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<document id="64850E460EE2222ACF66090BF91348E8" ID-DOI="10.11646/zootaxa.3925.4.1" ID-GBIF-Dataset="4ec28816-af9e-450f-a2b6-aa07fe6478d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244795" ID-ZooBank="69DADAD6-84DA-4CBE-B653-6527293BA560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004229777" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Louzada, Júlio &amp; Vaz-De-Mello, Fernando" docDate="2015" docId="03DE87D58522FF905991FAEE5191FC3D" docLanguage="en" docName="zt03925p504.pdf" docOrigin="Zootaxa 3925 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Aganhyboma) titovidaurrei Silva, Louzada &amp; Vaz-De-Mello, 2015, new species" docType="treatment" docVersion="10" lastPageNumber="468" masterDocId="FFE7FFAD8532FF815906FFF9554FFFD4" masterDocTitle="A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="504" masterPageNumber="451" pageNumber="467" updateTime="1698565278120" updateUser="plazi">
<mods:mods id="ECF21213F4BCF5DCE8A2B1B53F03725E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="03780A0F4BCFD5232313BE5F673B00E7">
<mods:title id="8F9D92DD1B17C27FD4C9B4F4788670A8">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="E8AEBE21818813972D5C5FC607DFE472" type="personal">
<mods:role id="51F87696BC1C49DCB5B3A6FDB90EBEA9">
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</mods:role>
<mods:namePart id="4397402C8D740BABA15475BC4DF31D6A">Silva, Fernando A. B.</mods:namePart>
</mods:name>
<mods:name id="BB361A62E2CA675F65B0D67A688343C4" type="personal">
<mods:role id="6592ACD8A8FE4BF9DE976B03F6D83E14">
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</mods:role>
<mods:namePart id="4A28268B67E91ADCD9E6826E84BA238E">Louzada, Júlio</mods:namePart>
</mods:name>
<mods:name id="0CB51244DD5FA6B675471E9676B2D1B3" type="personal">
<mods:role id="23D4E2B77C68F9318C684DF88CC76D63">
<mods:roleTerm id="05E4B044EE2E8C13D0754579A50321AC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="08DA8E8416F363596AEF5A29CC1264EB">Vaz-De-Mello, Fernando</mods:namePart>
</mods:name>
<mods:typeOfResource id="BC99DC23B512626E9212ADB3006386C2">text</mods:typeOfResource>
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<mods:title id="2D64EF08F009C8E1213BEA12000B7A3D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2E920FA2F3423EE9AD1A17D1FFE58231">
<mods:date id="6E9A71C4BC5C8F97834A805B671F45D8">2015</mods:date>
<mods:detail id="D46F12522F0F5749C02095B1DF27F2E3" type="volume">
<mods:number id="84D4464A15DBE86489BE524FD76C94F3">3925</mods:number>
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<mods:detail id="1CF978C8D85AE109C18CB2A16AFD9415" type="issue">
<mods:number id="7848E1D5F690430CDF74899947CD2C90">4</mods:number>
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<mods:start id="AFBE8BAA0185A1AC7E5D5410868EB203">451</mods:start>
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<mods:classification id="2F43830D6515698FA056216B9EE0C3F8">journal article</mods:classification>
<mods:identifier id="41CC9DE66CE941AE950EA7EA15B48D4A" type="DOI">10.11646/zootaxa.3925.4.1</mods:identifier>
<mods:identifier id="335DBA8E30CE6217DFBAE88BD7339DBC" type="GBIF-Dataset">4ec28816-af9e-450f-a2b6-aa07fe6478d8</mods:identifier>
<mods:identifier id="E34B23F93802C0F56B3C6BFEB1EB830D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2D9F49BF2BEC6A98DCA9A57B25525DCA" type="Zenodo-Dep">244795</mods:identifier>
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</mods:mods>
<treatment id="03DE87D58522FF905991FAEE5191FC3D" ID-DOI="http://doi.org/10.5281/zenodo.6115708" ID-GBIF-Taxon="119598954" ID-Zenodo-Dep="6115708" LSID="urn:lsid:plazi:treatment:03DE87D58522FF905991FAEE5191FC3D" httpUri="http://treatment.plazi.org/id/03DE87D58522FF905991FAEE5191FC3D" lastPageId="17" lastPageNumber="468" pageId="16" pageNumber="467">
<subSubSection id="C36D65488522FF915991FAEE5435FA87" pageId="16" pageNumber="467" type="nomenclature">
<paragraph id="8BC836C38522FF915991FAEE5652FAE6" blockId="16.[151,797,1303,1363]" box="[151,797,1303,1330]" pageId="16" pageNumber="467">
<heading id="D08081AF8522FF915991FAEE5652FAE6" bold="true" box="[151,797,1303,1330]" fontSize="11" level="1" pageId="16" pageNumber="467" reason="1">
<emphasis id="B903EAD18522FF915991FAEE5652FAE6" bold="true" box="[151,797,1303,1330]" pageId="16" pageNumber="467">
<taxonomicName id="4C774D408522FF915991FAEE57CAFAE5" ID-CoL="34KB4" box="[151,645,1303,1330]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="titovidaurrei" status="sp. nov." subGenus="Aganhyboma">
<emphasis id="B903EAD18522FF915991FAEE5462FAE5" bold="true" box="[151,301,1303,1329]" italics="true" pageId="16" pageNumber="467">Deltochilum</emphasis>
(
<emphasis id="B903EAD18522FF91583BFAEE5491FAE5" bold="true" box="[317,478,1303,1329]" italics="true" pageId="16" pageNumber="467">Aganhyboma</emphasis>
)
<emphasis id="B903EAD18522FF9158F6FAEE57CAFAE5" bold="true" box="[496,645,1303,1329]" italics="true" pageId="16" pageNumber="467">titovidaurrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057AA8522FF915B8BFAE15652FAE6" box="[653,797,1304,1330]" pageId="16" pageNumber="467" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836C38522FF915991FAC35435FA87" blockId="16.[151,797,1303,1363]" box="[151,378,1338,1363]" pageId="16" pageNumber="467">
(
<figureCitation id="134C2A468522FF915999FAC355A8FA87" box="[159,231,1338,1363]" captionStart="FIGURES 4 – 11" captionStartId="38.[151,264,1745,1768]" captionTargetBox="[181,1434,225,1688]" captionTargetId="figure@38.[151,1435,195,1688]" captionTargetPageId="38" captionText="FIGURES 4 – 11. Aedeagus in Deltochilum (Aganhyboma). 4. D. titovidaurrei new species. 5. D. cupreicolle. 6. D. viridescens new status. 7. D. viridicatum new species. 8. D. amandaarcanjoae new species. 9. D. trisignatum. 10. D. cangalha new species. and D. alpercata new species. 11. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244797/files/figure.png" pageId="16" pageNumber="467">Figs 4</figureCitation>
, 42, 90, 100)
</paragraph>
</subSubSection>
<subSubSection id="C36D65488522FF915991FA845062FA42" box="[151,1325,1405,1430]" pageId="16" pageNumber="467" type="etymology">
<paragraph id="8BC836C38522FF915991FA845062FA42" blockId="16.[151,1437,1405,2038]" box="[151,1325,1405,1430]" pageId="16" pageNumber="467">
<emphasis id="B903EAD18522FF915991FA845451FA42" bold="true" box="[151,286,1405,1430]" pageId="16" pageNumber="467">Etymology.</emphasis>
Named in honour of Tito Vidaurre Sánchez, a friend and collaborator in scarabaeine studies.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488522FF9059C1FA5854CDFE7F" lastPageId="17" lastPageNumber="468" pageId="16" pageNumber="467" type="description">
<paragraph id="8BC836C38522FF9159C1FA5850D3F822" blockId="16.[151,1437,1405,2038]" pageId="16" pageNumber="467">
<emphasis id="B903EAD18522FF9159C1FA5854E9FA6E" bold="true" box="[199,422,1441,1466]" pageId="16" pageNumber="467">Description. Body.</emphasis>
Head dark green with a weak sheen or opaque, pronotum in copper tone, with black or greenish spots, elytra dark brown, with an opaque sheen (
<figureCitation id="134C2A468522FF915A28FA3C56DCFA09" box="[814,915,1477,1502]" captionStart="FIGURES 95 – 102" captionStartId="48.[151,264,1951,1974]" captionTargetBox="[181,1434,202,1924]" captionTargetId="figure@48.[152,1434,193,1930]" captionTargetPageId="48" captionText="FIGURES 95 – 102. Species of D. (Aganhyboma). 95. D. trisignatum. 96. D. kolbei. 97. D. violaceum. 98. D. amandaarcanjoae new species. 99. D. cupreicolle. 100. D. titovidaurrei new species. 101. D. viridescens new status. 102. D. viridescens new status. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244807/files/figure.png" pageId="16" pageNumber="467">Fig. 100</figureCitation>
). Ventral surface black. Length
<quantity id="4C8F9B268522FF915C07FA3C50D7FA09" box="[1281,1432,1477,1501]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="16" pageNumber="467" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11 to 12 mm</quantity>
.
<emphasis id="B903EAD18522FF915991FA115591F9D5" bold="true" box="[151,222,1512,1537]" pageId="16" pageNumber="467">Head.</emphasis>
Dorsal surface with dense foveiform punctures, not confluent, separated at center of dorsal interocular surface by approximately twice their diameter. Punctures of variable size; largest around the eyes, and smallest on the anterolateral portion of the head, near the clypeus. Anterior margin of each puncture not defined (as in Fig. 60). Margin between clypeal teeth and genae curved outward approximately at halfway of its length.
<emphasis id="B903EAD18522FF915DD9F9AD5071F9B9" bold="true" box="[1247,1342,1620,1645]" pageId="16" pageNumber="467">Thorax.</emphasis>
Surface of pronotum with small punctures, apparently simple and uniformly distributed. Punctures on anterior angles larger, with a typical foveiform shape. Pronotum with a narrow greenish band extending the entire edge; Disc with transverse oval spot (
<figureCitation id="134C2A468522FF915889F94654BBF903" box="[399,500,1727,1752]" captionStart="FIGURES 95 – 102" captionStartId="48.[151,264,1951,1974]" captionTargetBox="[181,1434,202,1924]" captionTargetId="figure@48.[152,1434,193,1930]" captionTargetPageId="48" captionText="FIGURES 95 – 102. Species of D. (Aganhyboma). 95. D. trisignatum. 96. D. kolbei. 97. D. violaceum. 98. D. amandaarcanjoae new species. 99. D. cupreicolle. 100. D. titovidaurrei new species. 101. D. viridescens new status. 102. D. viridescens new status. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244807/files/figure.png" pageId="16" pageNumber="467">Fig. 100</figureCitation>
). Margin of pronotum (close to lateral angles) straight and emarginated, without depression. Margin between lateral and posterior angles approximately straight.
<emphasis id="B903EAD18522FF915D30F91A51C5F928" bold="true" box="[1078,1162,1763,1788]" pageId="16" pageNumber="467">Elytra.</emphasis>
Lateral margins evenly curved outward (
<figureCitation id="134C2A468522FF915852F8FF54F9F8CB" box="[340,438,1798,1823]" captionStart="FIGURES 95 – 102" captionStartId="48.[151,264,1951,1974]" captionTargetBox="[181,1434,202,1924]" captionTargetId="figure@48.[152,1434,193,1930]" captionTargetPageId="48" captionText="FIGURES 95 – 102. Species of D. (Aganhyboma). 95. D. trisignatum. 96. D. kolbei. 97. D. violaceum. 98. D. amandaarcanjoae new species. 99. D. cupreicolle. 100. D. titovidaurrei new species. 101. D. viridescens new status. 102. D. viridescens new status. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244807/files/figure.png" pageId="16" pageNumber="467">Fig. 100</figureCitation>
). Striae punctate, however faintly visible at a small magnification. Interstrial surface with small, dispersed foveiform punctures, amid small bright ocellate punctures, randomly distributed. First interstria slightly brighter than rest of elytra. Basal carina of seventh interstria short, length about twice the width of the seventh interestria (as in Fig. 72). Length of apical carina of seventh interstria with approximately 2.5 times that of apical carina of sixth interstria.
<emphasis id="B903EAD18522FF9158F9F86C5734F87A" bold="true" box="[511,635,1941,1966]" pageId="16" pageNumber="467">Abdomen.</emphasis>
Punctures large and conspicuous, concentrated anterolaterally on each sternite, reduced to a narrow belt along midline of abdomen (as in Fig. 73). Pygidium with dense foveiform punctures, transverse, elliptical and almost confluent at base. Apex of pygidium acuminate (as in Fig. 73).
<emphasis id="B903EAD18522FF915C5BF82450D3F822" bold="true" box="[1373,1436,2013,2038]" pageId="16" pageNumber="467">Legs.</emphasis>
</paragraph>
<paragraph id="8BC836C38523FF905991FF6E54CDFE7F" blockId="17.[151,1437,151,1002]" pageId="17" pageNumber="468">
Protibia not abruptly widened. Middle protibial tooth equidistant between apical and basal teeth. Protibial spur acuminate at apex. Apical third of metatibia only slightly curved inward (as in Fig. 68), with five clearly visible longitudinal carinae. Inner margin of metatibia without longitudinal row of small tubercles. Anterior edge of profemur completely emarginate, with abundant setae in two-thirds of its length. Foveiform punctures of profemur with transverse elliptical shape. Metatarsus approximately two-thirds as long as metatibia.
<emphasis id="B903EAD18523FF905DC1FEDF5453FEB7" bold="true" pageId="17" pageNumber="468">Secondary sexual characters.</emphasis>
<collectingCountry id="F36076538523FF905824FEB35413FEB7" box="[290,348,330,355]" name="Malta" pageId="17" pageNumber="468">Male</collectingCountry>
abdomen more longitudinally flattened than that of female. Males may possess medially between first and second ventrites, a small tubercle. Apical margin of pygidium longer in male. Protibial spur more acuminate in female.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488523FF9059C1FE4C568FFD1D" pageId="17" pageNumber="468" type="materials_examined">
<paragraph id="8BC836C38523FF9059C1FE4C51ADFE26" blockId="17.[151,1437,151,1002]" pageId="17" pageNumber="468">
<emphasis id="B903EAD18523FF9059C1FE4C57C0FE1A" bold="true" box="[199,655,437,462]" pageId="17" pageNumber="468">
Material studied.
<typeStatus id="54CC88618523FF9058A5FE4C575BFE1A" box="[419,532,437,462]" pageId="17" pageNumber="468" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F36076538523FF905B22FE4C57C0FE1A" box="[548,655,437,462]" name="Brazil" pageId="17" pageNumber="468">BRAZIL</collectingCountry>
</emphasis>
:
<collectionCode id="ED66AE068523FF905BA4FE4F57B3FE1A" box="[674,764,438,462]" pageId="17" pageNumber="468">MINAS</collectionCode>
GERAIS: Lavras, near Parque Ecológico Quedas do Rio Bonito, (
<geoCoordinate id="EE4350048523FF9059FFFE2354DEFE26" box="[249,401,474,498]" direction="south" orientation="latitude" pageId="17" pageNumber="468" precision="1" value="-21.33925">21°20'21.3&quot;S</geoCoordinate>
,
<geoCoordinate id="EE4350048523FF90589AFE20570FFE26" box="[412,576,473,498]" direction="west" orientation="longitude" pageId="17" pageNumber="468" precision="1" value="-44.977444">44°58'38.8&quot;W</geoCoordinate>
),
<date id="FFC910038523FF905B55FE235785FE26" box="[595,714,474,498]" pageId="17" pageNumber="468" value="2009-02-10">10.II.2009</date>
, Fernando Silva &amp; Tito Vidaurrei (
<specimenCount id="9D71FD4A8523FF905D52FE235133FE25" box="[1108,1148,474,498]" pageId="17" pageNumber="468" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068523FF905D85FE235199FE26" box="[1155,1238,474,498]" pageId="17" pageNumber="468">CEMT</collectionCode>
).
</paragraph>
<paragraph id="8BC836C38523FF9059C1FE045041FD8A" blockId="17.[151,1437,151,1002]" pageId="17" pageNumber="468">
<emphasis id="B903EAD18523FF9059C1FE0454A2FDC2" bold="true" box="[199,493,509,534]" pageId="17" pageNumber="468">
<typeStatus id="54CC88618523FF9059C1FE045470FDC2" box="[199,319,509,534]" pageId="17" pageNumber="468" type="paratype">Paratypes</typeStatus>
[5]:
<collectingCountry id="F36076538523FF90587EFE0454A8FDC2" box="[376,487,509,534]" name="Brazil" pageId="17" pageNumber="468">BRAZIL</collectingCountry>
:
</emphasis>
<collectionCode id="ED66AE068523FF9058F2FE045701FDC2" box="[500,590,509,534]" pageId="17" pageNumber="468">GOIÁS</collectionCode>
: Brazlândia,
<date id="FFC910038523FF905BE4FE075663FDC2" box="[738,812,510,534]" pageId="17" pageNumber="468" value="1994-01">I.1994</date>
, E. Grossi (
<specimenCount id="9D71FD4A8523FF905AB3FE075691FDC2" box="[949,990,510,534]" pageId="17" pageNumber="468" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068523FF905AE0FE075176FDC2" box="[998,1081,510,534]" pageId="17" pageNumber="468">CEMT</collectionCode>
); Jatahy,
<date id="FFC910038523FF905DA7FE045049FDC2" box="[1185,1286,509,534]" pageId="17" pageNumber="468" value="1895" valueMax="1896">1895–96</date>
,
<collectingCountry id="F36076538523FF905C14FE045079FDC2" box="[1298,1334,509,534]" name="Switzerland" pageId="17" pageNumber="468">Ch</collectingCountry>
. Pujol (2
<collectionCode id="ED66AE068523FF905991FDDB55BDFDEE" LSID="urn:lsid:biocol.org:col:34988" box="[151,242,546,570]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="468">MNHN</collectionCode>
);
<collectionCode id="ED66AE068523FF905802FDDB5411FDEE" box="[260,350,546,570]" pageId="17" pageNumber="468">MINAS</collectionCode>
GERAIS: Lavras,
<date id="FFC910038523FF905B37FDD857E7FDEE" box="[561,680,545,570]" pageId="17" pageNumber="468" value="1999-02-15">15.II.1999</date>
, Leila A.S. (
<specimenCount id="9D71FD4A8523FF905A30FDDB5611FDED" box="[822,862,546,570]" pageId="17" pageNumber="468" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068523FF905A62FDDB56F8FDEE" box="[868,951,546,570]" pageId="17" pageNumber="468">CEMT</collectionCode>
); Lavras, near Parque Ecológico Quedas do Rio Bonito, (
<geoCoordinate id="EE4350048523FF90582FFDBF548EFD8A" box="[297,449,582,606]" direction="south" orientation="latitude" pageId="17" pageNumber="468" precision="1" value="-21.33925">21°20'21.3&quot;S</geoCoordinate>
,
<geoCoordinate id="EE4350048523FF9058CAFDBC5720FD8A" box="[460,623,581,606]" direction="west" orientation="longitude" pageId="17" pageNumber="468" precision="1" value="-44.977444">44°58'38.8&quot;W</geoCoordinate>
),
<date id="FFC910038523FF905B84FDBF57B6FD8A" box="[642,761,582,606]" pageId="17" pageNumber="468" value="2009-02-10">10.II.2009</date>
, Fernando Silva &amp; Tito Vidaurrei (1 ♀
<collectionCode id="ED66AE068523FF905DB5FDBF504CFD8A" box="[1203,1283,582,606]" pageId="17" pageNumber="468">CREN</collectionCode>
).
</paragraph>
<paragraph id="8BC836C38523FF9059C1FD90516CFD56" blockId="17.[151,1437,151,1002]" box="[199,1059,617,642]" pageId="17" pageNumber="468">
<emphasis id="B903EAD18523FF9059C1FD9054E3FD56" bold="true" box="[199,428,617,642]" pageId="17" pageNumber="468">Sampling methods.</emphasis>
baited pitfall trap with injured diplopods [
<specimenCount id="9D71FD4A8523FF905A80FD935157FD56" box="[902,1048,617,642]" pageId="17" pageNumber="468" type="generic">2 specimens</specimenCount>
].
</paragraph>
<paragraph id="8BC836C38523FF9059C1FD74568FFD1D" blockId="17.[151,1437,151,1002]" pageId="17" pageNumber="468">
<emphasis id="B903EAD18523FF9059C1FD745465FD72" bold="true" box="[199,298,653,678]" pageId="17" pageNumber="468">Habitat.</emphasis>
<collectingCountry id="F36076538523FF905835FD745438FD72" box="[307,375,653,678]" name="Brazil" pageId="17" pageNumber="468">Brazil</collectingCountry>
(Goiás and Minas Gerais) (
<figureCitation id="134C2A468523FF905BBAFD74565DFD72" box="[700,786,653,678]" captionStart="FIGURES 89 – 92" captionStartId="46.[151,264,1748,1771]" captionTargetBox="[150,1406,193,1727]" captionTargetId="figure@46.[150,1437,193,1728]" captionTargetPageId="46" captionText="FIGURES 89 – 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex." httpUri="https://zenodo.org/record/244805/files/figure.png" pageId="17" pageNumber="468">Fig. 90</figureCitation>
), from available data this species inhabits Cerrado. Two individuals were collected in &quot;Campo Limpo&quot;, Cerrado,
<quantity id="4C8F9B268523FF905A16FD4B5623FD1D" box="[784,876,690,714]" metricMagnitude="0" metricUnit="m" metricValue="1.17" pageId="17" pageNumber="468" unit="m" value="1.17">1,170 m</quantity>
<emphasis id="B903EAD18523FF905A75FD4B56F6FD1D" box="[883,953,690,713]" italics="true" pageId="17" pageNumber="468">
<collectionCode id="ED66AE068523FF905A75FD4B56F6FD1D" box="[883,953,690,713]" pageId="17" pageNumber="468">AMSL</collectionCode>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488523FF9059C1FD2C5191FC3D" pageId="17" pageNumber="468" type="diagnosis">
<paragraph id="8BC836C38523FF9059C1FD2C55A3FC12" blockId="17.[151,1437,151,1002]" pageId="17" pageNumber="468">
<emphasis id="B903EAD18523FF9059C1FD2C5496FD3A" bold="true" box="[199,473,725,750]" pageId="17" pageNumber="468">Diagnosis and remarks</emphasis>
(within the
<taxonomicName id="4C774D408523FF905B66FD2F5792FD3A" box="[608,733,726,750]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="cupreicolle">
<emphasis id="B903EAD18523FF905B66FD2F5792FD3A" box="[608,733,726,750]" italics="true" pageId="17" pageNumber="468">cupreicolle</emphasis>
</taxonomicName>
complex): basal carina of seventh interstria short, length about twice the width of the seventh interestria (as in Fig. 72); elytral coloration can vary from dark brown to black; disc of pronotum with transverse oval spot (
<figureCitation id="134C2A468523FF905B4AFCE457E1FCE2" box="[588,686,797,822]" captionStart="FIGURES 95 – 102" captionStartId="48.[151,264,1951,1974]" captionTargetBox="[181,1434,202,1924]" captionTargetId="figure@48.[152,1434,193,1930]" captionTargetPageId="48" captionText="FIGURES 95 – 102. Species of D. (Aganhyboma). 95. D. trisignatum. 96. D. kolbei. 97. D. violaceum. 98. D. amandaarcanjoae new species. 99. D. cupreicolle. 100. D. titovidaurrei new species. 101. D. viridescens new status. 102. D. viridescens new status. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244807/files/figure.png" pageId="17" pageNumber="468">Fig. 100</figureCitation>
). The following aedeagus characters are also diagnostic: parameres shorter than phallobase, slightly rounded at apex, in lateral view forming an angle of approximately 30° with phallobase (
<figureCitation id="134C2A468523FF905818FC9C542CFCAA" box="[286,355,869,894]" captionStart="FIGURES 4 – 11" captionStartId="38.[151,264,1745,1768]" captionTargetBox="[181,1434,225,1688]" captionTargetId="figure@38.[151,1435,195,1688]" captionTargetPageId="38" captionText="FIGURES 4 – 11. Aedeagus in Deltochilum (Aganhyboma). 4. D. titovidaurrei new species. 5. D. cupreicolle. 6. D. viridescens new status. 7. D. viridicatum new species. 8. D. amandaarcanjoae new species. 9. D. trisignatum. 10. D. cangalha new species. and D. alpercata new species. 11. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244797/files/figure.png" pageId="17" pageNumber="468">Fig. 4</figureCitation>
). Lamella copulatrix: left lobe “J”-shaped (as in Fig. 33); right lobe baton-shaped (as in Fig. 33). Superior right peripheral sclerite (SRP) as in Fig. 42. Center of genital segment with a sclerotized portion and &quot;T&quot;- shaped.
</paragraph>
<paragraph id="8BC836C38523FF9059C1FC285191FC3D" blockId="17.[151,1437,151,1002]" box="[199,1246,977,1002]" pageId="17" pageNumber="468">
The specimens present the parameres and phallobase shorter or truncated than
<taxonomicName id="4C774D408523FF905D3EFC2B5198FC3E" box="[1080,1239,978,1002]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="cupreicolle">
<emphasis id="B903EAD18523FF905D3EFC2B5198FC3E" box="[1080,1239,978,1002]" italics="true" pageId="17" pageNumber="468">D. cupreicolle</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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