358 lines
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358 lines
42 KiB
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<document id="57B86FD0F9904E65758A62B43E1005EC" ID-DOI="10.1093/zoolinnean/zlad030" ID-ISSN="0024-4082" ID-Zenodo-Dep="10469776" ID-ZooBank="995541BE-533A-479B-8ED8-955F5EF30A0E" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1704716319055" checkinUser="plazi" docAuthor="Oliver, Adriana, Carro-Rodríguez, Patricia M., López-Guerrero, Paloma & Daxner-Höck, Gudrun" docDate="2023" docId="6B7C083CCA12893DFED6F8CCF581F9F7" docLanguage="en" docName="zlad030.pdf" docOrigin="Zoological Journal of the Linnean Society 199 (3)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad030" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Prodistylomys mongoliensis Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck 2023, sp.nov." docType="treatment" docVersion="3" lastPageNumber="643" masterDocId="97457044CA1A8937FFC2FFD7F57AFFFF" masterDocTitle="A new framework of the evolution of the ctenodactylids (Mammalia: Rodentia) in Asia: new species and phylogenetic status of distylomyins" masterLastPageNumber="655" masterPageNumber="633" pageNumber="641" updateTime="1705061696857" updateUser="jonas">
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<mods:title id="C8736B14A775267DE75780EA42D85784">A new framework of the evolution of the ctenodactylids (Mammalia: Rodentia) in Asia: new species and phylogenetic status of distylomyins</mods:title>
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<mods:namePart id="EC82BB5592BC4A24A1BEBE01F073AC25">Oliver, Adriana</mods:namePart>
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<mods:affiliation id="2C16F41EA64A504CB68270F17937486F">Department of Paleobiology, National Museum of Natural Sciences-Spanish National Research Council (MNCN-CSIC), C / José Gutiérrez Abascal 2, 28006 Madrid, Spain & Association Mujeres con los Pies en la Tierra, C / Jose Antonio Novais, 12, 28040 Madrid, Spain</mods:affiliation>
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<mods:namePart id="065A9BA0031CC0D2B4490F340E5256FD">Carro-Rodríguez, Patricia M.</mods:namePart>
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<mods:affiliation id="4F7ACDADC29867F6E0D70C044C334915">Association Mujeres con los Pies en la Tierra, C / Jose Antonio Novais, 12, 28040 Madrid, Spain & Department of Geodynamics, Stratigraphy and Palaeontology (GEODESPAL), Universidad Complutense de Madrid, C / Jose Antonio Novais, 12, 28040, Madrid, Spain & Sedimentary Geology and Environmental Change, Geosciences Institute (CSIC, UCM), C / del Dr Severo Ochoa, 7, 28040, Madrid, Spain</mods:affiliation>
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<mods:affiliation id="8897144D34553216FB1E1BD78B510602">Geological-Paleontological Department, Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<taxonomicName id="24D5C2A9CA12893FFED6F8CCF75FF8C9" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[276,549,1819,1846]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="species" species="mongoliensis" status="sp. nov.">
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<emphasis id="D1A16538CA12893FFED6F8CCF75FF8C9" bold="true" box="[276,549,1819,1846]" italics="true" pageId="8" pageNumber="641">Prodistylomys mongoliensis</emphasis>
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<paragraph id="E36AB92ACA12893FFF43F895F745F8A5" blockId="8.[129,575,1858,1882]" box="[129,575,1858,1882]" pageId="8" pageNumber="641">
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<figureCitation id="7BEEA5AFCA12893FFF43F895F5A2F8A5" box="[129,216,1858,1882]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStartId-0="9.[113,178,1607,1631]" captionStartId-1="10.[129,194,771,795]" captionTargetBox-0="[119,1449,1129,1569]" captionTargetBox-1="[141,1469,154,741]" captionTargetId-0="figure-192@9.[114,1458,1123,1579]" captionTargetId-1="figure-523@10.[129,1473,144,744]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figure 6. Prodistylomys mongoliensis sp. nov. from the type locality of Hotuliin Teeg, Valley of Lakes, Mongolia:(A) holotype of Prodistylomys mongoliensis sp. nov. from Hotuliin Teeg (HTE-012), (NHMW 2012/0051/0001). Right m2; (B) HTE-012 (NHMW 2012/0051/0002) M3 left. Left-side specimens are underlined. Scale bar: 1 mm." captionText-1="Figure 7. Upper and lower cheek teeth of Prodistylomys mongoliensis sp. nov. from the Valley of Lakes (Mongolia). From Hotuliin Teeg: (A) HTE-003 (NHMW 2012/0052/0002) M1/2 right; (B) HTE-003 (NHMW 2012/0052/0001) m2 left. All teeth are at the same magnification.Right-side specimens are underlined. Scale bar: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10469788" figureDoi-1="http://doi.org/10.5281/zenodo.10469790" httpUri-0="https://zenodo.org/record/10469788/files/figure.png" httpUri-1="https://zenodo.org/record/10469790/files/figure.png" pageId="8" pageNumber="641">Figs. 6-7</figureCitation>
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, Supporting Information Model S3
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</paragraph>
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<paragraph id="E36AB92ACA12893FFF43F8ACF48EF84D" blockId="8.[129,777,1915,2002]" pageId="8" pageNumber="641">
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<emphasis id="D1A16538CA12893FFF43F8ACF41AF86C" box="[129,352,1915,1939]" italics="true" pageId="8" pageNumber="641">Zoobank registration:</emphasis>
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urn:lsid:zoobank.org:pub:
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.
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<paragraph id="E36AB92ACA12893FFF5EF86DF7CEF82D" blockId="8.[129,777,1915,2002]" box="[156,692,1978,2002]" pageId="8" pageNumber="641">
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<emphasis id="D1A16538CA12893FFF5EF86DF46BF82D" box="[156,273,1978,2002]" italics="true" pageId="8" pageNumber="641">
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<taxonomicName id="24D5C2A9CA12893FFF5EF86DF47CF82D" box="[156,262,1978,2002]" class="Mammalia" family="Ctenodactylidae" genus="Distylomys" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="species" species="undetermined">Distylomys</taxonomicName>
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/
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<taxonomicName id="24D5C2A9CA12893FFED0F86DF4C1F82E" box="[274,443,1978,2002]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="D1A16538CA12893FFED0F86DF4E1F82D" box="[274,411,1978,2002]" italics="true" pageId="8" pageNumber="641">Prodistylomys</emphasis>
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sp.
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</taxonomicName>
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(
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<treatmentCitation id="62749F3BCA12893FFE05F86DF7D2F82D" author="Hock V & Daxner-Hock G & Schmid HP" box="[455,680,1978,2002]" page="118" pageId="8" pageNumber="641" year="1999">
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<bibRefCitation id="8744C4DBCA12893FFE05F86DF7D2F82D" author="Hock V & Daxner-Hock G & Schmid HP" box="[455,680,1978,2002]" pageId="8" pageNumber="641" pagination="83 - 125" refId="ref16320" refString="Hock V, Daxner-Hock G, Schmid HP et al. Oligocene-Miocene sediments, fossils and basalts from the Valley of Lakes (Central Mongolia) - an integrated study. Mitteilungen der Oesterreichischen Geologischen Gesellschass 1999; 90: 83 - 125." type="journal article" year="1999">
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Höck
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<emphasis id="D1A16538CA12893FFDCAF86DF74AF82D" box="[520,560,1978,2002]" italics="true" pageId="8" pageNumber="641">et al</emphasis>
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. 1999: 118
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).
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</paragraph>
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<paragraph id="E36AB92ACA12893FFC97FA85F0C7FA95" blockId="8.[825,1475,1362,1480]" box="[853,1469,1362,1386]" pageId="8" pageNumber="641">
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<taxonomicName id="24D5C2A9CA12893FFC97FA85F69BFA95" box="[853,993,1362,1386]" class="Mammalia" family="Ctenodactylidae" genus="Distylomys" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="D1A16538CA12893FFC97FA85F6BAFA95" box="[853,960,1362,1386]" italics="true" pageId="8" pageNumber="641">Distylomys</emphasis>
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sp.
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</taxonomicName>
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(
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<treatmentCitation id="62749F3BCA12893FFC2EFA85F0F3FA95" author="Daxner-Hock G & Badamgarav D" box="[1004,1417,1362,1386]" page="16" pageId="8" pageNumber="641" year="2007">
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<bibRefCitation id="8744C4DBCA12893FFC2EFA85F0F3FA95" author="Daxner-Hock G & Badamgarav D" box="[1004,1417,1362,1386]" pageId="8" pageNumber="641" pagination="1 - 24" refId="ref15459" refString="Daxner-Hock G, Badamgarav D. Oligocene-Miocene vertebrates from the Valley of Lakes (Central Mongolia): morphology, phylogenetic and stratigraphic implications - 1. Geological and stratigraphic seuing. Annalen des Naturhistorischen Museums in Wien 201 2007; 108 A: 1 - 24." type="journal article" year="2007">Daxner-Höck and Badamgarav 2007: 16</bibRefCitation>
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, 18).
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<paragraph id="E36AB92ACA12893FFC97FAA6F02FFA76" blockId="8.[825,1475,1362,1480]" box="[853,1365,1393,1418]" pageId="8" pageNumber="641">
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<treatmentCitationGroup id="C3C59E04CA12893FFC97FAA6F02FFA76" box="[853,1365,1393,1418]" pageId="8" pageNumber="641">
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<taxonomicName id="24D5C2A9CA12893FFC97FAA6F6A6FA76" box="[853,988,1393,1417]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="D1A16538CA12893FFC97FAA6F6A6FA76" box="[853,988,1393,1417]" italics="true" pageId="8" pageNumber="641">Prodistylomys</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CA92D843CA12893FFC23FAA5F13EFA76" box="[993,1092,1394,1417]" pageId="8" pageNumber="641" rank="species">nov. spec.</taxonomicNameLabel>
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1 (
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<treatmentCitation id="62749F3BCA12893FFBA0FAA6F032FA76" author="Oliver A & Sanisidro O & Pelaez-Campomanes P" box="[1122,1352,1393,1418]" page="61" pageId="8" pageNumber="641" year="2017">
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<bibRefCitation id="8744C4DBCA12893FFBA0FAA6F032FA76" author="Oliver A & Sanisidro O & Pelaez-Campomanes P" box="[1122,1352,1393,1418]" pageId="8" pageNumber="641" pagination="372 - 9" refId="ref17898" refString="Oliver A, Sanisidro O, Pelaez-Campomanes P et al. Wear reconstruction of the teeth of Yindirtemys deflexus (Ctenodactylidae, Rodentia) and evolution of its dental pauern. Hist Biol 2017 a; 30: 372 - 9." type="journal article" year="2017">
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Oliver
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<emphasis id="D1A16538CA12893FFB69FAA5F1A9FA76" box="[1195,1235,1393,1417]" italics="true" pageId="8" pageNumber="641">et al</emphasis>
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. 2017a: 61
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).
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<paragraph id="E36AB92ACA12893FFC97FA46F6DFFA37" blockId="8.[825,1475,1362,1480]" pageId="8" pageNumber="641">
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<treatmentCitationGroup id="C3C59E04CA12893FFC97FA46F0B9FA56" box="[853,1475,1425,1449]" pageId="8" pageNumber="641">
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<taxonomicName id="24D5C2A9CA12893FFC97FA46F6A6FA56" box="[853,988,1425,1449]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="D1A16538CA12893FFC97FA46F6A6FA56" box="[853,988,1425,1449]" italics="true" pageId="8" pageNumber="641">Prodistylomys</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CA92D843CA12893FFC23FA46F13FFA57" box="[993,1093,1425,1448]" pageId="8" pageNumber="641" rank="species">nov. spec.</taxonomicNameLabel>
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1 (
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<treatmentCitation id="62749F3BCA12893FFBA1FA46F0E9FA56" author="Daxner-Hock G & Badamgarav D & Barsbold D" box="[1123,1427,1425,1449]" page="130" pageId="8" pageNumber="641" year="2017">
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<bibRefCitation id="8744C4DBCA12893FFBA1FA46F0E9FA56" author="Daxner-Hock G & Badamgarav D & Barsbold D" box="[1123,1427,1425,1449]" pageId="8" pageNumber="641" pagination="1 - 108" refId="ref15507" refString="Daxner-Hock G, Badamgarav D, Barsbold D et al. Oligocene stratigraphy across the Eocene and Miocene boundaries in the Valley of Lakes (Mongolia). Palaeobiodivers Palaeoenviron 2017; 97: 1 - 108." type="journal article" year="2017">
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Daxner-Höck
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<emphasis id="D1A16538CA12893FFB35FA45F05CFA56" box="[1271,1318,1425,1449]" italics="true" pageId="8" pageNumber="641">et al.</emphasis>
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2017: 130
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, tab.
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1, fig. 47k)
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<subSubSection id="ABCFEAA1CA12893FFCFBFA3EF1EAF9DE" pageId="8" pageNumber="641" type="etymology">
|
|
<paragraph id="E36AB92ACA12893FFCFBFA3EF1EAF9DE" blockId="8.[825,1474,1513,1569]" pageId="8" pageNumber="641">
|
|
<emphasis id="D1A16538CA12893FFCFBFA3EF6D2F9FE" box="[825,936,1513,1537]" italics="true" pageId="8" pageNumber="641">Etymology:</emphasis>
|
|
The species name refers to
|
|
<collectingCountry id="9BC2F9BACA12893FFB1AFA3EF047F9FE" box="[1240,1341,1513,1537]" name="Mongolia" pageId="8" pageNumber="641">Mongolia</collectingCountry>
|
|
, the country where the new species was found.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="ABCFEAA1CA12893FFCFBF995F1AFF946" pageId="8" pageNumber="641" type="materials_examined">
|
|
<paragraph id="E36AB92ACA12893FFCFBF995F0B8F9A5" blockId="8.[825,1474,1602,1626]" box="[825,1474,1602,1626]" pageId="8" pageNumber="641">
|
|
<materialsCitation id="53BDB377CA12893FFCFBF995F0B8F9A5" box="[825,1474,1602,1626]" collectionCode="NHMW" pageId="8" pageNumber="641" specimenCode="NHMW 2012" specimenCount="1" typeStatus="holotype">
|
|
<emphasis id="D1A16538CA12893FFCFBF995F6E9F9A5" box="[825,915,1602,1626]" italics="true" pageId="8" pageNumber="641">
|
|
<typeStatus id="3C6E0788CA12893FFCFBF995F6F4F9A5" box="[825,910,1602,1626]" pageId="8" pageNumber="641" type="holotype">Holotype</typeStatus>
|
|
:
|
|
</emphasis>
|
|
<specimenCode id="B3731151CA12893FFC54F994F15EF9A5" box="[918,1060,1602,1626]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="641" type="Museum">NHMW 2012</specimenCode>
|
|
/0051/0001, Rightm2 (
|
|
<figureCitation id="7BEEA5AFCA12893FFACAF995F036F9A5" box="[1288,1356,1602,1626]" captionStart="Figure 6" captionStartId="9.[113,178,1607,1631]" captionTargetBox="[119,1449,1129,1569]" captionTargetId="figure-192@9.[114,1458,1123,1579]" captionTargetPageId="9" captionText="Figure 6. Prodistylomys mongoliensis sp. nov. from the type locality of Hotuliin Teeg, Valley of Lakes, Mongolia:(A) holotype of Prodistylomys mongoliensis sp. nov. from Hotuliin Teeg (HTE-012), (NHMW 2012/0051/0001). Right m2; (B) HTE-012 (NHMW 2012/0051/0002) M3 left. Left-side specimens are underlined. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10469788" httpUri="https://zenodo.org/record/10469788/files/figure.png" pageId="8" pageNumber="641">Fig.6A</figureCitation>
|
|
; ModelS3).
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<paragraph id="E36AB92ACA12893FFCFBF9A5F0B8F975" blockId="8.[825,1474,1649,1674]" box="[825,1474,1649,1674]" pageId="8" pageNumber="641">
|
|
<materialsCitation id="53BDB377CA12893FFCFBF9A5F0C7F975" box="[825,1469,1649,1674]" collectionCode="NHMW" pageId="8" pageNumber="641" specimenCode="NHMW 2012" specimenCount="1" typeStatus="paratype">
|
|
<emphasis id="D1A16538CA12893FFCFBF9A5F6E2F976" box="[825,920,1650,1673]" italics="true" pageId="8" pageNumber="641">
|
|
<typeStatus id="3C6E0788CA12893FFCFBF9A5F6E8F976" box="[825,914,1650,1673]" pageId="8" pageNumber="641" type="paratype">Paratype</typeStatus>
|
|
:
|
|
</emphasis>
|
|
HTE-012;
|
|
<specimenCode id="B3731151CA12893FFBD2F9A5F1D9F976" box="[1040,1187,1649,1673]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="641" type="Museum">NHMW 2012</specimenCode>
|
|
/0051/0002-0006 (
|
|
<figureCitation id="7BEEA5AFCA12893FFAAEF9A6F0C8F976" box="[1388,1458,1649,1673]" captionStart="Figure 6" captionStartId="9.[113,178,1607,1631]" captionTargetBox="[119,1449,1129,1569]" captionTargetId="figure-192@9.[114,1458,1123,1579]" captionTargetPageId="9" captionText="Figure 6. Prodistylomys mongoliensis sp. nov. from the type locality of Hotuliin Teeg, Valley of Lakes, Mongolia:(A) holotype of Prodistylomys mongoliensis sp. nov. from Hotuliin Teeg (HTE-012), (NHMW 2012/0051/0001). Right m2; (B) HTE-012 (NHMW 2012/0051/0002) M3 left. Left-side specimens are underlined. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10469788" httpUri="https://zenodo.org/record/10469788/files/figure.png" pageId="8" pageNumber="641">Fig. 6B</figureCitation>
|
|
)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="E36AB92ACA12893FFCFBF975F1AFF946" blockId="8.[825,1237,1697,1721]" box="[825,1237,1697,1721]" pageId="8" pageNumber="641">
|
|
<emphasis id="D1A16538CA12893FFCFBF975F6C1F946" box="[825,955,1697,1721]" italics="true" pageId="8" pageNumber="641">Type locality:</emphasis>
|
|
Hotuliin Teeg (HTE-012).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="ABCFEAA1CA12893EFCFBF906F169F920" lastPageId="9" lastPageNumber="642" pageId="8" pageNumber="641" type="diagnosis">
|
|
<paragraph id="E36AB92ACA12893FFCFBF906F6F7F8B9" blockId="8.[825,1474,1744,1862]" pageId="8" pageNumber="641">
|
|
<emphasis id="D1A16538CA12893FFCFBF906F6E5F917" box="[825,927,1745,1768]" italics="true" pageId="8" pageNumber="641">Diagnosis:</emphasis>
|
|
Large-sized species of
|
|
<taxonomicName id="24D5C2A9CA12893FFB4CF907F06FF917" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[1166,1301,1744,1768]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="genus">
|
|
<emphasis id="D1A16538CA12893FFB4CF907F06FF917" box="[1166,1301,1744,1768]" italics="true" pageId="8" pageNumber="641">Prodistylomys</emphasis>
|
|
</taxonomicName>
|
|
. Lower m2 with long metaflexid and posterolophid; upper M1 and M2 with metaflexus almost connected to the ectoflexus; M3 with a fossette.
|
|
</paragraph>
|
|
<paragraph id="E36AB92ACA12893EFCFBF88AF46EF8E1" blockId="8.[825,1475,1885,2003]" lastBlockId="9.[113,763,1735,1979]" lastPageId="9" lastPageNumber="642" pageId="8" pageNumber="641">
|
|
<emphasis id="D1A16538CA12893FFCFBF88AF059F88A" box="[825,1315,1885,1909]" italics="true" pageId="8" pageNumber="641">
|
|
Differential diagnosis:
|
|
<taxonomicName id="24D5C2A9CA12893FFBDBF88AF059F88A" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[1049,1315,1885,1909]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="species" species="mongoliensis" status="sp. nov.">Prodistylomys mongoliensis</taxonomicName>
|
|
</emphasis>
|
|
<taxonomicNameLabel id="CA92D843CA12893FFAE9F889F000F88A" box="[1323,1402,1886,1909]" pageId="8" pageNumber="641" rank="species">sp. nov.</taxonomicNameLabel>
|
|
differs from
|
|
<emphasis id="D1A16538CA12893FFCB0F8A9F177F86A" box="[882,1037,1917,1941]" italics="true" pageId="8" pageNumber="641">
|
|
<taxonomicName id="24D5C2A9CA12893FFCB0F8A9F6A9F86A" box="[882,979,1918,1941]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="species" species="wangae">P. wangae</taxonomicName>
|
|
,
|
|
<taxonomicName id="24D5C2A9CA12893FFC1CF8A9F173F86A" box="[990,1033,1917,1941]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="species" species="lii">P. lii</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
and
|
|
<taxonomicName id="24D5C2A9CA12893FFB82F8A9F1CEF86A" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[1088,1204,1918,1941]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="species" species="taatsinius">
|
|
<emphasis id="D1A16538CA12893FFB82F8A9F1CEF86A" box="[1088,1204,1918,1941]" italics="true" pageId="8" pageNumber="641">P. taatsinius</emphasis>
|
|
</taxonomicName>
|
|
by its larger size, m2 with metaflexid, and an M3 with a vertical fossette.
|
|
<taxonomicName id="24D5C2A9CA12893FFAF9F84BF6C9F82C" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="species" species="mongoliensis" status="sp. nov.">
|
|
<emphasis id="D1A16538CA12893FFAF9F84BF6C9F82C" italics="true" pageId="8" pageNumber="641">Prodistylomys mongoliensis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="CA92D843CA12893FFC7BF86BF17FF82C" box="[953,1029,1980,2003]" pageId="8" pageNumber="641" rank="species">sp. nov.</taxonomicNameLabel>
|
|
differs from
|
|
<taxonomicName id="24D5C2A9CA12893FFB4AF86BF060F82C" box="[1160,1306,1980,2003]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="species" species="xinjiangensis">
|
|
<emphasis id="D1A16538CA12893FFB4AF86BF060F82C" box="[1160,1306,1980,2003]" italics="true" pageId="8" pageNumber="641">P. xinjiangensis</emphasis>
|
|
</taxonomicName>
|
|
by its larger size and different proportions of the trigonid/talonid (
|
|
<taxonomicName id="24D5C2A9CA13893EFDA9F91FF780F920" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[619,762,1735,1759]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="642" phylum="Chordata" rank="species" species="mongoliensis" status="sp. nov.">
|
|
<emphasis id="D1A16538CA13893EFDA9F91FF780F920" box="[619,762,1735,1759]" italics="true" pageId="9" pageNumber="642">P. mongoliensis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="CA92D843CA13893EFFB3F930F5C5F901" box="[113,191,1767,1790]" pageId="9" pageNumber="642" rank="species">sp. nov.</taxonomicNameLabel>
|
|
longer and wider trigonid than talonid; the opposite in
|
|
<taxonomicName id="24D5C2A9CA13893EFFB3F8D0F479F8E2" box="[113,259,1798,1822]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="642" phylum="Chordata" rank="species" species="xinjiangensis">
|
|
<emphasis id="D1A16538CA13893EFFB3F8D0F479F8E2" box="[113,259,1798,1822]" italics="true" pageId="9" pageNumber="642">P. xinjiangensis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<caption id="B7AAE9A2CA13893EFFB3FC75F6AFFBD5" ID-DOI="http://doi.org/10.5281/zenodo.10469786" ID-Zenodo-Dep="10469786" httpUri="https://zenodo.org/record/10469786/files/figure.png" pageId="9" pageNumber="642" startId="9.[113,178,930,954]" targetBox="[121,1450,152,895]" targetPageId="9" targetType="figure">
|
|
<paragraph id="E36AB92ACA13893EFFB3FC75F6AFFBD5" blockId="9.[113,1460,930,1067]" pageId="9" pageNumber="642">
|
|
<emphasis id="D1A16538CA13893EFFB3FC75F5B3FC44" bold="true" box="[113,201,930,955]" pageId="9" pageNumber="642">Figure 5.</emphasis>
|
|
Upper and lower cheek teeth of
|
|
<taxonomicName id="24D5C2A9CA13893EFE38FC75F792FC45" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[506,744,930,954]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="642" phylum="Chordata" rank="species" species="taatsinius" status="sp. nov.">Prodistylomys taatsinius</taxonomicName>
|
|
<taxonomicNameLabel id="CA92D843CA13893EFD2FFC74F643FC45" box="[749,825,931,955]" pageId="9" pageNumber="642" rank="species">
|
|
<emphasis id="D1A16538CA13893EFD2FFC74F64EFC44" bold="true" box="[749,820,931,955]" pageId="9" pageNumber="642">sp. nov</emphasis>
|
|
.
|
|
</taxonomicNameLabel>
|
|
from the Valley of Lakes (Mongolia). From Unkheltseg: (A) UNCH-A/3 (NHMW 2012/0049/0003) P4 left; (B) UNCH-A/3 (NHMW 2012/0049/0005) M1/2 left; (C) UNCH-A/3 (NHMW 2012/0049/0001) m2 right; From Huch Teeg: (D) RHN-A/12 (NHMW 2012/0050/0001) DP4 right; (E) RHN-A/12 (NHMW 2012/0050/0003) M1/2 right; (F) RHN-A/12 (NHMW 2012/0050/0006) p4 right; (G) RHN-A/12 (NHMW 2012/0050/0002) m3 right. All teeth are at the same magnification. Left-side specimens are underlined. Scale bar: 1 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="B7AAE9A2CA13893EFFB3F990F612F968" ID-DOI="http://doi.org/10.5281/zenodo.10469788" ID-Zenodo-Dep="10469788" httpUri="https://zenodo.org/record/10469788/files/figure.png" pageId="9" pageNumber="642" startId="9.[113,178,1607,1631]" targetBox="[119,1449,1129,1569]" targetPageId="9" targetType="figure">
|
|
<paragraph id="E36AB92ACA13893EFFB3F990F612F968" blockId="9.[113,1389,1607,1687]" pageId="9" pageNumber="642">
|
|
<emphasis id="D1A16538CA13893EFFB3F990F4A3F9A0" bold="true" box="[113,473,1607,1631]" pageId="9" pageNumber="642">
|
|
Figure 6.
|
|
<taxonomicName id="24D5C2A9CA13893EFF0CF990F4A3F9A0" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[206,473,1607,1631]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="642" phylum="Chordata" rank="species" species="mongoliensis" status="sp. nov.">
|
|
<emphasis id="D1A16538CA13893EFF0CF990F4A3F9A0" bold="true" box="[206,473,1607,1631]" italics="true" pageId="9" pageNumber="642">Prodistylomys mongoliensis</emphasis>
|
|
</taxonomicName>
|
|
</emphasis>
|
|
<taxonomicNameLabel id="CA92D843CA13893EFE1FF990F753F9A0" box="[477,553,1607,1631]" pageId="9" pageNumber="642" rank="species">
|
|
<emphasis id="D1A16538CA13893EFE1FF990F75EF9A0" bold="true" box="[477,548,1607,1631]" pageId="9" pageNumber="642">sp. nov</emphasis>
|
|
.
|
|
</taxonomicNameLabel>
|
|
from the type locality of Hotuliin Teeg, Valley of Lakes, Mongolia: (A) holotype of
|
|
<taxonomicName id="24D5C2A9CA13893EFFB3F9B4F407F984" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[113,381,1635,1659]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="642" phylum="Chordata" rank="species" species="mongoliensis" status="sp. nov.">Prodistylomys mongoliensis</taxonomicName>
|
|
<taxonomicNameLabel id="CA92D843CA13893EFE43F9B4F4B7F984" box="[385,461,1635,1659]" pageId="9" pageNumber="642" rank="species">
|
|
<emphasis id="D1A16538CA13893EFE43F9B4F4B2F984" bold="true" box="[385,456,1635,1659]" pageId="9" pageNumber="642">sp. nov</emphasis>
|
|
.
|
|
</taxonomicNameLabel>
|
|
from Hotuliin Teeg (HTE-012), (NHMW 2012/0051/0001). Right m2; (B) HTE-012 (NHMW 2012/0051/0002) M3 left. Left-side specimens are underlined. Scale bar: 1 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="E36AB92ACA13893EFF4FF8F2F49CF883" blockId="9.[113,763,1735,1979]" pageId="9" pageNumber="642">
|
|
<taxonomicName id="24D5C2A9CA13893EFF4FF8F2F4E2F8C2" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[141,408,1829,1853]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="642" phylum="Chordata" rank="species" species="mongoliensis" status="sp. nov.">
|
|
<emphasis id="D1A16538CA13893EFF4FF8F2F4E2F8C2" box="[141,408,1829,1853]" italics="true" pageId="9" pageNumber="642">Prodistylomys mongoliensis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="CA92D843CA13893EFE60F8F1F488F8C2" box="[418,498,1830,1853]" pageId="9" pageNumber="642" rank="species">sp. nov.</taxonomicNameLabel>
|
|
differs from
|
|
<taxonomicName id="24D5C2A9CA13893EFDBDF8F1F781F8C2" box="[639,763,1830,1853]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="642" phylum="Chordata" rank="species" species="mengensis">
|
|
<emphasis id="D1A16538CA13893EFDBDF8F1F781F8C2" box="[639,763,1830,1853]" italics="true" pageId="9" pageNumber="642">P. mengensis</emphasis>
|
|
</taxonomicName>
|
|
by its larger size and upper molars, including M3 with a vertical fossette instead of a dentine isthmus.
|
|
</paragraph>
|
|
<paragraph id="E36AB92ACA13893EFFB3F854F5BFF844" blockId="9.[113,763,1735,1979]" pageId="9" pageNumber="642">
|
|
<emphasis id="D1A16538CA13893EFFB3F854F455F864" box="[113,303,1923,1947]" italics="true" pageId="9" pageNumber="642">Other fossil layers:</emphasis>
|
|
HTE-003; NHMW2012/0052/0001-0002 (
|
|
<figureCitation id="7BEEA5AFCA13893EFFBEF875F5CEF845" box="[124,180,1954,1978]" captionStart="Figure 7" captionStartId="10.[129,194,771,795]" captionTargetBox="[141,1469,154,741]" captionTargetId="figure-523@10.[129,1473,144,744]" captionTargetPageId="10" captionText="Figure 7. Upper and lower cheek teeth of Prodistylomys mongoliensis sp. nov. from the Valley of Lakes (Mongolia). From Hotuliin Teeg: (A) HTE-003 (NHMW 2012/0052/0002) M1/2 right; (B) HTE-003 (NHMW 2012/0052/0001) m2 left. All teeth are at the same magnification.Right-side specimens are underlined. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10469790" httpUri="https://zenodo.org/record/10469790/files/figure.png" pageId="9" pageNumber="642">Fig. 7</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="E36AB92ACA13893EFCE8F91FF169F920" blockId="9.[810,1043,1735,1759]" box="[810,1043,1735,1759]" pageId="9" pageNumber="642">
|
|
<emphasis id="D1A16538CA13893EFCE8F91FF6C6F920" box="[810,956,1736,1759]" italics="true" pageId="9" pageNumber="642">Measurements:</emphasis>
|
|
<tableCitation id="AE578C91CA13893EFC00F910F177F920" box="[962,1037,1735,1759]" captionStart="Table 4" captionStartId="12.[111,166,444,469]" captionTargetPageId="12" captionText="Table 4. Descriptive statistics of the upper and lower cheek teeth of Prodistylomys including the new species. All measurements are given in millimetres.Length and width represent the maximum antero-posterior and bucco-lingual distances. Abbreviations: Elem., element; N., number of teeth measured; min, minimum; max, maximum. For abbreviations of assemblages see Table 2." pageId="9" pageNumber="642">Table 4</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="ABCFEAA1CA13893EFCE8F925F185F8C9" pageId="9" pageNumber="642" type="biology_ecology">
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<paragraph id="E36AB92ACA13893EFCE8F925F009F8F5" blockId="9.[810,1395,1778,1802]" box="[810,1395,1778,1802]" pageId="9" pageNumber="642">
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<emphasis id="D1A16538CA13893EFCE8F925F690F8F5" box="[810,1002,1778,1802]" italics="true" pageId="9" pageNumber="642">Stratigraphic range:</emphasis>
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Early Miocene, Mongolian biozone D.
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</paragraph>
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<paragraph id="E36AB92ACA13893EFCE8F8C9F185F8C9" blockId="9.[810,1279,1822,1846]" box="[810,1279,1822,1846]" pageId="9" pageNumber="642">
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<emphasis id="D1A16538CA13893EFCE8F8C9F697F8CA" box="[810,1005,1822,1846]" italics="true" pageId="9" pageNumber="642">Geographical range:</emphasis>
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Valley of Lakes,
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<collectingCountry id="9BC2F9BACA13893EFB57F8C9F180F8C9" box="[1173,1274,1822,1846]" name="Mongolia" pageId="9" pageNumber="642">Mongolia</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="ABCFEAA1CA13893DFC85F888F714FA68" lastPageId="10" lastPageNumber="643" pageId="9" pageNumber="642" type="description">
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<paragraph id="E36AB92ACA13893EFC85F888F0ECF887" blockId="9.[839,1430,1886,1912]" box="[839,1430,1886,1912]" pageId="9" pageNumber="642">
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<emphasis id="D1A16538CA13893EFC85F888F0ECF887" box="[839,1430,1886,1912]" italics="true" pageId="9" pageNumber="642">Description of the new material flom Hotuliin Teeg (HTE-012)</emphasis>
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</paragraph>
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<paragraph id="E36AB92ACA13893DFCE8F852F410FB98" blockId="9.[810,1459,1924,1980]" lastBlockId="10.[129,778,915,1127]" lastPageId="10" lastPageNumber="643" pageId="9" pageNumber="642">
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<emphasis id="D1A16538CA13893EFCE8F852F604F862" box="[810,894,1925,1949]" italics="true" pageId="9" pageNumber="642">m1/m2:</emphasis>
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Bilophodont and hypsodont teeth. The specimens have two roots. The shape of the trigonid is fusiform (2/3) or rhomboidal, with the metaconid and protoconid narrower than the anterolophid (1/3). Both the metaconid and the protoconid are sharp. The trigonid is larger than the talonid in one (1/3) and smaller in two (2/3). The talonid is rhomboidal to subrhomboidal. The entoconid and the hypoconid are sharp. The mesoflexid and the ectoflexid are V-shaped, open, and deep. The metaflexid is deep.
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</paragraph>
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<caption id="B7AAE9A2CA10893DFF43FCD4F676FCAC" ID-DOI="http://doi.org/10.5281/zenodo.10469790" ID-Zenodo-Dep="10469790" httpUri="https://zenodo.org/record/10469790/files/figure.png" pageId="10" pageNumber="643" startId="10.[129,194,771,795]" targetBox="[141,1469,154,741]" targetPageId="10" targetType="figure">
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<paragraph id="E36AB92ACA10893DFF43FCD4F0EAFCE3" blockId="10.[129,1424,771,852]" box="[129,1424,771,796]" pageId="10" pageNumber="643">
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<emphasis id="D1A16538CA10893DFF43FCD4F5A2FCE3" bold="true" box="[129,216,771,796]" pageId="10" pageNumber="643">Figure 7.</emphasis>
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Upper and lower cheek teeth of
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<taxonomicName id="24D5C2A9CA10893DFDCBFCD4F66EFCE4" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[521,788,771,795]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="643" phylum="Chordata" rank="species" species="mongoliensis" status="sp. nov.">Prodistylomys mongoliensis</taxonomicName>
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<taxonomicNameLabel id="CA92D843CA10893DFCDBFCD3F61EFCE4" box="[793,868,772,796]" pageId="10" pageNumber="643" rank="species">
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<emphasis id="D1A16538CA10893DFCDBFCD3F61AFCE3" bold="true" box="[793,864,772,796]" pageId="10" pageNumber="643">sp. nov</emphasis>
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.
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</taxonomicNameLabel>
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from the Valley of Lakes (Mongolia). From Hotuliin Teeg:
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</paragraph>
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<paragraph id="E36AB92ACA10893DFF43FCF7F676FCAC" blockId="10.[129,1424,771,852]" pageId="10" pageNumber="643">(A) HTE-003 (NHMW 2012/0052/0002) M1/2 right; (B) HTE-003 (NHMW 2012/0052/0001) m2 left. All teeth are at the same magnification. Right-side specimens are underlined. Scale bar: 1 mm.</paragraph>
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</caption>
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<paragraph id="E36AB92ACA10893DFF43FB58F768FAFA" blockId="10.[129,778,1167,1285]" pageId="10" pageNumber="643">
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<emphasis id="D1A16538CA10893DFF43FB58F5A1FB58" box="[129,219,1167,1191]" italics="true" pageId="10" pageNumber="643">M1/M2:</emphasis>
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The teeth are hypsodont and rooted with one root. There is cement in the flexus. The trigon is kidney shaped and larger than the talon. The mesoflexus is deep, almost connected reaching the lingual border of the teeth.
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</paragraph>
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<paragraph id="E36AB92ACA10893DFF43FAF6F714FA68" blockId="10.[129,778,1313,1431]" pageId="10" pageNumber="643">
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<emphasis id="D1A16538CA10893DFF43FAF6F5D6FAC6" box="[129,172,1313,1337]" italics="true" pageId="10" pageNumber="643">M3:</emphasis>
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The only specimen is broken in the posterior border. It is hypsodont and rooted, with two roots. The flexus is filled with cement. The trigon is kidney shaped, whereas the talon is semicircular. The flexus is closed and forms a fossette.
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</paragraph>
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</subSubSection>
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<subSubSection id="ABCFEAA1CA10893DFF43FA66F581F9F7" pageId="10" pageNumber="643" type="discussion">
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<paragraph id="E36AB92ACA10893DFF43FA66F581F9F7" blockId="10.[129,778,1457,1544]" pageId="10" pageNumber="643">
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<emphasis id="D1A16538CA10893DFF43FA66F491FA36" box="[129,491,1457,1481]" italics="true" pageId="10" pageNumber="643">
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Remarks:
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<taxonomicName id="24D5C2A9CA10893DFF27FA66F491FA36" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[229,491,1457,1481]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="643" phylum="Chordata" rank="species" species="mongoliensis" status="sp. nov.">Prodistylomys mongoliensis</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="CA92D843CA10893DFE30FA65F745FA36" box="[498,575,1458,1481]" pageId="10" pageNumber="643" rank="species">sp. nov.</taxonomicNameLabel>
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occurs in two samples of Hotuliin Teeg (HTE-012 and HTE-003) in the region of Taatsin Gol.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |