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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.284.3983" ID-PMC="PMC3677377" ID-Pensoft-Pub="1313-2970-284-1" ID-Pensoft-UUID="192EFF970E2BFFDF3161FFE53C4BFFAA" ID-PubMed="23794843" ID-Zenodo-Dep="578323" ID-ZooBank="33B15A74746D4A82A865EA1E7E55A9BB" ModsDocID="1313-2970-284-1" checkinTime="1451247469560" checkinUser="pensoft" docAuthor="Shi, Hongliang, Zhou, Hongzhang &amp; Liang, Hongbin" docDate="2013" docId="E32A7D082F73A3D9688F9B4F3BBA73AB" docLanguage="en" docName="ZooKeys 284: 1-129" docOrigin="ZooKeys 284" docPubDate="2013-04-04" docSource="http://dx.doi.org/10.3897/zookeys.284.3983" docTitle="Paraphaea Bates 1873, status resurrected" docType="treatment" docVersion="5" id="192EFF970E2BFFDF3161FFE53C4BFFAA" lastPageNumber="12" masterDocId="192EFF970E2BFFDF3161FFE53C4BFFAA" masterDocTitle="Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera" masterLastPageNumber="129" masterPageNumber="1" pageNumber="10" updateTime="1668155586682" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shi, Hongliang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Graduate School of Chinese Academy of Sciences, Beijing 100039, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhou, Hongzhang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Hongbin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-04-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>284</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>129</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.284.3983</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.284.3983</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-284-1</mods:identifier>
<mods:identifier type="ZooBank">33B15A74746D4A82A865EA1E7E55A9BB</mods:identifier>
<mods:identifier type="Pensoft-UUID">192EFF970E2BFFDF3161FFE53C4BFFAA</mods:identifier>
<mods:identifier type="Zenodo-Dep">578323</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152042239" LSID="urn:lsid:plazi:treatment:E32A7D082F73A3D9688F9B4F3BBA73AB" httpUri="http://treatment.plazi.org/id/E32A7D082F73A3D9688F9B4F3BBA73AB" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
<subSubSection pageId="9" pageNumber="10" type="nomenclature">
<paragraph pageId="9" pageNumber="10">
Genus
<taxonomicName LSID="E32A7D08-2F73-A3D9-688F-9B4F3BBA73AB" authority="Bates, 1873" authorityName="Bates" authorityYear="1873" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paraphaea" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="status resurrected">Paraphaea Bates, 1873</taxonomicName>
<taxonomicNameLabel pageId="9" pageNumber="10">status resurrected</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="reference_group">
<paragraph pageId="9" pageNumber="10">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="status resurrected">Paraphaea</taxonomicName>
<bibRefCitation author="Bates, HW" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="85" pageNumber="86" pagination="229 - 322" refId="B23" refString="Bates, HW, 1873. On the Geodephagous Coleoptera of Japan. Fam. Carabidae. The Transactions of the Entomological Society of London 1873 (2): 229 - 322" title="On the Geodephagous Coleoptera of Japan. Fam. Carabidae." volume="1873" year="1873">Bates 1873</bibRefCitation>
: 312;
<bibRefCitation author="Chaudoir, BM" journalOrPublisher="Bulletin de la Societe Imperiale des Naturalistes de Moscou" pageId="86" pageNumber="87" pagination="188 - 268" refId="B35" refString="Chaudoir, BM, 1877. Genres nouveaux et especes inedites de la famille des Carabiques. Bulletin de la Societe Imperiale des Naturalistes de Moscou 52 (1): 188 - 268" title="Genres nouveaux et especes inedites de la famille des Carabiques." volume="52" year="1877">Chaudoir 1877</bibRefCitation>
: 236 (synonymized with
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="status resurrected">
<emphasis italics="true" pageId="9" pageNumber="10">Anchista</emphasis>
</taxonomicName>
Nietner);
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="87" pageNumber="88" pagination="269 - 579" refId="B58" refString="Jedlicka, A, 1963. Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden 28: 269 - 579" title="Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae)." volume="28" year="1963">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
1963
</bibRefCitation>
: 449 (in part, as a synonym of
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="status resurrected">
<emphasis italics="true" pageId="9" pageNumber="10">Anchista</emphasis>
</taxonomicName>
Nietner);
<bibRefCitation author="Habu, A" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="87" pageNumber="88" refId="B48" refString="Habu, A, 1967. Fauna Japonica, Carabidae Truncatipennes Group (Insecta: Coleoptera). Biogeographical Society of Japan. Japan, 338 pp." title="Fauna Japonica, Carabidae Truncatipennes Group (Insecta: Coleoptera). Biogeographical Society of Japan. Japan, 338 pp." year="1967">Habu 1967</bibRefCitation>
: 137 (in part, as a synonym of
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="status resurrected">
<emphasis italics="true" pageId="9" pageNumber="10">Anchista</emphasis>
</taxonomicName>
Nietner);
<bibRefCitation author="Habu, A" journalOrPublisher="Entomological Review of Japan" pageId="87" pageNumber="88" pagination="83 - 118" refId="B51" refString="Habu, A, 1982. Revised and supplementary notes on and descriptions of the Truncatipennes group of Japan (2) (Coleoptera, Carabidae). Entomological Review of Japan 37 (2): 83 - 118" title="Revised and supplementary notes on and descriptions of the Truncatipennes group of Japan (2) (Coleoptera, Carabidae)." volume="37" year="1982">Habu 1982</bibRefCitation>
: 102 (in part, as a synonym of
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="status resurrected">
<emphasis italics="true" pageId="9" pageNumber="10">Anchista</emphasis>
</taxonomicName>
Nietner).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="type-species">
<paragraph pageId="9" pageNumber="10">Type-species:</paragraph>
<paragraph pageId="9" pageNumber="10">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="signifera">
<emphasis italics="true" pageId="9" pageNumber="10">Paraphaea signifera</emphasis>
</taxonomicName>
Bates, 1873 [=
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Dejean" baseAuthorityYear="1825" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="binotata">
<emphasis italics="true" pageId="9" pageNumber="10">Paraphaea binotata</emphasis>
</taxonomicName>
(Dejean)], by monotypy.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="diagnosis">
<paragraph pageId="9" pageNumber="10">Diagnosis.</paragraph>
<paragraph pageId="9" pageNumber="10">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Paraphaea</emphasis>
</taxonomicName>
Bates can be readily distinguished from most genera of
<taxonomicName genus="Physoderina" lsidName="" pageId="9" pageNumber="10" rank="genus">Physoderina</taxonomicName>
except
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Anchista</emphasis>
</taxonomicName>
Nietner and
<taxonomicName authorityName="Shi &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Metallanchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Metallanchista</emphasis>
</taxonomicName>
gen. n. by the glabrous surface, moderately widened mandibles, and long setae around the aedeagal apical orifice.
</paragraph>
<paragraph pageId="9" pageNumber="10">
Differences between
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Anchista</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Paraphaea</emphasis>
</taxonomicName>
are: (1) pronotum lateral margins slightly angulate in middle in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Anchista</emphasis>
</taxonomicName>
, but completely rounded in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Paraphaea</emphasis>
</taxonomicName>
; (2)
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Anchista</emphasis>
</taxonomicName>
usually with distinct isodiametric microsculpture on the elytra, while
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Paraphaea</emphasis>
</taxonomicName>
has indistinct microsculpture; (3) in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Paraphaea</emphasis>
</taxonomicName>
, males with adhesive hairs on the 1st metatarsomere, but such hairs absent in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Anchista</emphasis>
</taxonomicName>
; (4) males with terminal sternum deeply emarginate in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Paraphaea</emphasis>
</taxonomicName>
, but only shallowly emarginate in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Anchista</emphasis>
</taxonomicName>
; (5) internal sac of aedeagus without main flagellum in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Anchista</emphasis>
</taxonomicName>
, but main flagellum well developed and sinuous in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Paraphaea</emphasis>
</taxonomicName>
; (6) in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Paraphaea</emphasis>
</taxonomicName>
, the spermathecal gland inserted near the middle of the spermatheca, spermatheca more or less bent near the middle, but in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Anchista</emphasis>
</taxonomicName>
the spermathecal gland inserted near the apex of spermatheca, spermatheca nearly straight.
</paragraph>
<paragraph pageId="9" pageNumber="10">
Comparison between
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Paraphaea</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Shi &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Metallanchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Metallanchista</emphasis>
</taxonomicName>
gen. n. is presented in the diagnosis part under
<taxonomicName authorityName="Shi &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Metallanchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="10">Metallanchista</emphasis>
</taxonomicName>
gen. n.
</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="generic characters">
<paragraph pageId="9" pageNumber="10">Generic characters.</paragraph>
<paragraph lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Dorsal side generally reddish-brown to dark brown, sometimes with faint metallic reflections; elytra unicolored or bicolored.
<emphasis bold="true" pageId="9" pageNumber="10">Head</emphasis>
glabrous; eyes hemispherical and strongly prominent; tempora shorter than half of eyes length, abruptly narrowed behind eyes; vertex flat. Antennae reaching elytral base; 1st antennomere slightly narrowed at base, 3rd slightly longer than 4th. Labrum smooth, without secondary setae; mandibles moderately widened, outer margin nearly straight (
<figureCitation captionStart="Figures 138–157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="9" pageNumber="10">Fig. 149</figureCitation>
), glabrous on outer scrobe and dorsal ridge; terminal maxillary palpomeres fusiform in males and females; terminal labial palpomeres strongly securiform and truncate apically in the males, narrower in the females; ligula with apex slightly projected, with four long setae; paraglossae membranous, not longer than ligula, adnate; mentum tooth simple, with two setae near base; submentum with two long setae; genae glabrous beneath eyes.
<emphasis bold="true" pageId="9" pageNumber="10">Pronotum</emphasis>
slightly wider than head, disc glabrous or sparsely pubescent; mid-lateral setae present; front angles more or less setose, hind angles generally with a few additional short setae; pronotal base briefly but distinctly lobed; lateral margins completely rounded in middle (
<figureCitation captionStart="Figures 138–157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="9" pageNumber="10">Fig. 155</figureCitation>
), more or less sinuate before hind angles; hind angles sharp, rectangular or subrectangular.
<emphasis bold="true" pageId="9" pageNumber="10">Elytra</emphasis>
wide,
<pageBreakToken pageId="10" pageNumber="11" start="start">apex</pageBreakToken>
truncate, sutural angles not projected, outer angles completely rounded; sides slightly depressed in anterior third, disc with an indistinct depression near apical two-fifths; intervals glabrous or only odd intervals with a few additional setae; umbilical pores of 9th interval placed in one row (
<figureCitation captionStart="Figures 138–157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="10" pageNumber="11">Fig. 147</figureCitation>
); basal margination nearly complete; basal pores well developed; 3rd interval with two to four setigerous pores, 5th interval with base slightly widened, with one setigerous pore; 7th and 8th intervals slightly tumid near apex.
<emphasis bold="true" pageId="10" pageNumber="11">Ventral side</emphasis>
nearly glabrous; males with terminal sternum deeply emarginate (
<figureCitation captionStart="Figures 138–157" captionStartId="F44" captionText="Figures 138 - 157. Characters of Physoderina. 138 - 139 Internal reproductive system of females, scale bars = 0.2 mm: 138 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 139 Orionella lewisii (Bates), a specimen from Zhejiang. 140 - 142 Right protibia, ventral view, showing the reduction of cleaning spur, scale bars = 0.5 mm: 140 Allocota aurata (Bates) (cleaning spur absent) 141 Allocota bicolor sp. n. (cleaning spur fine) 142 Diamella cupreomicans (Oberthuer) (cleaning spur well developed). 143 - 146 Terminal sternum, showing the setae and male emargination, scale bars = 0.5 mm: 143 Paraphaea binotata (Dejean), male (deeply emarginate, one seta on each side) 144 Paraphaea binotata (Dejean) (straight, two setae on each side), female 145 Allocota aurata (Bates), male (moderately emarginate, two setae on each side) 146 Allocota aurata (Bates), female (straight, two setae on each side, right side unusually with an additional seta). 147 - 148 Umbilical series of 9 th interval, right elytron, scale bars = 0.5 mm: 147 Paraphaea binotata (Dejean) (umbilical series placed in one row) 148 Metallanchista laticollis sp. n. (umbilical series placed in two rows). 149 - 150 Head, showing the different shape of mandibles, scale bars = 0.5 mm: 149 Paraphaea formosana (Jedlicka) (mandibles moderately widen) 150 Allocota bicolor sp. n. (mandibles strongly widen). 151 - 153 Pronotum, showing the different length of setae on front angle, scale bars = 0.5 mm: 151 Allocota viridipennis Motschulsky, (setae short and fine) 152 Allocota cyanipennis Heller, (setae relative long) 153 Allocota bicolor sp. n., (setae very long). 154 - 155 Pronotum, showing the difference on lateral margin, scale bars = 0.5 mm: 154 Anchista fenestrata fenestrata (Schmidt-Goebel), (lateral margin slightly angulated in middle) 155 Paraphaea formosana (Jedlicka), (lateral margin completely rounded in middle). 156 - 157 Pronotal hind angle, showing the geographical variation in Allocota bicolor sp. n., scale bars = 0.2 mm: 156 Allocota bicolor sp. n., a specimen from Guangdong, (hind angle sharp) 157 Allocota bicolor sp. n., holotype from Yunnan, (hind angle slightly rounded)." httpUri="https://binary.pensoft.net/fig/29722" pageId="10" pageNumber="11">Fig. 143</figureCitation>
), with one pair of setae; females with apex of terminal sternum straight or slightly emarginate, with two pairs of setae.
<emphasis bold="true" pageId="10" pageNumber="11">Legs</emphasis>
short; protibiae with cleaning spur well developed, distant from inner margin; tarsi widened, 4th tarsomere bifid, claws pectinate; males with adhesive hairs well developed (two whole rows) on 1st to 3rd pro-, 1st to 2nd meso- and 1st metatarsomeres, rudimentary (two rows but very weakly present near apex) on 3rd mesotarsomere.
<emphasis bold="true" pageId="10" pageNumber="11">Male genitalia</emphasis>
with median lobe of aedeagus twisted to left; apical orifice opened dorsally, strongly setose along basal margin; internal sac with main flagellum more or less sinuous, nearly reaching apical orifice, trumpet-form expansion small and strongly bent; apical bursa strongly sclerotized; three additional small sclerotized pieces placed near apical third; secondary flagellum short and indistinct (
<figureCitation captionStart="Figures 95–110" captionStartId="F40" captionText="Figures 95 - 110. Left and right parameres of male genitalia, scale bars = 0.2 mm: 95 Paraphaea binotata (Dejean), a specimen from Guangxi 96 Paraphaea minor sp. n., holotype 97 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir 98 Anchista pilosa sp. n., holotype 99 Metallanchista laticollis sp. n., holotype 100 Physodera amplicollis van de Poll, a specimen from Taiwan 101 Diamella cupreomicans (Oberthuer), a specimen from Yunnan 102 Diamella arrowi (Jedlicka), holotype 103 Allocota aurata (Bates), a specimen from Tonkin 104 Allocota aurata (Bates), a specimen from Shaanxi 105 Allocota bicolor sp. n., holotype 106 Lachnoderma asperum Bates, a specimen from Taiwan 107 Dasiosoma ivorense Basilewsky, a paratype from Cote d'Ivoire 108 Dasiosoma quadraticolle sp. n., holotype 109 Orionella kathmanduensis (Kirschenhofer), a specimen from Nepal 110 Endynomena pradieri (Fairmaire), a specimen from Fujian." httpUri="https://binary.pensoft.net/fig/29716" pageId="10" pageNumber="11">Fig. 67</figureCitation>
).
<emphasis bold="true" pageId="10" pageNumber="11">Female genitalia.</emphasis>
Spermatheca tubular, with distinct ring-sculpture, inserted on bursa copulatrix; spermathecal gland slender and long, inserted near middle of spermatheca; spermatheca more or less bent near middle. Apical segment of ovipositor scimitar-shaped, curved to outer side, inner margin slightly angulate near apex; with fine setae near apex; apex with elongate membranous extension.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="distribution">
<paragraph pageId="10" pageNumber="11">Distribution</paragraph>
<paragraph pageId="10" pageNumber="11">
(
<figureCitation captionStart="Map 1" captionStartId="F45" captionText="Map 1. Known distribution of genus Paraphaea I: ● Paraphaea binotata (Dejean)." httpUri="https://binary.pensoft.net/fig/29723" pageId="10" pageNumber="11">Maps 1</figureCitation>
,
<figureCitation captionStart="Map 2" captionStartId="F46" captionText="Map 2. Known distribution of genus Paraphaea II: ▲ Paraphaea minor sp. n. ◆ Paraphaea philippinensis (Jedlicka) ★ Paraphaea formosana (Jedlicka)." httpUri="https://binary.pensoft.net/fig/29724" pageId="10" pageNumber="11">2</figureCitation>
). This genus includes four species distributed in South and Southeast Asia. One of them (
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Dejean" baseAuthorityYear="1825" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="binotata">
<emphasis italics="true" pageId="10" pageNumber="11">Paraphaea binotata</emphasis>
</taxonomicName>
) has a rather wide distribution, from Japan and India to the western Pacific islands (
<figureCitation captionStart="Map 1" captionStartId="F45" captionText="Map 1. Known distribution of genus Paraphaea I: ● Paraphaea binotata (Dejean)." httpUri="https://binary.pensoft.net/fig/29723" pageId="10" pageNumber="11">Map 1</figureCitation>
), but the other three species are more restricted.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="monophyly and relationships">
<paragraph pageId="10" pageNumber="11">Monophyly and relationships.</paragraph>
<paragraph pageId="10" pageNumber="11">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="11">Paraphaea</emphasis>
</taxonomicName>
is presumed to be the sister group of
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="11">Anchista</emphasis>
</taxonomicName>
. The relationship is supported by these character states: (1) mandibles moderately widened; (2) terminal sternum with single seta on each side in males; (3) median lobe of aedeagus strongly setose around apical orifice; (4) apical segment of ovipositor with inner margin slightly angulate near apex.
</paragraph>
<paragraph pageId="10" pageNumber="11">
Monophyly of
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="11">Paraphaea</emphasis>
</taxonomicName>
is suggested by the following apomorphic character states: (1) males with adhesive hairs present on hind tarsi; (2) males with terminal sternum deeply emarginate; (3) median lobe of aedeagus twisted; (4) spermatheca more or less bent near middle.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="taxonomic comments">
<paragraph pageId="10" pageNumber="11">Taxonomic comments.</paragraph>
<paragraph pageId="10" pageNumber="11">
<bibRefCitation author="Bates, HW" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="85" pageNumber="86" pagination="229 - 322" refId="B23" refString="Bates, HW, 1873. On the Geodephagous Coleoptera of Japan. Fam. Carabidae. The Transactions of the Entomological Society of London 1873 (2): 229 - 322" title="On the Geodephagous Coleoptera of Japan. Fam. Carabidae." volume="1873" year="1873">Bates (1873)</bibRefCitation>
proposed the genus
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="11">Paraphaea</emphasis>
</taxonomicName>
without comparing it with
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="11">Anchista</emphasis>
</taxonomicName>
Nietner. Later,
<bibRefCitation author="Chaudoir, BM" journalOrPublisher="Bulletin de la Societe Imperiale des Naturalistes de Moscou" pageId="86" pageNumber="87" pagination="188 - 268" refId="B35" refString="Chaudoir, BM, 1877. Genres nouveaux et especes inedites de la famille des Carabiques. Bulletin de la Societe Imperiale des Naturalistes de Moscou 52 (1): 188 - 268" title="Genres nouveaux et especes inedites de la famille des Carabiques." volume="52" year="1877">Chaudoir (1877)</bibRefCitation>
synonymized these two genera without detailed explanation. Based on external characters, it is difficult to find significant differences between these two genera, but the genital and male secondary sexual characters mentioned above provide clear differences and justify the generic separation of
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="11">Paraphaea</emphasis>
</taxonomicName>
Bates.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="key to species of paraphaea bates">
<paragraph pageId="11" pageNumber="12">
<pageBreakToken pageId="11" pageNumber="12" start="start">Key</pageBreakToken>
to species of
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="11" pageNumber="12">Paraphaea</emphasis>
</taxonomicName>
Bates
</paragraph>
<paragraph pageId="11" pageNumber="12">
<table inLine="true" pageId="11" pageNumber="12">
<tr pageId="11" pageNumber="12">
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">1</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">
Pronotum more or less pubescent; pronotum reddish brown, elytra metallic blue; the Philippines (
<figureCitation captionStart="Figures 31–36" captionStartId="F6" captionText="Figures 31 - 36. Type materials, scale bars = 2.0 mm: 31 Lectotype of Plochionus binotatus Dejean [= Paraphaea binotata (Dejean)] 32 Lectotype of Calleida discophora Chaudoir [= Paraphaea binotata (Dejean)] 33 Lectotype of Paraphaea signifera Bates [= Paraphaea binotata (Dejean)] 34 Holotype of Parena formosana Jedlicka [= Paraphaea formosana (Jedlicka)] 35 Holotype of Paraphaea minor sp. n. 36 Holotype of Coptodera philippinensis Jedlicka [= Paraphaea philippinensis (Jedlicka)]." httpUri="https://binary.pensoft.net/fig/29679" pageId="11" pageNumber="12">Fig. 36</figureCitation>
)
</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Jedlicka" baseAuthorityYear="1935" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis italics="true" pageId="11" pageNumber="12">Paraphaea philippinensis</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
)
</td>
</tr>
<tr pageId="11" pageNumber="12">
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">-</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">Pronotum glabrous; elytra uniform brown or with bicolored pattern, not metallic</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">2</td>
</tr>
<tr pageId="11" pageNumber="12">
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">2</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">
Third interval with more than three setigerous pores, 3rd 5th and 7th intervals with some secondary setigerous pores in some specimens; Taiwan (
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus, scale bars = 2.0 mm: 1 Paraphaea binotata (Dejean), a specimen from Yunnan 2 Paraphaea formosana (Jedlicka), a specimen from Taiwan 3 Paraphaea minor sp. n., a paratype from Hainan 4 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir from India 5 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 6 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal." httpUri="https://binary.pensoft.net/fig/29670" pageId="11" pageNumber="12">Figs 2</figureCitation>
<figureCitation captionStart="Figures 31–36" captionStartId="F6" captionText="Figures 31 - 36. Type materials, scale bars = 2.0 mm: 31 Lectotype of Plochionus binotatus Dejean [= Paraphaea binotata (Dejean)] 32 Lectotype of Calleida discophora Chaudoir [= Paraphaea binotata (Dejean)] 33 Lectotype of Paraphaea signifera Bates [= Paraphaea binotata (Dejean)] 34 Holotype of Parena formosana Jedlicka [= Paraphaea formosana (Jedlicka)] 35 Holotype of Paraphaea minor sp. n. 36 Holotype of Coptodera philippinensis Jedlicka [= Paraphaea philippinensis (Jedlicka)]." httpUri="https://binary.pensoft.net/fig/29679" pageId="11" pageNumber="12">, 34</figureCitation>
)
</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Jedlicka" baseAuthorityYear="1946" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="formosana">
<emphasis italics="true" pageId="11" pageNumber="12">Paraphaea formosana</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
)
</td>
</tr>
<tr pageId="11" pageNumber="12">
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">-</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">Third interval with two setigerous pores, intervals without secondary setigerous pores</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">3</td>
</tr>
<tr pageId="11" pageNumber="12">
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">3</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">
Pronotum widest at apical third, lateral margins slightly sinuate before hind angles; elytra with background dark, each side with an elongate pale patch before middle (
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus, scale bars = 2.0 mm: 1 Paraphaea binotata (Dejean), a specimen from Yunnan 2 Paraphaea formosana (Jedlicka), a specimen from Taiwan 3 Paraphaea minor sp. n., a paratype from Hainan 4 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir from India 5 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 6 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal." httpUri="https://binary.pensoft.net/fig/29670" pageId="11" pageNumber="12">Fig. 1</figureCitation>
); widely distributed in southeast Asia
</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Dejean" baseAuthorityYear="1825" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="binotata">
<emphasis italics="true" pageId="11" pageNumber="12">Paraphaea binotata</emphasis>
</taxonomicName>
(Dejean)
</td>
</tr>
<tr pageId="11" pageNumber="12">
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">-</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">
Pronotum widest near middle, lateral margins distinctly sinuate before hind angles; elytra with disc yellowish, lateral margins and apex dark (
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus, scale bars = 2.0 mm: 1 Paraphaea binotata (Dejean), a specimen from Yunnan 2 Paraphaea formosana (Jedlicka), a specimen from Taiwan 3 Paraphaea minor sp. n., a paratype from Hainan 4 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir from India 5 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 6 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal." httpUri="https://binary.pensoft.net/fig/29670" pageId="11" pageNumber="12">Fig. 3</figureCitation>
); Hainan and Indo-China
</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">
<taxonomicName authorityName="Shi &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="minor">
<emphasis italics="true" pageId="11" pageNumber="12">Paraphaea minor</emphasis>
</taxonomicName>
sp. n.
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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