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<document id="146F3326354004B39A3BE43A81E4786C" ID-DOI="10.1163/18759866-20191416" ID-ISSN="1875-9866" ID-Zenodo-Dep="8343061" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1694662606570" checkinUser="felipe" docAuthor="Hyman, Isabel T. &amp; Köhler, Frank" docDate="2019" docId="03F32A40FF96B15251BC9801F637FE3A" docLanguage="en" docName="ContribZool.88.4.351-451.pdf" docOrigin="Contributions to Zoology 88 (4)" docSource="http://dx.doi.org/10.1163/18759866-20191416" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.2:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="2" docTitle="Fastosarion minerva" docType="treatment" docVersion="5" lastPageNumber="432" masterDocId="FFCA5238FFD9B100533A9C10F477FFFB" masterDocTitle="Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)" masterLastPageNumber="451" masterPageNumber="351" pageNumber="430" updateTime="1695298548213" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F40F590EFE6A0F01CAEC079B159C034C">Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)</mods:title>
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<taxonomicName id="4C5AE0D5FF96B14F51BC9801F055FBD0" ID-CoL="7RBRP" ID-ENA="2658031" authority="(Stanisic, 2010)" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[646,1058,1041,1067]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="minerva">
<emphasis id="B92E4744FF96B14F51BC9801F055FBD0" bold="true" box="[646,1058,1041,1067]" pageId="79" pageNumber="430">
<emphasis id="B92E4744FF96B14F51BC9801F708FBD0" bold="true" box="[646,895,1041,1067]" italics="true" pageId="79" pageNumber="430">Fastosarion minerva (</emphasis>
<bibRefCitation id="EFCBE6A7FF96B14F50BA9801F06EFBD0" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." box="[896,1049,1041,1067]" pageId="79" pageNumber="430" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic, 2010</bibRefCitation>
<emphasis id="B92E4744FF96B14F57229801F055FBD0" bold="true" box="[1048,1058,1041,1067]" italics="true" pageId="79" pageNumber="430">)</emphasis>
</emphasis>
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</paragraph>
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<paragraph id="8BE59B56FF96B14F51BC9825F7C3FBB4" blockId="79.[646,1156,1077,1175]" box="[646,948,1077,1104]" pageId="79" pageNumber="430">
<figureCitation id="136187D3FF96B14F51BC9825F6A1FBB4" box="[646,726,1077,1103]" captionStart="FIGURE 18" captionStartId="66.[129,202,1582,1603]" captionTargetBox="[129,1187,188,1550]" captionTargetId="figure-8@66.[129,1187,188,1550]" captionTargetPageId="66" captionText="FIGURE 18 Distribution of Clade B species. █ = F.alyssa, ▼ = F.deensis, × = F.ephelis, ♢ = F.hannianus, ▲ = F. insularis, ◻ = F.katatonos, ∆ = F.longimentula, ● = F.mcdonaldi, ♠ = F.minerva, ◆ = F.papillosus, ♣ = F.pustulosus, ♥ = F.rowani, Ǫ = F.sarina, ▽ = F.tuljun." figureDoi="http://doi.org/10.5281/zenodo.8343097" httpUri="https://zenodo.org/record/8343097/files/figure.png" pageId="79" pageNumber="430">
Figs.
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<figureCitation id="136187D3FF96B14F51DA9825F755FBAB" box="[736,802,1077,1104]" captionStart="FIGURE 15" captionStartId="60.[129,202,1418,1439]" captionTargetBox="[139,1187,188,1385]" captionTargetId="figure-123@60.[129,1187,188,1385]" captionTargetPageId="60" captionText="FIGURE 15 Genitalia of F.griseolus, MO55568, Bobby Range.A.Reproductive system. Arrow shows the junction of the penis and epiphallus.B. Penis interior. Scale bars 1 mm (B), 2 mm (A)." figureDoi="http://doi.org/10.5281/zenodo.8343091" httpUri="https://zenodo.org/record/8343091/files/figure.png" pageId="79" pageNumber="430">15EF</figureCitation>
,
<figureCitation id="136187D3FF96B14F50169825F73BFBB4" box="[812,844,1077,1103]" captionStart="FIGURE 19" captionStartId="67.[98,171,1480,1501]" captionTargetBox="[98,1104,171,1448]" captionTargetId="figure-115@67.[98,1104,170,1448]" captionTargetPageId="67" captionText="FIGURE 19 Shells of Clade B species. A. Fastosarion alyssa (type locality of D.peterbrocki), NMV F230748. B. Fastosarion deensis, MO85834 (holotype). C. Fastosarion ephelis, MO85835 (holotype). D. Fastosarion hannianus, MO56684.E. Fastosarion insularis sp. nov., MO76354. F. Fastosarion katatonos, MO85837 (holotype). G.Fastosarion longimentula, MO65313.H. Fastosarion mcdonaldi, MO50434.I. Fastosarion minerva, MO78358. J.Fastosarion papillosus, MO78896 (Mt Superbus).K.Fastosarion papillosus, MO62397 (Kroombit Tops). L.Fastosarion pustulosus, MO78894 (Kalpowar SF).M. Fastosarion rowani, MO85839 (holotype). N. Fastosarion sarina, MO85840 (holotype).O. Fastosarion tujun (Denham Range), MO76081 (holotype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8343099" httpUri="https://zenodo.org/record/8343099/files/figure.png" pageId="79" pageNumber="430">19I</figureCitation>
,
<figureCitation id="136187D3FF96B14F506C9825F7E7FBB4" box="[854,912,1077,1103]" captionStart="FIGURE 22" captionStartId="71.[98,171,1710,1731]" captionTargetBox="[98,887,184,1677]" captionTargetId="figure-39@71.[98,887,184,1677]" captionTargetPageId="71" captionText="FIGURE 22 Spermatophores of Clade B species. A-B. F.alyssa, MO77158.C-D.F.deensis, MO85834.E-F. F.hannianus, MO72277.G-H.F.mcdonaldi, MO50434.I-J. F.minerva, MO78358." figureDoi="http://doi.org/10.5281/zenodo.8343105" httpUri="https://zenodo.org/record/8343105/files/figure.png" pageId="79" pageNumber="430">22IJ</figureCitation>
,
<figureCitation id="136187D3FF96B14F50A09825F7C3FBB4" box="[922,948,1077,1103]" captionStart="FIGURE 29" captionStartId="81.[98,171,1163,1184]" captionTargetBox="[99,1156,188,1130]" captionTargetId="figure-217@81.[98,1156,188,1130]" captionTargetPageId="81" captionText="FIGURE 29 Genitalia of F.minerva, MO78358, Carnarvon NP.A.Reproductive system.Arrow shows the junction of the penis and epiphallus.B. Penis interior. C.Spermatophore. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8343119" httpUri="https://zenodo.org/record/8343119/files/figure.png" pageId="79" pageNumber="430">29</figureCitation>
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<taxonomicName id="4C5AE0D5FF96B14F519F9849F054FB88" ID-CoL="7R33V" authority="Stanisic, 2010" authorityName="Stanisic" authorityYear="2010" box="[677,1059,1113,1139]" class="Gastropoda" family="Helicarionidae" genus="Dimidarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="minerva">
<emphasis id="B92E4744FF96B14F519F9849F7F1FB88" box="[677,902,1113,1139]" italics="true" pageId="79" pageNumber="430">Dimidarion minerva</emphasis>
<bibRefCitation id="EFCBE6A7FF96B14F50B49849F054FB88" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." box="[910,1059,1113,1139]" pageId="79" pageNumber="430" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">
Stanisic,
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in
<treatmentCitation id="0AFBBD47FF96B14F57709849F7F3FB6C" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." page="316" pageId="79" pageNumber="430" year="2010">
<bibRefCitation id="EFCBE6A7FF96B14F57709849F7F3FB6C" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." pageId="79" pageNumber="430" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al., 2010: 316317</bibRefCitation>
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.
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<paragraph id="8BE59B56FF96B14F51BA98D9F723FB18" blockId="79.[640,1157,1225,1538]" box="[640,852,1225,1251]" pageId="79" pageNumber="430">Material examined</paragraph>
<paragraph id="8BE59B56FF96B14F51BC98FEF6AEFA6C" blockId="79.[640,1157,1225,1538]" pageId="79" pageNumber="430">
<emphasis id="B92E4744FF96B14F51BC98FEF6B2FAFC" box="[646,709,1262,1287]" italics="true" pageId="79" pageNumber="430">Types</emphasis>
:
<materialsCitation id="3B32910BFF96B14F51EC98FDF6AEFA6C" collectingDate="1995-09-20" collectionCode="QM" collectorName="Springsure, NW &amp; J. Stanisic &amp; D. Potter &amp; C. Eddie" latitude="-24.098612" location="Mt Zamia" longLatPrecision="21" longitude="148.01527" pageId="79" pageNumber="430" specimenCode="MO56626" specimenCount="1" typeStatus="holotype">
<typeStatus id="54E125F4FF96B14F51EC98FDF736FAFC" box="[726,833,1261,1287]" pageId="79" pageNumber="430">Holotype</typeStatus>
:
<collectionCode id="ED4B0393FF96B14F507598FDF70DFAFC" box="[847,890,1261,1287]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="79" pageNumber="430" type="Museum">QM</collectionCode>
<specimenCode id="DBFC332DFF96B14F50BF98FDF786FAF3" box="[901,1009,1261,1288]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="79" pageNumber="430" type="Museum">MO56626</specimenCode>
(
<collectorName id="26AFFE80FF96B14F573E98FDF6C6FAD0" pageId="79" pageNumber="430">Springsure, NW</collectorName>
at
<location id="8E85CD8DFF96B14F51DB9901F72FFAD0" LSID="urn:lsid:plazi:treatment:03F32A40FF96B15251BC9801F637FE3A:8E85CD8DFF96B14F51DB9901F72FFAD0" box="[737,856,1297,1323]" latitude="-24.098612" longLatPrecision="21" longitude="148.01527" name="Mt Zamia" pageId="79" pageNumber="430">Mt Zamia</location>
,
<taxonomicName id="4C5AE0D5FF96B14F50539901F0F6FAD0" authority="Hills NP, MEQ" authorityName="Hills NP, MEQ" box="[873,1153,1297,1323]" class="Gastropoda" family="Helicarionidae" genus="Dimidarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="minerva">Minerva Hills NP, MEQ</taxonomicName>
,
<geoCoordinate id="EE6EFD91FF96B14F51BC9924F777FAB4" box="[646,768,1332,1359]" degrees="24" direction="south" minutes="05" orientation="latitude" pageId="79" pageNumber="430" precision="15" seconds="55" value="-24.098612">24°0555” S</geoCoordinate>
,
<geoCoordinate id="EE6EFD91FF96B14F50309924F7E7FAB4" box="[778,912,1332,1359]" degrees="148" direction="east" minutes="00" orientation="longitude" pageId="79" pageNumber="430" precision="15" seconds="55" value="148.01527">148°0055” E</geoCoordinate>
, sevt, under rocks and logs, coll.
<date id="FFE4BD96FF96B14F51CF9949F70EFA88" box="[757,889,1368,1395]" pageId="79" pageNumber="430" value="1995-09-20">
<collectingDate id="EFA0447EFF96B14F51CF9949F70EFA88" box="[757,889,1368,1395]" pageId="79" pageNumber="430" value="1995-09-20">20 Sep 1995</collectingDate>
</date>
,
<collectorName id="26AFFE80FF96B14F50BE9949F785FA89" box="[900,1010,1368,1395]" pageId="79" pageNumber="430">J. Stanisic</collectorName>
,
<collectorName id="26AFFE80FF96B14F50C49949F016FA88" box="[1022,1121,1369,1395]" pageId="79" pageNumber="430">D. Potter</collectorName>
,
<collectorName id="26AFFE80FF96B14F57579949F6BAFA6C" pageId="79" pageNumber="430">C. Eddie</collectorName>
).
</materialsCitation>
</paragraph>
<paragraph id="8BE59B56FF96B14F519F99B1F752FA25" blockId="79.[640,1157,1225,1538]" pageId="79" pageNumber="430">
<materialsCitation id="3B32910BFF96B14F519F99B1F7BDFA41" box="[677,970,1440,1467]" collectionCode="QM" pageId="79" pageNumber="430" specimenCode="MO76088" specimenCount="1" typeStatus="paratype">
<typeStatus id="54E125F4FF96B14F519F99B1F763FA40" box="[677,788,1441,1467]" pageId="79" pageNumber="430">Paratypes</typeStatus>
:
<collectionCode id="ED4B0393FF96B14F501E99B0F738FA41" box="[804,847,1440,1466]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="79" pageNumber="430" type="Museum">QM</collectionCode>
<specimenCode id="DBFC332DFF96B14F506699B0F7BDFA41" box="[860,970,1440,1466]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="79" pageNumber="430" type="Museum">MO76088</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B32910BFF96B14F50E199B0F0F7FA41" box="[987,1152,1440,1466]" collectionCode="QM" pageId="79" pageNumber="430" specimenCode="MO76366" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED4B0393FF96B14F50E199B0F071FA41" box="[987,1030,1440,1466]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="79" pageNumber="430" type="Museum">QM</collectionCode>
<specimenCode id="DBFC332DFF96B14F572899B0F0F7FA41" box="[1042,1152,1440,1466]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="79" pageNumber="430" type="Museum">MO76366</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B32910BFF96B14F51BC99D4F756FA25" box="[646,801,1476,1502]" collectionCode="QM" pageId="79" pageNumber="430" specimenCode="MO78901" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED4B0393FF96B14F51BC99D4F6C6FA25" box="[646,689,1476,1502]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="79" pageNumber="430" type="Museum">QM</collectionCode>
<specimenCode id="DBFC332DFF96B14F518D99D4F756FA25" box="[695,801,1476,1502]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="79" pageNumber="430" type="Museum">MO78901</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE59B56FF96B14F519F99F9F798F9F9" blockId="79.[640,1157,1225,1538]" box="[677,1007,1512,1538]" pageId="79" pageNumber="430">
<emphasis id="B92E4744FF96B14F519F99F9F707F9F9" box="[677,880,1512,1538]" italics="true" pageId="79" pageNumber="430">Non-type material:</emphasis>
See
<tableCitation id="C6D8AEEDFF96B14F509899F8F79CF9F9" box="[930,1003,1512,1538]" captionStart="TABLE 1" captionStartId="3.[56,116,402,428]" captionTargetBox="[56,1638,477,1528]" captionTargetPageId="3" captionText="TABLE 1 Material examined. Abbreviations: AM, Australian Museum, Sydney; H, holotype; NMV,Museums Victoria, Melbourne; P, Paratype; QM, Queensland Museum, Brisbane; S, syntype; NHRM, Naturhistoriska Riksmuseet, Stockholm, Sweden" httpUri="http://table.plazi.org/id/DF25CBDEFFDAB10353029D82F71DFE37" pageId="79" pageNumber="430" tableUuid="DF25CBDEFFDAB10353029D82F71DFE37">table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C340C8DDFF96B15051BA9A20F781F923" lastPageId="80" lastPageNumber="431" pageId="79" pageNumber="430" type="diagnosis">
<paragraph id="8BE59B56FF96B14F51BA9A20F69AF9B1" blockId="79.[640,1156,1584,1754]" box="[640,749,1584,1610]" pageId="79" pageNumber="430">Diagnosis</paragraph>
<paragraph id="8BE59B56FF96B15051BC9A44F5B9F9B2" blockId="79.[640,1156,1584,1754]" lastBlockId="80.[129,641,1367,1752]" lastPageId="80" lastPageNumber="431" pageId="79" pageNumber="430">
<emphasis id="B92E4744FF96B14F51BC9A44F704F995" box="[646,883,1620,1646]" italics="true" pageId="79" pageNumber="430">External morphology</emphasis>
: Shell (figs. 15EF, 19I) small to medium-sized (
<quantity id="4CA236B3FF96B14F50999A68F04EF969" box="[931,1081,1656,1682]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.28" metricValueMin="0.86" pageId="79" pageNumber="430" unit="mm" value="10.7" valueMax="12.8" valueMin="8.6">8.612.8 mm</quantity>
), light golden amber, subglobose with a low spire, 3.23.5 whorls. Body
<quantity id="4CA236B3FF96B14F50409AD0F785F921" box="[890,1010,1728,1754]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="79" pageNumber="430" unit="mm" value="25.5" valueMax="27.0" valueMin="24.0">2427 mm</quantity>
long, colour (in ethanol) beige, darker brown on tail, mantle lobes large with black spots and mottling, shell lappets moderately large to large, finely pustulose with multiple rows of larger black warts, joined by a narrow collar; dark line along mantle edge. Tail not keeled, slime network weak, caudal horn large.
</paragraph>
<caption id="DF25CBDEFF89B15053BB98D7F00FFB00" ID-DOI="http://doi.org/10.5281/zenodo.8343117" ID-Zenodo-Dep="8343117" httpUri="https://zenodo.org/record/8343117/files/figure.png" pageId="80" pageNumber="431" startId="80.[129,202,1223,1244]" targetBox="[129,1167,188,1192]" targetPageId="80" targetType="figure">
<paragraph id="8BE59B56FF89B15053BB98D7F00FFB00" blockId="80.[129,1174,1223,1275]" pageId="80" pageNumber="431">
FIGURE 28 Genitalia of
<taxonomicName id="4C5AE0D5FF89B150525198D8F67DFB26" authority=", MO" authorityName="MO" box="[363,522,1223,1245]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="80" pageNumber="431" phylum="Mollusca" rank="species" species="mcdonaldi">
<emphasis id="B92E4744FF89B150525198D8F5A9FB27" box="[363,478,1223,1245]" italics="true" pageId="80" pageNumber="431">F. mcdonaldi</emphasis>
, MO
</taxonomicName>
504354, Eungella NP. A. Reproductive system. Arrow shows the junction of the penis and epiphallus. B. Spermatophore. C. Penis interior. Scale bars 1 mm (BC), 2 mm (A).
</paragraph>
</caption>
<paragraph id="8BE59B56FF89B15053A59A42F781F923" blockId="80.[129,641,1367,1752]" lastBlockId="80.[677,1188,1367,1752]" pageId="80" pageNumber="431">
<emphasis id="B92E4744FF89B15053A59A42F529F997" box="[159,350,1618,1644]" italics="true" pageId="80" pageNumber="431">Genital anatomy:</emphasis>
Genitalia (figs. 22IJ, 29) with short vagina; bursa copulatrix moderately short; duct very short, bursa large, oval shaped. Penis moderate size, cylindrical, inner penial wall sculptured with very fine diagonal ridges, penis interior with one large, raised longitudinal pilaster and three small, irregularly shaped pilasters (two distal, one proximal); 75% of penis contained in penial tunica. Epiphallus slightly longer than penis, entering penis through a simple pore; epiphallus 2 longer than epiphallus 1; epiphallic caecum short; flagellum moderately long, slender, coiled. Spermatophore with a single, finely toothed, spiraling ridge.
</paragraph>
</subSubSection>
<caption id="DF25CBDEFF88B1515358989BF7D2FB46" ID-DOI="http://doi.org/10.5281/zenodo.8343119" ID-Zenodo-Dep="8343119" httpUri="https://zenodo.org/record/8343119/files/figure.png" pageId="81" pageNumber="432" startId="81.[98,171,1163,1184]" targetBox="[99,1156,188,1130]" targetPageId="81" targetType="figure">
<paragraph id="8BE59B56FF88B1515358989BF7D2FB46" blockId="81.[98,1152,1162,1214]" pageId="81" pageNumber="432">
FIGURE 29 Genitalia of
<taxonomicName id="4C5AE0D5FF88B1515276989BF5A2FB5B" authority=", MO" authorityName="MO" box="[332,469,1162,1184]" class="Gastropoda" family="Helicarionidae" genus="Dimidarion" kingdom="Animalia" order="Stylommatophora" pageId="81" pageNumber="432" phylum="Mollusca" rank="species" species="minerva">
<emphasis id="B92E4744FF88B1515276989BF5DEFB5B" box="[332,425,1163,1184]" italics="true" pageId="81" pageNumber="432">F.minerva</emphasis>
, MO
</taxonomicName>
78358, Carnarvon NP. A. Reproductive system. Arrow shows the junction of the penis and epiphallus. B. Penis interior. C. Spermatophore. Scale bars 1 mm.
</paragraph>
</caption>
<subSubSection id="C340C8DDFF88B15253669907F637FE3A" lastPageId="82" lastPageNumber="433" pageId="81" pageNumber="432" type="discussion">
<paragraph id="8BE59B56FF88B15153669907F4C9FACA" blockId="81.[92,611,1303,1725]" box="[92,190,1303,1329]" pageId="81" pageNumber="432">Remarks</paragraph>
<paragraph id="8BE59B56FF88B1515358992CF058F9AA" blockId="81.[92,611,1303,1725]" lastBlockId="81.[646,1157,1303,1725]" pageId="81" pageNumber="432">
<taxonomicName id="4C5AE0D5FF88B1515358992CF533FAAE" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[98,324,1340,1365]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="81" pageNumber="432" phylum="Mollusca" rank="species" species="minerva">
<emphasis id="B92E4744FF88B1515358992CF533FAAE" box="[98,324,1340,1365]" italics="true" pageId="81" pageNumber="432">Fastosarion minerva</emphasis>
</taxonomicName>
(from
<taxonomicName id="4C5AE0D5FF88B15152AE992BF62DFAAD" authority="Hills NP" authorityName="Hills NP" box="[404,602,1339,1366]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="81" pageNumber="432" phylum="Mollusca" rank="species" species="minerva">Minerva Hills NP</taxonomicName>
) was originally described as part of the genus
<taxonomicName id="4C5AE0D5FF88B151535F9993F496FA66" authorityName="Stanisic" authorityYear="2010" box="[101,225,1411,1437]" class="Gastropoda" family="Helicarionidae" genus="Dimidarion" kingdom="Animalia" order="Stylommatophora" pageId="81" pageNumber="432" phylum="Mollusca" rank="genus">
<emphasis id="B92E4744FF88B151535F9993F496FA66" box="[101,225,1411,1437]" italics="true" pageId="81" pageNumber="432">Dimidarion</emphasis>
</taxonomicName>
, a group of small semislugs distributed in southern Qld, in habitats ranging from dry vine thicket to rainforest. Herein we show that
<taxonomicName id="4C5AE0D5FF88B15153AC99E0F57EF9F2" box="[150,265,1520,1545]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="81" pageNumber="432" phylum="Mollusca" rank="species" species="minerva">
<emphasis id="B92E4744FF88B15153AC99E0F57EF9F2" box="[150,265,1520,1545]" italics="true" pageId="81" pageNumber="432">F. minerva</emphasis>
</taxonomicName>
is not closely related to its former congeners (now synonymized to a single species,
<taxonomicName id="4C5AE0D5FF88B15153FC9A28F550F9AA" box="[198,295,1591,1617]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="81" pageNumber="432" phylum="Mollusca" rank="species" species="alyssa">
<emphasis id="B92E4744FF88B15153FC9A28F550F9AA" box="[198,295,1591,1617]" italics="true" pageId="81" pageNumber="432">F. alyssa</emphasis>
</taxonomicName>
) but instead groups most closely with
<taxonomicName id="4C5AE0D5FF88B15153C99A4BF52AF98E" box="[243,349,1627,1653]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="81" pageNumber="432" phylum="Mollusca" rank="species" species="deensis">
<emphasis id="B92E4744FF88B15153C99A4BF52AF98E" box="[243,349,1627,1653]" italics="true" pageId="81" pageNumber="432">F. deensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5AE0D5FF88B15152A29A4BF589F98F" box="[408,510,1627,1652]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="81" pageNumber="432" phylum="Mollusca" rank="species" species="rowani">
<emphasis id="B92E4744FF88B15152A29A4BF589F98F" box="[408,510,1627,1652]" italics="true" pageId="81" pageNumber="432">F. rowani</emphasis>
</taxonomicName>
, both of which are found near Rockhampton. A similar link between these distant locations is seen in helicarionid species
<taxonomicName id="4C5AE0D5FF88B15150579908F044FACA" authorityName="Stanisic" authorityYear="2010" box="[877,1075,1303,1329]" class="Gastropoda" family="Helicarionidae" genus="Tarocystis" kingdom="Animalia" order="Stylommatophora" pageId="81" pageNumber="432" phylum="Mollusca" rank="species" species="toomba">
<emphasis id="B92E4744FF88B15150579908F044FACA" box="[877,1075,1303,1329]" italics="true" pageId="81" pageNumber="432">Tarocystis toomba</emphasis>
</taxonomicName>
, which is distributed from Rockhampton to the Conway Range and west to Carnarvon Gorge (
<bibRefCitation id="EFCBE6A7FF88B15151AA9993F704FA66" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." box="[656,883,1411,1438]" pageId="81" pageNumber="432" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al., 2010</bibRefCitation>
), and camaenid genus
<taxonomicName id="4C5AE0D5FF88B15151BC99B7F7CBFA3A" authority="Stanisic, 2010" authorityName="Stanisic" authorityYear="2010" box="[646,956,1447,1473]" class="Gastropoda" family="Camaenidae" genus="Trachygenia" kingdom="Animalia" order="Stylommatophora" pageId="81" pageNumber="432" phylum="Mollusca" rank="genus">
<emphasis id="B92E4744FF88B15151BC99B7F779FA3A" box="[646,782,1447,1473]" italics="true" pageId="81" pageNumber="432">Trachygenia</emphasis>
<bibRefCitation id="EFCBE6A7FF88B151502499B7F7CBFA3A" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." box="[798,956,1447,1473]" pageId="81" pageNumber="432" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic, 2010</bibRefCitation>
</taxonomicName>
, which contains species ranging from from Dan Dan Scrub and Kroombit Tops (south of Rockhampton) to the Clarke Range and Conway Range, as well as one species from Carnarvon Gorge.
</paragraph>
<paragraph id="8BE59B56FF88B152519F9A4BF637FE3A" blockId="81.[646,1157,1303,1725]" lastBlockId="82.[129,640,171,449]" lastPageId="82" lastPageNumber="433" pageId="81" pageNumber="432">
The type locality of
<taxonomicName id="4C5AE0D5FF88B15150889A4BF058F98F" box="[946,1071,1627,1652]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="81" pageNumber="432" phylum="Mollusca" rank="species" species="minerva">
<emphasis id="B92E4744FF88B15150889A4BF058F98F" box="[946,1071,1627,1652]" italics="true" pageId="81" pageNumber="432">F. minerva</emphasis>
</taxonomicName>
is Mt Zamia in
<taxonomicName id="4C5AE0D5FF88B15151C89A6EF7C5F962" authority="Hills NP" authorityName="Hills NP" box="[754,946,1662,1689]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="81" pageNumber="432" phylum="Mollusca" rank="species" species="minerva">Minerva Hills NP</taxonomicName>
(see fig. 18). Populations from Carnarvon NP and Injune are genetically distinct from each other and from those from Mt Zamia. Material suitable for dissection was not available for the
<typeStatus id="54E125F4FF8BB15251309CE4F64CFEF5" box="[522,571,244,270]" pageId="82" pageNumber="433">type</typeStatus>
locality, but specimens from Carnarvon NP and Injune did not differ anatomically despite their genetic differentiation.We include all populations within
<taxonomicName id="4C5AE0D5FF8BB15252319D93F50BFE67" box="[267,380,387,412]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="82" pageNumber="433" phylum="Mollusca" rank="species" species="minerva">
<emphasis id="B92E4744FF8BB15252319D93F50BFE67" box="[267,380,387,412]" italics="true" pageId="82" pageNumber="433">F.minerva</emphasis>
</taxonomicName>
, pending further investigation once more material is available.
</paragraph>
</subSubSection>
</treatment>
</document>