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<document ID-DOI="http://dx.doi.org/10.3897/jhr.84.68709" ID-GBIF-Dataset="b9e61bc3-4158-4bee-8d57-a65655588f70" ID-Pensoft-Pub="1314-2607-84-145" ID-Pensoft-UUID="AEDB6967BB17574BB6988187C56EE6CA" ID-Zenodo-Dep="5349694" ID-ZooBank="5F83EC637C7947FDBD29E7A2B9328518" ModsDocID="1314-2607-84-145" checkinTime="1629847073322" checkinUser="pensoft" docAuthor="Okayasu, Juriya" docDate="2021" docId="C7BE87A15935594A9F4F197B961E885C" docLanguage="en" docName="JourHymenoptRes 84: 145-168" docOrigin="Journal of Hymenoptera Research 84" docPubDate="2021-08-24" docSource="http://dx.doi.org/10.3897/jhr.84.68709" docTitle="Arkaditilla bagrada Okayasu 2021, comb. nov." docType="treatment" docVersion="4" id="AEDB6967BB17574BB6988187C56EE6CA" lastPageNumber="145" masterDocId="AEDB6967BB17574BB6988187C56EE6CA" masterDocTitle="A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species" masterLastPageNumber="168" masterPageNumber="145" pageNumber="145" updateTime="1678756233000" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Okayasu, Juriya</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6307-2939</mods:nameIdentifier>
<mods:affiliation>Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566, Japan &amp; Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo, 060 - 8589, Japan</mods:affiliation>
<mods:nameIdentifier type="email">mutiphiidae@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-08-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>84</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>145</mods:start>
<mods:end>168</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.84.68709</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.84.68709</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-84-145</mods:identifier>
<mods:identifier type="ZooBank">5F83EC637C7947FDBD29E7A2B9328518</mods:identifier>
<mods:identifier type="Pensoft-UUID">AEDB6967BB17574BB6988187C56EE6CA</mods:identifier>
<mods:identifier type="Zenodo-Dep">5349694</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="184941293" LSID="urn:lsid:plazi:treatment:C7BE87A15935594A9F4F197B961E885C" httpUri="http://treatment.plazi.org/id/C7BE87A15935594A9F4F197B961E885C" lastPageNumber="145" pageId="0" pageNumber="145">
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="C7BE87A1-5935-594A-9F4F-197B961E885C" authority="(Cameron, 1902)" authorityName="Okayasu" authorityYear="2021" baseAuthorityName="Cameron" baseAuthorityYear="1902" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada" status="comb. nov.">Arkaditilla bagrada (Cameron, 1902)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures1-6" httpUri="https://binary.pensoft.net/fig/579480" pageId="0" pageNumber="145">Figures 1</figureCitation>
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures1-6" httpUri="https://binary.pensoft.net/fig/579480" pageId="0" pageNumber="145">, 2</figureCitation>
<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures13-18" httpUri="https://binary.pensoft.net/fig/579482" pageId="0" pageNumber="145">, 13</figureCitation>
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">, 19</figureCitation>
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">, 25</figureCitation>
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">, 26</figureCitation>
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">, 37</figureCitation>
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">, 38</figureCitation>
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">, 49-51</figureCitation>
<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">, 67</figureCitation>
<figureCitation captionStart="Figure 73–76" captionStartId="F8" captionText="Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle)." figureDoi="10.3897/jhr.84.68709.figure73-76" httpUri="https://binary.pensoft.net/fig/579487" pageId="0" pageNumber="145">, 73</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="reference_group">
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mutilla bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada" status="comb. nov.">Mutilla bagrada</taxonomicName>
Cameron, 1902: 76, ♂, holotype ♂ (Kuching, Sarawak, [Malaysia]) [Natural History Museum, London, UK].
</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Timulla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Timulla (Trogaspidia) bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada" status="comb. nov." subGenus="Trogaspidia">Timulla (Trogaspidia) bagrada</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" author="Mickel, CE" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="145" pagination="177 - 312" refId="B20" refString="Mickel, CE, 1935. The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae). Transactions of the Royal Entomological Society of London 83 (2): 177 - 312, DOI: https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" title="The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae)." url="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" volume="83" year="1935">Mickel 1935</bibRefCitation>
: 263, ♂; 1937: 449, ♂.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Krombeinidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krombeinidia bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada" status="comb. nov.">Krombeinidia bagrada</taxonomicName>
: Lelej, 1996a: 11; 2005: 65;
<bibRefCitation author="Lo Cascio, P" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="145" pagination="529 - 592" refId="B15" refString="Lo Cascio, P, 2015. Worldwide checklist of the island mutillid wasps (Hymenoptera Mutillidae). Biodiversity Journal 6 (2): 529 - 592" title="Worldwide checklist of the island mutillid wasps (Hymenoptera Mutillidae)." volume="6" year="2015">Lo Cascio 2015</bibRefCitation>
: 550;
<bibRefCitation author="Pagliano, G" journalOrPublisher="Bollettino del Museo Regionale di Scienze Naturali di Torino" pageId="0" pageNumber="145" pagination="5 - 425" refId="B24" refString="Pagliano, G, Brothers, DJ, Cambra, R, Lelej, AS, Lo Cascio, P, Matteini, Palmerini M, Scaramozzino, PL, Williams, KA, Romano, M, 2020. Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bollettino del Museo Regionale di Scienze Naturali di Torino 36 (1-2): 5 - 425" title="Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species." volume="36" year="2020">Pagliano et al. 2020</bibRefCitation>
: 250.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Petersenidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Petersenidia bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada" status="comb. nov.">Petersenidia bagrada</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.3759/tropics.6.91" author="Lelej, AS" journalOrPublisher="Tropics" pageId="0" pageNumber="145" pagination="91 - 104" refId="B11" refString="Lelej, AS, 1996b. Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae). Tropics 6 (1-2): 91 - 104, DOI: https://doi.org/10.3759/tropics.6.91" title="Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae)." url="https://doi.org/10.3759/tropics.6.91" volume="6" year="1996 b">Lelej 1996b</bibRefCitation>
: 95, ♂.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Clypeus dorsally elevated, sloping ventrally. Mandible with sharp subapical and large inner subbasal teeth (Fig.
<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures13-18" httpUri="https://binary.pensoft.net/fig/579482" pageId="0" pageNumber="145">13</figureCitation>
), ventrally deeply excised with large basal tooth (Fig.
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">19</figureCitation>
). Scape ventral carinae subparallel, mostly smooth between them (Fig.
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">25</figureCitation>
); F1 apical 1/3 and F2 basal 1/2 yellow, F2 apical 1/2 and F3-F11 with ventral reddish brown patch; F1 cylindrical (Fig.
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">26</figureCitation>
). T3 orange (Fig.
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures1-6" httpUri="https://binary.pensoft.net/fig/579480" pageId="0" pageNumber="145">2</figureCitation>
); cuspis strongly widened on posterior 1/3 with rounded apex (Figs
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">49</figureCitation>
,
<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">67</figureCitation>
); digitus posteriorly slender stick-like (Fig.
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">37</figureCitation>
); paracuspis stick-like (Fig.
<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">67</figureCitation>
); penis valve weakly expanded ventrally (Figs
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">50</figureCitation>
,
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">51</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown.
</paragraph>
<caption ID-Zenodo-Dep="5349700" doi="10.3897/jhr.84.68709.figures13-18" httpUri="https://binary.pensoft.net/fig/579482" pageId="0" pageNumber="145" start="Figures 13–18" startId="F3">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figures 13-18.</emphasis>
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla</emphasis>
</taxonomicName>
spp., ♂, face
<emphasis bold="true" pageId="0" pageNumber="145">13</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada">
<emphasis italics="true" pageId="0" pageNumber="145">A. bagrada</emphasis>
</taxonomicName>
(Cameron)
<emphasis bold="true" pageId="0" pageNumber="145">14</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. depressicornis" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="depressicornis">
<emphasis italics="true" pageId="0" pageNumber="145">A. depressicornis</emphasis>
</taxonomicName>
(Mickel)
<emphasis bold="true" pageId="0" pageNumber="145">15</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. frim" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="frim">
<emphasis italics="true" pageId="0" pageNumber="145">A. frim</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="145">16</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. gibba" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="gibba">
<emphasis italics="true" pageId="0" pageNumber="145">A. gibba</emphasis>
</taxonomicName>
sp. nov., paratype
<emphasis bold="true" pageId="0" pageNumber="145">17</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. leleji" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leleji">
<emphasis italics="true" pageId="0" pageNumber="145">A. leleji</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="145">18</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia">
<emphasis italics="true" pageId="0" pageNumber="145">A. nallinia</emphasis>
</taxonomicName>
(Zavattari).
</paragraph>
</caption>
<caption ID-Zenodo-Dep="5349702" doi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145" start="Figures 19–36" startId="F4">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figures 19-36.</emphasis>
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla</emphasis>
</taxonomicName>
spp., ♂
<emphasis bold="true" pageId="0" pageNumber="145">19-24</emphasis>
mandible, lateral view
<emphasis bold="true" pageId="0" pageNumber="145">25, 27, 29, 31, 33, 35</emphasis>
scape, ventral view
<emphasis bold="true" pageId="0" pageNumber="145">26, 28, 30, 32, 34, 36</emphasis>
pedicel, F1, and F2, lateral view
<emphasis bold="true" pageId="0" pageNumber="145">19, 25, 26</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada">
<emphasis italics="true" pageId="0" pageNumber="145">A. bagrada</emphasis>
</taxonomicName>
(Cameron)
<emphasis bold="true" pageId="0" pageNumber="145">20, 27, 28</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. depressicornis" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="depressicornis">
<emphasis italics="true" pageId="0" pageNumber="145">A. depressicornis</emphasis>
</taxonomicName>
(Mickel)
<emphasis bold="true" pageId="0" pageNumber="145">21, 29, 30</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. frim" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="frim">
<emphasis italics="true" pageId="0" pageNumber="145">A. frim</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="145">22, 31, 32</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. gibba" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="gibba">
<emphasis italics="true" pageId="0" pageNumber="145">A. gibba</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="145">23, 33, 34</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. leleji" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leleji">
<emphasis italics="true" pageId="0" pageNumber="145">A. leleji</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="145">24, 35, 36</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia">
<emphasis italics="true" pageId="0" pageNumber="145">A. nallinia</emphasis>
</taxonomicName>
(Zavattari).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="materials_examined">
<paragraph pageId="0" pageNumber="145">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="145">
<materialsCitation collectingDate="1993-03-25" collectingDateMax="1993-12-17" collectingDateMin="1993-03-25" collectingMethod="coarse Malaise trap, fine Malaise trap" collectorName="K. Konishi, K. Maeto" country="Malaysia" county="Pasoh Forest Reserve" location="Pasoh Forest Reserve" municipality="Pasoh Forest Reserve" specimenCount="48" specimenCount-male="48" stateProvince="Negeri Sembilan">
<specimenCount type="male">24♂</specimenCount>
.
<emphasis bold="true" pageId="0" pageNumber="145">
<collectingCountry name="Malaysia">Malaysia</collectingCountry>
</emphasis>
:
<collectingRegion country="Malaysia" name="Negeri Sembilan">Negeri Sembilan</collectingRegion>
:
<collectingCounty>Pasoh Forest Reserve</collectingCounty>
,
<collectingDate value="1993-03-25" valueMax="1993-04-02" valueMin="1993-03-25">25.III-2.IV.1993</collectingDate>
,
<collectorName>K. Konishi</collectorName>
&amp;
<collectorName>K. Maeto</collectorName>
leg.,
<collectingMethod>coarse Malaise trap</collectingMethod>
, plot 2 [
<specimenCount type="male">2♂</specimenCount>
FRIM, NARO],
<collectingDate value="1993-04-02" valueMax="1993-04-08" valueMin="1993-04-02">2-8.IV.1993</collectingDate>
[
<specimenCount type="male">1♂</specimenCount>
FRIM],
<collectingDate value="1993-04-08" valueMax="1993-04-15" valueMin="1993-04-08">8-15.IV.1993</collectingDate>
[
<specimenCount type="male">1♂</specimenCount>
NARO];
<collectingMunicipality>Pasoh Forest Reserve</collectingMunicipality>
,
<collectingDate value="1993-03-25" valueMax="1993-04-02" valueMin="1993-03-25">25.III-2.IV.1993</collectingDate>
,
<collectorName>K. Konishi</collectorName>
&amp;
<collectorName>K. Maeto</collectorName>
leg.,
<collectingMethod>fine Malaise trap</collectingMethod>
, plot 2 [
<specimenCount type="male">2♂</specimenCount>
FRIM, NARO];
<location LSID="urn:lsid:plazi:treatment:C7BE87A15935594A9F4F197B961E885C:1BE7470E606EBCBF3FF11BE2B150164D" country="Malaysia" county="Pasoh Forest Reserve" municipality="Pasoh Forest Reserve" name="Pasoh Forest Reserve" stateProvince="Negeri Sembilan">Pasoh Forest Reserve</location>
,
<collectingDate value="1993-03-26" valueMax="1993-04-03" valueMin="1993-03-26">26.III-3.IV.1993</collectingDate>
,
<collectorName>K. Konishi</collectorName>
&amp;
<collectorName>K. Maeto</collectorName>
leg.,
<collectingMethod>coarse Malaise trap</collectingMethod>
, plot 4 [
<specimenCount type="male">1♂</specimenCount>
FRIM],
<collectingDate value="1993-04-09" valueMax="1993-04-16" valueMin="1993-04-09">9-16.IV.1993</collectingDate>
[
<specimenCount type="male">1♂</specimenCount>
NARO];
<location LSID="urn:lsid:plazi:treatment:C7BE87A15935594A9F4F197B961E885C:94A9649040F7B8D43AA69869523D75F9" country="Malaysia" county="Pasoh Forest Reserve" municipality="Pasoh Forest Reserve" name="Pasoh Forest Reserve" stateProvince="Negeri Sembilan">Pasoh Forest Reserve</location>
,
<collectingDate value="1993-04-03" valueMax="1993-04-09" valueMin="1993-04-03">3-9.IV.1993</collectingDate>
,
<collectorName>K. Konishi</collectorName>
&amp;
<collectorName>K. Maeto</collectorName>
leg.,
<collectingMethod>fine Malaise trap</collectingMethod>
, plot 4 [
<specimenCount type="male">1♂</specimenCount>
FRIM];
<location LSID="urn:lsid:plazi:treatment:C7BE87A15935594A9F4F197B961E885C:3C8980E3BC00A708BC30FA0B31FC6B25" country="Malaysia" county="Pasoh Forest Reserve" municipality="Pasoh Forest Reserve" name="Pasoh Forest Reserve" stateProvince="Negeri Sembilan">Pasoh Forest Reserve</location>
,
<collectingDate value="1993-11-26" valueMax="1993-12-03" valueMin="1993-11-26">26.XI-3.XII.1993</collectingDate>
,
<collectorName>K. Maeto</collectorName>
leg.,
<collectingMethod>coarse Malaise trap</collectingMethod>
, plot 1 [
<specimenCount type="male">2♂</specimenCount>
FRIM, NARO];
<location LSID="urn:lsid:plazi:treatment:C7BE87A15935594A9F4F197B961E885C:2158D92A24B5E71EA0BB4FA0CCA2618C" country="Malaysia" county="Pasoh Forest Reserve" municipality="Pasoh Forest Reserve" name="Pasoh Forest Reserve" stateProvince="Negeri Sembilan">Pasoh Forest Reserve</location>
,
<collectingDate value="1993-11-26" valueMax="1993-12-03" valueMin="1993-11-26">26.XI-3.XII.1993</collectingDate>
,
<collectorName>K. Maeto</collectorName>
leg.,
<collectingMethod>coarse Malaise trap</collectingMethod>
, plot 4 [
<specimenCount type="male">2♂</specimenCount>
FRIM, NARO];
<location LSID="urn:lsid:plazi:treatment:C7BE87A15935594A9F4F197B961E885C:A2911BBDFCC9F1A59B85D81FE8176D3E" country="Malaysia" county="Pasoh Forest Reserve" municipality="Pasoh Forest Reserve" name="Pasoh Forest Reserve" stateProvince="Negeri Sembilan">Pasoh Forest Reserve</location>
,
<collectingDate value="1993-11-26" valueMax="1993-12-03" valueMin="1993-11-26">26.XI-3.XII.1993</collectingDate>
,
<collectorName>K. Maeto</collectorName>
leg.,
<collectingMethod>fine Malaise trap</collectingMethod>
, plot 1 [
<specimenCount type="male">2♂</specimenCount>
FRIM, NARO],
<collectingDate value="1993-12-10" valueMax="1993-12-17" valueMin="1993-12-10">10-17.XII.1993</collectingDate>
[
<specimenCount type="male">1♂</specimenCount>
FRIM];
<location LSID="urn:lsid:plazi:treatment:C7BE87A15935594A9F4F197B961E885C:4E0BBB4F026E46327BC6A1C9D06A6717" country="Malaysia" county="Pasoh Forest Reserve" municipality="Pasoh Forest Reserve" name="Pasoh Forest Reserve" stateProvince="Negeri Sembilan">Pasoh Forest Reserve</location>
,
<collectingDate value="1993-11-26" valueMax="1993-12-03" valueMin="1993-11-26">26.XI-3.XII.1993</collectingDate>
,
<collectorName>K. Maeto</collectorName>
leg.,
<collectingMethod>fine Malaise trap</collectingMethod>
, plot 3 [
<specimenCount type="male">1♂</specimenCount>
NARO],
<collectingDate value="1993-12-10" valueMax="1993-12-17" valueMin="1993-12-10">10-17.XII.1993</collectingDate>
[
<specimenCount type="male">1♂</specimenCount>
FRIM];
<location LSID="urn:lsid:plazi:treatment:C7BE87A15935594A9F4F197B961E885C:9BAC3D18B886620C1E5E585A7FB016B3" country="Malaysia" county="Pasoh Forest Reserve" municipality="Pasoh Forest Reserve" name="Pasoh Forest Reserve" stateProvince="Negeri Sembilan">Pasoh Forest Reserve</location>
,
<collectingDate value="1993-11-26" valueMax="1993-12-03" valueMin="1993-11-26">26.XI-3.XII.1993</collectingDate>
,
<collectorName>K. Maeto</collectorName>
leg.,
<collectingMethod>fine Malaise trap</collectingMethod>
, plot 4 [
<specimenCount type="male">1♂</specimenCount>
NARO],
<collectingDate value="1993-12-03" valueMax="1993-12-10" valueMin="1993-12-03">3-10.XII.1993</collectingDate>
[
<specimenCount type="male">1♂</specimenCount>
NARO];
<location LSID="urn:lsid:plazi:treatment:C7BE87A15935594A9F4F197B961E885C:3BA3560E470F4510B211894EB8AACD69" country="Malaysia" county="Pasoh Forest Reserve" municipality="Pasoh Forest Reserve" name="Pasoh Forest Reserve" stateProvince="Negeri Sembilan">Pasoh Forest Reserve</location>
,
<collectingDate value="1993-12-03" valueMax="1993-12-10" valueMin="1993-12-03">3-10.XII.1993</collectingDate>
,
<collectorName>K. Maeto</collectorName>
leg.,
<collectingMethod>coarse Malaise trap</collectingMethod>
, plot 6 [
<specimenCount type="male">1♂</specimenCount>
NARO];
<location LSID="urn:lsid:plazi:treatment:C7BE87A15935594A9F4F197B961E885C:97EB2F7B60F6C8CD758D2D459EE03837" country="Malaysia" county="Pasoh Forest Reserve" municipality="Pasoh Forest Reserve" name="Pasoh Forest Reserve" stateProvince="Negeri Sembilan">Pasoh Forest Reserve</location>
,
<collectingDate value="1993-12-10" valueMax="1993-12-17" valueMin="1993-12-10">10-17.XII.1993</collectingDate>
,
<collectorName>K. Maeto</collectorName>
leg.,
<collectingMethod>coarse Malaise trap</collectingMethod>
, plot 2 [
<specimenCount type="male">1♂</specimenCount>
FRIM];
<location LSID="urn:lsid:plazi:treatment:C7BE87A15935594A9F4F197B961E885C:AF25677E03D84412AF90060AB1F588BB" country="Malaysia" county="Pasoh Forest Reserve" municipality="Pasoh Forest Reserve" name="Pasoh Forest Reserve" stateProvince="Negeri Sembilan">Pasoh Forest Reserve</location>
,
<collectingDate value="1993-12-10" valueMax="1993-12-17" valueMin="1993-12-10">10-17.XII.1993</collectingDate>
,
<collectorName>K. Maeto</collectorName>
leg.,
<collectingMethod>fine Malaise trap</collectingMethod>
, plot 2 [
<specimenCount type="male">2♂</specimenCount>
FRIM, NARO]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="identification source">
<paragraph pageId="0" pageNumber="145">Identification source.</paragraph>
<paragraph pageId="0" pageNumber="145">
Keyed using
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" author="Mickel, CE" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="145" pagination="177 - 312" refId="B20" refString="Mickel, CE, 1935. The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae). Transactions of the Royal Entomological Society of London 83 (2): 177 - 312, DOI: https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" title="The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae)." url="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" volume="83" year="1935">Mickel (1935)</bibRefCitation>
and compared with the original description. No differences were found between the specimens examined herein and
<normalizedToken originalValue="Cameron’s">Cameron's</normalizedToken>
description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
Indonesia: Kalimantan Timur (
<bibRefCitation DOI="https://doi.org/10.3759/tropics.6.91" author="Lelej, AS" journalOrPublisher="Tropics" pageId="0" pageNumber="145" pagination="91 - 104" refId="B11" refString="Lelej, AS, 1996b. Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae). Tropics 6 (1-2): 91 - 104, DOI: https://doi.org/10.3759/tropics.6.91" title="Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae)." url="https://doi.org/10.3759/tropics.6.91" volume="6" year="1996 b">Lelej 1996b</bibRefCitation>
). Malaysia: Negeri Sembilan*, Sarawak (
<bibRefCitation author="Cameron, P" journalOrPublisher="Journal of the Straits Branch of the Royal Asiatic Society" pageId="0" pageNumber="145" pagination="29 - 131" refId="B2" refString="Cameron, P, 1902. On the Hymenoptera collected by Mr. Robert Shelford at Sarawak, and on the Hymenoptera of the Sarawak Museum. Journal of the Straits Branch of the Royal Asiatic Society 37: 29 - 131" title="On the Hymenoptera collected by Mr. Robert Shelford at Sarawak, and on the Hymenoptera of the Sarawak Museum." volume="37" year="1902">Cameron 1902</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1080/00222933708655284" author="Mickel, CE" journalOrPublisher="The Annals and Magazine of Natural History" pageId="0" pageNumber="145" pagination="441 - 456" refId="B21" refString="Mickel, CE, 1937. New species and records of Mutillidae (Hymenoptera) from Borneo and the Solomon Islands. The Annals and Magazine of Natural History 10 (19): 441 - 456, DOI: https://doi.org/10.1080/00222933708655284" title="New species and records of Mutillidae (Hymenoptera) from Borneo and the Solomon Islands." url="https://doi.org/10.1080/00222933708655284" volume="10" year="1937">Mickel 1937</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="remarks">
<paragraph pageId="0" pageNumber="145">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="145">
This is the only species in
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla</emphasis>
</taxonomicName>
with F1 cylindrical, like most species of
<taxonomicName authorityName="Latreille" authorityYear="1802" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Mutillinae">Mutillinae</taxonomicName>
. Similarly, the species of East Asian genera of the former
<taxonomicName authorityName="Lelej" authorityYear="1996" lsidName="" pageId="0" pageNumber="145" rank="tribe" tribe="Petersenidiini">Petersenidiini</taxonomicName>
, like
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Krombeinidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krombeinidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Krombeinidia</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Lelej" authorityYear="1996" class="Insecta" family="Mutillidae" genus="Orientidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Orientidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Orientidia</emphasis>
</taxonomicName>
Lelej, 1996,
<taxonomicName authorityName="Lelej" authorityYear="1996" class="Insecta" family="Mutillidae" genus="Pagdenidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pagdenidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Pagdenidia</emphasis>
</taxonomicName>
Lelej, 1996,
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Petersenidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Petersenidia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Petersenidia</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Tsuneki" authorityYear="1993" class="Insecta" family="Mutillidae" genus="Taiwanomyrme" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Taiwanomyrme" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Taiwanomyrme</emphasis>
</taxonomicName>
Tsuneki, 1993, and
<taxonomicName authorityName="Tsuneki" authorityYear="1993" class="Insecta" family="Mutillidae" genus="Zavatilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zavatilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Zavatilla</emphasis>
</taxonomicName>
Tsuneki, 1993, have the F1 cylindrical. I presume that this is a plesiomorphic state in
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla</emphasis>
</taxonomicName>
, and
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada">
<emphasis italics="true" pageId="0" pageNumber="145">A. bagrada</emphasis>
</taxonomicName>
is sister to the remaining members of the genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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