1FB83FD6A" blockId="24.[160,1136,668,728]" box="[160,1136,704,728]" pageId="24" pageNumber="2616">
|
||
This species is only known from the Tawau Hills Park, Sabah, Borneo Island (
|
||
<figureCitation id="15BF2A32587C130CFC77ABA1FB90FD6A" box="[1017,1123,704,728]" captionStart="Figure 18" captionStartId="25.[160,225,1675,1698]" captionTargetBox="[163,1153,977,1643]" captionTargetId="figure@25.[163,1153,977,1643]" captionTargetPageId="25" captionText="Figure 18. Map of Sabah and Brunei (Borneo) with collecting site of Grouvellinus species with new species record in red fonts." figureDoi="http://doi.org/10.5281/zenodo.3666562" httpUri="https://zenodo.org/record/3666562/files/figure.png" pageId="24" pageNumber="2616">Figure 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |