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<document ID-DOI="10.5733/afin.055.0203" ID-ISSN="2305-2562" ID-Zenodo-Dep="7918724" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1683583599153" checkinUser="felipe" docAuthor="Kvifte, Gunnar M." docDate="2014" docId="03DE811AA822B004FDAD6FE0531EFF50" docLanguage="en" docName="AfrInvertebr.55.2.413-417.pdf" docOrigin="African Invertebrates 55 (2)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.055.0203" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.4:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="4" docTitle="Potophila verrucosa Kvifte 2014, sp.n." docType="treatment" docVersion="3" lastPageNumber="416" masterDocId="FFE7F962A823B007FFAE6D36515DFFA9" masterDocTitle="Description of Potophila verrucosa, gen. n. et sp. n. (Diptera: Psychodidae: Psychodinae) from the West Usambara mountains, Tanzania" masterLastPageNumber="413" masterPageNumber="413" pageNumber="414" updateTime="1683701640121" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Description of Potophila verrucosa, gen. n. et sp. n. (Diptera: Psychodidae: Psychodinae) from the West Usambara mountains, Tanzania</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kvifte, Gunnar M.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-12-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>55</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>413</mods:start>
<mods:end>413</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://www.bioone.org/doi/abs/10.5733/afin.055.0203</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.055.0203</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7918724</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03DE811AA822B004FDAD6FE0531EFF50" httpUri="http://treatment.plazi.org/id/03DE811AA822B004FDAD6FE0531EFF50" lastPageId="3" lastPageNumber="416" pageId="1" pageNumber="414">
<subSubSection box="[515,808,726,753]" pageId="1" pageNumber="414" type="nomenclature">
<paragraph blockId="1.[515,808,726,753]" box="[515,808,726,753]" pageId="1" pageNumber="414">
<heading box="[515,808,726,753]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="414" reason="2">
<taxonomicName ID-CoL="8DGMF" authority="Kvifte, 2014" authorityName="Kvifte" authorityYear="2014" box="[515,748,726,753]" class="Insecta" family="Psychodidae" genus="Potophila" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="414" phylum="Arthropoda" rank="species" species="verrucosa" status="sp.n.">
<emphasis bold="true" box="[515,748,726,753]" italics="true" pageId="1" pageNumber="414">Potophila verrucosa</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[755,808,727,753]" pageId="1" pageNumber="414" rank="species">sp.n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="416" pageId="1" pageNumber="414" type="description">
<paragraph blockId="1.[629,694,769,795]" box="[629,694,769,795]" pageId="1" pageNumber="414">
<figureCitation box="[629,694,769,795]" captionStart="Fig" captionStartId="2.[144,177,1078,1100]" captionTargetBox="[174,1073,217,1061]" captionTargetId="figure-246@2.[163,1085,204,1075]" captionTargetPageId="2" captionText="Fig. 1. Potophila verrucosa, gen.­n­et­sp.­n,­hOlOtYpe­♂:­(A)­head­(dOrsal­view);­(B)­wing;­(C)­terminalia­ (laterodorsal view); (D) epandrium and proctiger (laterodorsal view).Abbreviations:a ap aedeagal apodeme; d distiphallus; gcx gonocoxite; gcx c gonocoxal condyles; gst gonostyle; hyp hypandrium; lk lateral knob; p paramere; pam parameral-aedeagal membrane; pp parameral projections; vepa ventral epandrial plate/aedeagal connection. Scale bars: 100 µm (A), 40 µm (C &amp; D)." figureDoi="http://doi.org/10.5281/zenodo.7918726" httpUri="https://zenodo.org/record/7918726/files/figure.png" pageId="1" pageNumber="414">Fig. 1</figureCitation>
</paragraph>
<paragraph blockId="1.[181,1132,811,837]" box="[181,1132,811,837]" pageId="1" pageNumber="414">
Etymology: From Latin
<emphasis box="[454,541,812,837]" italics="true" pageId="1" pageNumber="414">verruca</emphasis>
­(wart),­referring­tO­the­rugOse­field­On­the­paramere.
</paragraph>
<paragraph blockId="1.[181,587,854,879]" lastBlockId="1.[181,321,896,921]" pageId="1" pageNumber="414">Diagnostic characters: As for genus. Description:</paragraph>
<paragraph blockId="1.[181,247,938,964]" box="[181,247,938,964]" pageId="1" pageNumber="414">Male.</paragraph>
<paragraph blockId="1.[181,1149,980,1294]" pageId="1" pageNumber="414">
<emphasis box="[181,242,980,1006]" italics="true" pageId="1" pageNumber="414">Head</emphasis>
(
<figureCitation box="[261,353,980,1006]" captionStart="Fig" captionStartId="2.[144,177,1078,1100]" captionTargetBox="[174,1073,217,1061]" captionTargetId="figure-246@2.[163,1085,204,1075]" captionTargetPageId="2" captionText="Fig. 1. Potophila verrucosa, gen.­n­et­sp.­n,­hOlOtYpe­♂:­(A)­head­(dOrsal­view);­(B)­wing;­(C)­terminalia­ (laterodorsal view); (D) epandrium and proctiger (laterodorsal view).Abbreviations:a ap aedeagal apodeme; d distiphallus; gcx gonocoxite; gcx c gonocoxal condyles; gst gonostyle; hyp hypandrium; lk lateral knob; p paramere; pam parameral-aedeagal membrane; pp parameral projections; vepa ventral epandrial plate/aedeagal connection. Scale bars: 100 µm (A), 40 µm (C &amp; D)." figureDoi="http://doi.org/10.5281/zenodo.7918726" httpUri="https://zenodo.org/record/7918726/files/figure.png" pageId="1" pageNumber="414">Fig. 1A</figureCitation>
): Wider than long; vertex long,
<emphasis box="[730,742,981,1006]" italics="true" pageId="1" pageNumber="414">c</emphasis>
. 0.4 of total head length, tapering tOwards­ anteriOr­ end,­with­ small­ lateral­ knObs­(fig.­1A);­eYebridge­ Of­ three­ rOws­ Offacets, separated by one-facet diameter width; interocular suture present, obtuse; ocular bristles absent; frontal scar patch crown-shaped with median band of alveoli reaching interOcular­suture;­labellum­bulbOus,­setOse;­OnlY­first­palp­segment­present­On­specimen,­ 71­µm­lOng;­scape­cYlindrical,­pedicel­spherical,­0.8­×­length­Of­scape;­flagellOmereselOngate,­sYmmetricallY­nOdifOrm;­first­flagellOmere­with­nOde­verY­elOngate,­mOre­thanthree­times­as­lOng­as­internOde;­Other­flagellOmeres­with­internOde­as­lOng­as­nOde;­ each­flagellOmere­with­twO­ascOid­attachment­pOints,­ascOids­missing­frOm­specimen;­ sOme­flagellOmeres­with­spinifOrm­sensillae.
</paragraph>
<paragraph blockId="1.[181,1142,1310,1676]" pageId="1" pageNumber="414">
<emphasis box="[182,263,1310,1336]" italics="true" pageId="1" pageNumber="414">Thorax</emphasis>
: Anepisternum and dorsum densely setose; anepisternum with anterior spiracle present; anepimeron triangular without additional sclerite; laterotergite bare around margins; katepisternum and katepimeron bare; pronotum setose; metathoracal spiracle densely setose; coxae with longitudinal row of setae; legs missing.
</paragraph>
<paragraph blockId="1.[181,1142,1310,1676]" pageId="1" pageNumber="414">
<emphasis box="[182,240,1449,1474]" italics="true" pageId="1" pageNumber="414">Wing</emphasis>
(
<figureCitation box="[256,344,1448,1475]" captionStart="Fig" captionStartId="2.[144,177,1078,1100]" captionTargetBox="[174,1073,217,1061]" captionTargetId="figure-246@2.[163,1085,204,1075]" captionTargetPageId="2" captionText="Fig. 1. Potophila verrucosa, gen.­n­et­sp.­n,­hOlOtYpe­♂:­(A)­head­(dOrsal­view);­(B)­wing;­(C)­terminalia­ (laterodorsal view); (D) epandrium and proctiger (laterodorsal view).Abbreviations:a ap aedeagal apodeme; d distiphallus; gcx gonocoxite; gcx c gonocoxal condyles; gst gonostyle; hyp hypandrium; lk lateral knob; p paramere; pam parameral-aedeagal membrane; pp parameral projections; vepa ventral epandrial plate/aedeagal connection. Scale bars: 100 µm (A), 40 µm (C &amp; D)." figureDoi="http://doi.org/10.5281/zenodo.7918726" httpUri="https://zenodo.org/record/7918726/files/figure.png" pageId="1" pageNumber="414">Fig. 1B</figureCitation>
):
<quantity box="[368,484,1448,1475]" metricMagnitude="-3" metricUnit="m" metricValue="3.135" pageId="1" pageNumber="414" unit="mm" value="3.135">3.135 mm</quantity>
long ×
<quantity box="[574,676,1448,1475]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="414" unit="mm" value="1.26">1.26 mm</quantity>
wide; ovoid; basal costal node enlarged; membrane unpatterned, without vestiture; Sc terminating blind in the wing, reaching origin of R
<subScript attach="right" box="[300,308,1531,1545]" fontSize="6" pageId="1" pageNumber="414">5</subScript>
; R
<subScript attach="left" box="[339,364,1531,1545]" fontSize="6" pageId="1" pageNumber="414">2+3</subScript>
not connected to R
<subScript attach="right" box="[573,581,1531,1545]" fontSize="6" pageId="1" pageNumber="414">4</subScript>
; radial fork in distal half of the wing, medial fork in basal half; basal part of M
<subScript attach="left" box="[464,472,1563,1577]" fontSize="6" pageId="1" pageNumber="414">2</subScript>
weakened; r-m and m-cu faint; R
<subScript attach="left" box="[838,846,1563,1577]" fontSize="6" pageId="1" pageNumber="414">5</subScript>
terminating slightly below apex of wing; CuA
<subScript attach="left" box="[397,405,1595,1609]" fontSize="6" pageId="1" pageNumber="414">2</subScript>
terminating slightly distally to middle of wing but basal to radial fork; bases of M
<subScript attach="left" box="[366,391,1627,1641]" fontSize="6" pageId="1" pageNumber="414">1+2</subScript>
and CuA
<subScript attach="left" box="[499,507,1627,1641]" fontSize="6" pageId="1" pageNumber="414">2</subScript>
expanded; basal cells terminating close to base of wing.
<emphasis box="[181,287,1650,1676]" italics="true" pageId="1" pageNumber="414">Abdomen</emphasis>
without special features.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7918726" ID-Zenodo-Dep="7918726" httpUri="https://zenodo.org/record/7918726/files/figure.png" pageId="2" pageNumber="415" startId="2.[144,177,1078,1100]" targetBox="[174,1073,217,1061]" targetPageId="2" targetType="figure">
<paragraph blockId="2.[144,1110,1078,1219]" pageId="2" pageNumber="415">
Fig. 1.
<emphasis box="[209,401,1078,1099]" italics="true" pageId="2" pageNumber="415">
<taxonomicName authorityName="Kvifte" authorityYear="2014" box="[209,397,1078,1099]" class="Insecta" family="Psychodidae" genus="Potophila" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="verrucosa">Potophila verrucosa</taxonomicName>
,
</emphasis>
gen.­n­et­sp.­n,­hOlOtYpe­♂:­(A)­head­(dOrsal­view);­(B)­wing;­(C)­terminalia­ (laterodorsal view); (D) epandrium and proctiger (laterodorsal view).Abbreviations: a ap aedeagal apodeme; d distiphallus; gcx gonocoxite; gcx c gonocoxal condyles; gst gonostyle; hyp hypandrium; lk lateral knob; p paramere; pam parameral-aedeagal membrane; pp parameral projections; vepa ventral epandrial plate/aedeagal connection. Scale bars: 100 µm (A), 40 µm (C &amp; D).
</paragraph>
</caption>
<paragraph blockId="2.[144,1105,1265,1675]" pageId="2" pageNumber="415">
<taxonomicName box="[144,266,1265,1291]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="415" phylum="Tracheophyta" rank="genus">
<emphasis box="[144,266,1265,1291]" italics="true" pageId="2" pageNumber="415">Terminalia</emphasis>
</taxonomicName>
(
<figureCitation box="[281,399,1265,1291]" captionStart="Fig" captionStartId="2.[144,177,1078,1100]" captionTargetBox="[174,1073,217,1061]" captionTargetId="figure-246@2.[163,1085,204,1075]" captionTargetPageId="2" captionText="Fig. 1. Potophila verrucosa, gen.­n­et­sp.­n,­hOlOtYpe­♂:­(A)­head­(dOrsal­view);­(B)­wing;­(C)­terminalia­ (laterodorsal view); (D) epandrium and proctiger (laterodorsal view).Abbreviations:a ap aedeagal apodeme; d distiphallus; gcx gonocoxite; gcx c gonocoxal condyles; gst gonostyle; hyp hypandrium; lk lateral knob; p paramere; pam parameral-aedeagal membrane; pp parameral projections; vepa ventral epandrial plate/aedeagal connection. Scale bars: 100 µm (A), 40 µm (C &amp; D)." figureDoi="http://doi.org/10.5281/zenodo.7918726" httpUri="https://zenodo.org/record/7918726/files/figure.png" pageId="2" pageNumber="415">Fig. 1C, D</figureCitation>
): With hypandrium narrow, straight; gonostyle 1.5 × length of gonocoxite; gonostyle basally with a cluster of lateral setae, with scattered spiniform sensilla along length; gonocoxal condyles triangular in dorsal and ventral views; aedeagus asymmetrical; basiphallus broad in dorsal view, connected laterally to parameres by parameral-aedeagal membrane, medially to distiphallus, morphodorsally with pointed connection to ventral epandrial plate; distiphallus comprised of two lateral phallomeres and morphodorsal phallic membrane forming duct; distiphallus twisted 90° to the left; parameres triangular with a morphoventral projection carrying small, verruca-like rugae; epandrium (
<figureCitation box="[357,444,1521,1547]" captionStart="Fig" captionStartId="2.[144,177,1078,1100]" captionTargetBox="[174,1073,217,1061]" captionTargetId="figure-246@2.[163,1085,204,1075]" captionTargetPageId="2" captionText="Fig. 1. Potophila verrucosa, gen.­n­et­sp.­n,­hOlOtYpe­♂:­(A)­head­(dOrsal­view);­(B)­wing;­(C)­terminalia­ (laterodorsal view); (D) epandrium and proctiger (laterodorsal view).Abbreviations:a ap aedeagal apodeme; d distiphallus; gcx gonocoxite; gcx c gonocoxal condyles; gst gonostyle; hyp hypandrium; lk lateral knob; p paramere; pam parameral-aedeagal membrane; pp parameral projections; vepa ventral epandrial plate/aedeagal connection. Scale bars: 100 µm (A), 40 µm (C &amp; D)." figureDoi="http://doi.org/10.5281/zenodo.7918726" httpUri="https://zenodo.org/record/7918726/files/figure.png" pageId="2" pageNumber="415">Fig. 1D</figureCitation>
) broader than long; with single pseudospiracular opening; subepandrial plate large, semicircular in dorsal view; hypoproct broadly U-shaped, densely pilose, covering bases of epandrial claspers; epiproct small, oval; epandrial claspers curved, carrying three distal tenacula in one row; tenacula spatulate, 87 µm long, tips slightly expanded.
</paragraph>
<paragraph blockId="3.[181,1147,203,249]" pageId="3" pageNumber="416">
<materialsCitation collectingDate="1990-11-23" collectionCode="ZMB" collectorName="Tanzania Expedition" country="Tanzania" location="TANZANIA" pageId="3" pageNumber="416" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[181,268,203,225]" pageId="3" pageNumber="416">HOlOtYpe</typeStatus>
:­
<specimenCount box="[275,292,204,225]" pageId="3" pageNumber="416" type="male"></specimenCount>
­
<collectingCountry box="[296,414,204,225]" name="Tanzania" pageId="3" pageNumber="416">TANZANIA</collectingCountry>
:­Tanga­regiOn,­West­Usambara­Mts,­Mazumbai­fOrest­reserve,­1440­m,­
<date box="[1040,1138,203,225]" pageId="3" pageNumber="416" value="1990-11-23">
<collectingDate box="[1040,1138,203,225]" pageId="3" pageNumber="416" value="1990-11-23">23.Xi.1990</collectingDate>
</date>
,­
<collectorName box="[181,368,227,249]" pageId="3" pageNumber="416">Tanzania Expedition</collectorName>
leg,
<collectingMethod box="[413,503,228,249]" pageId="3" pageNumber="416">sweep net</collectingMethod>
(
<collectionCode box="[515,569,228,249]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="416" type="Museum">ZMB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>