153 lines
18 KiB
XML
153 lines
18 KiB
XML
<document id="8B01821161182060E4D2A403BEE6F9E5" ID-DOI="10.1163/18759866-20191416" ID-ISSN="1875-9866" ID-Zenodo-Dep="8343061" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1694662606570" checkinUser="felipe" docAuthor="Hyman, Isabel T. & Köhler, Frank" docDate="2019" docId="03F32A40FF8FB15653419E36F78EFCEB" docLanguage="en" docName="ContribZool.88.4.351-451.pdf" docOrigin="Contributions to Zoology 88 (4)" docSource="http://dx.doi.org/10.1163/18759866-20191416" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.2:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="2" docTitle="Fastosarion sarina Hyman & Köhler 2019, sp. nov." docType="treatment" docVersion="5" lastPageNumber="437" masterDocId="FFCA5238FFD9B100533A9C10F477FFFB" masterDocTitle="Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)" masterLastPageNumber="451" masterPageNumber="351" pageNumber="437" updateTime="1695298548213" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="F7BB54A9ABE54708EBED1FA2F2EC446A">
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<mods:title id="BB01D58538BF4225A96D9685982399B3">Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)</mods:title>
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<mods:name id="8769616909DDE5FED0B3C79B282B03B8" type="personal">
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<mods:namePart id="20F7B5C33FDF65ECFD3C2B1810E4EA9C">Hyman, Isabel T.</mods:namePart>
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<mods:name id="4D3DCE268772DAC3F96E16AF995A2ECD" type="personal">
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<mods:roleTerm id="76135A0BCD428C693729EF06D9F553BA">Author</mods:roleTerm>
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<mods:namePart id="69CDAC2320B5ED772C8D53AC432CB719">Köhler, Frank</mods:namePart>
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<mods:typeOfResource id="C40AF9364B3A79EBFE210B2CEC7A93A5">text</mods:typeOfResource>
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<mods:title id="C8B0D63A3BA106106ECF55CB55F1DA95">Contributions to Zoology</mods:title>
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</mods:titleInfo>
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<mods:part id="67BF0F50119EF31B1B85DDF421EFC6D6">
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<mods:date id="B2C651BD77E67D21CAC21E76E96CD788">2019</mods:date>
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<mods:detail id="792F34F9B97476D8C7BABC81E91342D6" type="pubDate">
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<mods:number id="7CA0BB2B6A234E736BB0CA925E9BC829">2019-10-03</mods:number>
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<mods:detail id="5A8FAEC9B87BB18C416647D4F38A704A" type="volume">
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<mods:number id="F791E01A9FCE286C17A2D153D8CDF145">88</mods:number>
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<mods:detail id="985E65891578BF13967687356E6790A0" type="issue">
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<mods:number id="7E03923C37077CA6E8547C67B1C86C9D">4</mods:number>
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<mods:location id="01463A6873899CE7999305A6611A3967">
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<mods:url id="9391B529B7CE0E99C90CF04CFEBAB0D7">http://dx.doi.org/10.1163/18759866-20191416</mods:url>
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<mods:classification id="8CA7FB1C594EBFFDB1DFEE1151AF362F">journal article</mods:classification>
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<mods:identifier id="0428166FB4C1833E5FE7301D2F782E7B" type="DOI">10.1163/18759866-20191416</mods:identifier>
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<mods:identifier id="4E7462F0A98869849B2EB0596604D8B1" type="ISSN">1875-9866</mods:identifier>
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<mods:identifier id="6406232BD2F58B9C1117368FAC5B47BF" type="Zenodo-Dep">8343061</mods:identifier>
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</mods:mods>
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<treatment id="03F32A40FF8FB15653419E36F78EFCEB" LSID="urn:lsid:plazi:treatment:03F32A40FF8FB15653419E36F78EFCEB" httpUri="http://treatment.plazi.org/id/03F32A40FF8FB15653419E36F78EFCEB" lastPageNumber="437" pageId="86" pageNumber="437">
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<subSubSection id="C340C8DDFF8FB15653419E36F5C7FDBA" box="[123,432,550,577]" pageId="86" pageNumber="437" type="nomenclature">
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<paragraph id="8BE59B56FF8FB15653419E36F5C7FDBA" blockId="86.[123,432,550,577]" box="[123,432,550,577]" pageId="86" pageNumber="437">
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<heading id="D0AD2C3AFF8FB15653419E36F5C7FDBA" box="[123,432,550,577]" fontSize="11" level="2" pageId="86" pageNumber="437" reason="2">
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<emphasis id="B92E4744FF8FB15653419E36F5C7FDBA" bold="true" box="[123,432,550,577]" pageId="86" pageNumber="437">
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<taxonomicName id="4C5AE0D5FF8FB15653419E36F522FDBB" ID-CoL="7RBRV" ID-ENA="2738849" authority="Hyman & Köhler, 2019" authorityName="Hyman & Köhler" authorityYear="2019" box="[123,341,550,576]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="86" pageNumber="437" phylum="Mollusca" rank="species" species="sarina" status="sp. nov.">
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<emphasis id="B92E4744FF8FB15653419E36F522FDBB" bold="true" box="[123,341,550,576]" italics="true" pageId="86" pageNumber="437">Fastosarion sarina</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21DFA3FFF8FB15652609E37F5C7FDBA" box="[346,432,551,577]" pageId="86" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C340C8DDFF8FB15653BA9E5AF78EFCEB" pageId="86" pageNumber="437" type="description">
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<paragraph id="8BE59B56FF8FB15653BA9E5AF526FD9F" blockId="86.[128,337,586,612]" box="[128,337,586,612]" pageId="86" pageNumber="437">
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<figureCitation id="136187D3FF8FB15653BA9E5AF4A7FD9F" box="[128,208,586,612]" captionStart="FIGURE 18" captionStartId="66.[129,202,1582,1603]" captionTargetBox="[129,1187,188,1550]" captionTargetId="figure-8@66.[129,1187,188,1550]" captionTargetPageId="66" captionText="FIGURE 18 Distribution of Clade B species. █ = F.alyssa, ▼ = F.deensis, × = F.ephelis, ♢ = F.hannianus, ▲ = F. insularis, ◻ = F.katatonos, ∆ = F.longimentula, ● = F.mcdonaldi, ♠ = F.minerva, ◆ = F.papillosus, ♣ = F.pustulosus, ♥ = F.rowani, Ǫ = F.sarina, ▽ = F.tuljun." figureDoi="http://doi.org/10.5281/zenodo.8343097" httpUri="https://zenodo.org/record/8343097/files/figure.png" pageId="86" pageNumber="437">
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Figs.
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<geoCoordinate id="EE6EFD91FF8FB15653829E5AF4A7FD9F" box="[184,208,586,612]" degrees="18, 19" direction="north" orientation="latitude" pageId="86" pageNumber="437" precision="555" value="18.19">18</geoCoordinate>
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</figureCitation>
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,
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<figureCitation id="136187D3FF8FB15653E09E5AF573FD9F" box="[218,260,586,612]" captionStart="FIGURE 19" captionStartId="67.[98,171,1480,1501]" captionTargetBox="[98,1104,171,1448]" captionTargetId="figure-115@67.[98,1104,170,1448]" captionTargetPageId="67" captionText="FIGURE 19 Shells of Clade B species. A. Fastosarion alyssa (type locality of D.peterbrocki), NMV F230748. B. Fastosarion deensis, MO85834 (holotype). C. Fastosarion ephelis, MO85835 (holotype). D. Fastosarion hannianus, MO56684.E. Fastosarion insularis sp. nov., MO76354. F. Fastosarion katatonos, MO85837 (holotype). G.Fastosarion longimentula, MO65313.H. Fastosarion mcdonaldi, MO50434.I. Fastosarion minerva, MO78358. J.Fastosarion papillosus, MO78896 (Mt Superbus).K.Fastosarion papillosus, MO62397 (Kroombit Tops). L.Fastosarion pustulosus, MO78894 (Kalpowar SF).M. Fastosarion rowani, MO85839 (holotype). N. Fastosarion sarina, MO85840 (holotype).O. Fastosarion tujun (Denham Range), MO76081 (holotype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8343099" httpUri="https://zenodo.org/record/8343099/files/figure.png" pageId="86" pageNumber="437">19N</figureCitation>
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,
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<figureCitation id="136187D3FF8FB15652379E5AF526FD9F" box="[269,337,586,612]" captionStart="FIGURE 31" captionStartId="85.[98,171,1229,1250]" captionTargetBox="[109,1149,188,1206]" captionTargetId="figure-6@85.[97,1155,188,1206]" captionTargetPageId="85" captionText="FIGURE 31 Genitalia.A–B. F.rowani, MO85839,Gladstone. C–E.F.sarina, MO85840, Sarina.A, C. Reproductive system.Arrows show the junction of the penis and epiphallus.B, D.Penis interior. E. spermatophore. Scale bars 1 mm (C–E), 2 mm (A–B)." figureDoi="http://doi.org/10.5281/zenodo.8343123" httpUri="https://zenodo.org/record/8343123/files/figure.png" pageId="86" pageNumber="437">31C–E</figureCitation>
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</paragraph>
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<paragraph id="8BE59B56FF8FB15653409E82F485FD57" blockId="86.[122,639,658,756]" box="[122,242,658,684]" pageId="86" pageNumber="437">Etymology</paragraph>
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<paragraph id="8BE59B56FF8FB15653BA9EA7F488FD0F" blockId="86.[122,639,658,756]" pageId="86" pageNumber="437">
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For Sarina, the
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<typeStatus id="54E125F4FF8FB15652679EA7F5F9FD2A" box="[349,398,695,721]" pageId="86" pageNumber="437">type</typeStatus>
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locality; noun in apposition.
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</paragraph>
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<paragraph id="8BE59B56FF8FB15653409F32F539FCC7" blockId="86.[122,639,802,1043]" box="[122,334,802,828]" pageId="86" pageNumber="437">Material examined</paragraph>
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<paragraph id="8BE59B56FF8FB15653BA9F56F5DEFC53" blockId="86.[122,639,802,1043]" pageId="86" pageNumber="437">
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<emphasis id="B92E4744FF8FB15653BA9F56F4C8FCA4" box="[128,191,838,863]" italics="true" pageId="86" pageNumber="437">Types</emphasis>
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:
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<materialsCitation id="3B32910BFF8FB15653F09F56F5DEFC53" collectingDate="2000-03-23" collectionCode="QM" collectorName="G. Monteith" latitude="-21.6" location="Blue Mts" longLatPrecision="1263" longitude="148.96666" pageId="86" pageNumber="437" specimenCode="MO85840" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54E125F4FF8FB15653F09F56F542FC9B" box="[202,309,838,864]" pageId="86" pageNumber="437">Holotype</typeStatus>
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:
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<collectionCode id="ED4B0393FF8FB15652049F56F51EFC9B" box="[318,361,838,864]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="86" pageNumber="437" type="Museum">QM</collectionCode>
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<specimenCode id="DBFC332DFF8FB15652559F55F5ACFC9B" box="[367,475,837,864]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="86" pageNumber="437" type="Museum">MO85840</specimenCode>
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(
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<taxonomicName id="4C5AE0D5FF8FB15652D39F56F615FC9B" authority=", SW" authorityName="SW" box="[489,610,838,864]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="86" pageNumber="437" phylum="Mollusca" rank="species" species="sarina">Sarina, SW</taxonomicName>
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at
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<location id="8E85CD8DFF8FB15653BA9F7AF493FC7F" LSID="urn:lsid:plazi:treatment:03F32A40FF8FB15653419E36F78EFCEB:8E85CD8DFF8FB15653BA9F7AF493FC7F" box="[128,228,874,900]" latitude="-21.6" longLatPrecision="1263" longitude="148.96666" name="Blue Mts" pageId="86" pageNumber="437">Blue Mts</location>
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, c.
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<quantity id="4CA236B3FF8FB156523E9F79F539FC7F" box="[260,334,873,900]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="86" pageNumber="437" unit="km" value="0.6">0.6 km</quantity>
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SE,
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<geoCoordinate id="EE6EFD91FF8FB15652419F7AF5A4FC7F" box="[379,467,873,900]" degrees="21" direction="south" minutes="36" orientation="latitude" pageId="86" pageNumber="437" precision="925" value="-21.6">21° 36'S</geoCoordinate>
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||
,
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<geoCoordinate id="EE6EFD91FF8FB15652E79F7AF630FC7F" box="[477,583,873,900]" degrees="148" direction="east" minutes="58" orientation="longitude" pageId="86" pageNumber="437" precision="925" value="148.96666">148° 58' E</geoCoordinate>
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, coll.
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<date id="FFE4BD96FF8FB15653459F9EF57EFC53" box="[127,265,910,936]" pageId="86" pageNumber="437" value="2000-03-23">
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<collectingDate id="EFA0447EFF8FB15653459F9EF57EFC53" box="[127,265,910,936]" pageId="86" pageNumber="437" value="2000-03-23">23 Mar 2000</collectingDate>
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</date>
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,
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<collectorName id="26AFFE80FF8FB15652289F9EF5E9FC53" box="[274,414,909,936]" pageId="86" pageNumber="437">G. Monteith</collectorName>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BE59B56FF8FB15653A49FA2F483FC0B" blockId="86.[122,639,802,1043]" pageId="86" pageNumber="437">
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<materialsCitation id="3B32910BFF8FB15653A49FA2F487FC0B" collectingDate="2000-03-23" collectionCode="QM" collectorName="G. Monteith" latitude="-21.6" location="Blue Mts" longLatPrecision="1263" longitude="148.96666" pageId="86" pageNumber="437" specimenCode="MO77305" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54E125F4FF8FB15653A49FA2F57AFC37" box="[158,269,946,972]" pageId="86" pageNumber="437">Paratypes</typeStatus>
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:
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<collectionCode id="ED4B0393FF8FB156521A9FA2F53CFC37" box="[288,331,946,972]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="86" pageNumber="437" type="Museum">QM</collectionCode>
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<specimenCode id="DBFC332DFF8FB15652609FA1F5B4FC37" box="[346,451,945,972]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="86" pageNumber="437" type="Museum">MO77305</specimenCode>
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(same data as holotype)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BE59B56FF8FB15653A49FEAF59EFBE8" blockId="86.[122,639,802,1043]" box="[158,489,1017,1043]" pageId="86" pageNumber="437">
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<emphasis id="B92E4744FF8FB15653A49FEAF51DFBE8" box="[158,362,1017,1043]" italics="true" pageId="86" pageNumber="437">Non-type material:</emphasis>
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See
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<tableCitation id="C6D8AEEDFF8FB15652A69FE9F593FBE8" box="[412,484,1017,1043]" captionStart="TABLE 1" captionStartId="3.[56,116,402,428]" captionTargetBox="[56,1638,477,1528]" captionTargetPageId="3" captionText="TABLE 1 Material examined. Abbreviations: AM, Australian Museum, Sydney; H, holotype; NMV,Museums Victoria, Melbourne; P, Paratype; QM, Queensland Museum, Brisbane; S, syntype; NHRM, Naturhistoriska Riksmuseet, Stockholm, Sweden" httpUri="http://table.plazi.org/id/DF25CBDEFFDAB10353029D82F71DFE37" pageId="86" pageNumber="437" tableUuid="DF25CBDEFFDAB10353029D82F71DFE37">table 1</tableCitation>
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.
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</paragraph>
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<paragraph id="8BE59B56FF8FB15653409851F490FBA0" blockId="86.[122,641,1089,1761]" box="[122,231,1089,1115]" pageId="86" pageNumber="437">Diagnosis</paragraph>
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<paragraph id="8BE59B56FF8FB15653BA9875F4BAF9F1" blockId="86.[122,641,1089,1761]" pageId="86" pageNumber="437">
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<emphasis id="B92E4744FF8FB15653BA9875F519FB84" box="[128,366,1125,1151]" italics="true" pageId="86" pageNumber="437">External morphology</emphasis>
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: Shell (fig. 19N) medium-sized (
|
||
<quantity id="4CA236B3FF8FB15653C39899F52AFB58" box="[249,349,1161,1187]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="86" pageNumber="437" unit="mm" value="17.7">17.7 mm</quantity>
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||
), amber, 3.2 whorls, flattened, last whorl very large. Body
|
||
<quantity id="4CA236B3FF8FB156513C98BDF609FB3C" box="[518,638,1197,1223]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="86" pageNumber="437" unit="mm" value="29.0" valueMax="30.0" valueMin="28.0">28–30 mm</quantity>
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long, colour (in ethanol) light grey-brown, sole orange-brown, keel cream, right mantle lobe moderately large, left and median lobes used to form a large cephalic shield, shell lappets moderately large, dark brown, paler at tips, with black markings, left lappet with three rows of warts, right lappet smooth. Tail strongly keeled, slime network moderately strong.
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</paragraph>
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<paragraph id="8BE59B56FF8FB15653A49A04F0D2FE82" blockId="86.[122,641,1089,1761]" lastBlockId="86.[676,1189,171,377]" pageId="86" pageNumber="437">
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||
<emphasis id="B92E4744FF8FB15653A49A04F52DF9D5" box="[158,346,1556,1582]" italics="true" pageId="86" pageNumber="437">Genital anatomy:</emphasis>
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Genitalia (fig.31C–E) with medium length vagina; bursa copulatrix long, duct broad, bursa elongate. Penis moderately long, base narrow, increasing in diameter, inner penial wall sculptured with transverse ridges, penis interior with three transversely ridged longitudinal pilasters; 85% of penis contained in penial tunica. Epiphallus equal in length to penis, entering penis through a simple pore; epiphallus 2 longer than epiphallus 1; epiphallic caecum long; flagellum long, slender. Spermatophore with simple tail-pipe.
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</paragraph>
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<paragraph id="8BE59B56FF8FB15651A49DA3F777FE36" blockId="86.[670,1187,435,784]" box="[670,768,435,461]" pageId="86" pageNumber="437">Remarks</paragraph>
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<paragraph id="8BE59B56FF8FB156519E9DC7F78EFCEB" blockId="86.[670,1187,435,784]" pageId="86" pageNumber="437">
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This species (identified above as ‘
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<taxonomicName id="4C5AE0D5FF8FB15657219DC7F6BBFDEF" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="86" pageNumber="437" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
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sp. Sarina’) is known only from a single location, in rainforest above
|
||
<quantity id="4CA236B3FF8FB15650DE9E0FF05EFDC2" box="[996,1065,543,569]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="86" pageNumber="437" unit="m" value="800.0">800 m</quantity>
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altitude at Blue Mountain southwest of Sarina (fig. 18).
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<taxonomicName id="4C5AE0D5FF8FB156519E9E77F719FD7B" box="[676,878,614,640]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="86" pageNumber="437" phylum="Mollusca" rank="species" species="alyssa">
|
||
<emphasis id="B92E4744FF8FB156519E9E77F719FD7B" box="[676,878,614,640]" italics="true" pageId="86" pageNumber="437">Fastosarion alyssa</emphasis>
|
||
</taxonomicName>
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and
|
||
<taxonomicName id="4C5AE0D5FF8FB15650939E77F058FD7B" box="[937,1071,615,640]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="86" pageNumber="437" phylum="Mollusca" rank="species" species="aquavitae">
|
||
<emphasis id="B92E4744FF8FB15650939E77F058FD7B" box="[937,1071,615,640]" italics="true" pageId="86" pageNumber="437">F. aquavitae</emphasis>
|
||
</taxonomicName>
|
||
are found nearby at Sarina. The former can easily be distinguished by its smaller size and the latter by its more globose shell; however, neither are recorded from Blue Mountain.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |