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468 lines
61 KiB
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<document id="B32CA245FB15E3E2A74EDA24811E0C1A" ID-CLB-Dataset="55761" ID-DOI="10.18590/euscorpius.2019.vol2019.iss277.1" ID-GBIF-Dataset="e9986ae3-7799-4a66-afc1-b1e1ecd827d8" ID-ISSN="1536-9307" ID-Zenodo-Dep="6511455" ID-ZooBank="64451868-1FB3-4D20-B88D-FBF7304EB775" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638261050631" checkinUser="felipe" docAuthor="Lowe, Graeme, Kovařík, František, Stockmann, Mark & Šťáhlavský, František" docDate="2019" docId="D81E87F7513AFF8E5E2CDBD7FAC9FB01" docLanguage="en" docName="Euscorpius.277.1-30.pdf" docOrigin="Euscorpius 277" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Trypanothacus buettikeri Lowe & Kovařík & Stockmann & Šťáhlavský 2019, comb. n." docType="treatment" docVersion="6" lastPageNumber="22" masterDocId="2427FF8F512DFF995E5DDA5DFF8BFFC1" masterDocTitle="Trypanothacus gen. n., a new genus of burrowing scorpion from the Arabian Peninsula (Scorpiones: Buthidae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="22" updateTime="1699471307476" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="6448E5EF6814522AF54D5AB3A72EDF0A">Trypanothacus gen. n., a new genus of burrowing scorpion from the Arabian Peninsula (Scorpiones: Buthidae)</mods:title>
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<mods:namePart id="DEF7770111E7AAEBAECF4F345EB6F1FA">Lowe, Graeme</mods:namePart>
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<mods:namePart id="5D7B34A077266A56B130D929E2ED8BC6">Kovařík, František</mods:namePart>
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<mods:affiliation id="EFA065D2EEC8ED61162B2E4438A29399">P. O. Box 27, CZ - 145 01 Praha 45, Czech Republic</mods:affiliation>
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<mods:nameIdentifier id="57F2C495FBD1749DE8EFD4E9C0CF0B4A" type="URL">http://www.scorpio.cz</mods:nameIdentifier>
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<mods:namePart id="4B63F9EB0483F8C73F4202040A6DBED4">Stockmann, Mark</mods:namePart>
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<mods:affiliation id="19D9A397E1168C0C8487AF3D1D5EA845">Im Hoek 20, D- 48477 Hörstel-Riesenbeck, Germany</mods:affiliation>
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<mods:namePart id="D60FF3F8B8458872B98AF47A714D07C8">Šťáhlavský, František</mods:namePart>
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<mods:affiliation id="FC8B86A52D668BD4D703354A1D6FD7C9">Department of Zoology, Charles University, Viničná 7, CZ- 128 44 Praha 2, Czech Republic</mods:affiliation>
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<treatment id="D81E87F7513AFF8E5E2CDBD7FAC9FB01" ID-DOI="http://doi.org/10.5281/zenodo.6511471" ID-GBIF-Taxon="194951821" ID-Zenodo-Dep="6511471" LSID="urn:lsid:plazi:treatment:D81E87F7513AFF8E5E2CDBD7FAC9FB01" httpUri="http://treatment.plazi.org/id/D81E87F7513AFF8E5E2CDBD7FAC9FB01" lastPageNumber="22" pageId="23" pageNumber="22">
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<taxonomicName id="97B74D62513AFF8E5E2CDBD7FD05FE64" ID-CoL="59FJV" authority="(Hendrixson, 2006)" authorityName="Lowe & Kovařík & Stockmann & Šťáhlavský" authorityYear="2019" baseAuthorityName="Hendrixson" baseAuthorityYear="2006" box="[113,654,394,421]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="buettikeri" status="comb. nov.">
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<emphasis id="62C3EAF3513AFF8E5E2CDBD7FE2AFE64" bold="true" box="[113,417,394,421]" italics="true" pageId="23" pageNumber="22">Trypanothacus buettikeri</emphasis>
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(
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<bibRefCitation id="34264B10513AFF8E5FF2DBD7FD0DFE64" author="Hendrixson" box="[431,646,394,421]" firstAuthor="Hendrixson" pageId="23" pageNumber="22" pagination="33 - 120" refId="ref14260" refString="HENDRIXSON, B. E. 2006. Buthid scorpions of Saudi Arabia, with notes on other families (Scorpiones: Buthidae, Liochelidae, Scorpionidae). Fauna of Arabia, 21: 33 - 120." type="journal article" year="2006">Hendrixson, 2006</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicNameLabel id="79F05788513AFF8E5CC4DBD7FC88FE64" box="[665,771,394,421]" pageId="23" pageNumber="22" rank="species">
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<emphasis id="62C3EAF3513AFF8E5CC4DBD7FD56FE65" bold="true" box="[665,733,394,420]" pageId="23" pageNumber="22">comb</emphasis>
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.
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<emphasis id="62C3EAF3513AFF8E5CB6DBD7FD70FE65" bold="true" box="[747,763,394,420]" pageId="23" pageNumber="22">n</emphasis>
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.
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<subSubSection id="18AD656A513AFF8E5F32DBF1FD86FE05" box="[367,525,428,452]" pageId="23" pageNumber="22" type="description">
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<paragraph id="500836E1513AFF8E5F32DBF1FD86FE05" blockId="23.[113,771,394,452]" box="[367,525,428,452]" pageId="23" pageNumber="22">
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(
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<figureCitation id="C88C2A64513AFF8E5F2ADBF1FD8EFE05" box="[375,517,428,452]" captionStart-0="Figures 90–93" captionStart-1="Figures 94–95" captionStart-2="Figures 96–106" captionStart-3="Figures 107–110" captionStartId-0="21.[113,190,1977,1999]" captionStartId-1="22.[113,190,1259,1281]" captionStartId-2="24.[113,187,1903,1925]" captionStartId-3="25.[113,189,1408,1430]" captionTargetBox-0="[131,1442,254,1923]" captionTargetBox-1="[113,1474,234,1247]" captionTargetBox-2="[113,1474,234,1891]" captionTargetBox-3="[121,1439,242,1379]" captionTargetId-0="figure-7@21.[110,1474,234,1945]" captionTargetId-1="figure-416@22.[113,1474,234,1248]" captionTargetId-2="figure-11@24.[113,1474,231,1891]" captionTargetId-3="figure-494@25.[113,1474,234,1386]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="24" captionTargetPageId-3="25" captionText-0="Figures 90–93: Trypanothacus buettikeri comb. n. Figures 90–91. Male holotype in dorsal (90) and ventral (91) views. Figures 92–93: Female paratype in dorsal (92) and ventral (93) views. Scale bars: 10 mm." captionText-1="Figures 94–95: Trypanothacus buettikeri comb. n. Male holotype, carapace and tergites I–IV (94), coxosternal area and sternites I–II (95). UV fluorescence. Scale bar: 2 mm." captionText-2="Figures 96–106: Trypanothacus buettikeri comb. n. Figures 96–101. Male holotype, lateral view of metasoma and telson (96), dorsal views of pedipalp chela (97) patella (98) and femur (99), external views of pedipalp chela (100) and patella (101). Figure 102. Male from Wadi Turabah (NHMB No. 509), dorsal view of pedipalp patella. Figures 103–106. Female paratype (NHMB 896), lateral views of metasoma (103) and pedipalp chela (104), dorsal view ofpedipalp patella (105) and femur (106).UV fluorescence.Scale bars:2 mm (96), 2 mm (97–101), 2 mm (102), 2 mm (103), 2 mm (104–106)." captionText-3="Figures 107–110: Morphometric scaling relations of the telson of fossorial ‘Buthus group’ scorpions. Figure 107. Double logarithmic scatterplot of telson vesicle length/ depth (L/ D) ratio vs. aculeus length normalized to vesicle length (aculeus L/ vesicle L), for large pelophiles/ semipsammophiles (4 genera, 53 species) (Buthus, Odontobuthus, Pantobuthus: orange circles; Trypanothacus gen. n.: blue circles) and large psammophiles/ ultrapsammophiles (6 genera, 29 species) (Buthacus: cyan circles; Vachoniolus: dark green circles; Apistobuthus, Buthiscus, Liobuthus, and Plesiobuthus: yellow circles). Dark line: linear least squares regression fit to all data shown (Pearson correlation coefficient, R = 0.57). Upper right inset: diagram of measurements estimated from inverted telson lateral profiles, with dorsal surface of vesicle approximately leveled; SA = ventral profile length of aculeus from base point to tip, LV = length of vesicle from anterior limit or inflection point to aculeus base point; HV = vesicle depth or height (distance between dorsal and ventral tangent planes). Base point was placed at local minimum of inverted ventral profile curve. SA was calculated by cubic spline fit (>10 anchor points). For a minority of broken tips, SA was estimated by intersecting extrapolated dorsal and ventral profile curves (<5%). Figure 108. Scatterplot of fossorial buthids in Figure 107, superposed upon larger scale plot of Order Scorpiones (gray circles). Gray line: linear least squares regression fit to Order Scorpiones data (including data in Figure 107) (Pearson correlation coefficient, R = –0.62).Also plotted are data from small ‘Buthus group’ psammophiles/ ultrapsammophiles (5 genera, 10 species: Compsobuthus arabicus, Lanzatus somalilandus, Neobuthus kutcheri, Picobuthus dundoni, P. wahibaensis, Anomalobuthus krivochatskyi, A. pavlovskyi, A. rickmersi, A. lowei, A. talebii) (dark yellow circles). Figure 109. Double logarithmic scatterplot of carapace length (L) vs. aculeus length normalized to vesicle length (i.e., aculeus L/ vesicle L) for psammophile/ ultrapsammophile members of the ‘Buthus group’, including large species in Figure 107, and small species in Figure 108 (dark yellow circles). Blue line: linear least squares regression fit to all data shown (Pearson correlation coefficient, R = 0.74). Figure 110. Scatterplot of psammophile buthid data in Figure 109 (all in one color, blue circles) superposed upon larger scale plot of Order Scorpiones (gray circles). Gray line: linear least squares regression to Order Scorpiones data (including data in Figure 109) (Pearson correlation coefficient, R = –0.01). Also plotted are data from bothriurid genus Brachistosternus (43 species) (magenta triangles), and corresponding linear least squares regression fit (R = 0.39). All plots in Figures 107-110 include data from both males and females. Data were assembled by mining taxonomic literature or directly measuring specimens. Included are 1854 species or subspecies (1331 ♂, 1201 ♀) representing all known extant genera (1841 species or subspecies) plus 7 fossil genera (13 species)." figureDoi-0="http://doi.org/10.5281/zenodo.6511505" figureDoi-1="http://doi.org/10.5281/zenodo.6511509" figureDoi-2="http://doi.org/10.5281/zenodo.6511511" figureDoi-3="http://doi.org/10.5281/zenodo.6511515" httpUri-0="https://zenodo.org/record/6511505/files/figure.png" httpUri-1="https://zenodo.org/record/6511509/files/figure.png" httpUri-2="https://zenodo.org/record/6511511/files/figure.png" httpUri-3="https://zenodo.org/record/6511515/files/figure.png" pageId="23" pageNumber="22">Figs. 90–108</figureCitation>
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)
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<taxonomicName id="97B74D62513AFF8E5E2CDBB1FDDEFDC5" authority="Hendrixson, 2006: 47 - 52" authorityName="Hendrixson" authorityPageNumber="47 - 52" authorityYear="2006" box="[113,597,492,516]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="buettikeri">
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<emphasis id="62C3EAF3513AFF8E5E2CDBB1FECBFDC5" box="[113,320,492,516]" italics="true" pageId="23" pageNumber="22">Buthacus buettikeri</emphasis>
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<bibRefCitation id="34264B10513AFF8E5F18DBB1FDDEFDC5" author="Hendrixson" box="[325,597,492,516]" firstAuthor="Hendrixson" page="47 - 52" pageId="23" pageNumber="22" pagination="33 - 120" refId="ref14260" refString="HENDRIXSON, B. E. 2006. Buthid scorpions of Saudi Arabia, with notes on other families (Scorpiones: Buthidae, Liochelidae, Scorpionidae). Fauna of Arabia, 21: 33 - 120." type="journal article" year="2006">Hendrixson, 2006: 47–52</bibRefCitation>
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, figs. 4–5, plates 3–4 (in part);
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;
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<treatmentCitation id="D11610F0513AFF8E5C2AD851FECFFD85" author="EL-HENNAWY, H. K." page="121, 126" pageId="23" pageNumber="22" year="2009">
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<bibRefCitation id="34264B10513AFF8E5C2AD851FE85FD85" author="EL-HENNAWY, H. K." journalOrPublisher="Serket" pageId="23" pageNumber="22" pagination="119 - 128" part="11" refId="ref14076" refString="EL-HENNAWY, H. K. 2009. Scorpions of Saudi Arabia. Serket, 11 (3 / 4): 119 - 128." title="Scorpions of Saudi Arabia" type="journal article" year="2009">El-Hennawy, 2009: 121</bibRefCitation>
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, 126
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;
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<treatmentCitation id="D11610F0513AFF8E5F12D871FD5AFD85" author="LOURENCO, W. R. & E. - A. LEGUIN" box="[335,721,556,580]" page="104, 107" pageId="23" pageNumber="22" year="2009">
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<bibRefCitation id="34264B10513AFF8E5F12D871FD17FD85" author="LOURENCO, W. R. & E. - A. LEGUIN" box="[335,668,556,580]" journalOrPublisher="Zoology in the Middle East" pageId="23" pageNumber="22" pagination="103 - 108" part="46" refId="ref15044" refString="LOURENCO, W. R. & E. - A. LEGUIN. 2009. A new species of the genus Buthacus Birula, 1908 from the United Arab Emirates (Scorpiones, Buthidae). Zoology in the Middle East, 46: 103 - 108." title="A new species of the genus Buthacus Birula, 1908 from the United Arab Emirates (Scorpiones, Buthidae)" type="journal article" year="2009">Lourenço & Leguin, 2009: 104</bibRefCitation>
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, 107
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, tab. 1;
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<treatmentCitation id="D11610F0513AFF8E5EE3D811FDFEFDA5" author="DESOUKY, M. M. A. & A. M. ALSHAMMARI" box="[190,629,588,612]" page="195-196" pageId="23" pageNumber="22" year="2011">
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<bibRefCitation id="34264B10513AFF8E5EE3D811FDFEFDA5" author="DESOUKY, M. M. A. & A. M. ALSHAMMARI" box="[190,629,588,612]" journalOrPublisher="Bulletin of the British Arachnological Society" pageId="23" pageNumber="22" pagination="193 - 200" part="15" refId="ref14018" refString="DESOUKY, M. M. A. & A. M. ALSHAMMARI. 2011. Scorpions of the Ha'il Region, northern Saudi Arabia, and molecular phylogenetics of two common species, Androctonus crassicauda and Scorpio maurus kruglovi. Bulletin of the British Arachnological Society, 15 (6): 193 - 200." title="Scorpions of the Ha'il Region, northern Saudi Arabia, and molecular phylogenetics of two common species, Androctonus crassicauda and Scorpio maurus kruglovi" type="journal article" year="2011">Desouky & Alshammari, 2011: 195–196</bibRefCitation>
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</treatmentCitation>
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, figs. 6–7 (?) (in part);
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<treatmentCitation id="D11610F0513AFF8E5F55D831FE77FD45" author="KOVARIK, F. & G. LOWE & J. PLISKOVA & F. STAHLAVSKY" box="[264,508,620,644]" page="3" pageId="23" pageNumber="22" year="2011">
|
||
<bibRefCitation id="34264B10513AFF8E5F55D831FE77FD45" author="KOVARIK, F. & G. LOWE & J. PLISKOVA & F. STAHLAVSKY" box="[264,508,620,644]" journalOrPublisher="Euscorpius" pageId="23" pageNumber="22" pagination="1 - 19" part="173" refId="ref14538" refString="KOVARIK, F., G. LOWE, J. PLISKOVA & F. STAHLAVSKY. 2013. A new scorpion genus, Gint gen. n., from the Horn of Africa (Scorpiones, Buthidae). Euscorpius, 173: 1 - 19." title="A new scorpion genus, Gint gen. n., from the Horn of Africa (Scorpiones, Buthidae)" type="journal article" year="2013">Kovařík et al., 2013: 3</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="D11610F0513AFF8E5C57D831FD74FD45" author="EL-HENNAWY, H. K." box="[522,767,620,644]" page="45" pageId="23" pageNumber="22" year="2014">
|
||
<bibRefCitation id="34264B10513AFF8E5C57D831FD74FD45" author="EL-HENNAWY, H. K." box="[522,767,620,644]" journalOrPublisher="Serket" pageId="23" pageNumber="22" pagination="22 - 58" part="14" refId="ref14102" refString="EL-HENNAWY, H. K. 2014. Preliminary list of spiders and other arachnids of Saudi Arabia (Except ticks and mites). Serket, 14 (1): 22 - 58." title="Preliminary list of spiders and other arachnids of Saudi Arabia (Except ticks and mites)" type="journal article" year="2014">El-Hennawy, 2014: 45</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="D11610F0513AFF8E5EFCD8D1FE15FD65" author="KOVARIK" box="[161,414,652,676]" page="2" pageId="23" pageNumber="22" year="2016">
|
||
<bibRefCitation id="34264B10513AFF8E5EFCD8D1FE15FD65" author="Kovarik" box="[161,414,652,676]" etAl="et al." firstAuthor="Kovarik" page="2" pageId="23" pageNumber="22" pagination="1 - 18" refId="ref14669" refString="KOVARIK, F., G. LOWE & F. STAHLAVSKY. 2016 b. Review of Northwestern African Buthacus, with description of Buthacus stockmanni sp. n. from Morocco and Western Sahara (Scorpiones, Buthidae). Euscorpius, 236: 1 - 18." type="journal article" year="2016" yearSuffix="b">Kovařík et al., 2016b: 2</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="D11610F0513AFF8E5FF0D8D1FDE2FD62" author="KOVARIK, F. " box="[429,617,652,676]" page="7" pageId="23" pageNumber="22" year="2018">
|
||
<bibRefCitation id="34264B10513AFF8E5FF0D8D1FDEEFD62" author="KOVARIK, F." box="[429,613,652,676]" journalOrPublisher="Euscorpius" pageId="23" pageNumber="22" pagination="1 - 12" part="269" refId="ref14419" refString="KOVARIK, F. 2018. Notes on the genera Buthacus, Compsobuthus, and Lanzatus with several synonymies and corrections of published characters (Scorpiones: Buthidae). Euscorpius, 269: 1 - 12." title="Notes on the genera Buthacus, Compsobuthus, and Lanzatus with several synonymies and corrections of published characters (Scorpiones: Buthidae)" type="journal article" year="2018">Kovařík, 2018: 7</bibRefCitation>
|
||
.
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="18AD656A513AFF8E5E2CD891FE99FBA5" pageId="23" pageNumber="22" type="materials_examined">
|
||
<paragraph id="500836E1513AFF8E5E2CD891FE90FCC2" blockId="23.[113,771,716,772]" pageId="23" pageNumber="22">
|
||
<typeStatus id="8F0C8843513AFF8E5E2CD891FF2CFD22" box="[113,167,716,739]" pageId="23" pageNumber="22">TYPE</typeStatus>
|
||
<collectionCode id="36A6AE24513AFF8E5EF3D891FEC3FD25" box="[174,328,716,740]" pageId="23" pageNumber="22">LOCALITY AND</collectionCode>
|
||
<typeStatus id="8F0C8843513AFF8E5F13D891FE0BFD22" box="[334,384,716,739]" pageId="23" pageNumber="22">TYPE</typeStatus>
|
||
<collectionCode id="36A6AE24513AFF8E5FD5D891FD8EFD25" box="[392,517,716,740]" pageId="23" pageNumber="22">DEPOSITORY</collectionCode>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5C53D891FD2EFD25" bold="true" box="[526,677,716,740]" pageId="23" pageNumber="22">
|
||
<collectingCountry id="28A07671513AFF8E5C53D891FD2EFD25" box="[526,677,716,740]" name="Saudi Arabia" pageId="23" pageNumber="22">Saudi Arabia</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<location id="5568603A513AFF8E5CEFD891FD51FD22" LSID="urn:lsid:plazi:treatment:D81E87F7513AFF8E5E2CDBD7FAC9FB01:5568603A513AFF8E5CEFD891FD51FD22" box="[690,730,716,739]" country="Saudi Arabia" municipality="Dabah" name="Um" pageId="23" pageNumber="22">Um</location>
|
||
ad-
|
||
<collectingMunicipality id="B06CAC9B513AFF8E5E2CD8B1FF32FCC5" box="[113,185,748,772]" pageId="23" pageNumber="22">Dabah</collectingMunicipality>
|
||
,
|
||
<collectionCode id="36A6AE24513AFF8E5E99D8B1FE9FFCC2" box="[196,276,748,771]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="23" pageNumber="22">NHMB</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="500836E1513AFF8E5E2CD971FE9AFBC5" blockId="23.[113,772,812,1124]" pageId="23" pageNumber="22">
|
||
<collectionCode id="36A6AE24513AFF8E5E2CD971FE03FC85" box="[113,392,812,836]" pageId="23" pageNumber="22">TYPE MATERIAL EXAMINED</collectionCode>
|
||
.
|
||
<materialsCitation id="E0DF3CBC513AFF8E5FCCD971FE4AFC45" ID-GBIF-Occurrence="3758760302" collectingDate="1980-05-13" collectionCode="NHMB" collectorName="W. Buttiker" country="Saudi Arabia" elevation="920" latitude="23.9" location="Um ad-Dabah" longLatPrecision="1254" longitude="45.016666" municipality="Um ad-Dabah" pageId="23" pageNumber="22" specimenCode="NHMB 0587" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="62C3EAF3513AFF8E5FCCD971FDA3FC85" bold="true" box="[401,552,812,836]" pageId="23" pageNumber="22">
|
||
<collectingCountry id="28A07671513AFF8E5FCCD971FDA3FC85" box="[401,552,812,836]" name="Saudi Arabia" pageId="23" pageNumber="22">Saudi Arabia</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="46B1FD68513AFF8E5C6BD971FDD3FC85" box="[566,600,812,836]" pageId="23" pageNumber="22" type="male">1♂</specimenCount>
|
||
(
|
||
<typeStatus id="8F0C8843513AFF8E5C3BD971FD42FC85" box="[614,713,812,836]" pageId="23" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
),
|
||
<collectingMunicipality id="B06CAC9B513AFF8E5C86D971FF52FCA5" pageId="23" pageNumber="22">Um ad-Dabah</collectingMunicipality>
|
||
[
|
||
<geoCoordinate id="35835026513AFF8E5EBBD911FECAFCA5" box="[230,321,844,868]" degrees="23" direction="north" minutes="54" orientation="latitude" pageId="23" pageNumber="22" precision="925" value="23.9">23°54'N</geoCoordinate>
|
||
<geoCoordinate id="35835026513AFF8E5F15D911FE2BFCA5" box="[328,416,844,868]" degrees="45" direction="east" minutes="01" orientation="longitude" pageId="23" pageNumber="22" precision="925" value="45.016666">45°01'E</geoCoordinate>
|
||
,
|
||
<elevation id="DB9AD1D2513AFF8E5FF6D911FDA3FCA5" box="[427,552,844,868]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="23" pageNumber="22" unit="m" value="920.0">
|
||
<quantity id="974F9B04513AFF8E5FF6D911FE64FCA2" box="[427,495,844,868]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="23" pageNumber="22" unit="m" value="920.0">920 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
],
|
||
<collectingDate id="344DE9C9513AFF8E5C60D911FD26FCA5" box="[573,685,844,868]" pageId="23" pageNumber="22">
|
||
<date id="24091021513AFF8E5C60D911FDD0FCA5" box="[573,603,844,868]" pageId="23" pageNumber="22" value="1980-05-13">13.</date>
|
||
V.1980
|
||
</collectingDate>
|
||
, leg.
|
||
<collectorName id="FD425337513AFF8E5CBBD911FF41FC45" pageId="23" pageNumber="22">W. Büttiker</collectorName>
|
||
,
|
||
<specimenCode id="00119E9A513AFF8E5E89D931FED4FC45" box="[212,351,876,900]" collectionCode="NHMB" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="23" pageNumber="22">NHMB 0587</specimenCode>
|
||
(=172a);
|
||
</materialsCitation>
|
||
<materialsCitation id="E0DF3CBC513AFF8E5F9AD931FEE5FC05" ID-GBIF-Occurrence="3758760311" collectingDate="1981-10-10" collectionCode="NHMB" collectorName="W. Buttiker" country="Saudi Arabia" elevation="750" latitude="23.833334" location="Khashm Dhibi" longLatPrecision="1254" longitude="45.083332" municipality="Khashm Dhibi" pageId="23" pageNumber="22" specimenCode="NHMB 0896" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="46B1FD68513AFF8E5F9AD931FE63FC45" box="[455,488,876,900]" pageId="23" pageNumber="22" type="female">1♀</specimenCount>
|
||
(
|
||
<typeStatus id="8F0C8843513AFF8E5FA8D931FDDDFC45" box="[501,598,876,900]" pageId="23" pageNumber="22" type="paratype">paratype</typeStatus>
|
||
),
|
||
<collectingMunicipality id="B06CAC9B513AFF8E5C35D931FC88FC45" box="[616,771,876,900]" country="Saudi Arabia" latitude="23.833334" longLatPrecision="1254" longitude="45.083332" name="Khashm Dhibi" pageId="23" pageNumber="22">Khashm Dhibi</collectingMunicipality>
|
||
[
|
||
<geoCoordinate id="35835026513AFF8E5E25D9D1FF58FC65" box="[120,211,908,932]" degrees="23" direction="north" minutes="50" orientation="latitude" pageId="23" pageNumber="22" precision="925" value="23.833334">23°50'N</geoCoordinate>
|
||
<geoCoordinate id="35835026513AFF8E5E87D9D1FEB9FC65" box="[218,306,908,932]" degrees="45" direction="east" minutes="05" orientation="longitude" pageId="23" pageNumber="22" precision="925" value="45.083332">45°05'E</geoCoordinate>
|
||
,
|
||
<elevation id="DB9AD1D2513AFF8E5F63D9D1FE36FC65" box="[318,445,908,932]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="23" pageNumber="22" unit="m" value="750.0">
|
||
<quantity id="974F9B04513AFF8E5F63D9D1FE08FC62" box="[318,387,908,932]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="23" pageNumber="22" unit="m" value="750.0">750 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
],
|
||
<date id="24091021513AFF8E5F8ED9D1FDCDFC65" box="[467,582,908,932]" pageId="23" pageNumber="22" value="1981-10-10">
|
||
<collectingDate id="344DE9C9513AFF8E5F8ED9D1FDCDFC65" box="[467,582,908,932]" pageId="23" pageNumber="22" value="1981-10-10">10.X.1981</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="FD425337513AFF8E5CDCD9D1FD74FC65" box="[641,767,908,932]" pageId="23" pageNumber="22">W. Büttiker</collectorName>
|
||
,
|
||
<specimenCode id="00119E9A513AFF8E5E2CD9F1FE8FFC05" box="[113,260,940,964]" collectionCode="NHMB" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="23" pageNumber="22">NHMB 0896</specimenCode>
|
||
(=172e);
|
||
</materialsCitation>
|
||
<materialsCitation id="E0DF3CBC513AFF8E5F21D9F1FE9AFBC5" ID-GBIF-Occurrence="3758760306" collectingDate="1981-01-30" collectionCode="NHMB" collectorName="W. Buttiker" country="Saudi Arabia" elevation="850" latitude="24.233334" location="Kushm Dibi" longLatPrecision="1252" longitude="46.083332" municipality="Kushm Dibi" pageId="23" pageNumber="22" specimenCode="NHMB 172cd" specimenCount="4" specimenCount-female="3" typeStatus="paratype">
|
||
<specimenCount id="46B1FD68513AFF8E5F21D9F1FE15FC05" box="[380,414,940,964]" pageId="23" pageNumber="22" type="female">3♀</specimenCount>
|
||
<specimenCount id="46B1FD68513AFF8E5FC3D9F1FE6BFC05" box="[414,480,940,964]" pageId="23" pageNumber="22" type="generic">1 juv.</specimenCount>
|
||
(
|
||
<typeStatus id="8F0C8843513AFF8E5FA9D9F1FDEBFC05" box="[500,608,940,964]" pageId="23" pageNumber="22" type="paratype">paratypes</typeStatus>
|
||
),
|
||
<collectingMunicipality id="B06CAC9B513AFF8E5C24D9F1FC88FC05" box="[633,771,940,964]" country="Saudi Arabia" latitude="24.233334" longLatPrecision="1252" longitude="46.083332" name="Kushm Dibi" pageId="23" pageNumber="22">Kushm Dibi</collectingMunicipality>
|
||
[
|
||
<geoCoordinate id="35835026513AFF8E5E25D991FF58FC25" box="[120,211,972,996]" degrees="24" direction="north" minutes="14" orientation="latitude" pageId="23" pageNumber="22" precision="925" value="24.233334">24°14'N</geoCoordinate>
|
||
<geoCoordinate id="35835026513AFF8E5E81D991FEB8FC25" box="[220,307,972,996]" degrees="46" direction="east" minutes="05" orientation="longitude" pageId="23" pageNumber="22" precision="925" value="46.083332">46°05'E</geoCoordinate>
|
||
,
|
||
<elevation id="DB9AD1D2513AFF8E5F1CD991FE49FC25" box="[321,450,972,996]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="23" pageNumber="22" unit="m" value="850.0">
|
||
<quantity id="974F9B04513AFF8E5F1CD991FE0CFC22" box="[321,391,972,996]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="23" pageNumber="22" unit="m" value="850.0">850 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
],
|
||
<date id="24091021513AFF8E5F87D991FDC8FC25" box="[474,579,972,996]" pageId="23" pageNumber="22" value="1981-01-30">
|
||
<collectingDate id="344DE9C9513AFF8E5F87D991FDC8FC25" box="[474,579,972,996]" pageId="23" pageNumber="22" value="1981-01-30">30.I.1981</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="FD425337513AFF8E5CDDD991FD74FC25" box="[640,767,972,996]" pageId="23" pageNumber="22">W. Büttiker</collectorName>
|
||
,
|
||
<specimenCode id="00119E9A513AFF8E5E2CD9B1FE87FBC5" box="[113,268,1004,1028]" pageId="23" pageNumber="22">NHMB 172cd</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="500836E1513AFF8E5E2CDE51FE99FBA5" blockId="23.[113,772,812,1124]" pageId="23" pageNumber="22">
|
||
<collectionCode id="36A6AE24513AFF8E5E2CDE51FE14FBE5" box="[113,415,1036,1060]" pageId="23" pageNumber="22">OTHER MATERIAL EXAMINED</collectionCode>
|
||
.
|
||
<materialsCitation id="E0DF3CBC513AFF8E5FFCDE51FE99FBA5" ID-GBIF-Occurrence="3758760313" collectingDate="1986-12" collectionCode="NHMB" collectorName="W. Buttiker" elevation="1430" latitude="25.5" location="Wadi Turabah" longLatPrecision="1246" longitude="41.283333" municipality="Wadi Turabah" pageId="23" pageNumber="22" specimenCode="NHMB Sc509" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="46B1FD68513AFF8E5FFCDE51FE49FBE5" box="[417,450,1036,1060]" pageId="23" pageNumber="22" type="male">1♂</specimenCount>
|
||
(fragmented),
|
||
<collectingMunicipality id="B06CAC9B513AFF8E5C34DE51FC88FBE5" box="[617,771,1036,1060]" latitude="25.5" longLatPrecision="1246" longitude="41.283333" name="Wadi Turabah" pageId="23" pageNumber="22">Wadi Turabah</collectingMunicipality>
|
||
[
|
||
<geoCoordinate id="35835026513AFF8E5E25DE71FF58FB85" box="[120,211,1068,1092]" degrees="25" direction="north" minutes="30" orientation="latitude" pageId="23" pageNumber="22" precision="925" value="25.5">25°30'N</geoCoordinate>
|
||
<geoCoordinate id="35835026513AFF8E5E87DE71FEBAFB85" box="[218,305,1068,1092]" degrees="41" direction="east" minutes="17" orientation="longitude" pageId="23" pageNumber="22" precision="925" value="41.283333">41°17'E</geoCoordinate>
|
||
,
|
||
<elevation id="DB9AD1D2513AFF8E5F60DE71FE44FB85" box="[317,463,1068,1092]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="23" pageNumber="22" unit="m" value="1430.0">
|
||
<quantity id="974F9B04513AFF8E5F60DE71FE1EFB82" box="[317,405,1068,1092]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="23" pageNumber="22" unit="m" value="1430.0">1,430 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
],
|
||
<date id="24091021513AFF8E5FB8DE71FDC3FB85" box="[485,584,1068,1092]" pageId="23" pageNumber="22" value="1986-12">
|
||
<collectingDate id="344DE9C9513AFF8E5FB8DE71FDC3FB85" box="[485,584,1068,1092]" pageId="23" pageNumber="22" value="1986-12">XII.1986</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="FD425337513AFF8E5CDCDE71FD74FB85" box="[641,767,1068,1092]" pageId="23" pageNumber="22">W. Büttiker</collectorName>
|
||
,
|
||
<specimenCode id="00119E9A513AFF8E5E2CDE11FE85FBA5" box="[113,270,1100,1124]" pageId="23" pageNumber="22">NHMB Sc509</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="18AD656A513AFF8E5E2CDED1FD6BFB05" pageId="23" pageNumber="22" type="distribution">
|
||
<paragraph id="500836E1513AFF8E5E2CDED1FD6BFB05" blockId="23.[113,771,1164,1220]" pageId="23" pageNumber="22">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="28A07671513AFF8E5F4DDED1FE44FB65" box="[272,463,1164,1188]" name="Saudi Arabia" pageId="23" pageNumber="22">SAUDI ARABIA</collectingCountry>
|
||
. Central plateau, Najd desert; east of Harrat Khaybar lava field, east of Hijaz mountains.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="18AD656A513AFF8E5E2CDEB1FE99FA45" pageId="23" pageNumber="22" type="diagnosis">
|
||
<paragraph id="500836E1513AFF8E5E2CDEB1FE99FA45" blockId="23.[113,771,1260,1412]" pageId="23" pageNumber="22">
|
||
DIAGNOSIS. A member of
|
||
<taxonomicName id="97B74D62513AFF8E5FDDDEB1FD94FAC5" authority="Lowe & Kovařík & Stockmann & Šťáhlavský, 2019" authorityName="Lowe & Kovařík & Stockmann & Šťáhlavský" authorityYear="2019" box="[384,543,1260,1284]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="62C3EAF3513AFF8E5FDDDEB1FD94FAC5" box="[384,543,1260,1284]" italics="true" pageId="23" pageNumber="22">Trypanothacus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="79F05788513AFF8E5C79DEB1FDE4FAC2" box="[548,623,1260,1284]" pageId="23" pageNumber="22" rank="genus">
|
||
<emphasis id="62C3EAF3513AFF8E5C79DEB1FDC7FAC5" bold="true" box="[548,588,1260,1284]" pageId="23" pageNumber="22">gen</emphasis>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5C04DEB1FDE3FAC5" bold="true" box="[601,616,1260,1284]" pageId="23" pageNumber="22">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differentiated from its congener as follows: pedipalp femur L/W
|
||
<specimenCount id="46B1FD68513AFF8E5CFCDF51FD3EFAE2" box="[673,693,1292,1315]" pageId="23" pageNumber="22" type="male">♂</specimenCount>
|
||
2.5,
|
||
<specimenCount id="46B1FD68513AFF8E5CB2DF51FC88FAE2" box="[751,771,1292,1315]" pageId="23" pageNumber="22" type="female">♀</specimenCount>
|
||
2.2; pedipalp patella L/W
|
||
<specimenCount id="46B1FD68513AFF8E5FC6DF71FE24FA82" box="[411,431,1324,1347]" pageId="23" pageNumber="22" type="male">♂</specimenCount>
|
||
2.1–2.2,
|
||
<specimenCount id="46B1FD68513AFF8E5C40DF71FDBAFA82" box="[541,561,1324,1347]" pageId="23" pageNumber="22" type="female">♀</specimenCount>
|
||
2.3, chela L/W
|
||
<specimenCount id="46B1FD68513AFF8E5CB2DF71FC88FA82" box="[751,771,1324,1347]" pageId="23" pageNumber="22" type="male">♂</specimenCount>
|
||
3.2–3.6,
|
||
<specimenCount id="46B1FD68513AFF8E5E92DF11FF68FAA2" box="[207,227,1356,1379]" pageId="23" pageNumber="22" type="female">♀</specimenCount>
|
||
4.0; metasoma IV L/D
|
||
<specimenCount id="46B1FD68513AFF8E5FBCDF11FE7EFAA2" box="[481,501,1356,1379]" pageId="23" pageNumber="22" type="male">♂</specimenCount>
|
||
1.8–2.0,
|
||
<specimenCount id="46B1FD68513AFF8E5C07DF11FDE5FAA2" box="[602,622,1356,1379]" pageId="23" pageNumber="22" type="female">♀</specimenCount>
|
||
1.9, V L/D
|
||
<specimenCount id="46B1FD68513AFF8E5CB2DF11FC88FAA2" box="[751,771,1356,1379]" pageId="23" pageNumber="22" type="male">♂</specimenCount>
|
||
2.4–2.6,
|
||
<specimenCount id="46B1FD68513AFF8E5E92DF31FF68FA42" box="[207,227,1388,1411]" pageId="23" pageNumber="22" type="female">♀</specimenCount>
|
||
2.4.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="18AD656A513AFF8E5E2CDFF1FAC9FB01" pageId="23" pageNumber="22" type="discussion">
|
||
<paragraph id="500836E1513AFF8E5E2CDFF1FC0EFEE0" blockId="23.[113,772,1452,2052]" lastBlockId="23.[816,1475,233,1153]" pageId="23" pageNumber="22">
|
||
COMMENTS. Hendrixson originally described
|
||
<taxonomicName id="97B74D62513AFF8E5CFDDFF1FF5CFA25" authorityName="Hendrixson" authorityYear="2006" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="buettikeri">
|
||
<emphasis id="62C3EAF3513AFF8E5CFDDFF1FF5CFA25" italics="true" pageId="23" pageNumber="22">Buthacus buettikeri</emphasis>
|
||
</taxonomicName>
|
||
on the basis of a series of specimens from several localities in
|
||
<collectingCountry id="28A07671513AFF8E5F5FDFB1FE18F9C5" box="[258,403,1516,1540]" name="Saudi Arabia" pageId="23" pageNumber="22">Saudi Arabia</collectingCountry>
|
||
(Hendrixon, 2006: 47, 51). We examined part of the type series and concluded that the
|
||
<typeStatus id="8F0C8843513AFF8E5E2CDC71FF45F985" box="[113,206,1580,1604]" pageId="23" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
male (No. 172a;
|
||
<figureCitation id="C88C2A64513AFF8E5FF4DC71FD81F985" box="[425,522,1580,1604]" captionStart="Figures 29–46" captionStartId="12.[115,192,1871,1893]" captionTargetBox="[131,1463,242,1849]" captionTargetId="figure-164@12.[113,1471,234,1854]" captionTargetPageId="12" captionText="Figures 29–46: Figures 29–42: Trypanothacus barnesi gen. et sp. n. Figures 29–40. Pedipalp segments. Figures 29–38. Male holotype, chela dorsal (29), external (30), and ventral (31) views. Patella dorsal (32), external (33) and ventral (34) views. Femur and trochanter internal (35), dorsal (36), and ventral (37) views. Movable finger dentition (38). Trichobothrial pattern is indicated in Figures 30–33 and 35–36. Figures 39– 40. Female paratype from the type locality, chela dorsal (39) and external (40) views. Figures 41–42. Telson, male holotype (41) and female paratype from type locality (42), lateral views. Figures 43–44: T. buettikeri comb. n., pedipalp chela dorsal, male holotype (43) and female paratype (44). Figures 45–46: Buthacus stockmanni Kovařík et al., 2016, telson, male holotype (45) and female paratype (46), lateral views." figureDoi="http://doi.org/10.5281/zenodo.6511485" httpUri="https://zenodo.org/record/6511485/files/figure.png" pageId="23" pageNumber="22">Figs. 43</figureCitation>
|
||
,
|
||
<figureCitation id="C88C2A64513AFF8E5C40DC71FDE9F985" box="[541,610,1580,1604]" captionStart="Figures 90–93" captionStartId="21.[113,190,1977,1999]" captionTargetBox="[131,1442,254,1923]" captionTargetId="figure-7@21.[110,1474,234,1945]" captionTargetPageId="21" captionText="Figures 90–93: Trypanothacus buettikeri comb. n. Figures 90–91. Male holotype in dorsal (90) and ventral (91) views. Figures 92–93: Female paratype in dorsal (92) and ventral (93) views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6511505" httpUri="https://zenodo.org/record/6511505/files/figure.png" pageId="23" pageNumber="22">90–91</figureCitation>
|
||
,
|
||
<figureCitation id="C88C2A64513AFF8E5C28DC71FD4DF985" box="[629,710,1580,1604]" captionStart-0="Figures 94–95" captionStart-1="Figures 96–106" captionStartId-0="22.[113,190,1259,1281]" captionStartId-1="24.[113,187,1903,1925]" captionTargetBox-0="[113,1474,234,1247]" captionTargetBox-1="[113,1474,234,1891]" captionTargetId-0="figure-416@22.[113,1474,234,1248]" captionTargetId-1="figure-11@24.[113,1474,231,1891]" captionTargetPageId-0="22" captionTargetPageId-1="24" captionText-0="Figures 94–95: Trypanothacus buettikeri comb. n. Male holotype, carapace and tergites I–IV (94), coxosternal area and sternites I–II (95). UV fluorescence. Scale bar: 2 mm." captionText-1="Figures 96–106: Trypanothacus buettikeri comb. n. Figures 96–101. Male holotype, lateral view of metasoma and telson (96), dorsal views of pedipalp chela (97) patella (98) and femur (99), external views of pedipalp chela (100) and patella (101). Figure 102. Male from Wadi Turabah (NHMB No. 509), dorsal view of pedipalp patella. Figures 103–106. Female paratype (NHMB 896), lateral views of metasoma (103) and pedipalp chela (104), dorsal view ofpedipalp patella (105) and femur (106).UV fluorescence.Scale bars:2 mm (96), 2 mm (97–101), 2 mm (102), 2 mm (103), 2 mm (104–106)." figureDoi-0="http://doi.org/10.5281/zenodo.6511509" figureDoi-1="http://doi.org/10.5281/zenodo.6511511" httpUri-0="https://zenodo.org/record/6511509/files/figure.png" httpUri-1="https://zenodo.org/record/6511511/files/figure.png" pageId="23" pageNumber="22">94–101</figureCitation>
|
||
) and
|
||
<typeStatus id="8F0C8843513AFF8E5E2CDC11FF46F9A5" box="[113,205,1612,1636]" pageId="23" pageNumber="22" type="paratype">paratype</typeStatus>
|
||
females from Khashm Dhibi (Nos. 172cd, NHMB 0896=172e;
|
||
<figureCitation id="C88C2A64513AFF8E5EAADC31FEC4F945" box="[247,335,1644,1668]" captionStart="Figures 29–46" captionStartId="12.[115,192,1871,1893]" captionTargetBox="[131,1463,242,1849]" captionTargetId="figure-164@12.[113,1471,234,1854]" captionTargetPageId="12" captionText="Figures 29–46: Figures 29–42: Trypanothacus barnesi gen. et sp. n. Figures 29–40. Pedipalp segments. Figures 29–38. Male holotype, chela dorsal (29), external (30), and ventral (31) views. Patella dorsal (32), external (33) and ventral (34) views. Femur and trochanter internal (35), dorsal (36), and ventral (37) views. Movable finger dentition (38). Trichobothrial pattern is indicated in Figures 30–33 and 35–36. Figures 39– 40. Female paratype from the type locality, chela dorsal (39) and external (40) views. Figures 41–42. Telson, male holotype (41) and female paratype from type locality (42), lateral views. Figures 43–44: T. buettikeri comb. n., pedipalp chela dorsal, male holotype (43) and female paratype (44). Figures 45–46: Buthacus stockmanni Kovařík et al., 2016, telson, male holotype (45) and female paratype (46), lateral views." figureDoi="http://doi.org/10.5281/zenodo.6511485" httpUri="https://zenodo.org/record/6511485/files/figure.png" pageId="23" pageNumber="22">Figs. 44</figureCitation>
|
||
,
|
||
<figureCitation id="C88C2A64513AFF8E5F04DC31FE15F945" box="[345,414,1644,1668]" captionStart="Figures 90–93" captionStartId="21.[113,190,1977,1999]" captionTargetBox="[131,1442,254,1923]" captionTargetId="figure-7@21.[110,1474,234,1945]" captionTargetPageId="21" captionText="Figures 90–93: Trypanothacus buettikeri comb. n. Figures 90–91. Male holotype in dorsal (90) and ventral (91) views. Figures 92–93: Female paratype in dorsal (92) and ventral (93) views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6511505" httpUri="https://zenodo.org/record/6511505/files/figure.png" pageId="23" pageNumber="22">92–93</figureCitation>
|
||
,
|
||
<figureCitation id="C88C2A64513AFF8E5FF4DC31FD82F945" box="[425,521,1644,1668]" captionStart="Figures 96–106" captionStartId="24.[113,187,1903,1925]" captionTargetBox="[113,1474,234,1891]" captionTargetId="figure-11@24.[113,1474,231,1891]" captionTargetPageId="24" captionText="Figures 96–106: Trypanothacus buettikeri comb. n. Figures 96–101. Male holotype, lateral view of metasoma and telson (96), dorsal views of pedipalp chela (97) patella (98) and femur (99), external views of pedipalp chela (100) and patella (101). Figure 102. Male from Wadi Turabah (NHMB No. 509), dorsal view of pedipalp patella. Figures 103–106. Female paratype (NHMB 896), lateral views of metasoma (103) and pedipalp chela (104), dorsal view ofpedipalp patella (105) and femur (106).UV fluorescence.Scale bars:2 mm (96), 2 mm (97–101), 2 mm (102), 2 mm (103), 2 mm (104–106)." figureDoi="http://doi.org/10.5281/zenodo.6511511" httpUri="https://zenodo.org/record/6511511/files/figure.png" pageId="23" pageNumber="22">103–106</figureCitation>
|
||
), from the Najd plateau region of central
|
||
<collectingCountry id="28A07671513AFF8E5F77DCD1FE3CF965" box="[298,439,1676,1700]" name="Saudi Arabia" pageId="23" pageNumber="22">Saudi Arabia</collectingCountry>
|
||
, belong to
|
||
<taxonomicName id="97B74D62513AFF8E5C72DCD1FD45F965" authority="Lowe & Kovařík & Stockmann & Šťáhlavský, 2019" authorityName="Lowe & Kovařík & Stockmann & Šťáhlavský" authorityYear="2019" box="[559,718,1676,1700]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="62C3EAF3513AFF8E5C72DCD1FD45F965" box="[559,718,1676,1700]" italics="true" pageId="23" pageNumber="22">Trypanothacus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="79F05788513AFF8E5C89DCD1FF0BF905" pageId="23" pageNumber="22" rank="genus">
|
||
<emphasis id="62C3EAF3513AFF8E5C89DCD1FD77F965" bold="true" box="[724,764,1676,1700]" pageId="23" pageNumber="22">gen</emphasis>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5E2CDCF1FF0BF905" bold="true" box="[113,128,1708,1732]" pageId="23" pageNumber="22">n</emphasis>
|
||
</taxonomicNameLabel>
|
||
.. However, several other specimens of the type series from widely disjunct sites along the western slopes of the
|
||
<collectingRegion id="9273F803513AFF8E5CD6DC91FD32F925" box="[651,697,1740,1764]" country="Saudi Arabia" name="'Asir" pageId="23" pageNumber="22">Asir</collectingRegion>
|
||
ranges (facing the Red Sea Coast), differed from the
|
||
<typeStatus id="8F0C8843513AFF8E5C0EDCB1FD3BF8C5" box="[595,688,1772,1796]" pageId="23" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
and are associated with the
|
||
<taxonomicName id="97B74D62513AFF8E5F10DD51FC88F8E5" authority="(Ehrenberg, 1829)" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" box="[333,771,1804,1828]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="leptochelys">
|
||
<emphasis id="62C3EAF3513AFF8E5F10DD51FDB8F8E5" box="[333,563,1804,1828]" italics="true" pageId="23" pageNumber="22">Buthacus leptochelys</emphasis>
|
||
(Ehrenberg, 1829)
|
||
</taxonomicName>
|
||
species complex of the genus
|
||
<taxonomicName id="97B74D62513AFF8E5FF3DD71FD9AF885" authorityName="Birula" authorityYear="1908" box="[430,529,1836,1860]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="62C3EAF3513AFF8E5FF3DD71FD9AF885" box="[430,529,1836,1860]" italics="true" pageId="23" pageNumber="22">Buthacus</emphasis>
|
||
</taxonomicName>
|
||
(i.e., female
|
||
<typeStatus id="8F0C8843513AFF8E5CC0DD71FC88F885" box="[669,771,1836,1860]" pageId="23" pageNumber="22" type="paratype">paratypes</typeStatus>
|
||
including NHMB 515=172b, 630=172f and 620=
|
||
<quantity id="974F9B04513AFF8E5CD0DD11FD49F8A5" box="[653,706,1868,1892]" metricMagnitude="-1" metricUnit="kg" metricValue="1.72" pageId="23" pageNumber="22" unit="g" value="172.0">172g</quantity>
|
||
; from Jabal al-Ghamdiyah, Wadi Maraum, Addar, Kijat (examined); also likely including MNHN RS8425 from
|
||
<collectingRegion id="9273F803513AFF8E5C1ADDD1FD15F865" box="[583,670,1932,1956]" country="Saudi Arabia" name="Makkah" pageId="23" pageNumber="22">Makkah</collectingRegion>
|
||
By-pass, based on specimen photographs on the MNHN website). In light of these misidentifications, the status of other MNHN
|
||
<typeStatus id="8F0C8843513AFF8E5E2CDDB1FF5CF7C5" box="[113,215,2028,2052]" pageId="23" pageNumber="22" type="paratype">paratypes</typeStatus>
|
||
from a widely disjunct site near the northern border of
|
||
<collectingCountry id="28A07671513AFF8E5D10DAB4FC53FEC0" box="[845,984,233,257]" name="Saudi Arabia" pageId="23" pageNumber="22">Saudi Arabia</collectingCountry>
|
||
(ca.
|
||
<quantity id="974F9B04513AFF8E5A51DAB4FBFAFEC0" box="[1036,1137,233,257]" metricMagnitude="6" metricUnit="m" metricValue="1.03" pageId="23" pageNumber="22" unit="km" value="1030.0">1,030 km</quantity>
|
||
from the type locality) remains unclear.
|
||
</paragraph>
|
||
<paragraph id="500836E1513AFF8E5D03DB74FA30FB41" blockId="23.[816,1475,233,1153]" pageId="23" pageNumber="22">
|
||
<bibRefCitation id="34264B10513AFF8E5D03DB74FB4BFE80" author="DESOUKY, M. M. A. & A. M. ALSHAMMARI" box="[862,1216,297,321]" journalOrPublisher="Bulletin of the British Arachnological Society" pageId="23" pageNumber="22" pagination="193 - 200" part="15" refId="ref14018" refString="DESOUKY, M. M. A. & A. M. ALSHAMMARI. 2011. Scorpions of the Ha'il Region, northern Saudi Arabia, and molecular phylogenetics of two common species, Androctonus crassicauda and Scorpio maurus kruglovi. Bulletin of the British Arachnological Society, 15 (6): 193 - 200." title="Scorpions of the Ha'il Region, northern Saudi Arabia, and molecular phylogenetics of two common species, Androctonus crassicauda and Scorpio maurus kruglovi" type="journal article" year="2011">Desouky & Alshammari (2011)</bibRefCitation>
|
||
recorded
|
||
<taxonomicName id="97B74D62513AFF8E5B64DB77FA49FE80" authorityName="Hendrixson" authorityYear="2006" box="[1337,1474,297,321]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="buettikeri">
|
||
<emphasis id="62C3EAF3513AFF8E5B64DB77FA49FE80" box="[1337,1474,297,321]" italics="true" pageId="23" pageNumber="22">B. buettikeri</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<collectingRegion id="9273F803513AFF8E5DC5DB14FB90FEA0" box="[920,1051,329,353]" country="Saudi Arabia" name="Ha'il" pageId="23" pageNumber="22">Ha’il region</collectingRegion>
|
||
in the central plateau of
|
||
<collectingCountry id="28A07671513AFF8E5B6BDB14FA49FEA0" box="[1334,1474,329,353]" name="Saudi Arabia" pageId="23" pageNumber="22">Saudi Arabia</collectingCountry>
|
||
(Hufair, Sarraa near Al-Ghazala), on “sandy desertˮ. In photographs, their adult female (carapace L
|
||
<quantity id="974F9B04513AFF8E5B55DBD4FAE2FE60" box="[1288,1385,393,417]" metricMagnitude="-3" metricUnit="m" metricValue="5.92" pageId="23" pageNumber="22" unit="mm" value="5.92">5.92 mm</quantity>
|
||
) (fig. 7) appears morphometrically consistent (e.g. patella L/W 2.57, metasoma I L/W 1.02) with females of the type series that we have examined, however their adult male (carapace L
|
||
<quantity id="974F9B04513AFF8E5BFCDBB4FCD0FDE0" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="23" pageNumber="22" unit="mm" value="6.4">6.4 mm</quantity>
|
||
) (fig. 6) has markedly different morphometrics with much more slender pedipalps and metasoma (e.g. pedipalp patella L/W 2.95, metasoma I L/W 1.38) than in the male types of
|
||
<taxonomicName id="97B74D62513AFF8E5BF6D817FC1FFD40" authorityName="Hendrixson" authorityYear="2006" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="buettikeri">
|
||
<emphasis id="62C3EAF3513AFF8E5BF6D817FA30FDA0" box="[1451,1467,586,609]" italics="true" pageId="23" pageNumber="22">B</emphasis>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5D6DD834FC1FFD40" box="[816,916,617,641]" italics="true" pageId="23" pageNumber="22">buettikeri</emphasis>
|
||
</taxonomicName>
|
||
. It is also more slender than all males of
|
||
<taxonomicName id="97B74D62513AFF8E5B68D837FA16FD40" authority="Lowe & Kovařík & Stockmann & Šťáhlavský, 2019" authorityName="Lowe & Kovařík & Stockmann & Šťáhlavský" authorityYear="2019" box="[1333,1437,617,641]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="barnesi" status="sp. nov.">
|
||
<emphasis id="62C3EAF3513AFF8E5B68D837FACFFD40" box="[1333,1348,618,641]" italics="true" pageId="23" pageNumber="22">T</emphasis>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5B12D834FA16FD40" box="[1359,1437,617,641]" italics="true" pageId="23" pageNumber="22">barnesi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="79F05788513AFF8E5BFED834FCCDFD61" pageId="23" pageNumber="22" rank="species">
|
||
<emphasis id="62C3EAF3513AFF8E5BFED834FA37FD40" bold="true" box="[1443,1468,617,641]" pageId="23" pageNumber="22">sp</emphasis>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5D6DD8D4FCB4FD60" bold="true" box="[816,831,649,673]" pageId="23" pageNumber="22">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
in our type series. This would seem to indicate large variation in male morphometrics of
|
||
<taxonomicName id="97B74D62513AFF8E5A14D8F7FB40FD00" authorityName="Hendrixson" authorityYear="2006" box="[1097,1227,681,705]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="buettikeri">
|
||
<emphasis id="62C3EAF3513AFF8E5A14D8F7FBD2FD00" box="[1097,1113,682,705]" italics="true" pageId="23" pageNumber="22">B</emphasis>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5A3AD8F4FB40FD00" box="[1127,1227,681,705]" italics="true" pageId="23" pageNumber="22">buettikeri</emphasis>
|
||
</taxonomicName>
|
||
, maybe reflecting local geographic races as
|
||
<collectingRegion id="9273F803513AFF8E5A51D894FBC9FD20" box="[1036,1090,713,737]" country="Saudi Arabia" name="Ha'il" pageId="23" pageNumber="22">Ha’il</collectingRegion>
|
||
is located some distance northwest of the type locality. However, a large adult male that we determined as
|
||
<taxonomicName id="97B74D62513AFF8E5D99D957FBCAFCE0" authorityName="Hendrixson" authorityYear="2006" box="[964,1089,777,801]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="buettikeri">
|
||
<emphasis id="62C3EAF3513AFF8E5D99D957FC58FCE0" box="[964,979,778,801]" italics="true" pageId="23" pageNumber="22">T</emphasis>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5D80D954FBCAFCE0" box="[989,1089,777,801]" italics="true" pageId="23" pageNumber="22">buettikeri</emphasis>
|
||
</taxonomicName>
|
||
, has morphometrics more robust than the
|
||
<typeStatus id="8F0C8843513AFF8E5D05D974FC3DFC80" box="[856,950,809,833]" pageId="23" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
, yet was collected from Wadi Turabah, relatively closer to the
|
||
<collectingRegion id="9273F803513AFF8E5DE5D914FBB0FCA0" box="[952,1083,841,865]" country="Saudi Arabia" name="Ha'il" pageId="23" pageNumber="22">Ha’il region</collectingRegion>
|
||
. As we have not studied
|
||
<collectingRegion id="9273F803513AFF8E5B1FD914FA49FCA0" box="[1346,1474,841,865]" country="Saudi Arabia" name="Ha'il" pageId="23" pageNumber="22">Ha’il region</collectingRegion>
|
||
material, we withhold judgement on the taxonomic status of those populations, but the large morphometric difference likely indicates a different species.
|
||
<bibRefCitation id="34264B10513AFF8E5AE4D9F4FC26FC20" author="DESOUKY, M. M. A. & A. M. ALSHAMMARI" journalOrPublisher="Bulletin of the British Arachnological Society" pageId="23" pageNumber="22" pagination="193 - 200" part="15" refId="ref14018" refString="DESOUKY, M. M. A. & A. M. ALSHAMMARI. 2011. Scorpions of the Ha'il Region, northern Saudi Arabia, and molecular phylogenetics of two common species, Androctonus crassicauda and Scorpio maurus kruglovi. Bulletin of the British Arachnological Society, 15 (6): 193 - 200." title="Scorpions of the Ha'il Region, northern Saudi Arabia, and molecular phylogenetics of two common species, Androctonus crassicauda and Scorpio maurus kruglovi" type="journal article" year="2011">Desouky & Alshammari (2011: 196)</bibRefCitation>
|
||
assumed
|
||
<taxonomicName id="97B74D62513AFF8E5A44D997FA35FC20" authority="Lourenco & Leguin, 2009" authorityName="Lourenco & Leguin" authorityYear="2009" box="[1049,1470,969,993]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="62C3EAF3513AFF8E5A44D997FBA2FC20" box="[1049,1065,970,993]" italics="true" pageId="23" pageNumber="22">B</emphasis>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5A64D994FB12FC20" box="[1081,1177,969,993]" italics="true" pageId="23" pageNumber="22">williamsi</emphasis>
|
||
<bibRefCitation id="34264B10513AFF8E5AFFD994FA35FC20" author="LOURENCO, W. R. & E. - A. LEGUIN" box="[1186,1470,969,993]" journalOrPublisher="Zoology in the Middle East" pageId="23" pageNumber="22" pagination="103 - 108" part="46" refId="ref15044" refString="LOURENCO, W. R. & E. - A. LEGUIN. 2009. A new species of the genus Buthacus Birula, 1908 from the United Arab Emirates (Scorpiones, Buthidae). Zoology in the Middle East, 46: 103 - 108." title="A new species of the genus Buthacus Birula, 1908 from the United Arab Emirates (Scorpiones, Buthidae)" type="journal article" year="2009">Lourenço & Leguin, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
, to be a junior synonym of
|
||
<taxonomicName id="97B74D62513AFF8E5A3CD9B7FB6CFBC0" authorityName="Hendrixson" authorityYear="2006" box="[1121,1255,1001,1025]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="buettikeri">
|
||
<emphasis id="62C3EAF3513AFF8E5A3CD9B7FBFAFBC0" box="[1121,1137,1002,1025]" italics="true" pageId="23" pageNumber="22">B</emphasis>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5ADED9B4FB6CFBC0" box="[1155,1255,1001,1025]" italics="true" pageId="23" pageNumber="22">buettikeri</emphasis>
|
||
</taxonomicName>
|
||
, but did not justify this by comparative study of the types of either species. They also claimed that the telson and aculeus in
|
||
<taxonomicName id="97B74D62513AFF8E5B56DE77FA04FB80" authorityName="Hendrixson" authorityYear="2006" box="[1291,1423,1065,1089]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="buettikeri">
|
||
<emphasis id="62C3EAF3513AFF8E5B56DE77FA90FB80" box="[1291,1307,1066,1089]" italics="true" pageId="23" pageNumber="22">B</emphasis>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5B76DE74FA04FB80" box="[1323,1423,1065,1089]" italics="true" pageId="23" pageNumber="22">buettikeri</emphasis>
|
||
</taxonomicName>
|
||
was “clearly more slender and elongated” than in
|
||
<taxonomicName id="97B74D62513AFF8E5B78DE17FA30FBA0" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" box="[1317,1467,1097,1121]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="leptochelys">
|
||
<emphasis id="62C3EAF3513AFF8E5B78DE17FABEFBA0" box="[1317,1333,1098,1121]" italics="true" pageId="23" pageNumber="22">B</emphasis>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5B1BDE14FA30FBA0" box="[1350,1467,1097,1121]" italics="true" pageId="23" pageNumber="22">leptochelys</emphasis>
|
||
</taxonomicName>
|
||
, misrepresenting
|
||
<bibRefCitation id="34264B10513AFF8E5D80DE34FB2DFB40" author="Hendrixson" box="[989,1190,1129,1153]" firstAuthor="Hendrixson" pageId="23" pageNumber="22" pagination="33 - 120" refId="ref14260" refString="HENDRIXSON, B. E. 2006. Buthid scorpions of Saudi Arabia, with notes on other families (Scorpiones: Buthidae, Liochelidae, Scorpionidae). Fauna of Arabia, 21: 33 - 120." type="journal article" year="2006">Hendrixson (2006)</bibRefCitation>
|
||
, when the opposite is true.
|
||
</paragraph>
|
||
<paragraph id="500836E1513AFF8E5D6DDEF4FAC9FB01" blockId="23.[816,1346,1193,1217]" box="[816,1346,1193,1217]" pageId="23" pageNumber="22">
|
||
<heading id="0B40818D513AFF8E5D6DDEF4FAC9FB01" box="[816,1346,1193,1217]" centered="true" fontSize="10" level="2" pageId="23" pageNumber="22" reason="4">
|
||
AFFINITIES. See comments under
|
||
<taxonomicName id="97B74D62513AFF8E5ACEDEF7FB74FB00" authority="Lowe & Kovařík & Stockmann & Šťáhlavský, 2019" authorityName="Lowe & Kovařík & Stockmann & Šťáhlavský" authorityYear="2019" box="[1171,1279,1193,1217]" class="Arachnida" family="Buthidae" genus="Trypanothacus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="barnesi" status="sp. nov.">
|
||
<emphasis id="62C3EAF3513AFF8E5ACEDEF7FB29FB00" box="[1171,1186,1194,1217]" italics="true" pageId="23" pageNumber="22">T</emphasis>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5AF2DEF4FB74FB00" box="[1199,1279,1193,1217]" italics="true" pageId="23" pageNumber="22">barnesi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="79F05788513AFF8E5B5BDEF4FAC9FB01" box="[1286,1346,1193,1217]" pageId="23" pageNumber="22" rank="species">
|
||
<emphasis id="62C3EAF3513AFF8E5B5BDEF4FA94FB00" bold="true" box="[1286,1311,1193,1217]" pageId="23" pageNumber="22">sp</emphasis>
|
||
.
|
||
<emphasis id="62C3EAF3513AFF8E5B70DEF4FAB7FB00" bold="true" box="[1325,1340,1193,1217]" pageId="23" pageNumber="22">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |