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<document id="375E6904428DAF1647EBB1D18B7CE273" ID-CLB-Dataset="28412" ID-DOI="10.11646/zootaxa.4565.1.1" ID-GBIF-Dataset="e9ffcfc7-9f7c-4212-ae6e-b842b31de6e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="2589489" ID-ZooBank="B9919693-2478-4089-B82E-C25EDFDB2A96" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552246080949" checkinUser="plazi" docAuthor="Chen, Hao, Zhang, Hui, Chen, Yongxia &amp; Freyhof, Jörg" docDate="2019" docId="5212F95FFFDBAA6EB5CAFF61FBB63185" docLanguage="en" docName="zootaxa.4565.1.1.pdf" docOrigin="Zootaxa 4565 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Barbatula toni" docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="AE2B8127FFDCAA67B55DFFF6FF9E324D" masterDocTitle="A review of the Barbatula loaches (Teleostei: Nemacheilidae) from north-eastern China, with the description of four new species" masterLastPageNumber="36" masterPageNumber="1" pageNumber="8" updateTime="1698701226448" updateUser="ExternalLinkService">
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<mods:title id="791BDE03FBBB22DF1C3F120C6AE2468E">A review of the Barbatula loaches (Teleostei: Nemacheilidae) from north-eastern China, with the description of four new species</mods:title>
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<mods:namePart id="0451DB12B3047414714C4EF75FF28ED0">Chen, Hao</mods:namePart>
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<mods:namePart id="C42FF60A27D5BAFDA300BC64D0D5BC7F">Zhang, Hui</mods:namePart>
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<mods:namePart id="129A8B0C2061DE89F4AF4A8E82D4A070">Chen, Yongxia</mods:namePart>
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<treatment id="5212F95FFFDBAA6EB5CAFF61FBB63185" ID-DOI="http://doi.org/10.5281/zenodo.5923813" ID-GBIF-Taxon="154366716" ID-Zenodo-Dep="5923813" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185" httpUri="http://treatment.plazi.org/id/5212F95FFFDBAA6EB5CAFF61FBB63185" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
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<paragraph id="DA044849FFDBAA60B5CAFF61FDB932FF" blockId="7.[151,551,151,211]" box="[151,551,151,178]" pageId="7" pageNumber="8">
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<taxonomicName id="1DBB33CAFFDBAA60B5CAFF61FDB932FF" ID-CoL="68C4Y" authority="(Dybowski, 1869)" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[151,551,151,178]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFDBAA60B5CAFF61FDB932FF" bold="true" box="[151,551,151,178]" pageId="7" pageNumber="8">
<emphasis id="E8CF945BFFDBAA60B5CAFF61FED932FC" bold="true" box="[151,327,151,177]" italics="true" pageId="7" pageNumber="8">Barbatula toni</emphasis>
(
<bibRefCitation id="BE2A35B8FFDBAA60B40AFF6EFD8132FF" author="Dybowski, B." box="[343,543,152,178]" pageId="7" pageNumber="8" pagination="1 - 25" refId="ref16083" refString="Dybowski, B. (1869) Fishes of the Baikal system. Izvestiya Sibirskogo Otdeleniya Russkogo Geografi cheskogo Obshchtva, 7, 1 - 25. [in Russian]" type="journal article" year="1869">Dybowski, 1869</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="92A11BC2FFDBAA60B5CAFF4CFE99329E" box="[151,263,186,211]" pageId="7" pageNumber="8" type="description">
<paragraph id="DA044849FFDBAA60B5CAFF4CFE99329E" blockId="7.[151,551,151,211]" box="[151,263,186,211]" pageId="7" pageNumber="8">
(
<figureCitation id="428054CCFFDBAA60B5C2FF4CFF61329E" box="[159,255,186,211]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStartId-0="7.[151,250,1920,1942]" captionStartId-1="8.[151,250,1588,1610]" captionStartId-2="9.[151,250,1397,1419]" captionTargetBox-0="[255,1364,808,1839]" captionTargetBox-1="[151,1436,223,1511]" captionTargetBox-2="[151,1426,1033,1374]" captionTargetId-0="figure@7.[151,1436,762,1898]" captionTargetId-1="figure@8.[151,1436,193,1566]" captionTargetId-2="figure@9.[151,1436,1023,1374]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 7. Barbatula toni, from the top: HU1608551, 74.9 mm SL; China: Heilongjiang prov.: Heilongjiang River drainage; IHB 58vii0183, 62.5 mm SL; China: Heilongjiang prov.: Heilongjiang River drainage; HU1609468, 65.9 mm SL; China: Inner Mongolia: Liao River drainage; IHB76ix9747, 71.7 mm SL; China: Inner Mongolia: Xilin River; IHB58v0058, 61.4 mm SL; China: Heilongjiang prov.: Heilongjiang River drainage." captionText-1="FIGURE 8. Barbatula toni, from the top: HU1608551, 74.9 mm SL; China: Heilongjiang prov.: Heilongjiang River drainage; HU1609468, 65.9 mm SL; China: Inner Mongolia: Liao River drainage; IHB76IX9747, 71.7 mm SL; China: Inner Mongolia: Xilin River; IHB58v0058, 61.4 mm SL; China: Heilongjiang prov.: Heilongjiang River drainage." captionText-2="FIGURE 9. Barbatula toni, IHB 58vii0183, 62.5 mm SL; China: Heilongjiang prov.: Heilongjiang River drainage. Scale bars: ab = 1 mm, c = 0.1 mm." httpUri-0="https://zenodo.org/record/2589503/files/figure.png" httpUri-1="https://zenodo.org/record/2589505/files/figure.png" httpUri-2="https://zenodo.org/record/2589507/files/figure.png" pageId="7" pageNumber="8">Fig. 79</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="DA044849FFDBAA60B5CAFEF4FB3B3355" blockId="7.[151,1189,258,280]" box="[151,1189,258,280]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="FAAB6F67FFDBAA60B5CAFEF4FB3B3355" box="[151,1189,258,280]" pageId="7" pageNumber="8">
<taxonomicName id="1DBB33CAFFDBAA60B5CAFEF4FE713355" authority="Dybowski, 1869: 957" authorityName="Dybowski" authorityPageNumber="957" authorityYear="1869" box="[151,495,258,280]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFDBAA60B5CAFEF4FE933355" box="[151,269,258,280]" italics="true" pageId="7" pageNumber="8">Cobitis toni</emphasis>
<treatmentCitation id="5B1A6E58FFDBAA60B449FEF4FE713355" author="Dybowski, B." box="[276,495,258,280]" page="957" pageId="7" pageNumber="8" year="1869">
<bibRefCitation id="BE2A35B8FFDBAA60B449FEF4FE213355" author="Dybowski, B." box="[276,447,258,280]" pageId="7" pageNumber="8" pagination="1 - 25" refId="ref16083" refString="Dybowski, B. (1869) Fishes of the Baikal system. Izvestiya Sibirskogo Otdeleniya Russkogo Geografi cheskogo Obshchtva, 7, 1 - 25. [in Russian]" type="journal article" year="1869">Dybowski, 1869</bibRefCitation>
: 957
</treatmentCitation>
</taxonomicName>
(
<collectingCountry id="A2AC08D9FFDBAA60B4A1FEF4FDFA3355" box="[508,612,258,280]" name="Mongolia" pageId="7" pageNumber="8">Mongolia</collectingCountry>
: Onon and Ingoda Rivers, Heilongjiang/Amur drainage).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="92A11BC2FFDBAA6FB5CAFEB3FEBA35CC" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="DA044849FFDBAA60B5CAFEB3FAB9308B" blockId="7.[151,1437,325,710]" pageId="7" pageNumber="8">
<emphasis id="E8CF945BFFDBAA60B5CAFEB3FE1C3313" bold="true" box="[151,386,325,350]" pageId="7" pageNumber="8">Material examined.</emphasis>
HU 160855052, 1608554, 160849798, 6,
<quantity id="1D43E5ACFFDBAA60B6DCFEB3FBBB3310" box="[897,1061,325,350]" metricMagnitude="-2" metricUnit="m" metricValue="7.59" metricValueMax="8.03" metricValueMin="7.15" pageId="7" pageNumber="8" unit="mm" value="75.9" valueMax="80.3" valueMin="71.5">71.580.3 mm</quantity>
SL; FSJF 4043, 1,
<quantity id="1D43E5ACFFDBAA60B05BFEB3FAF53310" box="[1286,1387,325,350]" metricMagnitude="-2" metricUnit="m" metricValue="7.63" pageId="7" pageNumber="8" unit="mm" value="76.3">76.3 mm</quantity>
SL;
<materialsCitation id="6AD34214FFDBAA60B5CAFE9FFC7033CC" ID-GBIF-Occurrence="2012822359" box="[151,1006,361,386]" country="China" location="Chenbarhu" municipality="Hailar River" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Nei Mongol">
<collectingCountry id="A2AC08D9FFDBAA60B5CAFE9FFF4133CF" box="[151,223,361,386]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
:
<collectingRegion id="187F86ABFFDBAA60B5B5FE9CFE0433CF" box="[232,410,361,386]" country="China" name="Nei Mongol" pageId="7" pageNumber="8">Inner Mongolia</collectingRegion>
:
<collectingMunicipality id="3A60D233FFDBAA60B4F9FE9FFDB033CF" box="[420,558,361,386]" pageId="7" pageNumber="8">Hailar River</collectingMunicipality>
<quantity id="1D43E5ACFFDBAA60B768FE9FFDF333CF" box="[565,621,361,386]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="km" value="5.0">5 km</quantity>
south of
<location id="DF641E92FFDBAA60B789FE9FFCD133CF" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFDBAA60B789FE9FFCD133CF" box="[724,847,361,386]" country="China" municipality="Hailar River" name="Chenbarhu" pageId="7" pageNumber="8" stateProvince="Nei Mongol">Chenbarhu</location>
banner, upper
</materialsCitation>
<materialsCitation id="6AD34214FFDBAA60B6A9FE9FFC2633EB" ID-GBIF-Occurrence="2012822343" country="China" latitude="49.483334" location="River" longLatPrecision="1103" longitude="119.76667" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Heilongjiang">
<collectingRegion id="187F86ABFFDBAA60B6A9FE9FFB1833CF" box="[1012,1158,361,386]" country="China" name="Heilongjiang" pageId="7" pageNumber="8">Heilongjiang</collectingRegion>
<location id="DF641E92FFDBAA60B1D1FE9FFB5433CF" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFDBAA60B1D1FE9FFB5433CF" box="[1164,1226,361,386]" country="China" latitude="49.483334" longLatPrecision="1103" longitude="119.76667" name="River" pageId="7" pageNumber="8" stateProvince="Heilongjiang">River</location>
drainage,
<geoCoordinate id="BF8F2E8EFFDBAA60B062FE9FFA0233CF" box="[1343,1436,361,386]" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="925" value="49.483334">49°29N</geoCoordinate>
<geoCoordinate id="BF8F2E8EFFDBAA60B5CAFE7BFF6033EB" box="[151,254,397,422]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="925" value="119.76667">119°46E</geoCoordinate>
(24).—HU1608484, 1608492, 1608495, 3,
<quantity id="1D43E5ACFFDBAA60B7B6FE7BFC1333E8" box="[747,909,397,422]" metricMagnitude="-2" metricUnit="m" metricValue="5.555" metricValueMax="6.13" metricValueMin="4.9799999999999995" pageId="7" pageNumber="8" unit="mm" value="55.55" valueMax="61.3" valueMin="49.8">49.861.3 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="6AD34214FFDBAA60B69EFE7BFE133387" ID-GBIF-Occurrence="2012822340" country="China" location="Harbin" municipality="Songhua River" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Heilongjiang">
<collectingCountry id="A2AC08D9FFDBAA60B69EFE7BFB9433EB" box="[963,1034,397,422]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
:
<collectingRegion id="187F86ABFFDBAA60B148FE7BFB3933EB" box="[1045,1191,397,422]" country="China" name="Heilongjiang" pageId="7" pageNumber="8">Heilongjiang</collectingRegion>
prov.:
<collectingMunicipality id="3A60D233FFDBAA60B1A8FE7BFA0233EB" box="[1269,1436,397,422]" pageId="7" pageNumber="8">Songhua River</collectingMunicipality>
west of
<location id="DF641E92FFDBAA60B5ACFE47FEDC3387" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFDBAA60B5ACFE47FEDC3387" box="[241,322,433,458]" country="China" municipality="Songhua River" name="Harbin" pageId="7" pageNumber="8" stateProvince="Heilongjiang">Harbin</location>
, lower
</materialsCitation>
<materialsCitation id="6AD34214FFDBAA60B4C8FE47FEDC33A3" ID-GBIF-Occurrence="2012822345" collectingDate="2017-04" country="China" location="River" longitude="126.78333" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Heilongjiang">
<collectingRegion id="187F86ABFFDBAA60B4C8FE47FDB93387" box="[405,551,433,458]" country="China" name="Heilongjiang" pageId="7" pageNumber="8">Heilongjiang</collectingRegion>
<location id="DF641E92FFDBAA60B772FE47FDF33387" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFDBAA60B772FE47FDF33387" box="[559,621,433,458]" country="China" longitude="126.78333" name="River" pageId="7" pageNumber="8" stateProvince="Heilongjiang">River</location>
drainage, 45°74N
<geoCoordinate id="BF8F2E8EFFDBAA60B614FE47FC2F3387" box="[841,945,433,458]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="925" value="126.78333">126°47E</geoCoordinate>
(23); Hao Chen,
<date id="AE056E89FFDBAA60B12AFE47FB773387" box="[1143,1257,433,458]" pageId="7" pageNumber="8" value="2017-04">
<collectingDate id="BE419761FFDBAA60B12AFE47FB773387" box="[1143,1257,433,458]" pageId="7" pageNumber="8" value="2017-04">Apr. 2017</collectingDate>
</date>
.—HU1608438, 1,
<quantity id="1D43E5ACFFDBAA60B5EEFE23FE8633A0" box="[179,280,469,494]" metricMagnitude="-2" metricUnit="m" metricValue="5.86" pageId="7" pageNumber="8" unit="mm" value="58.6">58.6 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="6AD34214FFDBAA60B413FE23FC7433A0" ID-GBIF-Occurrence="2012822353" box="[334,1002,469,494]" country="China" location="Tahe" municipality="Huma River" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Heilongjiang">
<collectingCountry id="A2AC08D9FFDBAA60B413FE23FE0B33A3" box="[334,405,469,494]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
:
<collectingRegion id="187F86ABFFDBAA60B4FDFE23FDAC33A3" box="[416,562,469,494]" country="China" name="Heilongjiang" pageId="7" pageNumber="8">Heilongjiang</collectingRegion>
prov.:
<collectingMunicipality id="3A60D233FFDBAA60B7DFFE20FC9233A3" box="[642,780,469,494]" pageId="7" pageNumber="8">Huma River</collectingMunicipality>
east of
<location id="DF641E92FFDBAA60B638FE23FC0033A3" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFDBAA60B638FE23FC0033A3" box="[869,926,469,494]" country="China" municipality="Huma River" name="Tahe" pageId="7" pageNumber="8" stateProvince="Heilongjiang">Tahe</location>
, upper
</materialsCitation>
<materialsCitation id="6AD34214FFDBAA60B6ACFE23FA06305C" ID-GBIF-Occurrence="2012822346" collectingDate="2017-05" country="China" latitude="52.533333" location="River" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Heilongjiang">
<collectingRegion id="187F86ABFFDBAA60B6ACFE23FB1D33A3" box="[1009,1155,469,494]" country="China" name="Heilongjiang" pageId="7" pageNumber="8">Heilongjiang</collectingRegion>
<location id="DF641E92FFDBAA60B1D7FE23FB5633A3" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFDBAA60B1D7FE23FB5633A3" box="[1162,1224,469,494]" country="China" latitude="52.533333" name="River" pageId="7" pageNumber="8" stateProvince="Heilongjiang">River</location>
drainage,
<geoCoordinate id="BF8F2E8EFFDBAA60B062FE23FA0233A3" box="[1343,1436,469,494]" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="925" value="52.533333">52°32N</geoCoordinate>
124°71E (25); Yongxia Chen,
<date id="AE056E89FFDBAA60B75BFE0CFDE1305F" box="[518,639,505,530]" pageId="7" pageNumber="8" value="2017-05">
<collectingDate id="BE419761FFDBAA60B75BFE0CFDE1305F" box="[518,639,505,530]" pageId="7" pageNumber="8" value="2017-05">May 2017</collectingDate>
</date>
.—HU1608907, 1608910, 1608913, 1609468, 4,
<quantity id="1D43E5ACFFDBAA60B19DFE0FFAF6305C" box="[1216,1384,505,530]" metricMagnitude="-2" metricUnit="m" metricValue="6.615" metricValueMax="7.12" metricValueMin="6.11" pageId="7" pageNumber="8" unit="mm" value="66.15" valueMax="71.2" valueMin="61.1">61.171.2 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="6AD34214FFDBAA60B5CAFDEBFF713033" ID-GBIF-Occurrence="2012822351" collectingDate="1958-07" collectingDateMax="2017-05" collectingDateMin="1958-07" collectionCode="IHB" country="China" county="Yin River" latitude="42.566666" location="upper Liao River" municipality="Chutoulangzhen" pageId="7" pageNumber="8" specimenCode="IHB 58" specimenCount="1" stateProvince="Nei Mongol">
<collectingCountry id="A2AC08D9FFDBAA60B5CAFDEBFF41307B" box="[151,223,541,566]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
:
<collectingRegion id="187F86ABFFDBAA60B5B3FDE8FE36307B" box="[238,424,541,566]" country="China" name="Nei Mongol" pageId="7" pageNumber="8">Inner Mongolia</collectingRegion>
:
<collectingCounty id="336530C5FFDBAA60B4E5FDEBFDB2307B" box="[440,556,541,566]" pageId="7" pageNumber="8">Yin River</collectingCounty>
<quantity id="1D43E5ACFFDBAA60B764FDEBFDE6307B" box="[569,632,541,566]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="km" value="5.0">5 km</quantity>
north-east of
<collectingMunicipality id="3A60D233FFDBAA60B67BFDEBFC7E307B" box="[806,992,541,566]" pageId="7" pageNumber="8">Chutoulangzhen</collectingMunicipality>
,
<location id="DF641E92FFDBAA60B6AFFDE8FB23307B" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFDBAA60B6AFFDE8FB23307B" box="[1010,1213,541,566]" country="China" county="Yin River" latitude="42.566666" municipality="Chutoulangzhen" name="upper Liao River" pageId="7" pageNumber="8" stateProvince="Nei Mongol">upper Liao River</location>
drainage,
<geoCoordinate id="BF8F2E8EFFDBAA60B062FDEBFA02307B" box="[1343,1436,541,566]" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="925" value="42.566666">42°34N</geoCoordinate>
118°72E (6);
<location id="DF641E92FFDBAA60B465FDB4FE333017" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFDBAA60B465FDB4FE333017" box="[312,429,577,602]" country="China" county="Yin River" latitude="42.566666" municipality="Chutoulangzhen" name="Hao Chen" pageId="7" pageNumber="8" stateProvince="Nei Mongol">Hao Chen</location>
,
<date id="AE056E89FFDBAA60B4E4FDB4FDAF3014" box="[441,561,577,602]" pageId="7" pageNumber="8" value="2017-05">
<collectingDate id="BE419761FFDBAA60B4E4FDB4FDAF3014" box="[441,561,577,602]" pageId="7" pageNumber="8" value="2017-05">May 2017</collectingDate>
</date>
.
<emphasis id="E8CF945BFFDBAA60B769FDB7FDCE3017" bold="true" box="[564,592,577,602]" pageId="7" pageNumber="8"></emphasis>
<specimenCode id="8A1DE032FFDBAA60B70DFDB4FD3B3017" box="[592,677,577,602]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="7" pageNumber="8">
IHB
<date id="AE056E89FFDBAA60B7D4FDB7FD3B3017" box="[649,677,577,602]" pageId="7" pageNumber="8" value="1958-07">58</date>
</specimenCode>
vii0180,
<date id="AE056E89FFDBAA60B657FDB7FCDA3017" box="[778,836,577,602]" pageId="7" pageNumber="8" value="1958-07">
<collectingDate id="BE419761FFDBAA60B657FDB7FCDA3017" box="[778,836,577,602]" pageId="7" pageNumber="8" value="1958-07">58vii</collectingDate>
</date>
0086,
<date id="AE056E89FFDBAA60B6D6FDB7FC5B3017" box="[907,965,577,602]" pageId="7" pageNumber="8" value="1958-07">
<collectingDate id="BE419761FFDBAA60B6D6FDB7FC5B3017" box="[907,965,577,602]" pageId="7" pageNumber="8" value="1958-07">58vii</collectingDate>
</date>
0183,
<date id="AE056E89FFDBAA60B151FDB7FBD83017" box="[1036,1094,577,602]" pageId="7" pageNumber="8" value="1958-07">
<collectingDate id="BE419761FFDBAA60B151FDB7FBD83017" box="[1036,1094,577,602]" pageId="7" pageNumber="8" value="1958-07">58vii</collectingDate>
</date>
0085,
<date id="AE056E89FFDBAA60B1D3FDB7FB593017" box="[1166,1223,577,602]" pageId="7" pageNumber="8" value="1958-07">
<collectingDate id="BE419761FFDBAA60B1D3FDB7FB593017" box="[1166,1223,577,602]" pageId="7" pageNumber="8" value="1958-07">58vii</collectingDate>
</date>
0586, 5,
<quantity id="1D43E5ACFFDBAA60B071FDB7FF5D3030" metricMagnitude="-2" metricUnit="m" metricValue="6.43" metricValueMax="6.81" metricValueMin="6.05" pageId="7" pageNumber="8" unit="mm" value="64.3" valueMax="68.1" valueMin="60.5">60.568.1 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="6AD34214FFDBAA60B5A6FD93FC9D3030" ID-GBIF-Occurrence="2012822349" box="[251,771,613,638]" country="China" location="Ergun River" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Heilongjiang">
<collectingCountry id="A2AC08D9FFDBAA60B5A6FD93FEDC3033" box="[251,322,613,638]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
:
<collectingRegion id="187F86ABFFDBAA60B413FD93FE7E3033" box="[334,480,613,638]" country="China" name="Heilongjiang" pageId="7" pageNumber="8">Heilongjiang</collectingRegion>
prov.:
<location id="DF641E92FFDBAA60B76CFD90FD253033" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFDBAA60B76CFD90FD253033" box="[561,699,613,638]" country="China" name="Ergun River" pageId="7" pageNumber="8" stateProvince="Heilongjiang">Ergun River</location>
upper
</materialsCitation>
<materialsCitation id="6AD34214FFDBAA60B656FD93FEFD30EF" ID-GBIF-Occurrence="2012822358" collectionCode="IHB" country="China" location="River" pageId="7" pageNumber="8" specimenCode="IHB 58" specimenCount="1" stateProvince="Heilongjiang">
<collectingRegion id="187F86ABFFDBAA60B656FD93FC033033" box="[779,925,613,638]" country="China" name="Heilongjiang" pageId="7" pageNumber="8">Heilongjiang</collectingRegion>
<location id="DF641E92FFDBAA60B6F8FD93FC7D3033" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFDBAA60B6F8FD93FC7D3033" box="[933,995,613,638]" country="China" name="River" pageId="7" pageNumber="8" stateProvince="Heilongjiang">River</location>
drainage (26).—
<specimenCode id="8A1DE032FFDBAA60B1F9FD93FB673033" box="[1188,1273,613,638]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="7" pageNumber="8">
IHB
<date id="AE056E89FFDBAA60B183FD93FB673033" box="[1246,1273,613,638]" pageId="7" pageNumber="8" value="1958-05">58</date>
</specimenCode>
v005776, 20, 41.071.0 mm SL
</materialsCitation>
;
<materialsCitation id="6AD34214FFDBAA60B433FD7FFC4F30EC" ID-GBIF-Occurrence="2012822348" box="[366,977,649,674]" country="China" location="Emur River" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Heilongjiang">
<collectingCountry id="A2AC08D9FFDBAA60B433FD7FFE2B30EF" box="[366,437,649,674]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
:
<collectingRegion id="187F86ABFFDBAA60B4E2FD7FFDCF30EF" box="[447,593,649,674]" country="China" name="Heilongjiang" pageId="7" pageNumber="8">Heilongjiang</collectingRegion>
prov.:
<location id="DF641E92FFDBAA60B7C3FD7CFCBF30EF" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFDBAA60B7C3FD7CFCBF30EF" box="[670,801,649,674]" country="China" name="Emur River" pageId="7" pageNumber="8" stateProvince="Heilongjiang">Emur River</location>
at Mohe, upper
</materialsCitation>
<materialsCitation id="6AD34214FFDBAA60B68AFD7FFD4C308B" ID-GBIF-Occurrence="2012822356" collectingDate="1976-09" collectionCode="IHB" country="China" location="River" pageId="7" pageNumber="8" specimenCode="IHB 76" specimenCount="1" stateProvince="Heilongjiang">
<collectingRegion id="187F86ABFFDBAA60B68AFD7FFBF730EF" box="[983,1129,649,674]" country="China" name="Heilongjiang" pageId="7" pageNumber="8">Heilongjiang</collectingRegion>
<location id="DF641E92FFDBAA60B132FD7FFB3330EF" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFDBAA60B132FD7FFB3330EF" box="[1135,1197,649,674]" country="China" name="River" pageId="7" pageNumber="8" stateProvince="Heilongjiang">River</location>
drainage (27).—
<specimenCode id="8A1DE032FFDBAA60B037FD7CFF2D308B" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="7" pageNumber="8">
IHB
<date id="AE056E89FFDBAA60B5CAFD5BFF2D308B" box="[151,179,685,710]" pageId="7" pageNumber="8" value="1976-09">76</date>
</specimenCode>
ix974348,
<date id="AE056E89FFDBAA60B464FD5BFEF5308B" box="[313,363,685,710]" pageId="7" pageNumber="8" value="1976-09">
<collectingDate id="BE419761FFDBAA60B464FD5BFEF5308B" box="[313,363,685,710]" pageId="7" pageNumber="8" value="1976-09">76ix</collectingDate>
</date>
975157, 13,
<quantity id="1D43E5ACFFDBAA60B759FD5BFD393088" box="[516,679,685,710]" metricMagnitude="-2" metricUnit="m" metricValue="6.215" metricValueMax="7.22" metricValueMin="5.21" pageId="7" pageNumber="8" unit="mm" value="62.15" valueMax="72.2" valueMin="52.1">52.172.2 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="6AD34214FFDBAA60B783FD5BFABD308B" ID-GBIF-Occurrence="2012822339" box="[734,1315,685,710]" country="China" location="Xilin River" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Nei Mongol">
<collectingCountry id="A2AC08D9FFDBAA60B783FD5BFCBB308B" box="[734,805,685,710]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
:
<collectingRegion id="187F86ABFFDBAA60B672FD58FC7C308B" box="[815,994,685,710]" country="China" name="Nei Mongol" pageId="7" pageNumber="8">Inner Mongolia</collectingRegion>
;
<location id="DF641E92FFDBAA60B6B0FD5BFBF5308B" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFDBAA60B6B0FD5BFBF5308B" box="[1005,1131,685,710]" country="China" name="Xilin River" pageId="7" pageNumber="8" stateProvince="Nei Mongol">Xilin River</location>
at Xilin Gol (8)
</materialsCitation>
.
</paragraph>
<caption id="8EC418C1FFDBAA60B5CAF876FD4D35BF" httpUri="https://zenodo.org/record/2589503/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1920,1942]" targetBox="[255,1364,808,1839]" targetPageId="7">
<paragraph id="DA044849FFDBAA60B5CAF876FD4D35BF" blockId="7.[151,1436,1920,2034]" pageId="7" pageNumber="8">
<emphasis id="E8CF945BFFDBAA60B5CAF876FE8D35D8" bold="true" box="[151,275,1920,1942]" pageId="7" pageNumber="8">FIGURE 7.</emphasis>
<taxonomicName id="1DBB33CAFFDBAA60B447F876FE3035DB" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[282,430,1920,1942]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFDBAA60B447F876FE3035DB" box="[282,430,1920,1942]" italics="true" pageId="7" pageNumber="8">Barbatula toni</emphasis>
</taxonomicName>
, from the top: HU1608551, 74.9 mm SL; China: Heilongjiang prov.: Heilongjiang River drainage; IHB 58vii0183, 62.5 mm SL; China: Heilongjiang prov.: Heilongjiang River drainage; HU1609468, 65.9 mm SL; China: Inner Mongolia: Liao River drainage; IHB76ix9747, 71.7 mm SL; China: Inner Mongolia: Xilin River; IHB58v0058, 61.4 mm SL; China: Heilongjiang prov.: Heilongjiang River drainage.
</paragraph>
</caption>
<caption id="8EC418C1FFD4AA6FB5CAF9C2FBF134CA" httpUri="https://zenodo.org/record/2589505/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1588,1610]" targetBox="[151,1436,223,1511]" targetPageId="8">
<paragraph id="DA044849FFD4AA6FB5CAF9C2FBF134CA" blockId="8.[151,1437,1588,1671]" pageId="8" pageNumber="9">
<emphasis id="E8CF945BFFD4AA6FB5CAF9C2FE8D3404" bold="true" box="[151,275,1588,1610]" pageId="8" pageNumber="9">FIGURE 8.</emphasis>
<taxonomicName id="1DBB33CAFFD4AA6FB447F9C2FE303407" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[282,430,1588,1610]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFD4AA6FB447F9C2FE303407" box="[282,430,1588,1610]" italics="true" pageId="8" pageNumber="9">Barbatula toni</emphasis>
</taxonomicName>
, from the top: HU1608551, 74.9 mm SL; China: Heilongjiang prov.: Heilongjiang River drainage; HU1609468, 65.9 mm SL; China: Inner Mongolia: Liao River drainage; IHB76IX9747, 71.7 mm SL; China: Inner Mongolia: Xilin River; IHB58v0058, 61.4 mm SL; China: Heilongjiang prov.: Heilongjiang River drainage.
</paragraph>
</caption>
<paragraph id="DA044849FFD4AA6FB59AF942FEBA35CC" blockId="8.[151,1437,1716,2029]" pageId="8" pageNumber="9">
<emphasis id="E8CF945BFFD4AA6FB59AF942FC8E3480" bold="true" box="[199,784,1716,1741]" pageId="8" pageNumber="9">Material used in the molecular genetic analysis.</emphasis>
HU1608484;
<collectingCountry id="A2AC08D9FFD4AA6FB6EBF942FC633480" box="[950,1021,1716,1741]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD4AA6FB156F942FB033480" box="[1035,1181,1716,1741]" country="China" name="Heilongjiang" pageId="8" pageNumber="9">Heilongjiang</collectingRegion>
prov.: Songhua River west of Harbin City, lower
<collectingRegion id="187F86ABFFD4AA6FB484F92EFDF534BC" box="[473,619,1752,1777]" country="China" name="Heilongjiang" pageId="8" pageNumber="9">Heilongjiang</collectingRegion>
River drainage, 45°74N
<geoCoordinate id="BF8F2E8EFFD4AA6FB6C5F92EFB9F34BC" box="[920,1025,1752,1777]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="126.78333">126°47E</geoCoordinate>
(23). (GenBank accession number:
<accessionNumber id="C5E8D5AAFFD4AA6FB5CAF90AFE883558" box="[151,278,1788,1813]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY451914" pageId="8" pageNumber="9">KY451914</accessionNumber>
).—FSJF 4043;
<collectingCountry id="A2AC08D9FFD4AA6FB48FF90AFD873558" box="[466,537,1788,1813]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD4AA6FB774F90BFD7C3558" box="[553,738,1788,1813]" country="China" name="Nei Mongol" pageId="8" pageNumber="9">Inner Mongolia</collectingRegion>
: Hailar River, upper
<collectingRegion id="187F86ABFFD4AA6FB6BCF90AFBED3558" box="[993,1139,1788,1813]" country="China" name="Heilongjiang" pageId="8" pageNumber="9">Heilongjiang</collectingRegion>
River drainage,
<geoCoordinate id="BF8F2E8EFFD4AA6FB062F90AFA023558" box="[1343,1436,1788,1813]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="49.483334">49°29N</geoCoordinate>
<geoCoordinate id="BF8F2E8EFFD4AA6FB5CAF8D6FF603574" box="[151,254,1824,1849]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="119.76667">119°46E</geoCoordinate>
(24). (GenBank accession number:
<accessionNumber id="C5E8D5AAFFD4AA6FB7CFF8D6FC8D3574" box="[658,787,1824,1849]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY451916" pageId="8" pageNumber="9">KY451916</accessionNumber>
).—
<materialsCitation id="6AD34214FFD4AA6FB666F8D6FEBE35CC" ID-GBIF-Occurrence="2012822342" country="China" county="Yin River" latitude="42.566666" location="upper Liao River" municipality="Chutoulangzhen" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Nei Mongol">
HU1608910;
<collectingCountry id="A2AC08D9FFD4AA6FB689F8D6FB853574" box="[980,1051,1824,1849]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD4AA6FB17BF8D7FB473574" box="[1062,1241,1824,1849]" country="China" name="Nei Mongol" pageId="8" pageNumber="9">Inner Mongolia</collectingRegion>
:
<collectingCounty id="336530C5FFD4AA6FB1BEF8D6FACE3574" box="[1251,1360,1824,1849]" pageId="8" pageNumber="9">Yin River</collectingCounty>
northeast of
<collectingMunicipality id="3A60D233FFD4AA6FB5A7F8B2FE2A3510" box="[250,436,1860,1885]" pageId="8" pageNumber="9">Chutoulangzhen</collectingMunicipality>
,
<location id="DF641E92FFD4AA6FB494F8B3FD053510" LSID="urn:lsid:plazi:treatment:5212F95FFFDBAA6EB5CAFF61FBB63185:DF641E92FFD4AA6FB494F8B3FD053510" box="[457,667,1860,1885]" country="China" county="Yin River" latitude="42.566666" municipality="Chutoulangzhen" name="upper Liao River" pageId="8" pageNumber="9" stateProvince="Nei Mongol">upper Liao River</location>
drainage,
<geoCoordinate id="BF8F2E8EFFD4AA6FB67EF8B2FC1E3510" box="[803,896,1860,1885]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="42.566666">42°34N</geoCoordinate>
118°72E (6). (GenBank accession number:
<accessionNumber id="C5E8D5AAFFD4AA6FB5CAF89EFE8635CC" box="[151,280,1896,1921]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY451915" pageId="8" pageNumber="9">KY451915</accessionNumber>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92A11BC2FFD4AA6EB59AF87AFA0E33B8" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="DA044849FFD4AA6EB59AF87AFF7A3350" blockId="8.[151,1437,1716,2029]" lastBlockId="9.[151,1436,151,968]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="E8CF945BFFD4AA6FB59AF87AFEDF35E8" bold="true" box="[199,321,1932,1957]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
<taxonomicName id="1DBB33CAFFD4AA6FB416F87AFE6C35E9" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[331,498,1932,1957]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFD4AA6FB416F87AFE6C35E9" box="[331,498,1932,1957]" italics="true" pageId="8" pageNumber="9">Barbatula toni</emphasis>
</taxonomicName>
is distinguished from the other species of
<taxonomicName id="1DBB33CAFFD4AA6FB6B9F87AFBC835E8" box="[996,1110,1932,1957]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="E8CF945BFFD4AA6FB6B9F87AFBC835E8" box="[996,1110,1932,1957]" italics="true" pageId="8" pageNumber="9">Barbatula</emphasis>
</taxonomicName>
in north-eastern
<collectingCountry id="A2AC08D9FFD4AA6FB040F87AFAFE35E8" box="[1309,1376,1932,1957]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
by a combination of characters, none of them unique: upper lip with a shallow median incision, its depth is 1020% of the width of the upper lip; lower lip with short lateral expansion, its length 3050% of the width of the upper lip (
<figureCitation id="428054CCFFD5AA6EB5C2FF61FF6D32FD" box="[159,243,151,176]" captionStart="FIGURE 9" captionStartId="9.[151,250,1397,1419]" captionTargetBox="[151,1426,1033,1374]" captionTargetId="figure@9.[151,1436,1023,1374]" captionTargetPageId="9" captionText="FIGURE 9. Barbatula toni, IHB 58vii0183, 62.5 mm SL; China: Heilongjiang prov.: Heilongjiang River drainage. Scale bars: ab = 1 mm, c = 0.1 mm." httpUri="https://zenodo.org/record/2589507/files/figure.png" pageId="9" pageNumber="10">Fig. 9a</figureCitation>
); upper jaw completely covered by the upper lip; snout blunt; nostrils widely spaced (
<figureCitation id="428054CCFFD5AA6EB18FFF61FAB632FD" box="[1234,1320,151,176]" captionStart="FIGURE 9" captionStartId="9.[151,250,1397,1419]" captionTargetBox="[151,1426,1033,1374]" captionTargetId="figure@9.[151,1436,1023,1374]" captionTargetPageId="9" captionText="FIGURE 9. Barbatula toni, IHB 58vii0183, 62.5 mm SL; China: Heilongjiang prov.: Heilongjiang River drainage. Scale bars: ab = 1 mm, c = 0.1 mm." httpUri="https://zenodo.org/record/2589507/files/figure.png" pageId="9" pageNumber="10">Fig. 9b</figureCitation>
); snout as long as the postorbital head length; dorsal profile convex; 7½ branched dorsal-fin rays; pelvic-fin origin in front of the vertical through the dorsal-fin origin, below the origin of the first dorsal-fin ray; and back and flank completely scaled.
</paragraph>
<paragraph id="DA044849FFD5AA6EB59AFED1FC05339D" blockId="9.[151,1436,151,968]" pageId="9" pageNumber="10">
<taxonomicName id="1DBB33CAFFD5AA6EB59AFED1FEF2330D" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[199,364,295,320]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFD5AA6EB59AFED1FEF2330D" box="[199,364,295,320]" italics="true" pageId="9" pageNumber="10">Barbatula toni</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="1DBB33CAFFD5AA6EB736FEDFFD5D330D" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[619,707,295,320]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nuda">
<emphasis id="E8CF945BFFD5AA6EB736FEDFFD5D330D" box="[619,707,295,320]" italics="true" pageId="9" pageNumber="10">B. nuda</emphasis>
</taxonomicName>
by having the nostrils widely spaced (vs. closely set), a shallow median incision in the upper lip, its depth 1020% of the width of the upper lip (vs. deep, 4060%), a short lateral expansion of the lower lip (vs. absent), upper jaw completely covered by the upper lip (vs. partly covered), pelvicfin origin in front of the vertical through the dorsal-fin origin (vs. below) and scales present on complete back and flank (vs. restricted to the back and flank behind the dorsal-fin base).
</paragraph>
<paragraph id="DA044849FFD5AA6EB59AFE2AFA0E33B8" blockId="9.[151,1436,151,968]" box="[199,1424,476,501]" pageId="9" pageNumber="10">
See
<tableCitation id="97397DF2FFD5AA6EB5ABFE2AFED733B9" box="[246,329,476,501]" captionStart="TABLE 3" captionStartId="31.[147,216,807,825]" captionText="TABLE 3. (Continued)" pageId="9" pageNumber="10">Table 3</tableCitation>
for the character states shown by the different species of
<taxonomicName id="1DBB33CAFFD5AA6EB690FE2AFBA133B8" box="[973,1087,476,501]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="E8CF945BFFD5AA6EB690FE2AFBA133B8" box="[973,1087,476,501]" italics="true" pageId="9" pageNumber="10">Barbatula</emphasis>
</taxonomicName>
found in north-eastern
<collectingCountry id="A2AC08D9FFD5AA6EB018FE2AFA1233B8" box="[1349,1420,476,501]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="92A11BC2FFD5AA6EB59AFE09FC5F30C8" pageId="9" pageNumber="10" type="distribution">
<paragraph id="DA044849FFD5AA6EB59AFE09FC5F30C8" blockId="9.[151,1436,151,968]" pageId="9" pageNumber="10">
<emphasis id="E8CF945BFFD5AA6EB59AFE09FEFE3055" bold="true" box="[199,352,511,536]" pageId="9" pageNumber="10">Distribution.</emphasis>
Widespread in the
<collectingRegion id="187F86ABFFD5AA6EB76BFE09FD563055" box="[566,712,511,536]" country="China" name="Heilongjiang" pageId="9" pageNumber="10">Heilongjiang</collectingRegion>
(Amur) River drainage in
<collectingCountry id="A2AC08D9FFD5AA6EB6ACFE09FBA93055" box="[1009,1079,511,536]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
,
<collectingCountry id="A2AC08D9FFD5AA6EB11CFE09FB123055" box="[1089,1164,511,536]" name="Russia" pageId="9" pageNumber="10">Russia</collectingCountry>
and
<collectingCountry id="A2AC08D9FFD5AA6EB19CFE09FAAC3055" box="[1217,1330,511,536]" name="Mongolia" pageId="9" pageNumber="10">Mongolia</collectingCountry>
. In
<collectingCountry id="A2AC08D9FFD5AA6EB004FE09FA023055" box="[1369,1436,511,536]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
known from the Ergun, Hailar, Songhua, and Huma Rivers in the
<collectingRegion id="187F86ABFFD5AA6EB6D6FDD2FB833070" box="[907,1053,548,573]" country="China" name="Heilongjiang" pageId="9" pageNumber="10">Heilongjiang</collectingRegion>
drainage as well as from the Yin River, which is a tributary of the Liao River in
<collectingRegion id="187F86ABFFD5AA6EB7F3FDBEFCC0302D" box="[686,862,583,608]" country="China" name="Nei Mongol" pageId="9" pageNumber="10">Inner Mongolia</collectingRegion>
which flow to the Bohai Sea and has its estuary at the Chinese city of Panjin (40°98N, 121°83E). See
<figureCitation id="428054CCFFD5AA6EB7BEFD9AFCDF30C9" box="[739,833,620,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1882,1904]" captionTargetBox="[173,1389,888,1818]" captionTargetId="figure@3.[151,1436,876,1861]" captionTargetPageId="3" captionText="FIGURE 2. Map showing the collection sites. Details for each number are listed in material examined." httpUri="https://zenodo.org/record/2589493/files/figure.png" pageId="9" pageNumber="10">Figure 2</figureCitation>
for details.
</paragraph>
</subSubSection>
<subSubSection id="92A11BC2FFD5AA6EB59AFD79FBB63185" pageId="9" pageNumber="10" type="discussion">
<paragraph id="DA044849FFD5AA6EB59AFD79FBA53110" blockId="9.[151,1436,151,968]" pageId="9" pageNumber="10">
<emphasis id="E8CF945BFFD5AA6EB59AFD79FEA530E5" bold="true" box="[199,315,655,680]" pageId="9" pageNumber="10">Remarks.</emphasis>
<taxonomicName id="1DBB33CAFFD5AA6EB41CFD79FE7B30E5" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[321,485,655,680]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFD5AA6EB41CFD79FE7B30E5" box="[321,485,655,680]" italics="true" pageId="9" pageNumber="10">Barbatula toni</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="BE2A35B8FFD5AA6EB7E9FD79FCE630E5" author="Dybowski, B." box="[692,888,655,680]" pageId="9" pageNumber="10" pagination="1 - 25" refId="ref16083" refString="Dybowski, B. (1869) Fishes of the Baikal system. Izvestiya Sibirskogo Otdeleniya Russkogo Geografi cheskogo Obshchtva, 7, 1 - 25. [in Russian]" type="journal article" year="1869">Dybowski (1869)</bibRefCitation>
from the Onon and Ingoda Rivers in the Russian
<collectingRegion id="187F86ABFFD5AA6EB5CAFD42FEB73080" box="[151,297,692,717]" country="China" name="Heilongjiang" pageId="9" pageNumber="10">Heilongjiang</collectingRegion>
River drainage. For a long period of time, this was the catch-all name for many Asian
<taxonomicName id="1DBB33CAFFD5AA6EB077FD42FA023080" box="[1322,1436,692,717]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="E8CF945BFFD5AA6EB077FD42FA023080" box="[1322,1436,692,717]" italics="true" pageId="9" pageNumber="10">Barbatula</emphasis>
</taxonomicName>
species.
<taxonomicName id="1DBB33CAFFD5AA6EB5AAFD21FE6830BD" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[247,502,727,752]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFD5AA6EB5AAFD21FE6830BD" box="[247,502,727,752]" italics="true" pageId="9" pageNumber="10">Barbatula pechiliensis</emphasis>
</taxonomicName>
,
<taxonomicName id="1DBB33CAFFD5AA6EB758FD2FFD3430BD" authorityName=": Nichols" authorityYear="1925" box="[517,682,727,752]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="subSpecies" species="toni" subSpecies="fowleri">
<emphasis id="E8CF945BFFD5AA6EB758FD2FFD3430BD" box="[517,682,727,752]" italics="true" pageId="9" pageNumber="10">B. toni fowleri</emphasis>
</taxonomicName>
,
<taxonomicName id="1DBB33CAFFD5AA6EB7E7FD2FFCD630BD" authorityName="Tchang" authorityYear="1932" box="[698,840,727,752]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="kirinensis">
<emphasis id="E8CF945BFFD5AA6EB7E7FD2FFCD630BD" box="[698,840,727,752]" italics="true" pageId="9" pageNumber="10">B. kirinensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="1DBB33CAFFD5AA6EB6D5FD2FFC7730BD" authorityName="Cao, Causse &amp; Zhang" authorityYear="2012" box="[904,1001,727,752]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="gibba">
<emphasis id="E8CF945BFFD5AA6EB6D5FD2FFC7730BD" box="[904,1001,727,752]" italics="true" pageId="9" pageNumber="10">B. gibba</emphasis>
</taxonomicName>
were all identified as synonyms of
<taxonomicName id="1DBB33CAFFD5AA6EB0D9FD2FFF5D3159" baseAuthorityName="Dybowski" baseAuthorityYear="1869" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFD5AA6EB0D9FD2FFF5D3159" italics="true" pageId="9" pageNumber="10">B. toni</emphasis>
</taxonomicName>
by earlier authors (
<bibRefCitation id="BE2A35B8FFD5AA6EB4C3FD0AFD913159" author="Zhu, S. Q." box="[414,527,764,789]" pageId="9" pageNumber="10" refId="ref17261" refString="Zhu, S. Q. (1989) The loaches of the subfamily Nemacheilinae in China (Cypriniformes: Cobitidae). Jiangsu Science and Technology Publishing House, Nanjing, 150 pp. [in Chinese]" type="book" year="1989">Zhu 1989</bibRefCitation>
,
<bibRefCitation id="BE2A35B8FFD5AA6EB746FD0AFD553159" author="Cao, L. &amp; Causse, R. &amp; Zhang, E." box="[539,715,764,789]" pageId="9" pageNumber="10" pagination="236 - 248" refId="ref16027" refString="Cao, L., Causse, R. &amp; Zhang, E. (2012) Revision of the loach species Barbatula nuda (Bleeker 1865) (Pisces: Balitoridae) from North China, with a description of a new species from Inner Mongolia. Zootaxa, 3586, 236 - 248." type="journal article" year="2012">
Cao
<emphasis id="E8CF945BFFD5AA6EB70DFD0BFD1F3158" box="[592,641,764,789]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
. 2012
</bibRefCitation>
,
<bibRefCitation id="BE2A35B8FFD5AA6EB78AFD0AFC143159" author="Prokofiev, A. M." box="[727,906,764,789]" pageId="9" pageNumber="10" pagination="243 - 248" refId="ref16631" refString="Prokofiev, A. M. (2014) Review on the book by M. Kottelat Conspectus cobitinum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). Uoprosy Ikhthiologii, 54 (2), 243 - 248." type="journal article" year="2014">Prokofiev 2014</bibRefCitation>
).
<bibRefCitation id="BE2A35B8FFD5AA6EB6C0FD0AFBD73158" author="Kottelat, M." box="[925,1097,764,789]" pageId="9" pageNumber="10" refId="ref16275" refString="Kottelat, M. (2006) Fishes of Mongolia: A check-list of the fishes known to occur in Mongolia with comments on systematics and nomenclature. The International Bank for Reconstruction and Development / The World Bank, Washington D. C., 183 pp." type="book" year="2006">Kottelat (2006)</bibRefCitation>
already suspected that
<taxonomicName id="1DBB33CAFFD5AA6EB00CFD0BFA023159" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[1361,1436,765,788]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFD5AA6EB00CFD0BFA023159" box="[1361,1436,765,788]" italics="true" pageId="9" pageNumber="10">B. toni</emphasis>
</taxonomicName>
is widely distributed in the upper reaches of the
<collectingRegion id="187F86ABFFD5AA6EB79CFCE9FCCD3175" box="[705,851,799,824]" country="China" name="Heilongjiang" pageId="9" pageNumber="10">Heilongjiang</collectingRegion>
(
<collectingRegion id="187F86ABFFD5AA6EB639FCD6FC373175" box="[868,937,800,824]" country="Russia" name="Amur" pageId="9" pageNumber="10">Amur</collectingRegion>
) River. We also report
<taxonomicName id="1DBB33CAFFD5AA6EB1E9FCD7FA9E3175" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[1204,1280,801,824]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFD5AA6EB1E9FCD7FA9E3175" box="[1204,1280,801,824]" italics="true" pageId="9" pageNumber="10">B. toni</emphasis>
</taxonomicName>
from the Yin River drainage, a small coastal river, and suspect that it might be more widespread.
</paragraph>
<paragraph id="DA044849FFD5AA6EB59AFC91FBB63185" blockId="9.[151,1436,151,968]" pageId="9" pageNumber="10">
The morphological character states of
<taxonomicName id="1DBB33CAFFD5AA6EB7DCFC9FFD5131CD" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[641,719,873,896]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFD5AA6EB7DCFC9FFD5131CD" box="[641,719,873,896]" italics="true" pageId="9" pageNumber="10">B. toni</emphasis>
</taxonomicName>
described here are based on the individuals examined for this study and agree well with those described by
<bibRefCitation id="BE2A35B8FFD5AA6EB7C7FC7AFCCA31E9" author="Prokofiev, A. M." box="[666,852,908,933]" pageId="9" pageNumber="10" pagination="695 - 707" refId="ref16562" refString="Prokofiev, A. M. (2003) Materials on the revision of common stone loaches (Balitoridae: Nemacheilinae: Orthrias Jordan and Fowler, 1903) from Mongolia and adjacent area: I. Two new species from Tuva and Mongolia. Journal of Ichthyology, 43 (9), 695 - 707. https: // doi. org / 10.1134 / S 0032945211060051" type="journal article" year="2003">Prokofiev (2003</bibRefCitation>
,
<bibRefCitation id="BE2A35B8FFD5AA6EB63DFC7AFC3831E8" author="Prokofiev, A. M." box="[864,934,908,933]" pageId="9" pageNumber="10" pagination="488 - 497" refId="ref16726" refString="Prokofiev, A. M. (2016 a) Redescription and systematic position of nominal loach species Nemacheilus compressirostris and N. sibiricus (Nemacheilidae). Journal of Ichthyology, 56 (4), 488 - 497. https: // doi. org / 10.1134 / S 0032945216040111" type="journal article" year="2016">2016a</bibRefCitation>
) based on fishes from the Onon River. They are also identical with
<taxonomicName id="1DBB33CAFFD5AA6EB4C9FC47FE413185" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[404,479,945,968]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFD5AA6EB4C9FC47FE413185" box="[404,479,945,968]" italics="true" pageId="9" pageNumber="10">B. toni</emphasis>
</taxonomicName>
from the Onon River shown by
<bibRefCitation id="BE2A35B8FFD5AA6EB617FC59FBBA3185" author="Kottelat, M." box="[842,1060,943,968]" pageId="9" pageNumber="10" refId="ref16275" refString="Kottelat, M. (2006) Fishes of Mongolia: A check-list of the fishes known to occur in Mongolia with comments on systematics and nomenclature. The International Bank for Reconstruction and Development / The World Bank, Washington D. C., 183 pp." type="book" year="2006">Kottelat (2006: 99)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>