259 lines
26 KiB
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259 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.198.79514" ID-Pensoft-Pub="1314-2003-198-1" ID-Pensoft-UUID="486F1F1B4F5854D2831AAA341B9A322C" ModsDocID="1314-2003-198-1" checkinTime="1654168732655" checkinUser="pensoft" docAuthor="Aubriot, Xavier & Knapp, Sandra" docDate="2022" docId="71B0B4525B4C6A5154CA1FAA3C357099" docLanguage="en" docName="PhytoKeys 198: 1-270" docOrigin="PhytoKeys 198" docPubDate="2022-06-01" docSource="http://dx.doi.org/10.3897/phytokeys.198.79514" docTitle="Solanum poka Dunal, Encycl. [J. Lamarck & al.] Suppl. 3: 768. 1814." docType="treatment" docVersion="1" id="486F1F1B4F5854D2831AAA341B9A322C" lastPageNumber="1" masterDocId="486F1F1B4F5854D2831AAA341B9A322C" masterDocTitle="A revision of the " spiny solanums " of Tropical Asia (Solanum, the Leptostemonum Clade, Solanaceae)" masterLastPageNumber="270" masterPageNumber="1" pageNumber="1" updateTime="1654168732655" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the " spiny solanums " of Tropical Asia (Solanum, the Leptostemonum Clade, Solanaceae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Aubriot, Xavier</mods:namePart>
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<mods:affiliation>Universite Paris-Saclay, CNRS, AgroParisTech, Ecologie Systematique et Evolution, 91190, Gif-sur-Yvette, France & The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Knapp, Sandra</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7698-3945</mods:nameIdentifier>
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<mods:affiliation>The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:nameIdentifier type="email">s.knapp@nhm.ac.uk</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-06-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>198</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>270</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.198.79514</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.198.79514</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-198-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">486F1F1B4F5854D2831AAA341B9A322C</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:71B0B4525B4C6A5154CA1FAA3C357099" httpUri="http://treatment.plazi.org/id/71B0B4525B4C6A5154CA1FAA3C357099" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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31.
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<taxonomicName authority="Dunal, Encycl. [J. Lamarck & al.] Suppl. 3: 768. 1814." authorityName="Dunal, Encycl. [J. Lamarck & al.] Suppl. 3: 768. 1814." class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum poka" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="poka">Solanum poka Dunal, Encycl. [J. Lamarck & al.] Suppl. 3: 768. 1814.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Solanum poka Dunal - Herbarium specimen (epitype) collected in Indonesia (Horsfield s. n., BM 000886306). Photograph credit: CC-BY, © copyright The Trustees of the Natural History Museum, London." figureDoi="10.3897/phytokeys.198.79514.figure51" httpUri="https://binary.pensoft.net/fig/689744" pageId="0" pageNumber="1">Fig. 51</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum torvum subsp. var. var. scabrescens" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="torvum" subSpecies="var." variety="scabrescens">Solanum torvum Sw. var. scabrescens</taxonomicName>
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Miq., Fl. Ned. Ind. 2: 648. 1857. Type. Indonesia. Sumatra: Sin. loc.,
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<emphasis italics="true" pageId="0" pageNumber="1">F.W. Junghuhn s.n.</emphasis>
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(lectotype, designated here: L [L.0403917]; isolectotype: U [U0113979]).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum torvum subsp. var. var. polyacanthum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="torvum" subSpecies="var." variety="polyacanthum">Solanum torvum Sw. var. polyacanthum</taxonomicName>
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Miq., Fl. Ned. Ind. 2: 648. 1857, as "
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<emphasis italics="true" pageId="0" pageNumber="1">polyacantha</emphasis>
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". Type. Indonesia. Java: sin. loc.,
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<emphasis italics="true" pageId="0" pageNumber="1">T. Horsfield s.n.</emphasis>
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(lectotype, designated here: BM [BM000886307]).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">
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<typeStatus>Type</typeStatus>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Based on an unpublished illustration of a Jean-Baptiste Leschenault de la Tour collection from Java kept in the
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<normalizedToken originalValue="Node-Véran">Node-Veran</normalizedToken>
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collection in Montpellier (
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<typeStatus>lectotype</typeStatus>
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, designated by
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<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.70.9758" author="Aubriot, X" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="97 - 110" refId="B9" refString="Aubriot, X, Loup, C, Knapp, S, 2016b. Confirming the identity of two enigmatic "spiny solanums" (Solanum subgenus Leptostemonum, Solanaceae) collected by Jean-Baptiste Leschenault in Java. PhytoKeys 70: 97 - 110, DOI: https://doi.org/10.3897/phytokeys.70.9758" title="Confirming the identity of two enigmatic " spiny solanums " (Solanum subgenus Leptostemonum, Solanaceae) collected by Jean-Baptiste Leschenault in Java." url="https://doi.org/10.3897/phytokeys.70.9758" volume="70" year="2016 b">Aubriot et al. 2016b</bibRefCitation>
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, pg. 105: [illustration] Sol. Tab. 55 [MPU028527]).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shrubs to 3 m, armed. Stems erect, terete, usually densely prickly distally, moderately pubescent, glabrescent; prickles to 3.5 mm long, to 2.5 mm wide at the base, straight, awl-shaped to deltate, conical, pale yellow, glabrescent; pubescence of mixed sessile and variously stalked porrect-stellate trichomes, the stalks to 0.2 mm long, the rays (4-)5-8, 0.1-0.25 mm long, the midpoints reduced to mere bumps; new growth moderately stellate-pubescent, black to dark brownish in dry material; bark of older stems brownish grey, sparsely stellate-pubescent. Sympodial units difoliate, the leaves geminate. Leaves simple, entire to deeply lobed, the blades 11-24 cm long, 4-13 cm wide, ca. 1.5-3 times longer than wide, elliptic to broadly ovate, chartaceous, slightly discolorous, unarmed or pricky with 1-6 prickles per leaf side, the prickles to 6 mm long, to 1.5 mm wide at the base, straight or slightly curved at the tip, awl-shaped, conical, pale yellow, glabrous; adaxial surface moderately stellate-pubescent with porrect, sessile and less often variously stalked trichomes, the stalks to 0.1 mm long, the rays 4-8, 0.1-0.4 mm long, the midpoints to 0.25 mm long; abaxial surface moderately stellate-pubescent with trichomes like those of the adaxial surface, but more often stalked; major veins 6-8 pairs drying yellow; base short-attenuate to truncate; margins entire or shallowly to deeply lobed, the lobes 1-5 on each side, 0.5-5 cm long, deltate, rounded to apically acute, the sinuses extending up to 2/3 of the distance to the midvein; apex acute; petiole 1.5-4 cm long, 1/10-1/5 of the leaf blade length, densely stellate-pubescent with porrect, sessile trichomes like those of the blades, unarmed or prickly with 1-5 prickles like those of the stems. Inflorescences 2-5 cm long, apparently lateral or leaf opposed, unbranched to up to 2 times branched, with ca. 5-20 flowers, 2-6 flowers open at any one time, moderately to densely stellate-pubescent with porrect-stellate trichomes like those of the stems, unarmed; peduncle 0.5-1.5 cm long, unarmed or with very few prickles; pedicels 0.5-1.2 cm long, ca. 1 mm in diameter at the base, ca. 1.5 mm in diameter at the apex, erect, unarmed, densely stellate-pubescent with porrect-stellate trichomes like those of the inflorescence axes, articulated at the base; pedicel scars spaced 2-4 mm apart. Flowers 5-merous, apparently all perfect. Calyx with the tube 1-3 mm long, conical, the lobes 3-5 mm long, ca. 1.5 mm wide, the lower part deltate and abruptly constricting to an elongate acumen, the acumen 3/4 of the total lobe length, the abaxial surface more or less strongly keeled along the midvein, unarmed, densely stellate-pubescent on the midvein with porrect-stellate trichomes like those of the pedicels. Corolla 1-2 cm in diameter, white, stellate, lobed ca. 1/2-2/3 of the way to the base, the lobes 5-8 mm long, 2-3.5 mm wide, deltate, spreading at anthesis, densely stellate-pubescent abaxially on parts exposed in bud. Stamens equal; anthers 5-6.5 mm long, ca. 0.75 mm wide, tapering, yellow, connivent, glabrous, poricidal at the tips, the pores not lengthening to slits with age; filament tube <0.5 mm long, glabrous; free portion of the filaments 0.75-1.5 mm long, glabrous. Ovary conical, minutely glandular-puberulent; style 0.6-1 cm long, slender, curved at the apex, with a few scattered hairs at the tip; stigma capitate, minutely papillate, stellate-pubescent. Fruit a globose berry, 8-18 per infructescence, 0.8-1.5 cm in diameter, the pericarp smooth, bluish green when young, turning to dark greyish yellow, glabrous; fruiting pedicels 1.2-2.5 cm long, ca. 1-1.5 mm in diameter at the base, 2-3 mm in diameter at the apex, woody, erect, unarmed; fruiting calyx lobes not expanding. Seeds 100-200 per berry, 1.75-2 mm long, 1.5-1.75 mm wide, flattened reniform, pale yellowish, the surface minutely pitted, the testal cells sinuate in outline. Chromosome number: not known.</paragraph>
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<caption doi="10.3897/phytokeys.198.79514.figure51" httpUri="https://binary.pensoft.net/fig/689744" pageId="0" pageNumber="1" start="Figure 51" startId="F51">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 51.</emphasis>
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<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum poka" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="poka">
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<emphasis italics="true" pageId="0" pageNumber="1">Solanum poka</emphasis>
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</taxonomicName>
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Dunal - Herbarium specimen (epitype) collected in Indonesia (
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<emphasis italics="true" pageId="0" pageNumber="1">Horsfield s.n.</emphasis>
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, BM000886306). Photograph credit: CC-BY, © copyright The Trustees of the Natural History Museum, London.
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</paragraph>
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</caption>
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<caption doi="10.3897/phytokeys.198.79514.figure52" httpUri="https://binary.pensoft.net/fig/689745" pageId="0" pageNumber="1" start="Figure 52" startId="F52">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 52.</emphasis>
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Distribution of
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<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. poka" order="Solanales" pageId="0" pageNumber="1" rank="species" species="poka">
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<emphasis italics="true" pageId="0" pageNumber="1">S. poka</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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(Fig.
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<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Distribution of S. poka." figureDoi="10.3897/phytokeys.198.79514.figure52" httpUri="https://binary.pensoft.net/fig/689745" pageId="0" pageNumber="1">52</figureCitation>
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).
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</emphasis>
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<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum poka" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="poka">
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<emphasis italics="true" pageId="0" pageNumber="1">Solanum poka</emphasis>
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</taxonomicName>
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is widely distributed in the Malay Archipelago, from western Sumatra to the Maluku Islands and across Sulawesi, northwards to the Talaud islands.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="ecology">
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<paragraph pageId="0" pageNumber="1">Ecology and habitat.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum poka" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="poka">
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<emphasis italics="true" pageId="0" pageNumber="1">Solanum poka</emphasis>
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</taxonomicName>
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is a plant of evergreen broadleaf forests, growing in open woodland, forest edges, degraded vegetation, usually on limestone or volcanic rocks; from sea level to 1,600 m elevation.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="common names and uses">
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<paragraph pageId="0" pageNumber="1">Common names and uses.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Indonesia. Java: pooka (
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<emphasis italics="true" pageId="0" pageNumber="1">Horsfield 786</emphasis>
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), daun
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<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum poka" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="poka">Solanum poka</taxonomicName>
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(
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<bibRefCitation author="Dunal, M-F" editor="Poiret, JLM" journalOrPublisher="Botanique, Supplement" pageId="0" pageNumber="1" pagination="738 - 780" refId="B71" refString="Dunal, M-F, 1814. Morelle, Solanum. In: Poiret, JLM, Ed., Encyclopedie methodique. Botanique, Supplement 3: 738 - 780" title="Morelle, Solanum." volume="3" volumeTitle="Encyclopedie methodique." year="1814">Dunal 1814</bibRefCitation>
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); North Sulawesi: poki poki (
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<emphasis italics="true" pageId="0" pageNumber="1">De La Savinierre 343bis</emphasis>
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), sangkirumanuwawi (
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<emphasis italics="true" pageId="0" pageNumber="1">Lam 2772</emphasis>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="preliminary conservation status">
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<paragraph pageId="0" pageNumber="1">Preliminary conservation status</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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(
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<bibRefCitation author="Hutchings, A" journalOrPublisher="AGRIVITA Journal of Agricultural Science" pageId="0" pageNumber="1" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B105" refString="2019. . http://www.iucnredlist.org/documents/RedListGuidelines.pdf" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2019">IUCN 2019</bibRefCitation>
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).
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</emphasis>
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Least Concern (LC). EOO (1,202,780 km2, LC); AOO (68 km2, EN). Although the EOO measurement indicates a status of least concern, the few collections coupled with the profound transformation in lowland Indonesian habitats where
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<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. poka" order="Solanales" pageId="0" pageNumber="1" rank="species" species="poka">
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<emphasis italics="true" pageId="0" pageNumber="1">S. poka</emphasis>
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</taxonomicName>
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is found (
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<bibRefCitation DOI="https://doi.org/10.1038/nclimate2277" author="Margono, BA" journalOrPublisher="Nature Climate Change" pageId="0" pageNumber="1" pagination="1 - 6" refId="B156" refString="Margono, BA, Potapov, PV, Turubanova, S, Stolle, F, Hansen, MC, 2014. Primary forest cover loss in Indonesia over 2000-2012. Nature Climate Change 4 (8): 1 - 6, DOI: https://doi.org/10.1038/nclimate2277" title="Primary forest cover loss in Indonesia over 2000 - 2012." url="https://doi.org/10.1038/nclimate2277" volume="4" year="2014">Margono et al. 2014</bibRefCitation>
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) suggest that the species is a priority for recollection and reassessment.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="discussion">
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<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum poka" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="poka">
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<emphasis italics="true" pageId="0" pageNumber="1">Solanum poka</emphasis>
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</taxonomicName>
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is a member of the Torva clade (
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<bibRefCitation DOI="https://doi.org/10.1111/boj.12412" author="Aubriot, X" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="199 - 223" refId="B8" refString="Aubriot, X, Singh, P, Knapp, S, 2016a. Tropical Asian species show the Old World clade of "spiny solanums" (subgenus Leptostemonum pro parte: Solanaceae) is not monophyletic. Botanical Journal of the Linnean Society 181 (2): 199 - 223, DOI: https://doi.org/10.1111/boj.12412" title="Tropical Asian species show the Old World clade of " spiny solanums " (subgenus Leptostemonum pro parte: Solanaceae) is not monophyletic." url="https://doi.org/10.1111/boj.12412" volume="181" year="2016 a">Aubriot et al. 2016a</bibRefCitation>
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) and is similar to other members of the group occurring in southeast Asia such as
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<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. pseudosaponaceum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="pseudosaponaceum">
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<emphasis italics="true" pageId="0" pageNumber="1">S. pseudosaponaceum</emphasis>
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</taxonomicName>
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and
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<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. torvoideum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="torvoideum">
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<emphasis italics="true" pageId="0" pageNumber="1">S. torvoideum</emphasis>
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</taxonomicName>
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. Morphologically,
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<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. poka" order="Solanales" pageId="0" pageNumber="1" rank="species" species="poka">
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<emphasis italics="true" pageId="0" pageNumber="1">S. poka</emphasis>
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</taxonomicName>
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most closely resembles
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<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. pseudosaponaceum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="pseudosaponaceum">
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<emphasis italics="true" pageId="0" pageNumber="1">S. pseudosaponaceum</emphasis>
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</taxonomicName>
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, a widespread species ranging from Taiwan and southern China to Indonesia but differs in having denser indumentum on the adaxial leaf surface, more numerous straight prickles on the upper stems, fewer, larger flowers with elongate strongly keeled calyx lobes, and much larger fruits. Flowers of
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<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. pseudosaponaceum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="pseudosaponaceum">
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<emphasis italics="true" pageId="0" pageNumber="1">S. pseudosaponaceum</emphasis>
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||
</taxonomicName>
|
||
are lilac or purplish-white while those of
|
||
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. poka" order="Solanales" pageId="0" pageNumber="1" rank="species" species="poka">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. poka</emphasis>
|
||
</taxonomicName>
|
||
are always described on labels as white.
|
||
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum poka" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="poka">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Solanum poka</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from the introduced
|
||
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. torvum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="torvum">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. torvum</emphasis>
|
||
</taxonomicName>
|
||
in its strongly keeled calyx lobes with elongate acumens (rather than deltate lobes), red (rather than green) berries, and in the lack of simple glandular trichomes in the inflorescence.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.70.9758" author="Aubriot, X" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="97 - 110" refId="B9" refString="Aubriot, X, Loup, C, Knapp, S, 2016b. Confirming the identity of two enigmatic "spiny solanums" (Solanum subgenus Leptostemonum, Solanaceae) collected by Jean-Baptiste Leschenault in Java. PhytoKeys 70: 97 - 110, DOI: https://doi.org/10.3897/phytokeys.70.9758" title="Confirming the identity of two enigmatic " spiny solanums " (Solanum subgenus Leptostemonum, Solanaceae) collected by Jean-Baptiste Leschenault in Java." url="https://doi.org/10.3897/phytokeys.70.9758" volume="70" year="2016 b">Aubriot et al. (2016b)</bibRefCitation>
|
||
recognised
|
||
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. torvum subsp. var. var. scabrescens" order="Solanales" pageId="0" pageNumber="1" rank="variety" species="torvum" subSpecies="var." variety="scabrescens">S. torvum var. scabrescens</taxonomicName>
|
||
as a synonym in their resurrection of the name
|
||
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. poka" order="Solanales" pageId="0" pageNumber="1" rank="species" species="poka">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. poka</emphasis>
|
||
</taxonomicName>
|
||
but erroneously referred to the Leiden duplicate (L.0403917) of the
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Junghuhn s.n.</emphasis>
|
||
collection cited in the protologue as
|
||
<normalizedToken originalValue="“holotype”">"holotype"</normalizedToken>
|
||
;
|
||
<bibRefCitation author="Miquel, FAW" journalOrPublisher="American Journal of Botany" pageId="0" pageNumber="1" refId="B176" refString="Miquel, FAW, 1857. Ordo CLXIV. Solanaeae Juss. Flora Nederlandsch Indie. Vol. 2. CG van der Post, Amsterdam, 633-672." title="Ordo CLXIV. Solanaeae Juss. Flora Nederlandsch Indie. Vol. 2. CG van der Post, Amsterdam, 633 - 672." year="1857">Miquel (1857)</bibRefCitation>
|
||
cited no herbaria in the protologue. They did not include
|
||
<normalizedToken originalValue="Miquel’s">Miquel's</normalizedToken>
|
||
var.
|
||
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum poka var. polyacanthum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="poka" variety="polyacanthum">Solanum poka polyacanthum</taxonomicName>
|
||
in the synonymy of
|
||
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. poka" order="Solanales" pageId="0" pageNumber="1" rank="species" species="poka">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. poka</emphasis>
|
||
</taxonomicName>
|
||
, but the minimal description with reference to pricklier stems and leaves approaching
|
||
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. giganteum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="giganteum">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. giganteum</emphasis>
|
||
</taxonomicName>
|
||
, however, suggest this varietal name refers to the taxon we recognise as
|
||
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. poka" order="Solanales" pageId="0" pageNumber="1" rank="species" species="poka">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. poka</emphasis>
|
||
</taxonomicName>
|
||
. The protologue states no specific locality, merely "β. Java (Horsfield)". We have therefore selected the specimen of
|
||
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. poka" order="Solanales" pageId="0" pageNumber="1" rank="species" species="poka">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. poka</emphasis>
|
||
</taxonomicName>
|
||
from Java collected by Thomas Horsfield that best matches the protologue at BM (BM000886307) as the lectotype for
|
||
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. torvum subsp. var. var. polycanthum" order="Solanales" pageId="0" pageNumber="1" rank="variety" species="torvum" subSpecies="var." variety="polycanthum">S. torvum var. polycanthum</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">See Suppl. materials 1-3.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |