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<document id="A2B05E24B9AAB02D3E4F658576BD9516" ID-DOI="10.11646/zootaxa.3609.5.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10097052" ID-ZooBank="D710A4F0-F816-4ECA-9BEB-92DBED2FC58A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629839427636" checkinUser="felipe" docAuthor="TRÝZNA, MILOŠ &amp; BAŇAŘ, PETR" docDate="2013" docId="6D0987D10755FFC61B8804BA3B2BFB52" docLanguage="en" docName="zt03609p512.pdf" docOrigin="Zootaxa 3609 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.3609.5.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Apatenia stysi TRÝZNA &amp; BAŇAŘ, 2013, sp. nov." docType="treatment" docVersion="4" lastPageNumber="507" masterDocId="9130FFA90754FFC51B1F001D387AFFF2" masterDocTitle="A new species of the genus Apatenia (Coleoptera: Anthribidae) from Madagascar with notes on female genitalia, redescription of the female of Apatenia quadristigma Frieser and list of Madagascan species" masterLastPageNumber="512" masterPageNumber="504" pageNumber="505" updateTime="1699582484059" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="206842DDE2BB86B1186323F7F29FA9A4">A new species of the genus Apatenia (Coleoptera: Anthribidae) from Madagascar with notes on female genitalia, redescription of the female of Apatenia quadristigma Frieser and list of Madagascan species</mods:title>
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<mods:namePart id="B20A1963D70CAE08D7B61D1CF6188CA0">TRÝZNA, MILOŠ</mods:namePart>
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<mods:namePart id="414F205A662690D95EA3FEEEB96237CC">BAŇAŘ, PETR</mods:namePart>
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<mods:title id="27F52B0C076224FC15E3A2DEDE9DFD60">Zootaxa</mods:title>
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<mods:date id="A1B6ED9384CE90DF847EB47D1F513B8C">2013</mods:date>
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<mods:number id="5C23ABFA1BA83FD26203974D3A8B5E3D">2013-02-01</mods:number>
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<treatment id="6D0987D10755FFC61B8804BA3B2BFB52" ID-DOI="http://doi.org/10.5281/zenodo.5462765" ID-Zenodo-Dep="5462765" LSID="urn:lsid:plazi:treatment:6D0987D10755FFC61B8804BA3B2BFB52" httpUri="http://treatment.plazi.org/id/6D0987D10755FFC61B8804BA3B2BFB52" lastPageId="3" lastPageNumber="507" pageId="1" pageNumber="505">
<subSubSection id="ADBA654C0755FFC41B8804BA39DEFB33" box="[151,420,1191,1217]" pageId="1" pageNumber="505" type="nomenclature">
<paragraph id="E51F36C70755FFC41B8804BA39DEFB33" blockId="1.[151,468,1191,1250]" box="[151,420,1191,1217]" pageId="1" pageNumber="505">
<heading id="BE5781AB0755FFC41B8804BA39DEFB33" bold="true" box="[151,420,1191,1217]" fontSize="11" level="1" pageId="1" pageNumber="505" reason="1">
<emphasis id="D7D4EAD50755FFC41B8804BA39DEFB33" bold="true" box="[151,420,1191,1217]" pageId="1" pageNumber="505">
<taxonomicName id="22A04D440755FFC41B8804BA3945FB33" ID-CoL="F8PV" authorityName="Tryzna &amp; Banar" authorityYear="2013" box="[151,319,1191,1217]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="505" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="D7D4EAD50755FFC41B8804BA3945FB33" bold="true" box="[151,319,1191,1217]" italics="true" pageId="1" pageNumber="505">Apatenia stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCE757AE0755FFC41A5904BA39DEFB33" box="[326,420,1191,1217]" pageId="1" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ADBA654C0755FFC41B8804D739AEFB10" box="[151,468,1226,1250]" pageId="1" pageNumber="505" type="description">
<paragraph id="E51F36C70755FFC41B8804D739AEFB10" blockId="1.[151,468,1191,1250]" box="[151,468,1226,1250]" pageId="1" pageNumber="505">
(
<figureCitation id="7D9B2A420755FFC41B8004D73970FB10" box="[159,266,1226,1250]" captionStart="FIGURES 14" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[159,1430,197,1943]" captionTargetId="figure-53@2.[151,1436,190,1950]" captionTargetPageId="2" captionText="FIGURES 14. Apatenia stysi sp. nov., female holotype, 1, habitus dorsal view; 2, head and pronotum, lateral view; 3, head, dorsal view; 4, habitus lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.10097054" httpUri="https://zenodo.org/record/10097054/files/figure.png" pageId="1" pageNumber="505">Figs. 14</figureCitation>
,
<figureCitation id="7D9B2A420755FFC41A0804D739B6FB10" box="[279,460,1226,1250]" captionStart-0="FIGURES 714" captionStart-1="FIGURES 1518" captionStartId-0="5.[151,263,1952,1975]" captionStartId-1="6.[151,264,1524,1547]" captionTargetBox-0="[207,1366,216,1906]" captionTargetBox-1="[161,1433,200,1504]" captionTargetId-0="figure-71@5.[203,1371,193,1911]" captionTargetId-1="figure-173@6.[151,1436,193,1533]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 714. 7, 9, 12, Apatenia quadristigma Frieser, 1981, female allotype; 8, 1011, 1314, Apatenia stysi sp. nov., female holotype; 78, head, dorsal view; 910, right antenna; 11, abdomen, ventral view; 1213, dorsal pronotal carina; 14, pygidium, posterior view. Scale bars in mm. Lettering: mcxc—metacoxal cavity; S1S5—sternites 1 to 5, respectively." captionText-1="FIGURES 1518. Apatenia stysi sp. nov., female holotype, genitalia; 15, hemisternites, ventral view; 16, segment 8, ventral view; 17, toothed plate of right hemisternite, ventral view; 18, bursa copulatrix and spermatheca. Scale bars in mm. Lettering: ap—apodeme; bc—bursa copulatrix; dl—dorsal lobe; lr—lateral rod; mo—median oviduct; mr—median rod; S8—sternite 8; sp—spermatheca; spd—spermathecal duct; spg—spermathecal gland; T8—tergite 8; vl—ventral lobe." figureDoi-0="http://doi.org/10.5281/zenodo.10097058" figureDoi-1="http://doi.org/10.5281/zenodo.10097060" httpUri-0="https://zenodo.org/record/10097058/files/figure.png" httpUri-1="https://zenodo.org/record/10097060/files/figure.png" pageId="1" pageNumber="505">8, 1011, 1318</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="ADBA654C0755FFC41B88050E3CE3F9F0" pageId="1" pageNumber="505" type="materials_examined">
<paragraph id="E51F36C70755FFC41B88050E3963FABC" blockId="1.[151,1437,1298,2006]" pageId="1" pageNumber="505">
<materialsCitation id="55C83C9A0755FFC41B88050E3963FABC" country="Madagascar" elevation="100" location="Andranofasika town" municipality="North-west" pageId="1" pageNumber="505" specimenCount="1" stateProvince="Mahajanga" typeStatus="holotype">
<emphasis id="D7D4EAD50755FFC41B88050E3943FAD9" bold="true" box="[151,313,1298,1323]" inLineHeading="true" pageId="1" pageNumber="505" reason="1">
<typeStatus id="3A1B88650755FFC41B88050E38A8FAD9" box="[151,210,1299,1323]" pageId="1" pageNumber="505">Type</typeStatus>
locality.
</emphasis>
<collectingMunicipality id="057BACBD0755FFC41A5B050F39B8FAD8" box="[324,450,1298,1322]" pageId="1" pageNumber="505">North-west</collectingMunicipality>
<collectingCountry id="9DB776570755FFC41AD2050F3A2CFAD8" box="[461,598,1298,1322]" name="Madagascar" pageId="1" pageNumber="505">Madagascar</collectingCountry>
,
<collectingRegion id="2764F8250755FFC41979050F3B29FAD8" box="[614,851,1298,1322]" country="Madagascar" name="Mahajanga" pageId="1" pageNumber="505">Mahajanga Province</collectingRegion>
,
<location id="E07F601C0755FFC4187C050F3C31FAD8" LSID="urn:lsid:plazi:treatment:6D0987D10755FFC61B8804BA3B2BFB52:E07F601C0755FFC4187C050F3C31FAD8" box="[867,1099,1298,1322]" country="Madagascar" municipality="North-west" name="Andranofasika town" pageId="1" pageNumber="505" stateProvince="Mahajanga">Andranofasika town</location>
surroundings, approximately
<elevation id="6E8DD1F40755FFC41B88052B3963FABC" box="[151,281,1334,1358]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="505" unit="m" value="100.0">
<quantity id="22589B220755FFC41B88052B38A4FABC" box="[151,222,1334,1358]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="505" unit="m" value="100.0">100 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
<paragraph id="E51F36C70755FFC41BD605473CE3F9F0" blockId="1.[151,1437,1298,2006]" pageId="1" pageNumber="505">
<emphasis id="D7D4EAD50755FFC41BD6054739B3FA81" bold="true" box="[201,457,1370,1395]" inLineHeading="true" pageId="1" pageNumber="505" reason="1">Material examined.</emphasis>
<materialsCitation id="55C83C9A0755FFC41AF905473BC9FA64" collectingDate="2003-01-02" collectorName="Z. Mracek" country="Madagascar" location="Mahajanga" pageId="1" pageNumber="505" specimenCount="1" stateProvince="Mahajanga" typeStatus="holotype">
<typeStatus id="3A1B88650755FFC41AF905473A34FA80" box="[486,590,1370,1394]" pageId="1" pageNumber="505" type="holotype">Holotype</typeStatus>
female, labelled: N
<collectingCountry id="9DB776570755FFC4188805473C1BFA80" box="[919,1121,1370,1394]" name="Madagascar" pageId="1" pageNumber="505">MADAGASCAR</collectingCountry>
, /
<collectingRegion id="2764F8250755FFC41FB805473D5EFA80" box="[1191,1316,1370,1394]" country="Madagascar" name="Mahajanga" pageId="1" pageNumber="505">Mahajanga</collectingRegion>
env., / ANDRANOFASIKA env. /
<collectorName id="485553110755FFC41ACE05623A3CFA64" box="[465,582,1406,1431]" pageId="1" pageNumber="505">Z. Mráček</collectorName>
leg.,
<date id="911E10070755FFC4199A05633A90FA64" box="[645,746,1406,1430]" pageId="1" pageNumber="505" value="2003-01-02">
<collectingDate id="815AE9EF0755FFC4199A05633A90FA64" box="[645,746,1406,1430]" pageId="1" pageNumber="505" value="2003-01-02">2.I. 2003</collectingDate>
</date>
[p, white label] //
</materialsCitation>
<materialsCitation id="55C83C9A0755FFC418A4056338B7FA2C" collectingDate="2003-01-02" collectionCode="BSNPC" collectorName="Z. Mracek" country="Madagascar" determinerName="M. Tryzna &amp; P. Banar" location="Mahajanga" pageId="1" pageNumber="505" specimenCount="2" stateProvince="Mahajanga" typeStatus="holotype">
<typeStatus id="3A1B88650755FFC418A405633C35FA64" box="[955,1103,1406,1430]" pageId="1" pageNumber="505" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="22A04D440755FFC41F7905633D7EFA64" authorityName="Tryzna &amp; Banar" authorityYear="2013" box="[1126,1284,1406,1430]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="505" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="D7D4EAD50755FFC41F7905633D7EFA64" box="[1126,1284,1406,1430]" italics="true" pageId="1" pageNumber="505">Apatenia stysi</emphasis>
</taxonomicName>
<emphasis id="D7D4EAD50755FFC41E1305623D1FFA65" bold="true" box="[1292,1381,1407,1431]" pageId="1" pageNumber="505">
<taxonomicNameLabel id="CCE757AE0755FFC41E1305623D1FFA65" box="[1292,1381,1407,1431]" pageId="1" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
/
<determinerName id="10C3DA030755FFC41E630562389CFA48" pageId="1" pageNumber="505">M. Trýzna</determinerName>
&amp;
<determinerName id="10C3DA030755FFC41A1605BE391CFA49" box="[265,358,1443,1467]" pageId="1" pageNumber="505">P. Baňař</determinerName>
det. [p, red label] (
<collectionCode id="83B1AE020755FFC4195905BF3ADFFA48" box="[582,677,1442,1466]" pageId="1" pageNumber="505">BSNPC</collectionCode>
);
<typeStatus id="3A1B88650755FFC419A605BE3B63FA49" box="[697,793,1443,1467]" pageId="1" pageNumber="505" type="paratype">paratype</typeStatus>
female, the same locality as holotype, specimen with red label
</materialsCitation>
<materialsCitation id="55C83C9A0755FFC41BFD05DB3CECF9F0" collectionCode="MMBC" country="Madagascar" determinerName="M. Tryzna &amp; P. Banar" pageId="1" pageNumber="505" specimenCount="3" stateProvince="Mahajanga" typeStatus="paratype">
<typeStatus id="3A1B88650755FFC41BFD05DB390AFA2C" box="[226,368,1478,1502]" pageId="1" pageNumber="505" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="22A04D440755FFC41A9305DB3A55FA2C" authorityName="Tryzna &amp; Banar" authorityYear="2013" box="[396,559,1478,1502]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="505" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="D7D4EAD50755FFC41A9305DB3A55FA2C" box="[396,559,1478,1502]" italics="true" pageId="1" pageNumber="505">Apatenia stysi</emphasis>
</taxonomicName>
<emphasis id="D7D4EAD50755FFC4192505DA3AE2FA2D" bold="true" box="[570,664,1479,1503]" pageId="1" pageNumber="505">
<taxonomicNameLabel id="CCE757AE0755FFC4192505DA3AE2FA2D" box="[570,664,1479,1503]" pageId="1" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
/
<determinerName id="10C3DA030755FFC419A805DA3B48FA2C" box="[695,818,1478,1502]" pageId="1" pageNumber="505">M. Trýzna</determinerName>
&amp;
<determinerName id="10C3DA030755FFC4184105DA3BBAFA2D" box="[862,960,1479,1503]" pageId="1" pageNumber="505">P. Baňař</determinerName>
det. (
<collectionCode id="83B1AE020755FFC41F0D05DB3C15FA2C" box="[1042,1135,1478,1502]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="1" pageNumber="505">MMBC</collectionCode>
). Both specimens are dry mounted on rectangular card,
<typeStatus id="3A1B88650755FFC41AF905F73A32F9F0" box="[486,584,1514,1538]" pageId="1" pageNumber="505" type="holotype">holotype</typeStatus>
without left fore leg,
<typeStatus id="3A1B88650755FFC4182305F63BE6F9F1" box="[828,924,1515,1539]" pageId="1" pageNumber="505" type="paratype">paratype</typeStatus>
without right fore leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="ADBA654C0755FFC61BD606133903FCAE" lastPageId="3" lastPageNumber="507" pageId="1" pageNumber="505" type="description">
<paragraph id="E51F36C70755FFC41BD606133A3BF90C" blockId="1.[151,1437,1298,2006]" pageId="1" pageNumber="505">
<emphasis id="D7D4EAD50755FFC41BD606133920F9D5" bold="true" box="[201,346,1550,1575]" inLineHeading="true" pageId="1" pageNumber="505" reason="1">Description.</emphasis>
Female
<typeStatus id="3A1B88650755FFC41ADD06133A52F9D4" box="[450,552,1550,1574]" pageId="1" pageNumber="505" type="holotype">holotype</typeStatus>
; (female
<typeStatus id="3A1B88650755FFC4198906133A86F9D4" box="[662,764,1550,1574]" pageId="1" pageNumber="505" type="paratype">paratype</typeStatus>
).
<emphasis id="D7D4EAD50755FFC4180E06123BC9F9D4" box="[785,947,1551,1574]" italics="true" pageId="1" pageNumber="505">Measurements</emphasis>
(in mm): Total body length—6.32 (6.67). Head: total length—1.33 (1.47); length of rostrum—0.91 (0.93); maximum width of rostrum—1.09 (1.11); length of eye—0.42 (0.53); maximum width across eyes—1.38 (1.44); minimum distance between eyes—0.76 (0.78). Antenna: length of segments: II—0.18 (0.22), III—0.36 (0.34), IV—0.26 (0.27), V—0.27 (0.25), VI—0.24 (0.22), VII—0.23 (0.22), VIII—0.16 (0.17), IX—0.29 (0.26), X—0.22 (0.18), XI—0.31 (0.28). Pronotum: maximum length—1.71 (1.82); width at carina—2.24 (2.49); minimum width—1.42 (1.44). Elytra: maximum length—2.92 (3.07); maximum width—2.42 (2.71).
</paragraph>
<paragraph id="E51F36C70755FFC41BD607173D70F8B4" blockId="1.[151,1437,1298,2006]" pageId="1" pageNumber="505">
<emphasis id="D7D4EAD50755FFC41BD607173920F8D1" bold="true" box="[201,346,1802,1827]" pageId="1" pageNumber="505">Colouration</emphasis>
of the cuticle generally black, meso- and metasternum partially dark brown; anterior-most part of pronotum somewhat paler brownish. Scape dark brown, rest of antennae black. Cuticle semi-lustrous.
</paragraph>
<paragraph id="E51F36C70755FFC61BD6074F3A44FF05" blockId="1.[151,1437,1298,2006]" lastBlockId="3.[151,1436,151,1184]" lastPageId="3" lastPageNumber="507" pageId="1" pageNumber="505">
<emphasis id="D7D4EAD50755FFC41BD6074F394EF899" bold="true" box="[201,308,1874,1899]" pageId="1" pageNumber="505">Vestiture</emphasis>
on head, pronotum and distal third of elytra light brown, setae longer and wider on anterior half of pronotum, giving pale brown colouration to this part (
<figureCitation id="7D9B2A420755FFC419EA076B3B18F87C" box="[757,866,1910,1934]" captionStart="FIGURES 14" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[159,1430,197,1943]" captionTargetId="figure-53@2.[151,1436,190,1950]" captionTargetPageId="2" captionText="FIGURES 14. Apatenia stysi sp. nov., female holotype, 1, habitus dorsal view; 2, head and pronotum, lateral view; 3, head, dorsal view; 4, habitus lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.10097054" httpUri="https://zenodo.org/record/10097054/files/figure.png" pageId="1" pageNumber="505">Figs. 12</figureCitation>
). Distal part of elytra similar, with setae forming a pale brown transverse strip. Proximal two thirds of elytra with blackish vestiture, except for four characteristic transverse spots, formed by long, wide, scale-like whitish setae (
<figureCitation id="7D9B2A420755FFC4187707A33BAFF824" box="[872,981,1982,2006]" captionStart="FIGURES 14" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[159,1430,197,1943]" captionTargetId="figure-53@2.[151,1436,190,1950]" captionTargetPageId="2" captionText="FIGURES 14. Apatenia stysi sp. nov., female holotype, 1, habitus dorsal view; 2, head and pronotum, lateral view; 3, head, dorsal view; 4, habitus lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.10097054" httpUri="https://zenodo.org/record/10097054/files/figure.png" pageId="1" pageNumber="505">Figs. 1, 4</figureCitation>
). Vestiture on antennae short, appressed, blackish. All legs covered with dense black setae, dorsal face of all femora, tibiae and tarsi with narrow, contrasting strips comprised of white setae (these narrow strips occur also on meso- and meta-sternum). Prosternum and venter of abdomen with whitish pubescence.
</paragraph>
<caption id="B1DF664F0756FFC71B8807AE3AC1F81A" ID-DOI="http://doi.org/10.5281/zenodo.10097054" ID-Zenodo-Dep="10097054" httpUri="https://zenodo.org/record/10097054/files/figure.png" pageId="2" pageNumber="506" startId="2.[151,264,1971,1994]" targetBox="[159,1430,197,1943]" targetPageId="2">
<paragraph id="E51F36C70756FFC71B8807AE3AC1F81A" blockId="2.[151,1436,1971,2024]" pageId="2" pageNumber="506">
<emphasis id="D7D4EAD50756FFC71B8807AE3941F838" bold="true" box="[151,315,1971,1994]" pageId="2" pageNumber="506">FIGURES 14.</emphasis>
<taxonomicName id="22A04D440756FFC71A5D07A939A8F83B" authorityName="Tryzna &amp; Banar" authorityYear="2013" box="[322,466,1972,1994]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="D7D4EAD50756FFC71A5D07A939A8F83B" box="[322,466,1972,1994]" italics="true" pageId="2" pageNumber="506">Apatenia stysi</emphasis>
</taxonomicName>
<emphasis id="D7D4EAD50756FFC71AC607A93A50F83B" bold="true" box="[473,554,1972,1993]" pageId="2" pageNumber="506">
<taxonomicNameLabel id="CCE757AE0756FFC71AC607A93A50F83B" box="[473,554,1972,1993]" pageId="2" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female holotype, 1, habitus dorsal view; 2, head and pronotum, lateral view; 3, head, dorsal view; 4, habitus lateral view. Scale bars in mm.
</paragraph>
</caption>
<paragraph id="E51F36C70757FFC61BD6011E3C27FE75" blockId="3.[151,1436,151,1184]" pageId="3" pageNumber="507">
<emphasis id="D7D4EAD50757FFC61BD6011E393BFEEE" bold="true" box="[201,321,259,284]" inLineHeading="true" pageId="3" pageNumber="507" reason="1">Structure.</emphasis>
Head long and narrow (
<figureCitation id="7D9B2A420757FFC6197301193ABAFEEE" box="[620,704,260,284]" captionStart="FIGURES 14" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[159,1430,197,1943]" captionTargetId="figure-53@2.[151,1436,190,1950]" captionTargetPageId="2" captionText="FIGURES 14. Apatenia stysi sp. nov., female holotype, 1, habitus dorsal view; 2, head and pronotum, lateral view; 3, head, dorsal view; 4, habitus lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.10097054" httpUri="https://zenodo.org/record/10097054/files/figure.png" pageId="3" pageNumber="507">Figs. 3</figureCitation>
,
<figureCitation id="7D9B2A420757FFC619CD01193A9BFEEE" box="[722,737,260,284]" captionStart="FIGURES 714" captionStartId="5.[151,263,1952,1975]" captionTargetBox="[207,1366,216,1906]" captionTargetId="figure-71@5.[203,1371,193,1911]" captionTargetPageId="5" captionText="FIGURES 714. 7, 9, 12, Apatenia quadristigma Frieser, 1981, female allotype; 8, 1011, 1314, Apatenia stysi sp. nov., female holotype; 78, head, dorsal view; 910, right antenna; 11, abdomen, ventral view; 1213, dorsal pronotal carina; 14, pygidium, posterior view. Scale bars in mm. Lettering: mcxc—metacoxal cavity; S1S5—sternites 1 to 5, respectively." figureDoi="http://doi.org/10.5281/zenodo.10097058" httpUri="https://zenodo.org/record/10097058/files/figure.png" pageId="3" pageNumber="507">8</figureCitation>
), rostrum flat, simple, without longitudinal carina, ratio of rostrum length to maximum width 0.83. Scrobes not visible dorsally, scape partly hidden in scrobe in dorsal view. Eyes small, ocular index 2.45. Ratio of maximum width across eyes to maximum width of rostrum 1.27. Antennae (
<figureCitation id="7D9B2A420757FFC61B8001723897FE75" box="[159,237,367,391]" captionStart="FIGURES 14" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[159,1430,197,1943]" captionTargetId="figure-53@2.[151,1436,190,1950]" captionTargetPageId="2" captionText="FIGURES 14. Apatenia stysi sp. nov., female holotype, 1, habitus dorsal view; 2, head and pronotum, lateral view; 3, head, dorsal view; 4, habitus lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.10097054" httpUri="https://zenodo.org/record/10097054/files/figure.png" pageId="3" pageNumber="507">Figs. 1</figureCitation>
,
<figureCitation id="7D9B2A420757FFC61BE50172396DFE75" box="[250,279,367,391]" captionStart="FIGURES 714" captionStartId="5.[151,263,1952,1975]" captionTargetBox="[207,1366,216,1906]" captionTargetId="figure-71@5.[203,1371,193,1911]" captionTargetPageId="5" captionText="FIGURES 714. 7, 9, 12, Apatenia quadristigma Frieser, 1981, female allotype; 8, 1011, 1314, Apatenia stysi sp. nov., female holotype; 78, head, dorsal view; 910, right antenna; 11, abdomen, ventral view; 1213, dorsal pronotal carina; 14, pygidium, posterior view. Scale bars in mm. Lettering: mcxc—metacoxal cavity; S1S5—sternites 1 to 5, respectively." figureDoi="http://doi.org/10.5281/zenodo.10097058" httpUri="https://zenodo.org/record/10097058/files/figure.png" pageId="3" pageNumber="507">10</figureCitation>
) sub-equal in length to head and pronotum together. Funicle and club thin.
</paragraph>
<paragraph id="E51F36C70757FFC61BD601893A11FE06" blockId="3.[151,1436,151,1184]" pageId="3" pageNumber="507">
Pronotum transverse, ratio of length to width at carina 0.76, gradually narrowed anteriorly, widest at dorsal transverse carina. Dorsal transverse carina bisinuate (
<figureCitation id="7D9B2A420757FFC619F301AA3B45FE3D" box="[748,831,439,463]" captionStart="FIGURES 714" captionStartId="5.[151,263,1952,1975]" captionTargetBox="[207,1366,216,1906]" captionTargetId="figure-71@5.[203,1371,193,1911]" captionTargetPageId="5" captionText="FIGURES 714. 7, 9, 12, Apatenia quadristigma Frieser, 1981, female allotype; 8, 1011, 1314, Apatenia stysi sp. nov., female holotype; 78, head, dorsal view; 910, right antenna; 11, abdomen, ventral view; 1213, dorsal pronotal carina; 14, pygidium, posterior view. Scale bars in mm. Lettering: mcxc—metacoxal cavity; S1S5—sternites 1 to 5, respectively." figureDoi="http://doi.org/10.5281/zenodo.10097058" httpUri="https://zenodo.org/record/10097058/files/figure.png" pageId="3" pageNumber="507">Fig. 13</figureCitation>
), conspicuously convex in the middle, lateral margins in obtuse arch directed anteriorly (
<figureCitation id="7D9B2A420757FFC6190701C13A27FE06" box="[536,605,476,500]" captionStart="FIGURES 14" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[159,1430,197,1943]" captionTargetId="figure-53@2.[151,1436,190,1950]" captionTargetPageId="2" captionText="FIGURES 14. Apatenia stysi sp. nov., female holotype, 1, habitus dorsal view; 2, head and pronotum, lateral view; 3, head, dorsal view; 4, habitus lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.10097054" httpUri="https://zenodo.org/record/10097054/files/figure.png" pageId="3" pageNumber="507">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="E51F36C70757FFC61BD601E23AACFDCE" blockId="3.[151,1436,151,1184]" pageId="3" pageNumber="507">
Elytra broadly oval (
<figureCitation id="7D9B2A420757FFC61AAC01E2398CFDE5" box="[435,502,511,535]" captionStart="FIGURES 14" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[159,1430,197,1943]" captionTargetId="figure-53@2.[151,1436,190,1950]" captionTargetPageId="2" captionText="FIGURES 14. Apatenia stysi sp. nov., female holotype, 1, habitus dorsal view; 2, head and pronotum, lateral view; 3, head, dorsal view; 4, habitus lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.10097054" httpUri="https://zenodo.org/record/10097054/files/figure.png" pageId="3" pageNumber="507">Fig. 1</figureCitation>
), conspicuously narrowed posteriorly from two thirds of their length. Ratio of the maximum length of elytra to maximum width 1.20.
</paragraph>
<paragraph id="E51F36C70757FFC61BD6025A38B5FD76" blockId="3.[151,1436,151,1184]" pageId="3" pageNumber="507">
Abdomen shorter than broad (
<figureCitation id="7D9B2A420757FFC61907025A3A12FDAD" box="[536,616,583,607]" captionStart="FIGURES 714" captionStartId="5.[151,263,1952,1975]" captionTargetBox="[207,1366,216,1906]" captionTargetId="figure-71@5.[203,1371,193,1911]" captionTargetPageId="5" captionText="FIGURES 714. 7, 9, 12, Apatenia quadristigma Frieser, 1981, female allotype; 8, 1011, 1314, Apatenia stysi sp. nov., female holotype; 78, head, dorsal view; 910, right antenna; 11, abdomen, ventral view; 1213, dorsal pronotal carina; 14, pygidium, posterior view. Scale bars in mm. Lettering: mcxc—metacoxal cavity; S1S5—sternites 1 to 5, respectively." figureDoi="http://doi.org/10.5281/zenodo.10097058" httpUri="https://zenodo.org/record/10097058/files/figure.png" pageId="3" pageNumber="507">Fig. 11</figureCitation>
), pygidium (
<figureCitation id="7D9B2A420757FFC619E7025A3B33FDAD" box="[760,841,583,607]" captionStart="FIGURES 714" captionStartId="5.[151,263,1952,1975]" captionTargetBox="[207,1366,216,1906]" captionTargetId="figure-71@5.[203,1371,193,1911]" captionTargetPageId="5" captionText="FIGURES 714. 7, 9, 12, Apatenia quadristigma Frieser, 1981, female allotype; 8, 1011, 1314, Apatenia stysi sp. nov., female holotype; 78, head, dorsal view; 910, right antenna; 11, abdomen, ventral view; 1213, dorsal pronotal carina; 14, pygidium, posterior view. Scale bars in mm. Lettering: mcxc—metacoxal cavity; S1S5—sternites 1 to 5, respectively." figureDoi="http://doi.org/10.5281/zenodo.10097058" httpUri="https://zenodo.org/record/10097058/files/figure.png" pageId="3" pageNumber="507">Fig. 14</figureCitation>
) robust, ratio of maximum width to maximum length 1.21.
</paragraph>
<paragraph id="E51F36C70757FFC61BD602923A36FCC5" blockId="3.[151,1436,151,1184]" pageId="3" pageNumber="507">
Female genitalia. Sternite of segment 8 with long apodeme (
<figureCitation id="7D9B2A420757FFC6186F02923BB9FD55" box="[880,963,655,679]" captionStart="FIGURES 1518" captionStartId="6.[151,264,1524,1547]" captionTargetBox="[161,1433,200,1504]" captionTargetId="figure-173@6.[151,1436,193,1533]" captionTargetPageId="6" captionText="FIGURES 1518. Apatenia stysi sp. nov., female holotype, genitalia; 15, hemisternites, ventral view; 16, segment 8, ventral view; 17, toothed plate of right hemisternite, ventral view; 18, bursa copulatrix and spermatheca. Scale bars in mm. Lettering: ap—apodeme; bc—bursa copulatrix; dl—dorsal lobe; lr—lateral rod; mo—median oviduct; mr—median rod; S8—sternite 8; sp—spermatheca; spd—spermathecal duct; spg—spermathecal gland; T8—tergite 8; vl—ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10097060" httpUri="https://zenodo.org/record/10097060/files/figure.png" pageId="3" pageNumber="507">Fig. 16</figureCitation>
), slightly widened apically. Hemisternites wide and robust (
<figureCitation id="7D9B2A420757FFC61A4602A939D1FD3E" box="[345,427,692,716]" captionStart="FIGURES 1518" captionStartId="6.[151,264,1524,1547]" captionTargetBox="[161,1433,200,1504]" captionTargetId="figure-173@6.[151,1436,193,1533]" captionTargetPageId="6" captionText="FIGURES 1518. Apatenia stysi sp. nov., female holotype, genitalia; 15, hemisternites, ventral view; 16, segment 8, ventral view; 17, toothed plate of right hemisternite, ventral view; 18, bursa copulatrix and spermatheca. Scale bars in mm. Lettering: ap—apodeme; bc—bursa copulatrix; dl—dorsal lobe; lr—lateral rod; mo—median oviduct; mr—median rod; S8—sternite 8; sp—spermatheca; spd—spermathecal duct; spg—spermathecal gland; T8—tergite 8; vl—ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10097060" httpUri="https://zenodo.org/record/10097060/files/figure.png" pageId="3" pageNumber="507">Fig. 15</figureCitation>
), apex of each with robust, strongly sclerotized toothed plate (
<figureCitation id="7D9B2A420757FFC61F7902A93C95FD3E" box="[1126,1263,692,716]" captionStart="FIGURES 1518" captionStartId="6.[151,264,1524,1547]" captionTargetBox="[161,1433,200,1504]" captionTargetId="figure-173@6.[151,1436,193,1533]" captionTargetPageId="6" captionText="FIGURES 1518. Apatenia stysi sp. nov., female holotype, genitalia; 15, hemisternites, ventral view; 16, segment 8, ventral view; 17, toothed plate of right hemisternite, ventral view; 18, bursa copulatrix and spermatheca. Scale bars in mm. Lettering: ap—apodeme; bc—bursa copulatrix; dl—dorsal lobe; lr—lateral rod; mo—median oviduct; mr—median rod; S8—sternite 8; sp—spermatheca; spd—spermathecal duct; spg—spermathecal gland; T8—tergite 8; vl—ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10097060" httpUri="https://zenodo.org/record/10097060/files/figure.png" pageId="3" pageNumber="507">Figs. 15, 17</figureCitation>
). Toothed plate without (or with extremely reduced?) stylus, with group of 56 strong setae on base. Bursa copulatrix simple (
<figureCitation id="7D9B2A420757FFC61E7102CA38CEFCE6" captionStart="FIGURES 1518" captionStartId="6.[151,264,1524,1547]" captionTargetBox="[161,1433,200,1504]" captionTargetId="figure-173@6.[151,1436,193,1533]" captionTargetPageId="6" captionText="FIGURES 1518. Apatenia stysi sp. nov., female holotype, genitalia; 15, hemisternites, ventral view; 16, segment 8, ventral view; 17, toothed plate of right hemisternite, ventral view; 18, bursa copulatrix and spermatheca. Scale bars in mm. Lettering: ap—apodeme; bc—bursa copulatrix; dl—dorsal lobe; lr—lateral rod; mo—median oviduct; mr—median rod; S8—sternite 8; sp—spermatheca; spd—spermathecal duct; spg—spermathecal gland; T8—tergite 8; vl—ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10097060" httpUri="https://zenodo.org/record/10097060/files/figure.png" pageId="3" pageNumber="507">Fig. 18</figureCitation>
) with small sclerites at insertion of spermathecal duct. Spermatheca strongly sclerotized, sickle-shaped, spermathecal gland spherical (
<figureCitation id="7D9B2A420757FFC61AF403023A44FCC5" box="[491,574,799,823]" captionStart="FIGURES 1518" captionStartId="6.[151,264,1524,1547]" captionTargetBox="[161,1433,200,1504]" captionTargetId="figure-173@6.[151,1436,193,1533]" captionTargetPageId="6" captionText="FIGURES 1518. Apatenia stysi sp. nov., female holotype, genitalia; 15, hemisternites, ventral view; 16, segment 8, ventral view; 17, toothed plate of right hemisternite, ventral view; 18, bursa copulatrix and spermatheca. Scale bars in mm. Lettering: ap—apodeme; bc—bursa copulatrix; dl—dorsal lobe; lr—lateral rod; mo—median oviduct; mr—median rod; S8—sternite 8; sp—spermatheca; spd—spermathecal duct; spg—spermathecal gland; T8—tergite 8; vl—ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10097060" httpUri="https://zenodo.org/record/10097060/files/figure.png" pageId="3" pageNumber="507">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="E51F36C70757FFC61BD603593903FCAE" blockId="3.[151,1436,151,1184]" box="[201,377,836,860]" pageId="3" pageNumber="507">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="ADBA654C0757FFC61BD603753A6FFC56" pageId="3" pageNumber="507" type="etymology">
<paragraph id="E51F36C70757FFC61BD603753A6FFC56" blockId="3.[151,1436,151,1184]" pageId="3" pageNumber="507">
<emphasis id="D7D4EAD50757FFC61BD603753935FC72" bold="true" box="[201,335,872,896]" inLineHeading="true" pageId="3" pageNumber="507" reason="1">Etymology.</emphasis>
Patronym, dedicated to our teacher and friend, Professor Pavel Štys, eminent scholar of many groups of true bugs (Heteroptera).
</paragraph>
</subSubSection>
<subSubSection id="ADBA654C0757FFC61BD603B23A9FFC3A" box="[201,741,943,968]" pageId="3" pageNumber="507" type="distribution">
<paragraph id="E51F36C70757FFC61BD603B23A9FFC3A" blockId="3.[151,1436,151,1184]" box="[201,741,943,968]" pageId="3" pageNumber="507">
<emphasis id="D7D4EAD50757FFC61BD603B23918FC3A" bold="true" box="[201,354,943,968]" inLineHeading="true" pageId="3" pageNumber="507" reason="1">Distribution.</emphasis>
North-west
<collectingCountry id="9DB776570757FFC61AF103B23A0FFC35" box="[494,629,943,967]" name="Madagascar" pageId="3" pageNumber="507">Madagascar</collectingCountry>
(
<figureCitation id="7D9B2A420757FFC6199B03B23AACFC35" box="[644,726,943,967]" captionStart="FIGURE 19" captionStartId="7.[151,250,2002,2025]" captionTargetBox="[299,1302,195,1959]" captionTargetId="figure-35@7.[299,1316,195,1959]" captionTargetPageId="7" captionText="FIGURE 19. Distribution of Apatenia stysi sp. nov. and A. quadristigma Frieser, 1981 in Madagascar." figureDoi="http://doi.org/10.5281/zenodo.10097062" httpUri="https://zenodo.org/record/10097062/files/figure.png" pageId="3" pageNumber="507">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="ADBA654C0757FFC61BD603CE3B2BFB52" pageId="3" pageNumber="507" type="diagnosis">
<paragraph id="E51F36C70757FFC61BD603CE3B2BFB52" blockId="3.[151,1436,151,1184]" pageId="3" pageNumber="507">
<emphasis id="D7D4EAD50757FFC61BD603CE39ABFC1E" bold="true" box="[201,465,979,1004]" inLineHeading="true" pageId="3" pageNumber="507" reason="1">Differential diagnosis.</emphasis>
Female
<taxonomicName id="22A04D440757FFC6192B03C93AAEFC1E" authorityName="Tryzna &amp; Banar" authorityYear="2013" box="[564,724,980,1004]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="D7D4EAD50757FFC6192B03C93AAEFC1E" box="[564,724,980,1004]" italics="true" pageId="3" pageNumber="507">Apatenia stysi</emphasis>
</taxonomicName>
<emphasis id="D7D4EAD50757FFC619C303C93B4DFC1E" bold="true" box="[732,823,980,1004]" pageId="3" pageNumber="507">
<taxonomicNameLabel id="CCE757AE0757FFC619C303C93B4DFC1E" box="[732,823,980,1004]" pageId="3" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from female
<taxonomicName id="22A04D440757FFC61F3B03C93D0DFC1E" authority="Frieser, 1981" authorityName="Frieser" authorityYear="1981" box="[1060,1399,980,1004]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="quadristigma">
<emphasis id="D7D4EAD50757FFC61F3B03C93CA0FC1E" box="[1060,1242,980,1004]" italics="true" pageId="3" pageNumber="507">A. quadristigma</emphasis>
<bibRefCitation id="81314B360757FFC61FFC03C93D0DFC1E" author="Frieser" box="[1251,1399,980,1004]" firstAuthor="Frieser" pageId="3" pageNumber="507" pagination="249 - 258" refId="ref3433" refString="Frieser, R. (1981) Beitrag zur Kenntnis der Anthribiden (Coleoptera, Anthribidae). Entomologische Arbeiten aus dem Museum G. Frey, 29, 249 - 258." type="journal article" year="1981">Frieser, 1981</bibRefCitation>
</taxonomicName>
by larger body length (more than
<quantity id="22589B220757FFC61AE703E53A44FBFD" box="[504,574,1016,1040]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="507" unit="mm" value="6.0">6 mm</quantity>
;
<quantity id="22589B220757FFC6195303E53AC8FBFD" box="[588,690,1016,1040]" metricMagnitude="-3" metricUnit="m" metricValue="4.33" pageId="3" pageNumber="507" unit="mm" value="4.33">4.33 mm</quantity>
in
<typeStatus id="3A1B88650757FFC619C403E53B4FFBE2" box="[731,821,1016,1040]" pageId="3" pageNumber="507" type="allotype">allotype</typeStatus>
of
<taxonomicName id="22A04D440757FFC6184003E53C6CFBE2" box="[863,1046,1016,1040]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="quadristigma">
<emphasis id="D7D4EAD50757FFC6184003E53C6CFBE2" box="[863,1046,1016,1040]" italics="true" pageId="3" pageNumber="507">A. quadristigma</emphasis>
</taxonomicName>
); much smaller eye (ocular index 2.45;
<quantity id="22589B220757FFC61BC604013953FBC6" box="[217,297,1052,1076]" metricMagnitude="-2" metricUnit="m" metricValue="1.4986" pageId="3" pageNumber="507" unit="in" value="0.59">0.59 in</quantity>
<taxonomicName id="22A04D440757FFC61A2E0401399DFBC6" box="[305,487,1052,1076]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="quadristigma">
<emphasis id="D7D4EAD50757FFC61A2E0401399DFBC6" box="[305,487,1052,1076]" italics="true" pageId="3" pageNumber="507">A. quadristigma</emphasis>
</taxonomicName>
); shape of dorsal pronotal carina (
<figureCitation id="7D9B2A420757FFC6186B04013B86FBC6" box="[884,1020,1052,1076]" captionStart="FIGURES 714" captionStartId="5.[151,263,1952,1975]" captionTargetBox="[207,1366,216,1906]" captionTargetId="figure-71@5.[203,1371,193,1911]" captionTargetPageId="5" captionText="FIGURES 714. 7, 9, 12, Apatenia quadristigma Frieser, 1981, female allotype; 8, 1011, 1314, Apatenia stysi sp. nov., female holotype; 78, head, dorsal view; 910, right antenna; 11, abdomen, ventral view; 1213, dorsal pronotal carina; 14, pygidium, posterior view. Scale bars in mm. Lettering: mcxc—metacoxal cavity; S1S5—sternites 1 to 5, respectively." figureDoi="http://doi.org/10.5281/zenodo.10097058" httpUri="https://zenodo.org/record/10097058/files/figure.png" pageId="3" pageNumber="507">Figs. 1213</figureCitation>
) and different colour patterns of the dorsum of body (
<figureCitation id="7D9B2A420757FFC61A44045D39E7FBAA" box="[347,413,1088,1112]" captionStart="FIGURES 14" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[159,1430,197,1943]" captionTargetId="figure-53@2.[151,1436,190,1950]" captionTargetPageId="2" captionText="FIGURES 14. Apatenia stysi sp. nov., female holotype, 1, habitus dorsal view; 2, head and pronotum, lateral view; 3, head, dorsal view; 4, habitus lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.10097054" httpUri="https://zenodo.org/record/10097054/files/figure.png" pageId="3" pageNumber="507">Fig. 1</figureCitation>
).
<taxonomicName id="22A04D440757FFC61AA8045D3A2CFBA5" authorityName="Tryzna &amp; Banar" authorityYear="2013" box="[439,598,1088,1112]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="D7D4EAD50757FFC61AA8045D3A2CFBA5" box="[439,598,1088,1112]" italics="true" pageId="3" pageNumber="507">Apatenia stysi</emphasis>
</taxonomicName>
<emphasis id="D7D4EAD50757FFC61941045D3AC3FBAA" bold="true" box="[606,697,1088,1112]" pageId="3" pageNumber="507">
<taxonomicNameLabel id="CCE757AE0757FFC61941045D3AC3FBAA" box="[606,697,1088,1112]" pageId="3" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all other Madagascan species of the genus
<taxonomicName id="22A04D440757FFC61E28045D3DE6FBAA" authorityName="Pascoe" authorityYear="1859" box="[1335,1436,1088,1112]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="genus">
<emphasis id="D7D4EAD50757FFC61E28045D3DE6FBAA" box="[1335,1436,1088,1112]" italics="true" pageId="3" pageNumber="507">Apatenia</emphasis>
</taxonomicName>
(with an exception of
<taxonomicName id="22A04D440757FFC61AAB04793A08FB8E" box="[436,626,1124,1148]" class="Insecta" family="Anthribidae" genus="Apatenia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="quadristigma">
<emphasis id="D7D4EAD50757FFC61AAB04793A08FB8E" box="[436,626,1124,1148]" italics="true" pageId="3" pageNumber="507">A. quadristigma</emphasis>
</taxonomicName>
) by its totally different colour pattern composed mainly of four characteristics transverse white spots on the elytra (
<figureCitation id="7D9B2A420757FFC619C804953B39FB52" box="[727,835,1160,1184]" captionStart="FIGURES 14" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[159,1430,197,1943]" captionTargetId="figure-53@2.[151,1436,190,1950]" captionTargetPageId="2" captionText="FIGURES 14. Apatenia stysi sp. nov., female holotype, 1, habitus dorsal view; 2, head and pronotum, lateral view; 3, head, dorsal view; 4, habitus lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.10097054" httpUri="https://zenodo.org/record/10097054/files/figure.png" pageId="3" pageNumber="507">Figs. 1, 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>