207 lines
34 KiB
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207 lines
34 KiB
XML
<document id="DD14E9B9F083559AE94FD22EABFD4F66" ID-DOI="10.11646/zootaxa.991.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5046283" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625062394442" checkinUser="felipe" docAuthor="Blahnik Table Of Contents, Roger J." docDate="2005" docId="EC4BE952FFCB6807FEBDCB578E59E9E5" docLanguage="en" docName="zt00991.pdf" docOrigin="Zootaxa 991 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.991.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Alterosa Blahnik Table Of Contents 2005, new genus" docType="treatment" docVersion="3" lastPageNumber="13" masterDocId="1072912AFFC2680BFFB5CA3E8D48EC73" masterDocTitle="Alterosa, a new caddisfly genus from Brazil (Trichoptera: Philopotamidae)" masterLastPageNumber="60" masterPageNumber="1" pageNumber="10" updateTime="1698979645207" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="365DCCF25B5A38A4559AFF64EA867227">Alterosa, a new caddisfly genus from Brazil (Trichoptera: Philopotamidae)</mods:title>
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<mods:namePart id="860A2471644FD1427B93773500B019AC">Blahnik Table Of Contents, Roger J.</mods:namePart>
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<mods:title id="44C9E2C4702ED51EB801D5A7DBCBF630">Zootaxa</mods:title>
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<mods:date id="76B59304DA946E1F110EE4BA4392CE53">2005</mods:date>
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<mods:number id="98D86B6B6A3F8FF0172EAE4E0054ED21">991</mods:number>
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<treatment id="EC4BE952FFCB6807FEBDCB578E59E9E5" LSID="urn:lsid:plazi:treatment:EC4BE952FFCB6807FEBDCB578E59E9E5" httpUri="http://treatment.plazi.org/id/EC4BE952FFCB6807FEBDCB578E59E9E5" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
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<subSubSection id="2CF80BCFFFCB6802FEBDCB578CB6EDF0" box="[264,510,361,387]" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="645D5844FFCB6802FEBDCB578CB6EDF0" blockId="9.[264,510,361,387]" box="[264,510,361,387]" pageId="9" pageNumber="10">
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<heading id="3F15EF28FFCB6802FEBDCB578CB6EDF0" bold="true" box="[264,510,361,387]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
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<emphasis id="56968456FFCB6802FEBDCB578CB6EDF0" bold="true" box="[264,510,361,387]" pageId="9" pageNumber="10">
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<taxonomicName id="A3E223C7FFCB6802FEBDCB578C25EDF0" authority="Blahnik Table Of Contents, 2005" authorityName="Blahnik Table Of Contents" authorityYear="2005" box="[264,365,361,387]" class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="56968456FFCB6802FEBDCB578C25EDF0" bold="true" box="[264,365,361,387]" italics="true" pageId="9" pageNumber="10">Alterosa</emphasis>
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,
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<taxonomicNameLabel id="4DA5392DFFCB6802FEC9CB578CB6EDF0" box="[380,510,361,387]" pageId="9" pageNumber="10" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="645D5844FFCB6802FEBDCB818E65EDAA" blockId="9.[264,813,447,473]" box="[264,813,447,473]" pageId="9" pageNumber="10">
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<typeStatus id="BB59E6E6FFCB6802FEBDCB818C0AEDAA" box="[264,322,447,473]" pageId="9" pageNumber="10">Type</typeStatus>
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species:
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<taxonomicName id="A3E223C7FFCB6802FE00CB818FC2EDAB" authority="Blahnik Table Of Contents, 2005" authorityName="Blahnik Table Of Contents" authorityYear="2005" box="[437,650,447,472]" class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bocainae" status="sp. nov.">
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<emphasis id="56968456FFCB6802FE00CB818FC2EDAB" box="[437,650,447,472]" italics="true" pageId="9" pageNumber="10">Alterosa bocainae</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="4DA5392DFFCB6802FD2CCB818E61EDAA" box="[665,809,447,473]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
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.
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</paragraph>
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<paragraph id="645D5844FFCB6802FEBDC82A8F11E8DD" blockId="9.[264,1325,532,1878]" pageId="9" pageNumber="10">
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The single character most useful for diagnosing
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<taxonomicName id="A3E223C7FFCB6802FC89C82A8ED6EE5E" authorityName="Blahnik Table Of Contents" authorityYear="2005" box="[828,926,532,557]" class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
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<emphasis id="56968456FFCB6802FC89C82A8ED6EE5E" box="[828,926,532,557]" italics="true" pageId="9" pageNumber="10">Alterosa</emphasis>
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</taxonomicName>
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, and which may be uniquely apomorphic, is the structure of the phallobase of the male (
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<figureCitation id="FCD944C1FFCB6802FC3FC8028EBAEE25" box="[906,1010,572,598]" captionStart="FIGURE 6" captionStartId="15.[264,367,1579,1603]" captionTargetBox="[391,1214,447,1560]" captionTargetId="figure-132@15.[385,1224,427,1567]" captionTargetPageId="15" captionText="FIGURE 6. Alterosa bocainae, new species. Male genitalia: 6A—lateral view; 6B—dorsal view of segments IX and X; 6C—inferior appendage, dorsal view; 6D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046299" httpUri="https://zenodo.org/record/5046299/files/figure.png" pageId="9" pageNumber="10">Figs. 6D</figureCitation>
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,
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<figureCitation id="FCD944C1FFCB6802FC4BC802896CEE25" box="[1022,1060,572,598]" captionStart="FIGURE 8" captionStartId="19.[264,368,1665,1689]" captionTargetBox="[321,1264,430,1621]" captionTargetId="figure-61@19.[305,1282,409,1638]" captionTargetPageId="19" captionText="FIGURE 8. Alterosa caparaonensis, new species. Male genitalia: 8A—lateral view; 8B—dorsal view of segments IX and X; 8C—inferior appendage, dorsal view; 8D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046303" httpUri="https://zenodo.org/record/5046303/files/figure.png" pageId="9" pageNumber="10">8D</figureCitation>
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,
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<figureCitation id="FCD944C1FFCB6802FB84C8028910EE25" box="[1073,1112,572,598]" captionStart="FIGURE 9" captionStartId="21.[264,367,1473,1497]" captionTargetBox="[350,1237,291,1446]" captionTargetId="figure-108@21.[335,1252,281,1449]" captionTargetPageId="21" captionText="FIGURE 9. Alterosa escova, new species. Male genitalia: 9A—lateral view; 9B—dorsal view of segments IX and X; 9C—inferior appendage, dorsal view; 9D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046305" httpUri="https://zenodo.org/record/5046305/files/figure.png" pageId="9" pageNumber="10">9D</figureCitation>
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), which is tubular basally and lacks the basodorsal expansion common in most philopotamids, including species in the subfamily
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<taxonomicName id="A3E223C7FFCB6802FDB5C8B28FDEEED5" authorityName="P Rambur" authorityYear="1842" box="[512,662,652,678]" class="Insecta" family="Philopotamidae" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subFamily" subFamily="Chimarrinae">Chimarrinae</taxonomicName>
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. Other characters of the male genitalia useful, in combination, to diagnose the genus include: intermediate appendages positioned mesally to the preanal appendages (
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<figureCitation id="FCD944C1FFCB6802FE48C8E28FC9EE85" box="[509,641,732,758]" captionStart="FIGURE 6" captionStartId="15.[264,367,1579,1603]" captionTargetBox="[391,1214,447,1560]" captionTargetId="figure-132@15.[385,1224,427,1567]" captionTargetPageId="15" captionText="FIGURE 6. Alterosa bocainae, new species. Male genitalia: 6A—lateral view; 6B—dorsal view of segments IX and X; 6C—inferior appendage, dorsal view; 6D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046299" httpUri="https://zenodo.org/record/5046299/files/figure.png" pageId="9" pageNumber="10">Figs. 6A,B</figureCitation>
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;
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<figureCitation id="FCD944C1FFCB6802FD38C8E28F95EE85" box="[653,733,732,758]" captionStart="FIGURE 10" captionStartId="22.[264,367,1796,1820]" captionTargetBox="[324,1279,540,1756]" captionTargetId="figure-93@22.[305,1282,529,1769]" captionTargetPageId="22" captionText="FIGURE 10. Alterosa falcata, new species. Male genitalia: 10A—lateral view; 10B—dorsal view of segments IX and X; 10C—inferior appendage, dorsal view; 10D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046307" httpUri="https://zenodo.org/record/5046307/files/figure.png" pageId="9" pageNumber="10">10A,B</figureCitation>
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), absence of ventral processes on any of the terminal segments of the male genitalia, and sternum IX with the posteroventral margin distinctly produced and widely truncate ventrally (
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<figureCitation id="FCD944C1FFCB6802FC89C9128E89EF35" box="[828,961,812,838]" captionStart="FIGURE 9" captionStartId="21.[264,367,1473,1497]" captionTargetBox="[350,1237,291,1446]" captionTargetId="figure-108@21.[335,1252,281,1449]" captionTargetPageId="21" captionText="FIGURE 9. Alterosa escova, new species. Male genitalia: 9A—lateral view; 9B—dorsal view of segments IX and X; 9C—inferior appendage, dorsal view; 9D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046305" httpUri="https://zenodo.org/record/5046305/files/figure.png" pageId="9" pageNumber="10">Figs. 9A,B</figureCitation>
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;
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<figureCitation id="FCD944C1FFCB6802FC7AC9128955EF35" box="[975,1053,812,838]" captionStart="FIGURE 10" captionStartId="22.[264,367,1796,1820]" captionTargetBox="[324,1279,540,1756]" captionTargetId="figure-93@22.[305,1282,529,1769]" captionTargetPageId="22" captionText="FIGURE 10. Alterosa falcata, new species. Male genitalia: 10A—lateral view; 10B—dorsal view of segments IX and X; 10C—inferior appendage, dorsal view; 10D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046307" httpUri="https://zenodo.org/record/5046307/files/figure.png" pageId="9" pageNumber="10">10A,B</figureCitation>
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) and the posterodorsal margin
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<emphasis id="56968456FFCB6802FED4C96A8CFEEF1E" box="[353,438,852,877]" italics="true" pageId="9" pageNumber="10">acutely</emphasis>
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articulating with the tergum (
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<figureCitation id="FCD944C1FFCB6802FCA5C96A8E3FEF1D" box="[784,887,852,878]" captionStart="FIGURE 6" captionStartId="15.[264,367,1579,1603]" captionTargetBox="[391,1214,447,1560]" captionTargetId="figure-132@15.[385,1224,427,1567]" captionTargetPageId="15" captionText="FIGURE 6. Alterosa bocainae, new species. Male genitalia: 6A—lateral view; 6B—dorsal view of segments IX and X; 6C—inferior appendage, dorsal view; 6D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046299" httpUri="https://zenodo.org/record/5046299/files/figure.png" pageId="9" pageNumber="10">Figs. 6A</figureCitation>
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,
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<figureCitation id="FCD944C1FFCB6802FC36C96A8EE3EF1D" box="[899,939,852,878]" captionStart="FIGURE 9" captionStartId="21.[264,367,1473,1497]" captionTargetBox="[350,1237,291,1446]" captionTargetId="figure-108@21.[335,1252,281,1449]" captionTargetPageId="21" captionText="FIGURE 9. Alterosa escova, new species. Male genitalia: 9A—lateral view; 9B—dorsal view of segments IX and X; 9C—inferior appendage, dorsal view; 9D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046305" httpUri="https://zenodo.org/record/5046305/files/figure.png" pageId="9" pageNumber="10">9A</figureCitation>
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). Preanal appendages arise from the base of tergum X (
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<figureCitation id="FCD944C1FFCB6802FDACC9428F31EFE5" box="[537,633,892,918]" captionStart="FIGURE 6" captionStartId="15.[264,367,1579,1603]" captionTargetBox="[391,1214,447,1560]" captionTargetId="figure-132@15.[385,1224,427,1567]" captionTargetPageId="15" captionText="FIGURE 6. Alterosa bocainae, new species. Male genitalia: 6A—lateral view; 6B—dorsal view of segments IX and X; 6C—inferior appendage, dorsal view; 6D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046299" httpUri="https://zenodo.org/record/5046299/files/figure.png" pageId="9" pageNumber="10">Fig. 6A</figureCitation>
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,
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<figureCitation id="FCD944C1FFCB6802FD32C9428FF5EFE5" box="[647,701,892,918]" captionStart="FIGURE 10" captionStartId="22.[264,367,1796,1820]" captionTargetBox="[324,1279,540,1756]" captionTargetId="figure-93@22.[305,1282,529,1769]" captionTargetPageId="22" captionText="FIGURE 10. Alterosa falcata, new species. Male genitalia: 10A—lateral view; 10B—dorsal view of segments IX and X; 10C—inferior appendage, dorsal view; 10D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046307" httpUri="https://zenodo.org/record/5046307/files/figure.png" pageId="9" pageNumber="10">10A</figureCitation>
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), rather than the dorsal margin of sternum IX (as is typical in
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<taxonomicName id="A3E223C7FFCB6802FE35C99A8C91EFCE" authorityName="Navas" authorityYear="1918" box="[384,473,932,957]" class="Insecta" family="Philopotamidae" genus="Sortosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
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<emphasis id="56968456FFCB6802FE35C99A8C91EFCE" box="[384,473,932,957]" italics="true" pageId="9" pageNumber="10">Sortosa</emphasis>
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</taxonomicName>
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, see
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<bibRefCitation id="007325B5FFCB6802FDADC99A8FDEEFCD" author="Ross, H. H." box="[536,662,932,958]" pageId="9" pageNumber="10" refId="ref24382" refString="Ross, H. H. (1956) Evolution and Classification of the Mountain Caddisflies. Urbana, University of Illinois Press, 213 pp." type="book" year="1956">Ross 1956</bibRefCitation>
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, figs. 36A, 37A, 38A). In species in which the preanal appendages are not secondarily modified (
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<figureCitation id="FCD944C1FFCB6802FD4CC9F28E1FEF95" box="[761,855,972,998]" captionStart="FIGURE 9" captionStartId="21.[264,367,1473,1497]" captionTargetBox="[350,1237,291,1446]" captionTargetId="figure-108@21.[335,1252,281,1449]" captionTargetPageId="21" captionText="FIGURE 9. Alterosa escova, new species. Male genitalia: 9A—lateral view; 9B—dorsal view of segments IX and X; 9C—inferior appendage, dorsal view; 9D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046305" httpUri="https://zenodo.org/record/5046305/files/figure.png" pageId="9" pageNumber="10">Fig. 9A</figureCitation>
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), they are relatively simple in structure, elongate and fingerlike (
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<figureCitation id="FCD944C1FFCB6802FD98C9CA8FDEE87D" box="[557,662,1012,1038]" captionStart="FIGURE 6" captionStartId="15.[264,367,1579,1603]" captionTargetBox="[391,1214,447,1560]" captionTargetId="figure-132@15.[385,1224,427,1567]" captionTargetPageId="15" captionText="FIGURE 6. Alterosa bocainae, new species. Male genitalia: 6A—lateral view; 6B—dorsal view of segments IX and X; 6C—inferior appendage, dorsal view; 6D—phallic apparatus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046299" httpUri="https://zenodo.org/record/5046299/files/figure.png" pageId="9" pageNumber="10">Figs. 6A</figureCitation>
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,
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<figureCitation id="FCD944C1FFCB6802FD14C9CA8F9EE87D" box="[673,726,1012,1038]" captionStart="FIGURE 11" captionStartId="24.[264,367,1816,1840]" captionTargetBox="[370,1232,583,1804]" captionTargetId="figure-102@24.[348,1240,578,1808]" captionTargetPageId="24" captionText="FIGURE 11. Alterosa fimbriata, new species. Male genitalia: 11A—lateral view; 11B—dorsal view of segments IX and X; 11C—inferior appendage, dorsal view; 11D—phallic apparatus, lateral view; 11E—dorsal view of segment VIII." figureDoi="http://doi.org/10.5281/zenodo.5046309" httpUri="https://zenodo.org/record/5046309/files/figure.png" pageId="9" pageNumber="10">11A</figureCitation>
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). Venation (
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<figureCitation id="FCD944C1FFCB6802FCDDC9CA8EE3E87D" box="[872,939,1012,1038]" captionStart="FIGURE 1" captionStartId="10.[264,368,1571,1595]" captionTargetBox="[281,1312,547,1481]" captionTargetId="figure-52@10.[273,1319,509,1564]" captionTargetPageId="10" captionText="FIGURE 1. Alterosa bocainae, new species. Wings: 1A—forewing; 1B—hind wing." figureDoi="http://doi.org/10.5281/zenodo.5046285" httpUri="https://zenodo.org/record/5046285/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
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) is primitive for
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<taxonomicName id="A3E223C7FFCB6802FBC7C9CA8862E87D" box="[1138,1322,1012,1038]" class="Insecta" family="Philopotamidae" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Philopotamidae</taxonomicName>
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and very similar to
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<taxonomicName id="A3E223C7FFCB6802FE4CCE228F1AE846" authorityName="Navas" authorityYear="1918" box="[505,594,1052,1077]" class="Insecta" family="Philopotamidae" genus="Sortosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
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<emphasis id="56968456FFCB6802FE4CCE228F1AE846" box="[505,594,1052,1077]" italics="true" pageId="9" pageNumber="10">Sortosa</emphasis>
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</taxonomicName>
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and
|
||
<taxonomicName id="A3E223C7FFCB6802FD27CE228E71E846" authorityName="Tillyard" authorityYear="1924" box="[658,825,1052,1077]" class="Insecta" family="Philopotamidae" genus="Hydrobiosella" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56968456FFCB6802FD27CE228E71E846" box="[658,825,1052,1077]" italics="true" pageId="9" pageNumber="10">Hydrobiosella</emphasis>
|
||
</taxonomicName>
|
||
. Female genitalia (
|
||
<figureCitation id="FCD944C1FFCB6802FB9ACE228932E845" box="[1071,1146,1052,1078]" captionStart="FIGURE 4" captionStartId="12.[264,368,1854,1878]" captionTargetBox="[302,1285,1489,1828]" captionTargetId="figure-372@12.[302,1285,1489,1828]" captionTargetPageId="12" captionText="FIGURE 4. Alterosa bocainae, new species. Female genitalia—lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046289" httpUri="https://zenodo.org/record/5046289/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
|
||
), like those of most genera of
|
||
<taxonomicName id="A3E223C7FFCB6802FE0BCE7A8F31E82D" authorityName=", Ross" authorityYear="1956" box="[446,633,1092,1118]" class="Insecta" family="Philopotamidae" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subFamily" subFamily="Philopotaminae">Philopotaminae</taxonomicName>
|
||
, excluding
|
||
<taxonomicName id="A3E223C7FFCB6802FD4ACE7A8ED5E82E" authorityName="Ulmer" authorityYear="1909" box="[767,925,1092,1117]" class="Insecta" family="Philopotamidae" genus="Dolophilodes" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56968456FFCB6802FD4ACE7A8ED5E82E" box="[767,925,1092,1117]" italics="true" pageId="9" pageNumber="10">Dolophilodes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A3E223C7FFCB6802FC1ECE7A895DE82E" authorityName="Ross" authorityYear="1956" box="[939,1045,1092,1117]" class="Insecta" family="Philopotamidae" genus="Fumonta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56968456FFCB6802FC1ECE7A895DE82E" box="[939,1045,1092,1117]" italics="true" pageId="9" pageNumber="10">Fumonta</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="A3E223C7FFCB6802FBE0CE7A89B0E82E" authorityName="Stephens" authorityYear="1829" box="[1109,1272,1092,1117]" class="Insecta" family="Philopotamidae" genus="Philopotamus" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56968456FFCB6802FBE0CE7A89B0E82E" box="[1109,1272,1092,1117]" italics="true" pageId="9" pageNumber="10">Philopotamus</emphasis>
|
||
</taxonomicName>
|
||
, are elongate and tapered, with very long, articulated apodemes from the anterior margin of sternum VIII and tergum IX.
|
||
</paragraph>
|
||
<paragraph id="645D5844FFCB6801FE8DCE828F12ED12" blockId="9.[264,1325,532,1878]" lastBlockId="10.[264,1322,284,353]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||
Adult. Color of pinned, dry specimens dark brown overall with contrasting small light brown or yellowish spots on forewings, these sometimes arranged in irregularly linear arrays near costal margins; legs, palps, and antennae usually paler in color. Three ocelli present, prominent. Eyes with short projecting setae between the ommatidia. Maxillary palps (
|
||
<figureCitation id="FCD944C1FFCB6802FEE9CF628CE0E905" box="[348,424,1372,1398]" captionStart="FIGURE 2" captionStartId="11.[264,367,1662,1686]" captionTargetBox="[290,1296,289,1643]" captionTargetId="figure-70@11.[290,1296,289,1644]" captionTargetPageId="11" captionText="FIGURE 2. Alterosa bocainae, new species. Head and palps: 2A—lateral view; 2B—dorsal view of maxillary palp segments 1–3. FIGURE 3. Alterosa bocainae, new species. Setal warts of head and thorax—dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5046287" httpUri="https://zenodo.org/record/5046287/files/figure.png" pageId="9" pageNumber="10">Fig. 2</figureCitation>
|
||
) 5segmented, relatively short: 1st segment very short, usually broadly joined (often apparently fused) to 2nd segment, usually with a few setae on apicomesal margin; 2nd segment short, globose, with numerous elongate setae on apicomesal surface; 3rd segment relatively short, about 3 or 4x length of segment 2; 4th segment very short, about 2x as long as wide; 5th segment longer than segment 3, variable in length among species. Setal warts of head (Fig. 3) well developed; posterolateral warts large, typical of
|
||
<taxonomicName id="A3E223C7FFCB6802FEBDCC728C8CEA15" box="[264,452,1612,1638]" class="Insecta" family="Philopotamidae" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Philopotamidae</taxonomicName>
|
||
; posterior setal warts rounded, separated mesally; anteromesal setal wart large, subtriangular, notched mesally to accommodate anterior ocellus, laterally fused to anterior setal warts; fused anterior and anteromesal warts roughly Mshaped. Prothorax with rounded mesal setal warts, slightly separated mesally, laterally with indistinctly defined setal areas. Wing venation (
|
||
<figureCitation id="FCD944C1FFCB6802FD71CCD28E43EB75" box="[708,779,1772,1798]" captionStart="FIGURE 1" captionStartId="10.[264,368,1571,1595]" captionTargetBox="[281,1312,547,1481]" captionTargetId="figure-52@10.[273,1319,509,1564]" captionTargetPageId="10" captionText="FIGURE 1. Alterosa bocainae, new species. Wings: 1A—forewing; 1B—hind wing." figureDoi="http://doi.org/10.5281/zenodo.5046285" httpUri="https://zenodo.org/record/5046285/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
|
||
) primitive (forewing with forks 1–5 present, hind wing with fork 4 missing, anal veins of hind wing reaching wing margin); fork 1 of forewing branching just proximal to discal crossvein;
|
||
<emphasis id="56968456FFCB6802FCCACD028EC2EB26" box="[895,906,1852,1877]" italics="true" pageId="9" pageNumber="10">s</emphasis>
|
||
,
|
||
<emphasis id="56968456FFCB6802FC2CCD028E8BEB26" box="[921,963,1852,1877]" italics="true" pageId="9" pageNumber="10">rm</emphasis>
|
||
, and m crossveins of forewing nearly linearly arranged, these plus
|
||
<emphasis id="56968456FFC86801FD0ECB228FBDED46" box="[699,757,284,309]" italics="true" pageId="10" pageNumber="11">mcu</emphasis>
|
||
, base of fork 3, and apex of Cu
|
||
<subScript id="F8665A01FFC86801FB35CB1689C2ED4A" attach="left" box="[1152,1162,296,313]" fontSize="7" pageId="10" pageNumber="11">2</subScript>
|
||
clear, unpigmented. Spur formula 2:4:4.
|
||
</paragraph>
|
||
<caption id="309D08CCFFC86801FEBDCC1D89E2EA49" ID-DOI="http://doi.org/10.5281/zenodo.5046285" ID-Zenodo-Dep="5046285" box="[264,1194,1571,1595]" httpUri="https://zenodo.org/record/5046285/files/figure.png" pageId="10" pageNumber="11" startId="10.[264,368,1571,1595]" targetBox="[281,1312,547,1481]" targetPageId="10">
|
||
<paragraph id="645D5844FFC86801FEBDCC1D89E2EA49" blockId="10.[264,1194,1571,1595]" box="[264,1194,1571,1595]" pageId="10" pageNumber="11">
|
||
<emphasis id="56968456FFC86801FEBDCC1D8CC2EA48" bold="true" box="[264,394,1571,1595]" pageId="10" pageNumber="11">FIGURE 1.</emphasis>
|
||
<taxonomicName id="A3E223C7FFC86801FE2DCC1D8F12EA49" authority="Blahnik Table Of Contents, 2005" authorityName="Blahnik Table Of Contents" authorityYear="2005" box="[408,602,1571,1594]" class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bocainae" status="sp. nov.">
|
||
<emphasis id="56968456FFC86801FE2DCC1D8F12EA49" box="[408,602,1571,1594]" italics="true" pageId="10" pageNumber="11">Alterosa bocainae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="4DA5392DFFC86801FDD2CC1D8FA2EA49" box="[615,746,1571,1594]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
|
||
. Wings: 1A—forewing; 1B—hind wing.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="309D08CCFFC96800FEBDCC408F5DEAA9" ID-DOI="http://doi.org/10.5281/zenodo.5046287" ID-Zenodo-Dep="5046287" httpUri="https://zenodo.org/record/5046287/files/figure.png" pageId="11" pageNumber="12" startId="11.[264,367,1662,1686]" targetBox="[290,1296,289,1643]" targetPageId="11">
|
||
<paragraph id="645D5844FFC96800FEBDCC408F5DEAA9" blockId="11.[264,1324,1662,1754]" pageId="11" pageNumber="12">
|
||
<emphasis id="56968456FFC96800FEBDCC408CC2EAE5" bold="true" box="[264,394,1662,1686]" pageId="11" pageNumber="12">FIGURE 2.</emphasis>
|
||
<taxonomicName id="A3E223C7FFC96800FE2DCC408F12EAE6" authority="Blahnik Table Of Contents, 2005" authorityName="Blahnik Table Of Contents" authorityYear="2005" box="[408,602,1662,1685]" class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bocainae" status="sp. nov.">
|
||
<emphasis id="56968456FFC96800FE2DCC408F12EAE6" box="[408,602,1662,1685]" italics="true" pageId="11" pageNumber="12">Alterosa bocainae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="4DA5392DFFC96800FDD2CC408FA4EAE6" box="[615,748,1662,1685]" pageId="11" pageNumber="12" rank="species">new species</taxonomicNameLabel>
|
||
. Head and palps: 2A—lateral view; 2B—dorsal view of maxillary palp segments 1–3.
|
||
<emphasis id="56968456FFC96800FDDFCC9E8FA5EACB" bold="true" box="[618,749,1696,1720]" pageId="11" pageNumber="12">FIGURE 3.</emphasis>
|
||
<taxonomicName id="A3E223C7FFC96800FD49CC9F8EF7EACB" authority="Blahnik Table Of Contents, 2005" authorityName="Blahnik Table Of Contents" authorityYear="2005" box="[764,959,1697,1720]" class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bocainae" status="sp. nov.">
|
||
<emphasis id="56968456FFC96800FD49CC9F8EF7EACB" box="[764,959,1697,1720]" italics="true" pageId="11" pageNumber="12">Alterosa bocainae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="4DA5392DFFC96800FC78CC9F8919EACB" box="[973,1105,1697,1720]" pageId="11" pageNumber="12" rank="species">new species</taxonomicNameLabel>
|
||
. Setal warts of head and thorax—dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="645D5844FFC96800FE8DCD2D8863EB26" blockId="11.[264,1323,1811,1877]" pageId="11" pageNumber="12">Male Genitalia. At least some abdominal terga, through and including tergum VIII, with unpigmented spots surrounding elongate setae on posterior margins. Abdominal</paragraph>
|
||
<paragraph id="645D5844FFCE6807FEBDCB228EA1EFAD" blockId="12.[264,1323,284,1430]" pageId="12" pageNumber="13">
|
||
sterna VII, VIII, and IX without ventromesal processes. Tergum VIII usually with distinct posteromesal invagination (
|
||
<figureCitation id="FCD944C1FFCE6807FDE6CB7A8FF7ED2D" box="[595,703,324,350]" captionStart="FIGURE 11" captionStartId="24.[264,367,1816,1840]" captionTargetBox="[370,1232,583,1804]" captionTargetId="figure-102@24.[348,1240,578,1808]" captionTargetPageId="24" captionText="FIGURE 11. Alterosa fimbriata, new species. Male genitalia: 11A—lateral view; 11B—dorsal view of segments IX and X; 11C—inferior appendage, dorsal view; 11D—phallic apparatus, lateral view; 11E—dorsal view of segment VIII." figureDoi="http://doi.org/10.5281/zenodo.5046309" httpUri="https://zenodo.org/record/5046309/files/figure.png" pageId="12" pageNumber="13">Fig. 11E</figureCitation>
|
||
), but sometimes absent or very weakly developed. Sternum IX with posteroventral margin greatly produced; as viewed ventrally, broadly truncate, usually with slight mesal invagination, with distinct mesal suture line extending from anterior to posterior margin; posterodorsal margin acutely articulated with tergum, and below dorsal development of tergum X. Tergum IX greatly reduced or fused to base of tergum X, sometimes developed into mesal or lateral projections extending over base of tergum X. Tergum X moderately to greatly elongate, entire, apically with numerous short sensilla. Intermediate appendage mesal to preanal appendage, very differently developed in different species, often with apical spines or setae. Preanal appendage inserted membranously or apparently partially fused to base of tergum X, elongate linear and basally constricted in most species, highly modified in shape, with apical armature of spines or setae in others. Inferior appendages elongate linear, with 2 articles, varying in relative length among species; apex of 2nd article with pad of short, stiff setae on mesal surface. Phallic apparatus with phallobase tubular, without basodorsal expansion, usually somewhat curved, sometimes distinctly bent; endotheca highly variable, usually with included spines, these varying greatly in size, number and position among different species; phallotremal sclerite usually small, weakly sclerotized and indistinct.
|
||
</paragraph>
|
||
<paragraph id="645D5844FFCE6807FE8DC9D28E59E9E5" blockId="12.[264,1323,284,1430]" pageId="12" pageNumber="13">
|
||
Female Genitalia. Genitalia (
|
||
<figureCitation id="FCD944C1FFCE6807FD17C9D28FA5E875" box="[674,749,1004,1030]" captionStart="FIGURE 4" captionStartId="12.[264,368,1854,1878]" captionTargetBox="[302,1285,1489,1828]" captionTargetId="figure-372@12.[302,1285,1489,1828]" captionTargetPageId="12" captionText="FIGURE 4. Alterosa bocainae, new species. Female genitalia—lateral view." figureDoi="http://doi.org/10.5281/zenodo.5046289" httpUri="https://zenodo.org/record/5046289/files/figure.png" pageId="12" pageNumber="13">Fig. 4</figureCitation>
|
||
) elongate and tapered. Segment VII distinctly elongate, pleural region more or less pleated. Intersegmental region between segments VII and VIII membranous and relatively elongate. Segment VIII with tergum reduced, tapered posteriorly; sternum with elongate, rodlike, articulated apodemes from anterolateral margins. Segment IX reduced and tapered; tergum often more sclerotized along anterior and ventral margins, anterolaterally with elongate, rodlike, articulated apodemes; sternum less sclerotized, often nearly membranous. Segment X comprised of pair of bulbous, setose lobes, each terminating in short apical cercus. Vaginal apparatus usually membranous and often more or less indistinct, anteriorly with cupped sclerite. Etymology.
|
||
<taxonomicName id="A3E223C7FFCE6807FE64CF6A8F7BE91E" authorityName="Blahnik Table Of Contents" authorityYear="2005" box="[465,563,1364,1389]" class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56968456FFCE6807FE64CF6A8F7BE91E" box="[465,563,1364,1389]" italics="true" pageId="12" pageNumber="13">Alterosa</emphasis>
|
||
</taxonomicName>
|
||
is a name, feminine in gender, traditionally used to refer to the mountains of southern
|
||
<collectingRegion id="A62696A6FFCE6807FDA0CF428FFDE9E5" box="[533,693,1404,1430]" country="Brazil" name="Minas Gerais" pageId="12" pageNumber="13">Minas Gerais</collectingRegion>
|
||
,
|
||
<collectingCountry id="1CF518D4FFCE6807FD74CF428E44E9E5" box="[705,780,1404,1430]" name="Brazil" pageId="12" pageNumber="13">Brazil</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |