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<document id="B12842088BF8B7C06E6992A84F80CDF3" ID="10.11646/zootaxa.4272.1.7SLASH11154" ID-CLB-Dataset="32967" ID-DOI="10.11646/zootaxa.4272.1.7" ID-GBIF-Dataset="4e8156a4-20ef-4643-ae15-89ed6e02f867" ID-ISSN="1175-5326" ID-Zenodo-Dep="583406" ID-ZooBank="5B053AE7-4871-4E79-9FCA-F0D71FC64078" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495786457860" checkinUser="plazi" docAuthor="Ng, Peter K. L., Devi, Suvarna &amp; Kumar, Appukuttannair Biju" docDate="2017" docId="FC4987B1F20FFFE5E38CAA8955CA04CB" docLanguage="en" docName="zootaxa.4272.1.7SLASH11154.pdf" docOrigin="Zootaxa 4272 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Typhlocarcinus kerala Ng, Devi &amp; Kumar, 2017, sp. nov." docType="treatment" docVersion="9" lastPageNumber="138" masterDocId="0070FFC9F20EFFE2E31BAE0257320504" masterDocTitle="Typhlocarcinus kerala, a new species of rhizopine crab from southwestern India, and the identity of T. craterifer Rathbun, 1914 (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="141" masterPageNumber="131" pageNumber="132" updateTime="1698439939152" updateUser="ExternalLinkService">
<mods:mods id="C185F8860F65A97B61D8FFF29AC981EC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="85F5F606F8B1FCD436DE438DBB3F1FDF">
<mods:title id="8F23C2D8AF3BA3709940D703579ED6ED">Typhlocarcinus kerala, a new species of rhizopine crab from southwestern India, and the identity of T. craterifer Rathbun, 1914 (Crustacea: Brachyura: Pilumnidae)</mods:title>
</mods:titleInfo>
<mods:name id="80F2473EB851CFE005AA6896FABB39B4" type="personal">
<mods:role id="ACE27B9F87B0C6B9BA680C5D87CD4911">
<mods:roleTerm id="D679C953F2CFBD0BEEB18DC3B2EFC2C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F2E91A6D0863F490C2E93A5295834FE1">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="E96C55F8FBF137299457F1FE084AF484" type="personal">
<mods:role id="D5F46D834B62AC8A5F34E44F07614EDB">
<mods:roleTerm id="14332FFC42DBC629A21B251D0682C9A0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6529CD208968A6D0C62B2EBFC622E2D1">Devi, Suvarna</mods:namePart>
</mods:name>
<mods:name id="D5A854196F2FFC6367777AE2EC560CDE" type="personal">
<mods:role id="A8767F034C33BBBBF98696665EA87DD9">
<mods:roleTerm id="6B1BCC6D5CE48FB22C0677F11C27754C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="88693F1F300715B223F96C2821701ECE">Kumar, Appukuttannair Biju</mods:namePart>
</mods:name>
<mods:typeOfResource id="204F1BAAE7B1336615ED9DACB27D3C67">text</mods:typeOfResource>
<mods:relatedItem id="A894A28B4553EA6A0E201A4B4065316A" type="host">
<mods:titleInfo id="DE7C14FBB729100ABB883A140370DB5C">
<mods:title id="DD74499B63773875E32D782A4C19E3F3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C357B59E7C70F5FAB95520937FEFDC63">
<mods:date id="26D81632FDB04622BE59B15BB1637F77">2017</mods:date>
<mods:detail id="C922D52D529DA7D1E53270BFC3F5ED24" type="volume">
<mods:number id="57D0DB4F4B29DEB4BA1B854B185E8FA2">4272</mods:number>
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<mods:detail id="C8B4C1A232B9D9AAF7CCD00CCE959F36" type="issue">
<mods:number id="D51D3ABFD3318AF07A25E44EC5D1E728">1</mods:number>
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<mods:start id="132AA611AE8E41F869202F225FF8CDBC">131</mods:start>
<mods:end id="7C058AE354315AA45EC6D631E12E8ED5">141</mods:end>
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<mods:classification id="BC81B7B60EFE4F8123309DE5B7ABE4A4">journal article</mods:classification>
<mods:identifier id="AD351753986CAA5BB39B5A0BEDFDF0C2" type="CLB-Dataset">32967</mods:identifier>
<mods:identifier id="A9CD5956DDD66C58C638938276F98EDA" type="DOI">10.11646/zootaxa.4272.1.7</mods:identifier>
<mods:identifier id="5D4C49DBEDD4BB908EE30F27F076D048" type="GBIF-Dataset">4e8156a4-20ef-4643-ae15-89ed6e02f867</mods:identifier>
<mods:identifier id="14596C5F82C1463D20E7E291D615E9C6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A1925BEB84BF15A606CB6F56A08094C6" type="Zenodo-Dep">583406</mods:identifier>
<mods:identifier id="AC14E9BE23B1F412F98948A7B7B3A389" type="ZooBank">5B053AE7-4871-4E79-9FCA-F0D71FC64078</mods:identifier>
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<treatment id="FC4987B1F20FFFE5E38CAA8955CA04CB" ID-DOI="http://doi.org/10.5281/zenodo.6034996" ID-GBIF-Taxon="130845163" ID-Zenodo-Dep="6034996" LSID="urn:lsid:plazi:treatment:FC4987B1F20FFFE5E38CAA8955CA04CB" httpUri="http://treatment.plazi.org/id/FC4987B1F20FFFE5E38CAA8955CA04CB" lastPageId="7" lastPageNumber="138" pageId="1" pageNumber="132">
<subSubSection id="3CFA652CF20FFFE3E38CAA89566B01C2" pageId="1" pageNumber="132" type="nomenclature">
<paragraph id="745F36A7F20FFFE3E38CAA89553901A1" blockId="1.[151,523,1163,1222]" box="[151,523,1163,1189]" pageId="1" pageNumber="132">
<heading id="2F1781CBF20FFFE3E38CAA89553901A1" bold="true" box="[151,523,1163,1189]" fontSize="11" level="1" pageId="1" pageNumber="132" reason="1">
<emphasis id="4694EAB5F20FFFE3E38CAA89553901A1" bold="true" box="[151,523,1163,1189]" pageId="1" pageNumber="132">
<taxonomicName id="B3E04D24F20FFFE3E38CAA89569401A1" ID-CoL="59VHK" box="[151,422,1163,1189]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">
<emphasis id="4694EAB5F20FFFE3E38CAA89569401A1" bold="true" box="[151,422,1163,1189]" italics="true" pageId="1" pageNumber="132">Typhlocarcinus kerala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DA757CEF20FFFE3E2B5AA89553901A1" box="[430,523,1163,1189]" pageId="1" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="745F36A7F20FFFE3E38CAAAF566B01C2" blockId="1.[151,523,1163,1222]" box="[151,345,1197,1222]" pageId="1" pageNumber="132">
(
<figureCitation id="ECDB2A22F20FFFE3E384AAAF563901C2" box="[159,267,1197,1222]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="4.[151,250,1944,1966]" captionStartId-1="5.[151,250,1839,1861]" captionStartId-2="6.[151,250,1620,1642]" captionStartId-3="8.[151,250,1862,1884]" captionTargetBox-0="[162,1422,193,1915]" captionTargetBox-1="[151,1436,193,1816]" captionTargetBox-2="[151,1436,193,1597]" captionTargetBox-3="[151,1436,367,1842]" captionTargetId-0="figure@4.[162,1422,193,1915]" captionTargetId-1="figure@5.[151,1436,193,1816]" captionTargetId-2="figure@6.[151,1436,193,1597]" captionTargetId-3="figure@8.[151,1436,367,1842]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionText-0="FIGURE 3. Typhlocarcinus kerala sp. nov. A, holotype male (17.0 × 11.4 mm) (ZSI / WGRC / IR-INV 8237); Kerala, India; B, D, paratype male (18.3 × 13.0 mm) (ZSI), Kerala, India; C, paratype female (19.6 × 13.0 mm) (DABFUK), Kerala, India. A – C, overall dorsal view; D, dorsal view of carapace right side (denuded)." captionText-1="FIGURE 4. Typhlocarcinus kerala sp. nov. A, E, paratype male (18.3 × 13.0 mm) (DABFUK), Kerala, India; B – D, F, paratype female (19.6 × 13.0 mm) (DABFUK), Kerala, India. A, C, D, frontal view of cephalothorax; B, right dorsal surface of carapace (denuded); E, F, outer view of chelae." captionText-2="FIGURE 5. Typhlocarcinus kerala sp. nov., paratype male (18.3 × 13.0 mm) (DABFUK), Kerala, India. A, right last ambulatory leg; B, right second or third ambulatory leg; C, ventro-mesial view of second and fourth ambulatory dactyli and propodi; D, dorsal view of right cheliped; E, outer view of merus and carpus of right cheliped." captionText-3="FIGURE 6. Typhlocarcinus kerala sp. nov. A – C, paratype male (18.3 × 13.0 mm) (DABFUK), Kerala, India; D – G, paratype female (19.6 × 13.0 mm) (DABFUK), Kerala, India; G, H, holotype male (17.0 × 11.4 mm) (ZSI / WGRC / IR-INV 8237), Kerala, India. A, anterior thoracic sternum, telson, and pleonal somites 5 and 6; B, E, thoracic sternum and pleonal somites 1 – 6; C, posterior thoracic sternum and pleonal somites 1 – 5; F, thoracic sternum and vulvae; G, lateral side of carapace showing granules; H, posterior thoracic sternites and sternopleonal cavity showing G 1, G 2 and pleonal locking tubercle on sternite 5." httpUri-0="https://zenodo.org/record/583409/files/figure.png" httpUri-1="https://zenodo.org/record/583410/files/figure.png" httpUri-2="https://zenodo.org/record/583411/files/figure.png" httpUri-3="https://zenodo.org/record/583412/files/figure.png" pageId="1" pageNumber="132">Figs. 3–6</figureCitation>
,
<figureCitation id="ECDB2A22F20FFFE3E20DAAAC561101C2" box="[278,291,1198,1222]" captionStart="FIGURE 7" captionStartId="9.[151,250,1306,1328]" captionTargetBox="[167,1419,210,1287]" captionTargetId="figure@9.[151,1436,193,1303]" captionTargetPageId="9" captionText="FIGURE 7. A – C, Typhlocarcinus craterifer Rathbun, 1914, holotype female (13.4 × 9.5 mm) (USNM 46397), Philippines; D – I, Typhlocarcinus kerala sp. nov., paratype male (18.3 × 13.0 mm) (DABFUK), Kerala, India. A, right side of carapace (denuded); B, frontal margin (denuded); C, left third maxilliped (denuded); D, right third maxilliped (denuded); E, ventral view of left G 1; F, dorsal view of left G 1; G, ventral view of distal part of left G 1; H, dorsal view of distal part of left G 1; I, left G 2. Scales: A = 1.0 mm; B – I = 0.5 mm." httpUri="https://zenodo.org/record/583413/files/figure.png" pageId="1" pageNumber="132">7</figureCitation>
D–I)
</paragraph>
</subSubSection>
<subSubSection id="3CFA652CF20FFFE3E38CAAED5264004B" pageId="1" pageNumber="132" type="materials_examined">
<paragraph id="745F36A7F20FFFE3E38CAAED5264004B" blockId="1.[151,1437,1263,2033]" pageId="1" pageNumber="132">
<materialsCitation id="C4883CFAF20FFFE3E38CAAED56B70028" ID-GBIF-Occurrence="1503742116" collectionCode="WGRC" collectorName="Material" location="Kerala" pageId="1" pageNumber="132" specimenCount="1" stateProvince="Kerala" typeStatus="holotype">
<emphasis id="4694EAB5F20FFFE3E38CAAED564B000C" bold="true" box="[151,377,1263,1288]" pageId="1" pageNumber="132">
<collectorName id="D9155371F20FFFE3E38CAAED57CD000C" box="[151,255,1263,1288]" pageId="1" pageNumber="132">Material</collectorName>
examined
</emphasis>
.
<typeStatus id="AB5B8805F20FFFE3E293AAED56C7000C" box="[392,501,1263,1288]" pageId="1" pageNumber="132" type="holotype">Holotype</typeStatus>
: male (17.0 ×
<quantity id="B3189B42F20FFFE3E18CAAF255CE000C" box="[663,764,1264,1288]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="1" pageNumber="132" unit="mm" value="11.4">11.4 mm</quantity>
) (ZSI/
<collectionCode id="12F1AE62F20FFFE3E05EAAF254A8000C" box="[837,922,1264,1288]" pageId="1" pageNumber="132">WGRC</collectionCode>
/IR-INV 8237) off
<taxonomicName id="B3E04D24F20FFFE3E769AAED538F000C" box="[1138,1213,1263,1288]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="141" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">
<collectingRegion id="B624F845F20FFFE3E769AAED538F000C" box="[1138,1213,1263,1288]" country="India" name="Kerala" pageId="1" pageNumber="132">Kerala</collectingRegion>
</taxonomicName>
coast, Kollam port, Thiruvananthapuram
</materialsCitation>
,
<materialsCitation id="C4883CFAF20FFFE3E28AAB1156F5004B" ID-GBIF-Occurrence="1503742118" collectingDate="2013-02-22" collectorName="A. B. Kumar" country="India" location="Kerala" pageId="1" pageNumber="132" specimenCount="2" specimenCount-male="2" stateProvince="Kerala" typeStatus="paratype">
<taxonomicName id="B3E04D24F20FFFE3E28AAB1156EC0028" box="[401,478,1299,1324]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="141" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">
<collectingRegion id="B624F845F20FFFE3E28AAB1156EC0028" box="[401,478,1299,1324]" country="India" name="Kerala" pageId="1" pageNumber="132">Kerala</collectingRegion>
</taxonomicName>
,
<collectingCountry id="0CF77637F20FFFE3E2F1AB1155140028" box="[490,550,1299,1324]" name="India" pageId="1" pageNumber="132">India</collectingCountry>
, from trawls, coll.
<collectorName id="D9155371F20FFFE3E1E5AB1654B8002F" box="[766,906,1300,1324]" pageId="1" pageNumber="132">A.B. Kumar</collectorName>
,
<collectingDate id="101AE98FF20FFFE3E08DAB1653510028" box="[918,1123,1299,1324]" pageId="1" pageNumber="132" value="2013-02-22">
<date id="005E1067F20FFFE3E08DAB1653510028" box="[918,1123,1299,1324]" pageId="1" pageNumber="132" value="2013-02-22">22 February 2013</date>
</collectingDate>
.
<typeStatus id="AB5B8805F20FFFE3E775AB1653ED0028" box="[1134,1247,1300,1324]" pageId="1" pageNumber="132" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="62E6FD2EF20FFFE3E7F0AB1652700028" box="[1259,1346,1299,1324]" pageId="1" pageNumber="132" type="male">2 males</specimenCount>
(19.4 × 14.0 mm, 18.3 × 13.0 mm)
</materialsCitation>
,
<materialsCitation id="C4883CFAF20FFFE3E2C9AB355264004B" ID-GBIF-Occurrence="1503742119" box="[466,1366,1334,1359]" collectionCode="WGRC" collectorName="Material" location="Kerala" pageId="1" pageNumber="132" specimenCount="2" specimenCount-female="2" stateProvince="Kerala" typeStatus="paratype">
<specimenCount id="62E6FD2EF20FFFE3E2C9AB35550C004B" box="[466,574,1334,1359]" pageId="1" pageNumber="132" type="female">2 females</specimenCount>
(19.6 × 13.0 mm, 13.2 × 9.0 mm) (DABFUK), same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3CFA652CF20FFFE3E3DCAB58557702F5" pageId="1" pageNumber="132" type="diagnosis">
<paragraph id="745F36A7F20FFFE3E3DCAB58557702F5" blockId="1.[151,1437,1263,2033]" pageId="1" pageNumber="132">
<emphasis id="4694EAB5F20FFFE3E3DCAB5856080077" bold="true" box="[199,314,1370,1395]" pageId="1" pageNumber="132">Diagnosis</emphasis>
. Carapace transversely subovate, male width 1.38–1.51 times length, female width 1.51 times length; dorsal surfaces smooth, covered with short pubescence on surface, with longer plumose setae on lateral and frontal margins, regions demarcated, H-shaped gastric and cardiac grooves deep, distinct, surface adjacent appears crenulated in adults; anterolateral margin strongly convex, lined with small granules, appears entire throughout length, granules at junction with posterolateral margin appear separated by very short groove but no clear tooth visible; posterolateral margins subparallel, straight, margin and surface evenly covered with small rounded granules (
<figureCitation id="ECDB2A22F20FFFE3E21DA82D56600343" box="[262,338,1583,1608]" captionStart="FIGURE 3" captionStartId="4.[151,250,1944,1966]" captionTargetBox="[162,1422,193,1915]" captionTargetId="figure@4.[162,1422,193,1915]" captionTargetPageId="4" captionText="FIGURE 3. Typhlocarcinus kerala sp. nov. A, holotype male (17.0 × 11.4 mm) (ZSI / WGRC / IR-INV 8237); Kerala, India; B, D, paratype male (18.3 × 13.0 mm) (ZSI), Kerala, India; C, paratype female (19.6 × 13.0 mm) (DABFUK), Kerala, India. A – C, overall dorsal view; D, dorsal view of carapace right side (denuded)." httpUri="https://zenodo.org/record/583409/files/figure.png" pageId="1" pageNumber="132">Figs. 3</figureCitation>
A–D, 4A–D). Frontal margin deflexed downwards, barely visible from dorsal view; bilobed with distinct median cleft, margin of each lobe distinctly convex (
<figureCitation id="ECDB2A22F20FFFE3E053A85054A7036F" box="[840,917,1618,1643]" captionStart="FIGURE 3" captionStartId="4.[151,250,1944,1966]" captionTargetBox="[162,1422,193,1915]" captionTargetId="figure@4.[162,1422,193,1915]" captionTargetPageId="4" captionText="FIGURE 3. Typhlocarcinus kerala sp. nov. A, holotype male (17.0 × 11.4 mm) (ZSI / WGRC / IR-INV 8237); Kerala, India; B, D, paratype male (18.3 × 13.0 mm) (ZSI), Kerala, India; C, paratype female (19.6 × 13.0 mm) (DABFUK), Kerala, India. A – C, overall dorsal view; D, dorsal view of carapace right side (denuded)." httpUri="https://zenodo.org/record/583409/files/figure.png" pageId="1" pageNumber="132">Figs. 3</figureCitation>
D, 4A–D). Orbit ovate; eyes small, with stout peduncle, small cornea, fused to carapace, immovable in male, mobile in female, tip with small spot of pigment (
<figureCitation id="ECDB2A22F20FFFE3E384A89B57D003B6" box="[159,226,1689,1714]" captionStart="FIGURE 4" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[151,1436,193,1816]" captionTargetId="figure@5.[151,1436,193,1816]" captionTargetPageId="5" captionText="FIGURE 4. Typhlocarcinus kerala sp. nov. A, E, paratype male (18.3 × 13.0 mm) (DABFUK), Kerala, India; B – D, F, paratype female (19.6 × 13.0 mm) (DABFUK), Kerala, India. A, C, D, frontal view of cephalothorax; B, right dorsal surface of carapace (denuded); E, F, outer view of chelae." httpUri="https://zenodo.org/record/583410/files/figure.png" pageId="1" pageNumber="132">Fig. 4</figureCitation>
A–C). Posterior margin of epistome gently convex, not clearly divided into lobes (
<figureCitation id="ECDB2A22F20FFFE3E762A89B538F03B6" box="[1145,1213,1689,1714]" captionStart="FIGURE 4" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[151,1436,193,1816]" captionTargetId="figure@5.[151,1436,193,1816]" captionTargetPageId="5" captionText="FIGURE 4. Typhlocarcinus kerala sp. nov. A, E, paratype male (18.3 × 13.0 mm) (DABFUK), Kerala, India; B – D, F, paratype female (19.6 × 13.0 mm) (DABFUK), Kerala, India. A, C, D, frontal view of cephalothorax; B, right dorsal surface of carapace (denuded); E, F, outer view of chelae." httpUri="https://zenodo.org/record/583410/files/figure.png" pageId="1" pageNumber="132">Fig. 4</figureCitation>
D). Outer surface of ischium, merus and exopod of third maxilliped with numerous small granules; anteroexternal angle of merus rounded; exopod slender (
<figureCitation id="ECDB2A22F20FFFE3E2A5A8E2553103FD" box="[446,515,1760,1785]" captionStart="FIGURE 7" captionStartId="9.[151,250,1306,1328]" captionTargetBox="[167,1419,210,1287]" captionTargetId="figure@9.[151,1436,193,1303]" captionTargetPageId="9" captionText="FIGURE 7. A – C, Typhlocarcinus craterifer Rathbun, 1914, holotype female (13.4 × 9.5 mm) (USNM 46397), Philippines; D – I, Typhlocarcinus kerala sp. nov., paratype male (18.3 × 13.0 mm) (DABFUK), Kerala, India. A, right side of carapace (denuded); B, frontal margin (denuded); C, left third maxilliped (denuded); D, right third maxilliped (denuded); E, ventral view of left G 1; F, dorsal view of left G 1; G, ventral view of distal part of left G 1; H, dorsal view of distal part of left G 1; I, left G 2. Scales: A = 1.0 mm; B – I = 0.5 mm." httpUri="https://zenodo.org/record/583413/files/figure.png" pageId="1" pageNumber="132">Fig. 7</figureCitation>
D). Chelipeds stout, inner angle of carpus with low or sharp granules; margins of carpus and merus with numerous granules and dense setae; chela stout, dorsal and ventral margins with numerous rounded granules and dense setae; cutting edges of propodal finger with prominent broad teeth (
<figureCitation id="ECDB2A22F20FFFE3E7D1A92552240244" box="[1226,1302,1831,1856]" captionStart="FIGURE 3" captionStartId="4.[151,250,1944,1966]" captionTargetBox="[162,1422,193,1915]" captionTargetId="figure@4.[162,1422,193,1915]" captionTargetPageId="4" captionText="FIGURE 3. Typhlocarcinus kerala sp. nov. A, holotype male (17.0 × 11.4 mm) (ZSI / WGRC / IR-INV 8237); Kerala, India; B, D, paratype male (18.3 × 13.0 mm) (ZSI), Kerala, India; C, paratype female (19.6 × 13.0 mm) (DABFUK), Kerala, India. A – C, overall dorsal view; D, dorsal view of carapace right side (denuded)." httpUri="https://zenodo.org/record/583409/files/figure.png" pageId="1" pageNumber="132">Figs. 3</figureCitation>
A–C, 4E, F, 5D, E). Ambulatory legs moderately short; propodus of last leg relatively short; dactylus of last leg straight from dorsal view, slightly curved on mesial view (
<figureCitation id="ECDB2A22F20FFFE3E1BFA96C55C70283" box="[676,757,1902,1927]" captionStart="FIGURE 3" captionStartId="4.[151,250,1944,1966]" captionTargetBox="[162,1422,193,1915]" captionTargetId="figure@4.[162,1422,193,1915]" captionTargetPageId="4" captionText="FIGURE 3. Typhlocarcinus kerala sp. nov. A, holotype male (17.0 × 11.4 mm) (ZSI / WGRC / IR-INV 8237); Kerala, India; B, D, paratype male (18.3 × 13.0 mm) (ZSI), Kerala, India; C, paratype female (19.6 × 13.0 mm) (DABFUK), Kerala, India. A – C, overall dorsal view; D, dorsal view of carapace right side (denuded)." httpUri="https://zenodo.org/record/583409/files/figure.png" pageId="1" pageNumber="132">Figs. 3</figureCitation>
B, C, 5A–C). Male pleonal locking tubercle peg-like, near anterior edge of sternite 5. Male pleon relatively narrow, telson triangular (
<figureCitation id="ECDB2A22F20FFFE3E0EDA993530902AE" box="[1014,1083,1937,1962]" captionStart="FIGURE 6" captionStartId="8.[151,250,1862,1884]" captionTargetBox="[151,1436,367,1842]" captionTargetId="figure@8.[151,1436,367,1842]" captionTargetPageId="8" captionText="FIGURE 6. Typhlocarcinus kerala sp. nov. A – C, paratype male (18.3 × 13.0 mm) (DABFUK), Kerala, India; D – G, paratype female (19.6 × 13.0 mm) (DABFUK), Kerala, India; G, H, holotype male (17.0 × 11.4 mm) (ZSI / WGRC / IR-INV 8237), Kerala, India. A, anterior thoracic sternum, telson, and pleonal somites 5 and 6; B, E, thoracic sternum and pleonal somites 1 – 6; C, posterior thoracic sternum and pleonal somites 1 – 5; F, thoracic sternum and vulvae; G, lateral side of carapace showing granules; H, posterior thoracic sternites and sternopleonal cavity showing G 1, G 2 and pleonal locking tubercle on sternite 5." httpUri="https://zenodo.org/record/583412/files/figure.png" pageId="1" pageNumber="132">Fig. 6</figureCitation>
A). Female pleon ovate; telson broadly triangular (
<figureCitation id="ECDB2A22F20FFFE3E274A9B7568102CA" box="[367,435,1973,1998]" captionStart="FIGURE 6" captionStartId="8.[151,250,1862,1884]" captionTargetBox="[151,1436,367,1842]" captionTargetId="figure@8.[151,1436,367,1842]" captionTargetPageId="8" captionText="FIGURE 6. Typhlocarcinus kerala sp. nov. A – C, paratype male (18.3 × 13.0 mm) (DABFUK), Kerala, India; D – G, paratype female (19.6 × 13.0 mm) (DABFUK), Kerala, India; G, H, holotype male (17.0 × 11.4 mm) (ZSI / WGRC / IR-INV 8237), Kerala, India. A, anterior thoracic sternum, telson, and pleonal somites 5 and 6; B, E, thoracic sternum and pleonal somites 1 – 6; C, posterior thoracic sternum and pleonal somites 1 – 5; F, thoracic sternum and vulvae; G, lateral side of carapace showing granules; H, posterior thoracic sternites and sternopleonal cavity showing G 1, G 2 and pleonal locking tubercle on sternite 5." httpUri="https://zenodo.org/record/583412/files/figure.png" pageId="1" pageNumber="132">Fig. 6</figureCitation>
D). G1 distinctly sinuous, distal part slender, curved, tapering to rounded tip (
<figureCitation id="ECDB2A22F20FFFE3E60DA9B7526B02CA" box="[1302,1369,1973,1998]" captionStart="FIGURE 7" captionStartId="9.[151,250,1306,1328]" captionTargetBox="[167,1419,210,1287]" captionTargetId="figure@9.[151,1436,193,1303]" captionTargetPageId="9" captionText="FIGURE 7. A – C, Typhlocarcinus craterifer Rathbun, 1914, holotype female (13.4 × 9.5 mm) (USNM 46397), Philippines; D – I, Typhlocarcinus kerala sp. nov., paratype male (18.3 × 13.0 mm) (DABFUK), Kerala, India. A, right side of carapace (denuded); B, frontal margin (denuded); C, left third maxilliped (denuded); D, right third maxilliped (denuded); E, ventral view of left G 1; F, dorsal view of left G 1; G, ventral view of distal part of left G 1; H, dorsal view of distal part of left G 1; I, left G 2. Scales: A = 1.0 mm; B – I = 0.5 mm." httpUri="https://zenodo.org/record/583413/files/figure.png" pageId="1" pageNumber="132">Fig. 7</figureCitation>
E–H). Vulva relatively large, ovate (
<figureCitation id="ECDB2A22F20FFFE3E2F8A9DA551402F5" box="[483,550,2008,2033]" captionStart="FIGURE 6" captionStartId="8.[151,250,1862,1884]" captionTargetBox="[151,1436,367,1842]" captionTargetId="figure@8.[151,1436,367,1842]" captionTargetPageId="8" captionText="FIGURE 6. Typhlocarcinus kerala sp. nov. A – C, paratype male (18.3 × 13.0 mm) (DABFUK), Kerala, India; D – G, paratype female (19.6 × 13.0 mm) (DABFUK), Kerala, India; G, H, holotype male (17.0 × 11.4 mm) (ZSI / WGRC / IR-INV 8237), Kerala, India. A, anterior thoracic sternum, telson, and pleonal somites 5 and 6; B, E, thoracic sternum and pleonal somites 1 – 6; C, posterior thoracic sternum and pleonal somites 1 – 5; F, thoracic sternum and vulvae; G, lateral side of carapace showing granules; H, posterior thoracic sternites and sternopleonal cavity showing G 1, G 2 and pleonal locking tubercle on sternite 5." httpUri="https://zenodo.org/record/583412/files/figure.png" pageId="1" pageNumber="132">Fig. 6</figureCitation>
F).
</paragraph>
</subSubSection>
<caption id="209F662FF20CFFE0E38CA99255E002E0" httpUri="https://zenodo.org/record/583407/files/figure.png" pageId="2" pageNumber="133" targetBox="[162,1422,193,1910]" targetPageId="2">
<paragraph id="745F36A7F20CFFE0E38CA99255E002E0" blockId="2.[151,1436,1935,2020]" pageId="2" pageNumber="133">
<emphasis id="4694EAB5F20CFFE0E38CA992562302A1" bold="true" box="[151,273,1936,1958]" pageId="2" pageNumber="133">FIGURE 1</emphasis>
.
<taxonomicName id="B3E04D24F20CFFE0E23AA99255FF02A2" authority="Rathbun, 1914" authorityName="Rathbun" authorityYear="1914" box="[289,717,1936,1958]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="craterifer">
<emphasis id="4694EAB5F20CFFE0E23AA992551502A2" box="[289,551,1936,1958]" italics="true" pageId="2" pageNumber="133">Typhlocarcinus craterifer</emphasis>
Rathbun, 1914
</taxonomicName>
, holotype female (13.4 × 9.5 mm) (USNM 46397), Philippines. A, overall view; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax (partially denuded); D, ventral view of cephalothorax showing buccal cavity and pleon.
</paragraph>
</caption>
<caption id="209F662FF20DFFE1E38CA8FF55390274" httpUri="https://zenodo.org/record/583408/files/figure.png" pageId="3" pageNumber="134" targetBox="[151,1436,193,1765]" targetPageId="3">
<paragraph id="745F36A7F20DFFE1E38CA8FF55390274" blockId="3.[151,1436,1788,1904]" pageId="3" pageNumber="134">
<emphasis id="4694EAB5F20DFFE1E38CA8FF56230216" bold="true" box="[151,273,1789,1811]" pageId="3" pageNumber="134">FIGURE 2</emphasis>
.
<taxonomicName id="B3E04D24F20DFFE1E23AA8FE55FF0216" authority="Rathbun, 1914" authorityName="Rathbun" authorityYear="1914" box="[289,717,1788,1810]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="craterifer">
<emphasis id="4694EAB5F20DFFE1E23AA8FE55150216" box="[289,551,1788,1810]" italics="true" pageId="3" pageNumber="134">Typhlocarcinus craterifer</emphasis>
Rathbun, 1914
</taxonomicName>
, holotype female (13.4 × 9.5 mm) (USNM 46397), Philippines. A, posterior view of thoracic sternum and pleonal somites 1–4; thoracic sternum and pleonal somites 1–6; C, thoracic sternum and vulvae; D, dorsal view of right cheliped; E, outer view of right chela; F, outer view of left chela; G, right last ambulatory leg; H, right second or third ambulatory leg.
</paragraph>
</caption>
<caption id="209F662FF20AFFE6E38CA99A547C02EB" httpUri="https://zenodo.org/record/583409/files/figure.png" pageId="4" pageNumber="135" targetBox="[162,1422,193,1915]" targetPageId="4">
<paragraph id="745F36A7F20AFFE6E38CA99A547C02EB" blockId="4.[151,1436,1943,2031]" pageId="4" pageNumber="135">
<emphasis id="4694EAB5F20AFFE6E38CA99A563E02A9" bold="true" box="[151,268,1944,1966]" pageId="4" pageNumber="135">FIGURE 3</emphasis>
.
<taxonomicName id="B3E04D24F20AFFE6E201A99A56CE02AA" box="[282,508,1944,1966]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">
<emphasis id="4694EAB5F20AFFE6E201A99A56CE02AA" box="[282,508,1944,1966]" italics="true" pageId="4" pageNumber="135">Typhlocarcinus kerala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DA757CEF20AFFE6E118A99A556102A9" box="[515,595,1944,1965]" pageId="4" pageNumber="135" rank="species">
<emphasis id="4694EAB5F20AFFE6E118A99A556102A9" bold="true" box="[515,595,1944,1965]" pageId="4" pageNumber="135">sp. nov.</emphasis>
</taxonomicNameLabel>
A, holotype male (17.0 × 11.4 mm) (ZSI/WGRC/IR-INV 8237);
<taxonomicName id="B3E04D24F20AFFE6E7E9A99A520A02AA" box="[1266,1336,1944,1966]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">Kerala</taxonomicName>
, India; B, D, paratype male (18.3 × 13.0 mm) (ZSI),
<taxonomicName id="B3E04D24F20AFFE6E15FA9BB55B802CB" box="[580,650,1977,1999]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">Kerala</taxonomicName>
, India; C, paratype female (19.6 × 13.0 mm) (DABFUK),
<taxonomicName id="B3E04D24F20AFFE6E7CDA9BB522E02CB" box="[1238,1308,1977,1999]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">Kerala</taxonomicName>
, India. A–C, overall dorsal view; D, dorsal view of carapace right side (denuded).
</paragraph>
</caption>
<caption id="209F662FF20BFFE7E38CA92D55270282" httpUri="https://zenodo.org/record/583410/files/figure.png" pageId="5" pageNumber="136" targetBox="[151,1436,193,1816]" targetPageId="5">
<paragraph id="745F36A7F20BFFE7E38CA92D55270282" blockId="5.[151,1436,1838,1926]" pageId="5" pageNumber="136">
<emphasis id="4694EAB5F20BFFE7E38CA92D56390240" bold="true" box="[151,267,1839,1861]" pageId="5" pageNumber="136">FIGURE 4</emphasis>
.
<taxonomicName id="B3E04D24F20BFFE7E203A92D56CB0241" box="[280,505,1839,1861]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">
<emphasis id="4694EAB5F20BFFE7E203A92D56CB0241" box="[280,505,1839,1861]" italics="true" pageId="5" pageNumber="136">Typhlocarcinus kerala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DA757CEF20BFFE7E2E4A92D557D0240" box="[511,591,1839,1860]" pageId="5" pageNumber="136" rank="species">
<emphasis id="4694EAB5F20BFFE7E2E4A92D557D0240" bold="true" box="[511,591,1839,1860]" pageId="5" pageNumber="136">sp. nov.</emphasis>
</taxonomicNameLabel>
A, E, paratype male (18.3 × 13.0 mm) (DABFUK),
<taxonomicName id="B3E04D24F20BFFE7E77BA92D53940241" box="[1120,1190,1839,1861]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="141" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">Kerala</taxonomicName>
, India; B–D, F, paratype female (19.6 ×13.0 mm) (DABFUK),
<taxonomicName id="B3E04D24F20BFFE7E100A95255530262" box="[539,609,1872,1894]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="141" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">Kerala</taxonomicName>
, India. A, C, D, frontal view of cephalothorax; B, right dorsal surface of carapace (denuded); E, F, outer view of chelae.
</paragraph>
</caption>
<caption id="209F662FF208FFE4E38CA856536303AD" httpUri="https://zenodo.org/record/583411/files/figure.png" pageId="6" pageNumber="137" targetBox="[151,1436,193,1597]" targetPageId="6">
<paragraph id="745F36A7F208FFE4E38CA856536303AD" blockId="6.[151,1435,1619,1705]" pageId="6" pageNumber="137">
<emphasis id="4694EAB5F208FFE4E38CA8565620036E" bold="true" box="[151,274,1620,1642]" pageId="6" pageNumber="137">FIGURE 5</emphasis>
.
<taxonomicName id="B3E04D24F208FFE4E23FA856553E036E" box="[292,524,1620,1642]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">
<emphasis id="4694EAB5F208FFE4E23FA856553E036E" box="[292,524,1620,1642]" italics="true" pageId="6" pageNumber="137">Typhlocarcinus kerala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DA757CEF208FFE4E10CA857555C036E" box="[535,622,1621,1642]" pageId="6" pageNumber="137" rank="species">
<emphasis id="4694EAB5F208FFE4E10CA857555C036E" bold="true" box="[535,622,1621,1642]" pageId="6" pageNumber="137">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype male (18.3 × 13.0 mm) (DABFUK),
<taxonomicName id="B3E04D24F208FFE4E767A85653F0036E" box="[1148,1218,1620,1642]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">Kerala</taxonomicName>
, India. A, right last ambulatory leg; B, right second or third ambulatory leg; C, ventro-mesial view of second and fourth ambulatory dactyli and propodi; D, dorsal view of right cheliped; E, outer view of merus and carpus of right cheliped.
</paragraph>
</caption>
<subSubSection id="3CFA652CF208FFE5E3DCA8D755CA04CB" lastPageId="7" lastPageNumber="138" pageId="6" pageNumber="137" type="description">
<paragraph id="745F36A7F208FFE5E3DCA8D756E904A8" blockId="6.[151,1437,1749,2026]" lastBlockId="7.[151,1436,151,464]" lastPageId="7" lastPageNumber="138" pageId="6" pageNumber="137">
<emphasis id="4694EAB5F208FFE4E3DCA8D7560703EA" bold="true" box="[199,309,1749,1774]" pageId="6" pageNumber="137">Variation</emphasis>
. The carapace proportions of the five specimens vary rather substantially. The holotype male has a width to length ratio of 1.49, which is similar to that of two paratype females (1.46 and 1.51). The two paratype males (both larger than the holotype) are distinctly less broad, with a width to length ratio of 1.38 and 1.41, respectively. They, however, agree in all other characters, with the two males possessing the same structures of the front, anterolateral margin, pleon and G1. As such, we are confident they are conspecific. The H-shaped gastric and cardiac grooves are deep in the adult specimens but in the smallest female (13.2 × 9.0 mm) which is still subadult, these grooves are relatively shallower and less distinct. The condition of the eyes has been described as fused in many rhizopines but it is not easy to determine the condition in some. In
<taxonomicName id="B3E04D24F208FFE4E0DFA9D0531402EE" box="[964,1062,2001,2026]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">
<emphasis id="4694EAB5F208FFE4E0DFA9D0531402EE" box="[964,1062,2001,2026]" italics="true" pageId="6" pageNumber="137">T. kerala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DA757CEF208FFE4E736A9D353B402EE" box="[1069,1158,2001,2026]" pageId="6" pageNumber="137" rank="species">
<emphasis id="4694EAB5F208FFE4E736A9D353B402EE" bold="true" box="[1069,1158,2001,2026]" pageId="6" pageNumber="137">sp. nov.</emphasis>
</taxonomicNameLabel>
, the eyes are immovable in one paratype male (18.3´13.0 mm) but slightly mobile in the holotype and other paratypes, and we are not sure if its due to the preservation as the specimens had been preserved in formalin which has made the carapaces relatively softer. The tip of the cornea, however, appears to have a tiny spot of pigment in all the specimens.
<emphasis id="4694EAB5F209FFE5E3DCAF0656280419" bold="true" box="[199,282,260,285]" pageId="7" pageNumber="138">Colour</emphasis>
. In life, the carapace and appendages are white, with the fine pubescence and setae light brown.
<emphasis id="4694EAB5F209FFE5E3DCAF2556740444" bold="true" box="[199,326,295,320]" pageId="7" pageNumber="138">Etymology</emphasis>
. The species is named after the state of
<taxonomicName id="B3E04D24F209FFE5E01DAF2554630444" box="[774,849,295,320]" class="Malacostraca" family="Pilumnidae" genus="Typhlocarcinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="141" phylum="Arthropoda" rank="species" species="kerala" status="sp. nov.">Kerala</taxonomicName>
in India. The name is used as a noun in apposition.
<emphasis id="4694EAB5F209FFE5E3DCAF4E56060461" bold="true" box="[199,308,332,357]" pageId="7" pageNumber="138">Remarks</emphasis>
. The specimens collected were light brown and covered with a fine brown sediment; suggesting that like many rhizopines, they are luteophilous in habits, i.e. living in mud and fine sediment, with the waters usually turbid (see Ng 1987b, 1990).
</paragraph>
<paragraph id="745F36A7F209FFE5E3DCAFBA55CA04CB" blockId="7.[151,1436,151,464]" box="[199,760,439,464]" pageId="7" pageNumber="138">See General Discussion for taxonomic comments.</paragraph>
</subSubSection>
</treatment>
</document>
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