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<document id="9E255E7CAC9731710084E69F974A0836" ID-DOI="10.1163/18759866-20191416" ID-GBIF-Dataset="6388d839-a72d-4598-a141-053897bfc900" ID-ISSN="1875-9866" ID-Zenodo-Dep="8343061" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="diego" IM.tables_requiresApprovalFor="operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1694662606570" checkinUser="felipe" docAuthor="Hyman, Isabel T. &amp; Köhler, Frank" docDate="2019" docId="03F32A40FF84B15E53589D85F523FAC8" docLanguage="en" docName="ContribZool.88.4.351-451.pdf" docOrigin="Contributions to Zoology 88 (4)" docSource="http://dx.doi.org/10.1163/18759866-20191416" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.3:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="3" docTitle="Stanisicarion aquila" docType="treatment" docVersion="9" lastPageNumber="444" masterDocId="FFCA5238FFD9B100533A9C10F477FFFB" masterDocTitle="Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)" masterLastPageNumber="451" masterPageNumber="351" pageNumber="444" updateTime="1731578815876" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:title id="216FA84890C542C609FE1C0DBABF0E36">Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)</mods:title>
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<mods:namePart id="7AC82000DCB9DB08FEB851764E731FF1">Hyman, Isabel T.</mods:namePart>
<mods:affiliation id="2173830D2EA68620A2F8C4A7D595ABAD">Australian Museum, 1 William St, Sydney NSW 2010, Australia Isabel. Hyman @ austmus. gov. au</mods:affiliation>
<mods:nameIdentifier id="58F98DCE1FC3373C1EFCB2079117A436" type="email">Isabel.Hyman@austmus.gov.au</mods:nameIdentifier>
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<mods:namePart id="E1BC6D4C54ECB784250E62A2F2D64EA9">Köhler, Frank</mods:namePart>
<mods:affiliation id="E813962101B2CE03F382B4C823F7FD29">Australian Museum, 1 William St, Sydney NSW 2010, Australia</mods:affiliation>
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<mods:title id="91F82D32A2FEB091591C9C2609D09D2A">Contributions to Zoology</mods:title>
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<mods:date id="CDBE02CEFB5C7344AD879B95AC72803D">2019</mods:date>
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<mods:number id="DFF8C0358001657D76463DC8A420794A">2019-10-03</mods:number>
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<mods:number id="AD7BA91912EF12921E35EC314919DEA6">88</mods:number>
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<treatment id="03F32A40FF84B15E53589D85F523FAC8" ID-DOI="http://doi.org/10.5281/zenodo.8366933" ID-GBIF-Taxon="213408610" ID-Zenodo-Dep="8366933" LSID="urn:lsid:plazi:treatment:03F32A40FF84B15E53589D85F523FAC8" httpUri="http://treatment.plazi.org/id/03F32A40FF84B15E53589D85F523FAC8" lastPageId="94" lastPageNumber="444" pageId="93" pageNumber="444">
<subSubSection id="C340C8DDFF84B15D53589D85F5B8FE54" box="[98,463,405,432]" pageId="93" pageNumber="444" type="nomenclature">
<paragraph id="8BE59B56FF84B15D53589D85F5B8FE54" blockId="93.[98,463,405,432]" box="[98,463,405,432]" pageId="93" pageNumber="444">
<heading id="D0AD2C3AFF84B15D53589D85F5B8FE54" box="[98,463,405,432]" fontSize="11" level="2" pageId="93" pageNumber="444" reason="2">
<taxonomicName id="4C5AE0D5FF84B15D53589D85F5B8FE54" ID-CoL="7T7J7" authority="(Cox, 1868)" baseAuthorityName="Cox" baseAuthorityYear="1868" box="[98,463,405,432]" class="Gastropoda" family="Helicarionidae" genus="Stanisicarion" kingdom="Animalia" order="Stylommatophora" pageId="93" pageNumber="444" phylum="Mollusca" rank="species" species="aquila">
<emphasis id="B92E4744FF84B15D53589D85F5B8FE54" bold="true" box="[98,463,405,432]" pageId="93" pageNumber="444">
<emphasis id="B92E4744FF84B15D53589D85F52FFE54" bold="true" box="[98,344,405,432]" italics="true" pageId="93" pageNumber="444">Stanisicarion aquila (</emphasis>
<bibRefCitation id="EFCBE6A7FF84B15D52639D85F5B1FE54" author="Cox, J. C." box="[345,454,405,431]" pageId="93" pageNumber="444" refId="ref40612" refString="Cox, J. C. (1868) A Monograph of Australian Land Shells. Maddock, Sydney." type="book" year="1868">Cox, 1868</bibRefCitation>
<emphasis id="B92E4744FF84B15D52FF9D85F5B8FE54" bold="true" box="[453,463,405,431]" italics="true" pageId="93" pageNumber="444">)</emphasis>
</emphasis>
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</paragraph>
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<subSubSection id="C340C8DDFF84B15D53589DA9F676FE28" box="[98,513,441,467]" pageId="93" pageNumber="444" type="description">
<paragraph id="8BE59B56FF84B15D53589DA9F676FE28" blockId="93.[98,607,441,611]" box="[98,513,441,467]" pageId="93" pageNumber="444">
<figureCitation id="136187D3FF84B15D53589DA9F4AFFE28" box="[98,216,441,467]" captionStart="FIGURE 14" captionStartId="59.[98,171,1179,1200]" captionTargetBox="[98,1157,188,1147]" captionTargetId="figure-6@59.[98,1157,188,1147]" captionTargetPageId="59" captionText="FIGURE 14 Shell microsculpture. A–C.Fastosarion griseolus, MO55568 (A = protoconch, B = early teleoconch, C = late teleoconch). D. Fastosarion alyssa, F230784 (protoconch). E–F.Fastosarion minerva, MO65418 (E = protoconch, F = teleoconch). G–I.Fastosarion papillosus, MO78896 (G = protoconch, H = early teleoconch, I = late teleoconch).J–L.Stanisicarion aquila, MO49825 (J = protoconch, K = early teleoconch, L = late teleoconch). Scale bars 20 µm (A, D, E, I), 100 µm (B, F, L), 50 µm (C, G-H, J-K)." figureDoi="http://doi.org/10.5281/zenodo.8343089" httpUri="https://zenodo.org/record/8343089/files/figure.png" pageId="93" pageNumber="444">Figs. 14J–L</figureCitation>
,
<figureCitation id="136187D3FF84B15D53DB9DA9F48BFE28" box="[225,252,441,467]" captionStart="FIGURE 33" captionStartId="90.[129,202,1218,1239]" captionTargetBox="[129,1161,188,1193]" captionTargetId="figure-9@90.[129,1187,188,1193]" captionTargetPageId="90" captionText="FIGURE 33 Distribution of Clade C species. ▲ = S.aquila, ● = S. freycineyti, ◻ = S.wolvi." figureDoi="http://doi.org/10.5281/zenodo.8343127" httpUri="https://zenodo.org/record/8343127/files/figure.png" pageId="93" pageNumber="444">33</figureCitation>
,
<figureCitation id="136187D3FF84B15D523C9DA9F547FE28" box="[262,304,441,467]" captionStart="FIGURE 34" captionStartId="91.[98,171,1078,1099]" captionTargetBox="[107,1157,189,1046]" captionTargetId="figure-6@91.[98,1157,188,1046]" captionTargetPageId="91" captionText="FIGURE 34 Shells of Stanisicarion. A. Stanisicarion freycineti, MO70449. B. Stanisicarion aquila, MO49825. C. Stanisicarion wolvi sp. nov., AM C.581912.Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8343129" httpUri="https://zenodo.org/record/8343129/files/figure.png" pageId="93" pageNumber="444">34B</figureCitation>
,
<figureCitation id="136187D3FF84B15D52019DA9F5F1FE28" box="[315,390,441,467]" captionStart="FIGURE 36" captionStartId="92.[128,201,1171,1192]" captionTargetBox="[130,641,189,1147]" captionTargetId="figure-331@92.[130,641,189,1147]" captionTargetPageId="92" captionText="FIGURE 36 Stanisicarion live images (Clade C). A. Stanisicarion freycineti, Emu heights, Sydney (image J.Wradjan).B.Stanisicarion freycineti, AM C.506245, Oxley Wild Rivers NP.C. Stanisicarion aquila, C.512654, Broken Head.D.Stanisicarion aquila, C.512596, Mt Tamborine.E. Stanisicarion aquila, C.269856, Cicada Glen Creek." figureDoi="http://doi.org/10.5281/zenodo.8343135" httpUri="https://zenodo.org/record/8343135/files/figure.png" pageId="93" pageNumber="444">36C–E</figureCitation>
,
<figureCitation id="136187D3FF84B15D52B59DA9F5DDFE28" box="[399,426,441,467]" captionStart="FIGURE 38" captionStartId="94.[129,202,1115,1136]" captionTargetBox="[138,1186,189,1082]" captionTargetId="figure-193@94.[128,1186,189,1082]" captionTargetPageId="94" captionText="FIGURE 38 Genitalia of S.aquila, MO23337,McDonald NP.A.Reproductive system.Arrow shows the junction of the penis and epiphallus.B. Penis interior. C.Spermatophore. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8343137" httpUri="https://zenodo.org/record/8343137/files/figure.png" pageId="93" pageNumber="444">38</figureCitation>
,
<figureCitation id="136187D3FF84B15D528F9DA9F676FE28" box="[437,513,441,467]" captionStart="FIGURE 39" captionStartId="95.[98,171,1411,1432]" captionTargetBox="[98,1157,188,1379]" captionTargetId="figure-6@95.[98,1157,188,1379]" captionTargetPageId="95" captionText="FIGURE 39 Stanisicarion spermatophores (Clade C).A–B. S.aquila, MO77158.C–D.S.aquila, AM C.269856.E–F. S.wolvi, AM C.581912." figureDoi="http://doi.org/10.5281/zenodo.8343139" httpUri="https://zenodo.org/record/8343139/files/figure.png" pageId="93" pageNumber="444">39A–D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C340C8DDFF84B15D53BB9DCEF57FFD98" pageId="93" pageNumber="444" type="reference_group">
<paragraph id="8BE59B56FF84B15D53BB9DCEF5CAFE0C" blockId="93.[98,607,441,611]" box="[129,445,477,503]" pageId="93" pageNumber="444">
<treatmentCitationGroup id="AB4ABC78FF84B15D53BB9DCEF5CAFE0C" box="[129,445,477,503]" pageId="93" pageNumber="444">
<treatmentCitation id="0AFBBD47FF84B15D53BB9DCEF5CEFE0C" author="Cox, J. C." box="[129,441,477,503]" page="109" pageId="93" pageNumber="444" year="1868">
<taxonomicName id="4C5AE0D5FF84B15D53BB9DCEF5CEFE0C" authority="Cox, 1868: 109" authorityName="Cox" authorityPageNumber="109" authorityYear="1868" box="[129,441,477,503]" class="Gastropoda" family="Vitrinidae" genus="Vitrina" kingdom="Animalia" order="Stylommatophora" pageId="93" pageNumber="444" phylum="Mollusca" rank="species" species="aquila">
<emphasis id="B92E4744FF84B15D53BB9DCEF56EFE0C" box="[129,281,477,503]" italics="true" pageId="93" pageNumber="444">Vitrina aquila</emphasis>
<bibRefCitation id="EFCBE6A7FF84B15D52259DCDF5CEFE0C" author="Cox, J. C." box="[287,441,477,503]" pageId="93" pageNumber="444" refId="ref40612" refString="Cox, J. C. (1868) A Monograph of Australian Land Shells. Maddock, Sydney." type="book" year="1868">Cox, 1868: 109</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE59B56FF84B15D53BB9E11F660FDE0" blockId="93.[98,607,441,611]" box="[129,535,513,539]" pageId="93" pageNumber="444">
<treatmentCitationGroup id="AB4ABC78FF84B15D53BB9E11F660FDE0" box="[129,535,513,539]" pageId="93" pageNumber="444">
<treatmentCitation id="0AFBBD47FF84B15D53BB9E11F663FDE0" author="Iredale, T." box="[129,532,513,539]" page="7" pageId="93" pageNumber="444" year="1941">
<taxonomicName id="4C5AE0D5FF84B15D53BB9E11F663FDE0" authority=": Iredale, 1941: 6 - 7" authorityName="Iredale" authorityPageNumber="6 - 7" authorityYear="1941" box="[129,532,513,539]" class="Gastropoda" family="Helicarionidae" genus="Vercularion" kingdom="Animalia" order="Stylommatophora" pageId="93" pageNumber="444" phylum="Mollusca" rank="species" species="aquila">
<emphasis id="B92E4744FF84B15D53BB9E11F527FDE0" box="[129,336,513,539]" italics="true" pageId="93" pageNumber="444">Vercularion aquila:</emphasis>
<bibRefCitation id="EFCBE6A7FF84B15D526C9E11F663FDE0" author="Iredale, T." box="[342,532,513,539]" pageId="93" pageNumber="444" pagination="1 - 8" refId="ref41676" refString="Iredale, T. (1941) Guide to the land shells of New South Wales - part III. Aust. Zool., 11, 1 - 8." type="journal article" year="1941">Iredale, 1941: 6–7</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE59B56FF84B15D53BB9E35F57FFD98" blockId="93.[98,607,441,611]" pageId="93" pageNumber="444">
<treatmentCitationGroup id="AB4ABC78FF84B15D53BB9E35F57FFD98" pageId="93" pageNumber="444">
<taxonomicName id="4C5AE0D5FF84B15D53BB9E35F59EFDC4" ID-CoL="7S235" authority=": Stanisic, 2010" authorityName=": Stanisic" authorityYear="2010" box="[129,489,549,575]" class="Gastropoda" family="Helicarionidae" genus="Macularion" kingdom="Animalia" order="Stylommatophora" pageId="93" pageNumber="444" phylum="Mollusca" rank="species" species="aquila">
<emphasis id="B92E4744FF84B15D53BB9E35F53CFDC4" box="[129,331,549,575]" italics="true" pageId="93" pageNumber="444">Macularion aquila</emphasis>
:
<bibRefCitation id="EFCBE6A7FF84B15D526C9E35F59EFDC4" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." box="[342,489,549,575]" pageId="93" pageNumber="444" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">
Stanisic,
<quantity id="4CA236B3FF84B15D52839E35F59EFDC4" box="[441,489,549,575]" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" pageId="93" pageNumber="444" unit="in" value="2010.0">2010</quantity>
</bibRefCitation>
</taxonomicName>
in
<treatmentCitation id="0AFBBD47FF84B15D51329E35F573FD98" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." page="312" pageId="93" pageNumber="444" year="2010">
<bibRefCitation id="EFCBE6A7FF84B15D51329E35F573FD98" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." pageId="93" pageNumber="444" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al., 2010: 312</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C340C8DDFF84B15D53669E80F695FF23" pageId="93" pageNumber="444" type="materials_examined">
<paragraph id="8BE59B56FF84B15D53669E80F547FD51" blockId="93.[92,608,656,791]" box="[92,304,656,682]" pageId="93" pageNumber="444">Material examined</paragraph>
<paragraph id="8BE59B56FF84B15D53589EA5F57DFD08" blockId="93.[92,608,656,791]" pageId="93" pageNumber="444">
<emphasis id="B92E4744FF84B15D53589EA5F4D6FD35" box="[98,161,693,718]" italics="true" pageId="93" pageNumber="444">Types</emphasis>
:
<materialsCitation id="3B32910BFF84B15D53899EA4F57DFD08" ID-GBIF-Occurrence="4413347360" collectionCode="AM" pageId="93" pageNumber="444" specimenCount="1" typeStatus="holotype">
<typeStatus id="54E125F4FF84B15D53899EA4F569FD35" box="[179,286,692,718]" pageId="93" pageNumber="444" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED4B0393FF84B15D52149EA5F520FD34" box="[302,343,693,719]" country="Australia" name="Australian Museum" pageId="93" pageNumber="444" type="Museum">AM</collectionCode>
C.101185 (Eagle Scrub, Brisbane, Qld).
</materialsCitation>
</paragraph>
<paragraph id="8BE59B56FF84B15D53BB9EEDF5BCFCED" blockId="93.[92,608,656,791]" box="[129,459,764,791]" pageId="93" pageNumber="444">
<emphasis id="B92E4744FF84B15D53BB9EEDF530FCED" box="[129,327,764,790]" italics="true" pageId="93" pageNumber="444">Non-type material</emphasis>
: See
<tableCitation id="C6D8AEEDFF84B15D52449EECF5B0FCED" box="[382,455,764,790]" captionStart="TABLE 1" captionStartId="3.[56,116,402,428]" captionTargetBox="[56,1638,477,1528]" captionTargetPageId="3" captionText="TABLE 1 Material examined. Abbreviations: AM, Australian Museum, Sydney; H, holotype; NMV,Museums Victoria, Melbourne; P, Paratype; QM, Queensland Museum, Brisbane; S, syntype; NHRM, Naturhistoriska Riksmuseet, Stockholm, Sweden" httpUri="http://table.plazi.org/id/DF25CBDEFFDAB10353029D82F71DFE37" pageId="93" pageNumber="444" tableUuid="DF25CBDEFFDAB10353029D82F71DFE37">table 1</tableCitation>
.
</paragraph>
<paragraph id="8BE59B56FF84B15D53669F54F497FCA5" blockId="93.[92,610,836,1760]" box="[92,224,836,862]" pageId="93" pageNumber="444">Description</paragraph>
<paragraph id="8BE59B56FF84B15D53589F78F65FFAF6" blockId="93.[92,610,836,1760]" pageId="93" pageNumber="444">
<emphasis id="B92E4744FF84B15D53589F78F539FC79" box="[98,334,872,898]" italics="true" pageId="93" pageNumber="444">External morphology</emphasis>
: Shell (figs. 14J–L, 34B) medium-sized (
<quantity id="4CA236B3FF84B15D522C9F9CF5D6FC5D" box="[278,417,908,934]" metricMagnitude="-2" metricUnit="m" metricValue="1.6149999999999998" metricValueMax="1.83" metricValueMin="1.4" pageId="93" pageNumber="444" unit="mm" value="16.15" valueMax="18.3" valueMin="14.0">14–18.3 mm</quantity>
), amber, 3.9–4.2 whorls, subglobose with a low spire. Body (figs. 36C–E)
<quantity id="4CA236B3FF84B15D52389FC3F509FC15" box="[258,382,979,1006]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="93" pageNumber="444" unit="mm" value="42.5" valueMax="45.0" valueMin="40.0">40–45 mm</quantity>
long, light orange-brown to grey, tail often darker, depressions of slime network darker (giving tail a diagonally striped appearance), eyestalks dark grey, foot border sometimes orange. Mantle lobes and shell lappets moderately large, pustulose, with numerous, discrete, raised black spots or ridges. Dark line along mantle edge. Tail keeled at tip, keel paler, slime network prominent.
</paragraph>
<paragraph id="8BE59B56FF84B15D53BB9907F67DF91B" blockId="93.[92,610,836,1760]" pageId="93" pageNumber="444">
<emphasis id="B92E4744FF84B15D53BB9907F54BFACA" box="[129,316,1303,1329]" italics="true" pageId="93" pageNumber="444">Genital anatomy:</emphasis>
Genitalia (figs. 38, 39A–D) with moderately long vagina; bursa copulatrix moderately long, duct distinct, bursa oval. Penis long, slender, internally pustulose; 50–80% of penis contained in penial tunica. Epiphallus longer than penis; epiphallus enters penis through a short, pointed verge, partially fused to the penis wall; epiphallus 2 much longer than epiphallus 1; epiphallic caecum very short; flagellum moderately long, slender, irregularly coiled or twisted. Spermatophore a soft capsule with a simple tail-pipe; terminal end hooked with several small spines.
</paragraph>
<paragraph id="8BE59B56FF84B15D51BA9CAEF695FF23" blockId="93.[640,1159,190,1760]" box="[640,738,190,216]" pageId="93" pageNumber="444">Remarks</paragraph>
</subSubSection>
<subSubSection id="C340C8DDFF84B15D51BC9CF2F796FACA" pageId="93" pageNumber="444" type="reference_group">
<paragraph id="8BE59B56FF84B15D51BC9CF2F069FBA2" blockId="93.[640,1159,190,1760]" pageId="93" pageNumber="444">
<taxonomicName id="4C5AE0D5FF84B15D51BC9CF2F719FF00" ID-CoL="7T7J7" baseAuthorityName="Cox" baseAuthorityYear="1868" box="[646,878,225,252]" class="Gastropoda" family="Helicarionidae" genus="Stanisicarion" kingdom="Animalia" order="Stylommatophora" pageId="93" pageNumber="444" phylum="Mollusca" rank="species" species="aquila">
<emphasis id="B92E4744FF84B15D51BC9CF2F719FF00" box="[646,878,225,252]" italics="true" pageId="93" pageNumber="444">Stanisicarion aquila</emphasis>
</taxonomicName>
is currently recorded from Mullumbimby in northeastern NSW to Kenilworth in southeastern Qld (
<bibRefCitation id="EFCBE6A7FF84B15D57369D39F6AAFE9C" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." pageId="93" pageNumber="444" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al., 2010</bibRefCitation>
), where it is found in rainforest habitats under logs, rocks and fallen palm fronds (fig. 33). This relatively broad range and some variation in external morphology (Stanisic, pers. comm.) led
<bibRefCitation id="EFCBE6A7FF84B15D506A9DCDF045FE0C" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." box="[848,1074,477,503]" pageId="93" pageNumber="444" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al. (2010)</bibRefCitation>
to suggest that
<taxonomicName id="4C5AE0D5FF84B15D51CE9E11F722FDE0" ID-CoL="7T7J7" baseAuthorityName="Cox" baseAuthorityYear="1868" box="[756,853,513,539]" class="Gastropoda" family="Helicarionidae" genus="Stanisicarion" kingdom="Animalia" order="Stylommatophora" pageId="93" pageNumber="444" phylum="Mollusca" rank="species" species="aquila">
<emphasis id="B92E4744FF84B15D51CE9E11F722FDE0" box="[756,853,513,539]" italics="true" pageId="93" pageNumber="444">S. aquila</emphasis>
</taxonomicName>
may represent more than one species. Semislugs from northern NSW have brown bodies with orange markings and discrete raised spots on the shell lappets, while those from Mt Tamborine in southern Qld have cream bodies with orange markings and raised short ridges rather than spots on their shell lappets (fig. 34A–C). Further north still, semislugs from Conondale and the Sunshine Coast are grey rather than cream, with no orange colouring and much less visible spots on the shell lappets (fig. 34A–C). These three morphs form separate clades in the phylogenetic tree, but sampling across the entire range has revealed little anatomical variation and a degree of genetic variation consistent with other widespread species (COI: 1.4%, 16S:2.2%).
</paragraph>
<paragraph id="8BE59B56FF84B15D519F9873F796FACA" blockId="93.[640,1159,190,1760]" pageId="93" pageNumber="444">A population of this species was also discovered in Cicada Glen in northern Sydney and is assumed to have been introduced, perhaps in plants being brought in to a nearby nursery. It matches the Conondale population in its body colour and genetics.</paragraph>
</subSubSection>
<subSubSection id="C340C8DDFF84B15E519F992BF523FAC8" lastPageId="94" lastPageNumber="445" pageId="93" pageNumber="444" type="description">
<paragraph id="8BE59B56FF84B15E519F992BF523FAC8" blockId="93.[640,1159,190,1760]" lastBlockId="94.[128,638,1232,1331]" lastPageId="94" lastPageNumber="445" pageId="93" pageNumber="444">
Both introduced and native populations of
<taxonomicName id="4C5AE0D5FF84B15D51BD994FF690FA83" baseAuthorityName="Cox" baseAuthorityYear="1868" box="[647,743,1374,1401]" class="Gastropoda" family="Helicarionidae" genus="Stanisicarion" kingdom="Animalia" order="Stylommatophora" pageId="93" pageNumber="444" phylum="Mollusca" rank="species" species="aquila">
<emphasis id="B92E4744FF84B15D51BD994FF690FA83" box="[647,743,1374,1401]" italics="true" pageId="93" pageNumber="444">S. aquila</emphasis>
</taxonomicName>
occur within the range of congener
<taxonomicName id="4C5AE0D5FF84B15D51BC9993F773FA66" box="[646,772,1411,1437]" class="Gastropoda" family="Helicarionidae" genus="Stanisicarion" kingdom="Animalia" order="Stylommatophora" pageId="93" pageNumber="444" phylum="Mollusca" rank="species" species="freycineti">
<emphasis id="B92E4744FF84B15D51BC9993F773FA66" box="[646,772,1411,1437]" italics="true" pageId="93" pageNumber="444">S. freycineti</emphasis>
</taxonomicName>
, but
<taxonomicName id="4C5AE0D5FF84B15D50019993F7CEFA66" box="[827,953,1411,1437]" class="Gastropoda" family="Helicarionidae" genus="Stanisicarion" kingdom="Animalia" order="Stylommatophora" pageId="93" pageNumber="444" phylum="Mollusca" rank="species" species="freycineti">
<emphasis id="B92E4744FF84B15D50019993F7CEFA66" box="[827,953,1411,1437]" italics="true" pageId="93" pageNumber="444">S. freycineti</emphasis>
</taxonomicName>
is generally found in dry sclerophyll forest and open woodland rather than rainforest and the two species have not been recorded in sympatry.
<taxonomicName id="4C5AE0D5FF84B15D570899FFF76FF9D7" baseAuthorityName="Cox" baseAuthorityYear="1868" class="Gastropoda" family="Helicarionidae" genus="Stanisicarion" kingdom="Animalia" order="Stylommatophora" pageId="93" pageNumber="444" phylum="Mollusca" rank="species" species="aquila">
<emphasis id="B92E4744FF84B15D570899FFF76FF9D7" italics="true" pageId="93" pageNumber="444">Stanisicarion aquila</emphasis>
</taxonomicName>
can be distinguished by its larger, more highly whorled but less globose shell and more highly pustulose shell lappets, usually with visible spots or ridges. It is also sympatric with arboreal semislug
<taxonomicName id="4C5AE0D5FF84B15D50F79AB3F6BCF91B" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="93" pageNumber="444" phylum="Mollusca" rank="species" species="papillosus">
<emphasis id="B92E4744FF84B15D50F79AB3F6BCF91B" italics="true" pageId="93" pageNumber="444">Fastosarion papillosus</emphasis>
</taxonomicName>
, but can be distinguished by its larger size and evenly pustulose lappets, in contrast to the single row of papillae on the shell lappet of
<taxonomicName id="4C5AE0D5FF87B15E53F29909F539FAC9" box="[200,334,1304,1330]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="94" pageNumber="445" phylum="Mollusca" rank="species" species="papillosus">
<emphasis id="B92E4744FF87B15E53F29909F539FAC9" box="[200,334,1304,1330]" italics="true" pageId="94" pageNumber="445">F. papillosus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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