218 lines
28 KiB
XML
218 lines
28 KiB
XML
<document id="BA18032CF51941D16073C4E8B7E0A3A0" ID-DOI="10.1649/0010-065x-63.sp7.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="4912048" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623157183841" checkinUser="carolina" docAuthor="Seago, Ainsley E." docDate="2009" docId="039887BAFFCDC06192EA83A3EED69690" docLanguage="en" docName="TheColeopBull.63.mo7.1-73.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 7) 63" docStyle="DocumentStyle:A431C8F32F3D86E670C18C13DA9B5F2B.4:TheColeopBull.2000-2009.monograph" docStyleId="A431C8F32F3D86E670C18C13DA9B5F2B" docStyleName="TheColeopBull.2000-2009.monograph" docStyleVersion="4" docTitle="Agyrtodes variabilis Seago 2009, new species" docType="treatment" docVersion="5" lastPageNumber="45" masterDocId="FFA1FFC2FFE7C04D936D8227ED609758" masterDocTitle="Revision Of Agyrtodes Portevin (Coleoptera: Leiodidae)" masterLastPageNumber="73" masterPageNumber="1" pageNumber="43" updateTime="1698957853775" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="EC2010E9D94BDF3A8B0357B68B2F5BBE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D986D8952A32AC6155CE7B2F8A590999">
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<mods:title id="5A011A3111173DB08EE7AFCCEFDDB15E">Revision Of Agyrtodes Portevin (Coleoptera: Leiodidae)</mods:title>
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</mods:titleInfo>
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||
<mods:name id="F70F5D83C454D5D2A83723E55D4107AB" type="personal">
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<mods:role id="3C2C52FF8E1642115F35736254716669">
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<mods:roleTerm id="872230652BC554AC5F20D8692902ADEF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="58306E2253EC7EF2D0394024FAEA9F7C">Seago, Ainsley E.</mods:namePart>
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<mods:affiliation id="912228B051C6B53DE9A538FB41562A2B">ESPM- Div. of Insect Biology 137 Mulford Hall, University of California Berkeley, CA 94720 - 3114</mods:affiliation>
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<mods:nameIdentifier id="2998CF79AEBA371BE6E139F1C99757CD" type="email">seago@nature.berkeley.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="9E08B900D3700E2F34E354F6D36D036C">text</mods:typeOfResource>
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<mods:relatedItem id="C649AA4C634D16A946720060211D475D" type="host">
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<mods:titleInfo id="CFB35DE2D6702C62423B45E267179D15">
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<mods:title id="CE0F9CC17F9AA7E7A8A94FA78417FD35">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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||
<mods:part id="B5F78A37BC85AF112297FE735010EA7C">
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||
<mods:date id="9A5D59778893FCF4E553EDEB9D453699">2009</mods:date>
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<mods:detail id="B652C2490AE8AD357B960E653C587C04" type="series">
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<mods:title id="23473048024D47274E58C5EC714ABC53">mo 7</mods:title>
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</mods:detail>
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||
<mods:detail id="1B62D6792275BD2CFBB45C414C9C4EC5" type="pubDate">
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<mods:number id="4EA9872F8F7A553E65395020C90F4EEB">2009-04-22</mods:number>
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</mods:detail>
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||
<mods:detail id="21076E97A5B477A76260160363C107A9" type="volume">
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<mods:number id="9E88944C253931452F05AB2821A36AF6">63</mods:number>
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</mods:detail>
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<mods:extent id="923B56D9A9AFCEDB011059A59A71831D" unit="page">
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<mods:start id="73A3EA1010385752ABC004B0AEC2EE60">1</mods:start>
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<mods:end id="CD9F3F7F0EA3D2F4DA39568E03CBE23C">73</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="5938A692C82C3DF2363F33AE3CE8E7B9">journal article</mods:classification>
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<mods:identifier id="45BAE00F9D7ABCCEC34EE8E9301C7E7D" type="DOI">10.1649/0010-065x-63.sp7.1</mods:identifier>
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<mods:identifier id="0F0256FFCE9A296DD175A505FFF7188C" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="D1F0AF3DC72857F52EA0EFD41AB56CF0" type="Zenodo-Dep">4912048</mods:identifier>
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<mods:identifier id="4663E03215F682BBA838FB959B73F3B5" type="ZooBank">B58B2216-0CFA-41C6-8141-7024E65ECF85</mods:identifier>
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</mods:mods>
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<treatment id="039887BAFFCDC06192EA83A3EED69690" ID-DOI="http://doi.org/10.5281/zenodo.4912303" ID-Zenodo-Dep="4912303" LSID="urn:lsid:plazi:treatment:039887BAFFCDC06192EA83A3EED69690" httpUri="http://treatment.plazi.org/id/039887BAFFCDC06192EA83A3EED69690" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43">
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<subSubSection id="C32B6527FFCDC06792EA83A3EF8B96ED" pageId="42" pageNumber="43" type="nomenclature">
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<paragraph id="8B8E36ACFFCDC06792EA83A3EE7596C2" blockId="42.[391,789,388,437]" box="[391,789,388,410]" pageId="42" pageNumber="43">
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<heading id="D0C681C0FFCDC06792EA83A3EE7596C2" box="[391,789,388,410]" centered="true" fontSize="8" level="2" pageId="42" pageNumber="43" reason="2">
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<emphasis id="B945EABEFFCDC06792EA83A3EE7596C2" bold="true" box="[391,789,388,410]" pageId="42" pageNumber="43">
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||
<taxonomicName id="4C314D2FFFCDC06792EA83A3EFF596C2" ID-CoL="BBW5" authority="Seago" authorityName="Seago" authorityYear="2009" box="[391,661,388,410]" class="Insecta" family="Leiodidae" genus="Agyrtodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
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<emphasis id="B945EABEFFCDC06792EA83A3EF3096C2" bold="true" box="[391,592,388,410]" italics="true" pageId="42" pageNumber="43">Agyrtodes variabilis</emphasis>
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Seago
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27657C5FFCDC06791CF83A3EE7596C2" box="[674,789,388,410]" pageId="42" pageNumber="43" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8E36ACFFCDC06792DC83B8EF8B96ED" blockId="42.[391,789,388,437]" box="[433,747,415,437]" pageId="42" pageNumber="43">(Figs. 13, 30a, 50, 70, 90, 100)</paragraph>
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</subSubSection>
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<subSubSection id="C32B6527FFCDC06793AA83F1EFD19565" pageId="42" pageNumber="43" type="materials_examined">
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<paragraph id="8B8E36ACFFCDC06793AA83F1EF809579" blockId="42.[175,1005,470,1570]" pageId="42" pageNumber="43">
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<materialsCitation id="3B593CF1FFCDC06793AA83F1EF809579" collectingDate="1987-11-05" collectingDateMax="1987-12-07" collectingDateMin="1987-11-05" collectionCode="MDPI, QDPI, QMBA" collectorName="Tully Falls S. F. & Storey & Dickinson & Trap & Site No." country="Australia" elevation="1000" location="Qld" municipality="Tully Falls" pageId="42" pageNumber="43" specimenCount="1" typeStatus="holotype">
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<emphasis id="B945EABEFFCDC06793AA83F1EC4A96B4" bold="true" box="[199,298,470,492]" pageId="42" pageNumber="43">
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<typeStatus id="548A880EFFCDC06793AA83F1EC4696B4" box="[199,294,470,492]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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male, labeled ‘‘
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<collectingCountry id="F326763CFFCDC06792A183F1EF0696B4" box="[460,614,470,492]" name="Australia" pageId="42" pageNumber="43">AUSTRALIA</collectingCountry>
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: n.
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<location id="8EEE6077FFCDC06791E283F1EFD896B4" LSID="urn:lsid:plazi:treatment:039887BAFFCDC06192EA83A3EED69690:8EEE6077FFCDC06791E283F1EFD896B4" box="[655,696,470,492]" country="Australia" municipality="Tully Falls" name="Qld" pageId="42" pageNumber="43">Qld</location>
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,
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<collectorName id="26C4537AFFCDC06791A983F1EE1096B4" box="[708,880,470,492]" pageId="42" pageNumber="43">
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<collectingMunicipality id="6BEAACD6FFCDC06791A983F1EE5596B4" box="[708,821,470,492]" pageId="42" pageNumber="43">Tully Falls</collectingMunicipality>
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S. F.
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</collectorName>
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<quantity id="4CC99B49FFCDC067901483F1EEAD96B4" box="[889,973,470,492]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="43" unit="m" value="1000.0">
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||
<elevation id="001CD19FFFCDC067901483F1EEAD96B4" box="[889,973,470,492]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="43" unit="m" value="1000.0">1,000 m</elevation>
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</quantity>
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,
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<locationDeviation id="9A3C2F8EFFCDC06790B783F1ECFA955F" location="Ravenshoe" pageId="42" pageNumber="43">
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<quantity id="4CC99B49FFCDC06790B783F1ED84955F" metricMagnitude="3" metricUnit="m" metricValue="9.5" pageId="42" pageNumber="43" unit="km" value="9.5">9. 5 km</quantity>
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SSW Ravenshoe
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</locationDeviation>
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,
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<date id="FF8F106CFFCDC06792C883D6EF04955F" box="[421,612,497,519]" pageId="42" pageNumber="43" value="1987-11-05" valueMax="1987-12-07" valueMin="1987-11-05">
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<collectingDate id="EFCBE984FFCDC06792C883D6EF04955F" box="[421,612,497,519]" pageId="42" pageNumber="43" value="1987-11-05" valueMax="1987-12-07" valueMin="1987-11-05">5. XI-7. XII. 1987</collectingDate>
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</date>
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,
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<collectorName id="26C4537AFFCDC067911D83D6EFD2955F" box="[624,690,497,519]" pageId="42" pageNumber="43">Storey</collectorName>
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&
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<collectorName id="26C4537AFFCDC06791B883D6EE5F955F" box="[725,831,497,519]" pageId="42" pageNumber="43">Dickinson</collectorName>
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/
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<collectionCode id="ED20AE69FFCDC067902A83D6EEE8955F" box="[839,904,497,519]" pageId="42" pageNumber="43">MDPI</collectionCode>
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<collectingMethod id="52704EBBFFCDC06790FD83D6EE8C955F" box="[912,1004,497,519]" pageId="42" pageNumber="43">Intercept</collectingMethod>
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<collectorName id="26C4537AFFCDC06793C2802CED849579" box="[175,228,523,545]" pageId="42" pageNumber="43">Trap</collectorName>
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,
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<collectorName id="26C4537AFFCDC067939D802CEC269579" box="[240,326,523,545]" pageId="42" pageNumber="43">Site No.</collectorName>
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29A’’ (ex
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<collectionCode id="ED20AE69FFCDC06792DE802CEC939579" box="[435,499,523,545]" country="Australia" httpUri="http://grbio.org/cool/221v-j740" name="Queensland Department of Primary Industries" pageId="42" pageNumber="43">QDPI</collectionCode>
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; deposited in
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<collectionCode id="ED20AE69FFCDC06791E9802CEFB59579" box="[644,725,523,545]" pageId="42" pageNumber="43">QMBA</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8B8E36ACFFCDC06793AA8000EFD19565" blockId="42.[175,1005,470,1570]" box="[199,689,551,573]" pageId="42" pageNumber="43">
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||
<emphasis id="B945EABEFFCDC06793AA8000EC529565" bold="true" box="[199,306,551,573]" pageId="42" pageNumber="43">
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<typeStatus id="548A880EFFCDC06793AA8000EC4E9565" box="[199,302,551,573]" pageId="42" pageNumber="43" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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listed in Appendix 1 (
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<specimenCount id="9D37FD25FFCDC06791778000EFC69565" box="[538,678,551,573]" pageId="42" pageNumber="43" type="generic">40 specimens</specimenCount>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B6527FFCDC06793AA8064EE8D9501" box="[199,1005,579,601]" pageId="42" pageNumber="43" type="distribution">
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<paragraph id="8B8E36ACFFCDC06793AA8064EE8D9501" blockId="42.[175,1005,470,1570]" box="[199,1005,579,601]" pageId="42" pageNumber="43">
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<emphasis id="B945EABEFFCDC06793AA8064EC269501" bold="true" box="[199,326,579,601]" pageId="42" pageNumber="43">Distribution.</emphasis>
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<collectingCountry id="F326763CFFCDC06792208064ECCF9501" box="[333,431,579,601]" name="Australia" pageId="42" pageNumber="43">Australia</collectingCountry>
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: Northeast
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<collectingRegion id="49F5F84EFFCDC067914A8064EFC19501" box="[551,673,579,601]" country="Australia" name="Queensland" pageId="42" pageNumber="43">Queensland</collectingRegion>
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, Atherton tablelands (
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<figureCitation id="130A2A29FFCDC06790E48064EE829501" box="[905,994,579,601]" captionStart="Fig" captionStartId="23.[167,198,1205,1224]" captionTargetBox="[166,955,194,1183]" captionTargetId="figure-134@23.[166,955,194,1183]" captionTargetPageId="23" captionText="Fig. 100. Distribution of Agyrtodes spp. in Australia, part 2. A. tasmanicus, A. koebelei, A. variabilis, new species." figureDoi="http://doi.org/10.5281/zenodo.4912072" httpUri="https://zenodo.org/record/4912072/files/figure.png" pageId="42" pageNumber="43">Fig. 100</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B6527FFCDC06793AA8079EEE29413" pageId="42" pageNumber="43" type="diagnosis">
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<paragraph id="8B8E36ACFFCDC06793AA8079EEE29413" blockId="42.[175,1005,470,1570]" pageId="42" pageNumber="43">
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<emphasis id="B945EABEFFCDC06793AA8079EC4F952C" bold="true" box="[199,303,606,628]" pageId="42" pageNumber="43">Diagnosis.</emphasis>
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The coloration of this species is highly variable, ranging from entirely dark brown to light reddish brown with dark fasciae. The similarly sized
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<taxonomicName id="4C314D2FFFCDC06793C280B3EC4D95F1" authorityName="Seago" authorityYear="2009" box="[175,301,660,681]" class="Insecta" family="Leiodidae" genus="Agyrtodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="monteithi" status="sp. nov.">
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<emphasis id="B945EABEFFCDC06793C280B3EC4D95F1" box="[175,301,660,681]" italics="true" pageId="42" pageNumber="43">A. monteithi</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27657C5FFCDC067925180B3ECD595F2" box="[316,437,660,682]" pageId="42" pageNumber="43" rank="species">new species</taxonomicNameLabel>
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, has three distinct transverse elytral stripes marked by dark pubescence and an enlarged, bright yellow apical antennal segment, whereas
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<taxonomicName id="4C314D2FFFCDC06793C280EDEC4B9587" authorityName="Seago" authorityYear="2009" box="[175,299,714,735]" class="Insecta" family="Leiodidae" genus="Agyrtodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="variabilis">
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<emphasis id="B945EABEFFCDC06793C280EDEC4B9587" box="[175,299,714,735]" italics="true" pageId="42" pageNumber="43">A. variabilis</emphasis>
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</taxonomicName>
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has no more than two complete elytral fasciae and antennae that are uniformly brown.
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<taxonomicName id="4C314D2FFFCDC067920880C2EF7F95A2" baseAuthorityName="Blackburn" baseAuthorityYear="1903" box="[357,543,741,762]" class="Insecta" family="Leiodidae" genus="Agyrtodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="koebelei">
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<emphasis id="B945EABEFFCDC067920880C2EF7F95A2" box="[357,543,741,762]" italics="true" pageId="42" pageNumber="43">Agyrtodes koebelei</emphasis>
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</taxonomicName>
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also has two transverse dark brown fasciae on the elytra, but has a markedly smaller body size and long antennae with terminal segment light brown;
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<taxonomicName id="4C314D2FFFCDC06792E4813CEF609468" authorityName="Seago" authorityYear="2009" box="[393,512,795,816]" class="Insecta" family="Leiodidae" genus="Agyrtodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="variabilis">
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<emphasis id="B945EABEFFCDC06792E4813CEF609468" box="[393,512,795,816]" italics="true" pageId="42" pageNumber="43">A. variabilis</emphasis>
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</taxonomicName>
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has a uniformly brown antennal club and a very wide eighth antennal segment, scarcely narrower than segments 7 and 9.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B6527FFCDC06193AA8176EC1396CA" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43" type="description">
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||
<paragraph id="8B8E36ACFFCDC06793AA8176ECBA94DA" blockId="42.[175,1005,470,1570]" pageId="42" pageNumber="43">
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<emphasis id="B945EABEFFCDC06793AA8176EC20943F" bold="true" box="[199,320,849,871]" pageId="42" pageNumber="43">Description.</emphasis>
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TBL
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<emphasis id="B945EABEFFCDC06792108176ECF1943F" bold="true" box="[381,401,849,871]" pageId="42" pageNumber="43">5</emphasis>
|
||
<quantity id="4CC99B49FFCDC06792F58176EC88943F" box="[408,488,849,871]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="42" pageNumber="43" unit="mm" value="1.9">1.9 mm</quantity>
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, EW
|
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<emphasis id="B945EABEFFCDC067914C8176EF55943F" bold="true" box="[545,565,849,871]" pageId="42" pageNumber="43">5</emphasis>
|
||
<quantity id="4CC99B49FFCDC06791568176EFEC943F" box="[571,652,849,871]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="42" pageNumber="43" unit="mm" value="1.1">1.1 mm</quantity>
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||
, PNW
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<emphasis id="B945EABEFFCDC06791BB8176EF8A943F" bold="true" box="[726,746,849,871]" pageId="42" pageNumber="43">5</emphasis>
|
||
1.0 mm, HW
|
||
<emphasis id="B945EABEFFCDC06790138176EEF2943F" bold="true" box="[894,914,849,871]" pageId="42" pageNumber="43">5</emphasis>
|
||
<quantity id="4CC99B49FFCDC06790F58176EE89943F" box="[920,1001,849,871]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="42" pageNumber="43" unit="mm" value="0.5">0.5 mm</quantity>
|
||
. (TBL variation: 1.8–2.0 mm)
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||
</paragraph>
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<paragraph id="8B8E36ACFFCDC06793AA81A0EE5D932C" blockId="42.[175,1005,470,1570]" pageId="42" pageNumber="43">Small body size, body shape narrow-sided and strongly convex (Fig. 13). Color pattern of variable appearance depending on melanization of integument and angle of lighting; most individuals appear dark reddish-brown in dorsal view, with paired round paler red spots flanking elytral suture immediately posterior to scutellum and paired sloping pale red fasciae at elytral midpoint; pronotum uniformly dark reddish-brown, ferrugineous. Dorsal vestiture dark gold on pronotum, gold on pale regions of elytra, darker brown on dark regions. In heavily melanized individuals, fasciae at elytral midpoint appear only as paired, sloping ‘‘stripes’’ of gold setae against dark brown integument.</paragraph>
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<paragraph id="8B8E36ACFFCDC06793AA865DEE36917A" blockId="42.[175,1005,470,1570]" pageId="42" pageNumber="43">
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Head short, broad, eyes produced, narrowly emarginate behind; integument matte with very fine strigulose microsculpture, punctures imperceptible. Epistomal suture absent, clypeus short and trapezoidal, tapering anteriad; labrum extremely short, transverse, scarcely excavate. Mandibles very large, outer edges strongly curved. Maxillae (
|
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<figureCitation id="130A2A29FFCDC067920686C2ECA093A3" box="[363,448,1253,1275]" captionStart="Figs" captionStartId="7.[167,206,1266,1285]" captionTargetBox="[156,966,194,1240]" captionTargetId="figure-35@7.[151,969,194,1243]" captionTargetPageId="7" captionText="Figs. 29–37. Agyrtodes spp. 29) Agyrtodes ovatus, left mandible, dorsal view; 30a) Agyrtodes ovatus, left maxilla, dorsal view; 30b) A. variabilis, new species, left maxilla, dorsal view; 31) labium, ventral view; 32) male terminalia in situ, dorsal view; 33) male genital segment, ventral view; 34) aedeagus, ventral view; 35) Agyrtodes eucalypti, female reproductive tract, in situ, ventral view; 36) Agyrtodes ovatus, female reproductive tract, partially dissected, with partially sclerotized spermatheca; 37) Agyrtodes lescheni, new species, female reproductive tract, partially dissected, with unsclerotized spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4912052" httpUri="https://zenodo.org/record/4912052/files/figure.png" pageId="42" pageNumber="43">Fig. 30a</figureCitation>
|
||
) with greatly enlarged, apically pointed hyaline galea bearing a large, loose cluster of long setae externally and a dense brush of minute setae on internal face; lacinia small, narrow, tapering distally, crowned with small apical ‘‘brush’’ comprising 3–4 short rows of minute, curving spines. Maxillary palpi with terminal segment long, acuminate, weakly produced at apex; penultimate segment very faintly wider than base of terminal segment, less than half as long. Labium with anterior margin of ligula short, hyaline, bearing only two small subapical setae. Labial palps with terminal segments elongate, slender, tapering apically, curved inward, with sparse array of short digitiform setae and many minute setae at base; penltimate palpomere wider and less than half as long as terminal segment, bearing two extremely long, curving, stiff setae, extending past apex of palp; first palpomere with single long seta on interior face.
|
||
</paragraph>
|
||
<paragraph id="8B8E36ACFFCCC06693C782E5EEF396E8" blockId="43.[146,976,194,1570]" pageId="43" pageNumber="44">
|
||
Antennae (
|
||
<figureCitation id="130A2A29FFCCC066927682E5EC069780" box="[283,358,194,216]" captionStart="Figs" captionStartId="9.[167,206,1301,1320]" captionTargetBox="[197,877,194,1279]" captionTargetId="figure-57@9.[197,924,194,1279]" captionTargetPageId="9" captionText="Figs. 48–57. Antennae of Agyrtodes spp. 48) A. atropos; 49) A. koebelei; 50) A. variabilis, new species; 51) A. bicolor; 52) A. nebulosus; 53) A. lescheni, new species; 54) A. labralis; 55) A. monticola; 56) A. nemoralis; 57) A. hunuensis." figureDoi="http://doi.org/10.5281/zenodo.4912129" httpUri="https://zenodo.org/record/4912129/files/figure.png" pageId="43" pageNumber="44">Fig. 50</figureCitation>
|
||
) robust, segments 1–5 light yellowish-brown, segments 6–11 medium to dark brown, 11 pale brown in distal half. Segments 1 and 2 weakly enlarged, cylindrical, elongate (3
|
||
<emphasis id="B945EABEFFCCC06692B082DFEC919656" bold="true" box="[477,497,248,270]" pageId="43" pageNumber="44">3</emphasis>
|
||
as long as wide). Segment 3 slender, 4 and 5 of equal size and shape, just over twice as long as wide; segment 6 cylindrical, stouter than 3, width subequal to base of 7. Segment 7 broadly cylindrical, robust, more than twice as long as wide, apex as wide as base. Segment 8 subquadrate, shorter than any other segment, scarcely narrowed: width subequal to basal width of 7 and 9. Segment 9 nearly as long as 7, slightly wider; 10 rectangular, less than 1.5
|
||
<emphasis id="B945EABEFFCCC06690358358EE0C96CD" bold="true" box="[856,876,383,405]" pageId="43" pageNumber="44">3</emphasis>
|
||
as long as wide; 11 elongate, oval, weakly produced distally, with bluntly pointed apex.
|
||
</paragraph>
|
||
<paragraph id="8B8E36ACFFCCC06693C78391EEFF9560" blockId="43.[146,976,194,1570]" pageId="43" pageNumber="44">Pronotum transverse, weakly convex, reddish-brown; integument matte with shallow punctures at setal insertions and fine, confused strigulose microsculpture. Pronotal vestiture dark brown, semirecumbent. Lateral margins of pronotum very weakly curved anteriorly, straight posteriorly; hind angles sharply pointed, acutely produced posteriad. Narrow region of posterior pronotal margin transparent.</paragraph>
|
||
<paragraph id="8B8E36ACFFCCC06693C7801AEF3F9582" blockId="43.[146,976,194,1570]" pageId="43" pageNumber="44">Elytra convex at base, weakly convex in posterior half; posterior margins rounded, bluntly pointed at apices. Elytral integument shining, transverse strigae deeply impressed, disc without longitudinal strial impressions. Elytral vestiture light gold on scutellum, paired subscutellar pale spots and discal transverse fasciae, dark brown on darker reddish-brown regions of elytron. Scutellum large in relation to body size, pale reddish-brown.</paragraph>
|
||
<paragraph id="8B8E36ACFFCCC06693C780C7ECC29496" blockId="43.[146,976,194,1570]" pageId="43" pageNumber="44">Prosternum pale orange; hypomeron weakly concave posterior to coxal insertions, strongly concave immediately anterior to hind margin of pronotum. Coxal cavities large, globular. Mesosternum reddish-brown, darker than prosternum, with pronounced median carina smoothly curved in lateral view. Mesepimera broad, subrectangular, bluntly pointed at mesocoxal articulations. Metasternum with pronounced median bulge with long setae, flatter and sparsely setose toward lateral margins, integument finely textured with shallow isodiametric sculpticells. Metepisternum narrow, elongately oval, rounded anteriorly and posteriorly.</paragraph>
|
||
<paragraph id="8B8E36ACFFCCC06693C781F3ECB19378" blockId="43.[146,976,194,1570]" pageId="43" pageNumber="44">Abdominal sterna 3–6 normal, male sternum VII broadly, shallowly emarginate and elevated medially, sternum VIII extremely narrow and transverse, shallowly emarginate medially.</paragraph>
|
||
<paragraph id="8B8E36ACFFCCC06693C78602EFBE9386" blockId="43.[146,976,194,1570]" pageId="43" pageNumber="44">
|
||
Legs short, robust, protibial apices armed with short, broad, curving spines. Male protarsi with first three segments broadly expanded, cordate, bearing tenent setae; first segment as wide as tibial apex, subquadrate, longer than segment 2; segment 4 short, globular, segment 5 short, narrowly ovoid, shorter than segments 2–4 combined. Male mesotarsi with first three segments expanded, each with thick ventral pad of tenent setae; first segment ovate, approximately 2/3
|
||
<emphasis id="B945EABEFFCCC06690D6868BEEAF939A" bold="true" box="[955,975,1196,1218]" pageId="43" pageNumber="44">3</emphasis>
|
||
width of tibial apex, segments 2 and 3 weakly expanded.
|
||
</paragraph>
|
||
<paragraph id="8B8E36ACFFCCC06693C786C4EF2E917A" blockId="43.[146,976,194,1570]" pageId="43" pageNumber="44">
|
||
Male genital segment (
|
||
<figureCitation id="130A2A29FFCCC06692F986C4ECBE93A1" box="[404,478,1251,1273]" captionStart="Figs" captionStartId="12.[196,235,1299,1318]" captionTargetBox="[194,986,194,1276]" captionTargetId="figure-1@12.[194,986,194,1276]" captionTargetPageId="12" captionText="Figs. 68–77. Male genital segment of Agyrtodes spp. 68) A. atropos; 69) A. koebelei; 70) A. variabilis, new species; 71) A. bicolor; 72) A. nebulosus; 73) A. lescheni, new species; 74) A. labralis; 75) A. monticola; 76) A. nemoralis.; 77) A. hunuensis." figureDoi="http://doi.org/10.5281/zenodo.4912062" httpUri="https://zenodo.org/record/4912062/files/figure.png" pageId="43" pageNumber="44">Fig. 70</figureCitation>
|
||
) subquadrate, bluntly pointed at apex, pleurites truncate and densely setose apically, tergite sparsely setose, with minute thumbnail-shaped heavily sclerotized region at apex. Anterior apophysis produced, broad, weakly expanded and truncate at apex. Pleurites very weakly fused ventrally. Aedeagus (
|
||
<figureCitation id="130A2A29FFCCC06692D48768EF63923D" box="[441,515,1359,1381]" captionStart="Figs" captionStartId="14.[196,235,1332,1351]" captionTargetBox="[182,999,194,1309]" captionTargetId="figure-1@14.[180,1000,194,1309]" captionTargetPageId="14" captionText="Figs. 88–97. Aedeagus (median lobe and parameres) of Agyrtodes spp. 88) A. atropos; 89) A. koebelei; 90) A. variabilis, new species; 91) A. bicolor; 92) A. nebulosus; 93) A. lescheni, new species; 94) A. labralis; 95) A. monticola; 96) A. nemoralis; 97) A. hunuensis." figureDoi="http://doi.org/10.5281/zenodo.4912066" httpUri="https://zenodo.org/record/4912066/files/figure.png" pageId="43" pageNumber="44">Fig. 90</figureCitation>
|
||
) with median lobe short, stout, gradually tapering to a blunt triangular point at apex. Basal piece short, broad, barely longer than penis. Parameres slender, cylindrical, scarcely extending past apex of median lobe; interor faces of paramere apices with only sparse, minute setae. In repose, parameres lie parallel and adjacent to each other, dorsal to the penis; during eversion, the parameres splay outward. Endophallus with two small patches of short hyaline spines or asperities, as well as 3–4 large, heavily sclerotized, curving, asymmetrical sclerites.
|
||
</paragraph>
|
||
<paragraph id="8B8E36ACFFCBC06193AA82E5EE2A9655" blockId="44.[175,1004,193,456]" pageId="44" pageNumber="45">
|
||
Female genitalia with coxites broad at base, tapering distally,.4
|
||
<emphasis id="B945EABEFFCBC061901F82E6EEE6978F" bold="true" box="[882,902,193,215]" pageId="44" pageNumber="45">3</emphasis>
|
||
length of stylus. Styli extremely minute, weakly attenuate and rounded apically, broad at base, bearing terminal seta approximately 3/4
|
||
<emphasis id="B945EABEFFCBC06191EB82D0EFFA9655" bold="true" box="[646,666,247,269]" pageId="44" pageNumber="45">3</emphasis>
|
||
length of coxite.
|
||
</paragraph>
|
||
<paragraph id="8B8E36ACFFCBC06193AA8335EC1396CA" blockId="44.[175,1004,193,456]" pageId="44" pageNumber="45">
|
||
<emphasis id="B945EABEFFCBC06193AA8335EC0E9670" bold="true" box="[199,366,274,296]" pageId="44" pageNumber="45">Natural History.</emphasis>
|
||
The majority of specimens examined were collected in malaise traps and flight intercept traps in March–June, some were collected by pyrethrum knockdown and litter sifting; the pinned material examined for this study was also more than 75% female, perhaps suggesting a difference in activity levels between males and females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6527FFCBC06193AA83B0EED69690" pageId="44" pageNumber="45" type="etymology">
|
||
<paragraph id="8B8E36ACFFCBC06193AA83B0EED69690" blockId="44.[175,1004,193,456]" pageId="44" pageNumber="45">
|
||
<emphasis id="B945EABEFFCBC06193AA83B0EC5A96F5" bold="true" box="[199,314,407,429]" pageId="44" pageNumber="45">Etymology.</emphasis>
|
||
The specific epithet is based on the Latin
|
||
<taxonomicName id="4C314D2FFFCBC061907083B0EE1896F4" authorityName="Seago" authorityYear="2009" box="[797,888,407,428]" class="Insecta" family="Leiodidae" genus="Agyrtodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B945EABEFFCBC061907083B0EE1896F4" box="[797,888,407,428]" italics="true" pageId="44" pageNumber="45">variabilis</emphasis>
|
||
</taxonomicName>
|
||
(variable), referring to the variation between light and dark coloration in this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |