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<document id="96CA91B626A2FF89F7FE29CF80C85E5B" ID-DOI="10.1016/j.sajb.2018.11.021" ID-ISSN="1727-9321" ID-Zenodo-Dep="10496596" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1705062987214" checkinUser="felipe" docAuthor="Elliott, T. L., Barrett, R. L. &amp; Muasya, A. M." docDate="2019" docId="68465A3B6B781D4D0071D789F696FBBE" docLanguage="en" docName="SAfrJBot.121.519-535.pdf" docOrigin="South African Journal of Botany 121" docSource="http://dx.doi.org/10.1016/j.sajb.2018.11.021" docStyle="DocumentStyle:AD72C92759D651943977BAEA6E2218B4.1:SAfrJBot.2014-.journal_article" docStyleId="AD72C92759D651943977BAEA6E2218B4" docStyleName="SAfrJBot.2014-.journal_article" docStyleVersion="1" docTitle="Schoenus quartziticus T. L. Elliott &amp; Muasya 2019, sp. nov." docType="treatment" docVersion="1" lastPageNumber="532" masterDocId="947F22436B741D40032DD230F45EFFF9" masterDocTitle="A taxonomic revision of Schoenus cuspidatus and allies (Cyperaceae, tribe Schoeneae) - Part 1 *" masterLastPageNumber="535" masterPageNumber="519" pageNumber="531" updateTime="1705673609269" updateUser="guilherme">
<mods:mods id="C75B509A4092959A556461B15C2EB77E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="22FCBF0315CCB60D8FFD564FF56F7005">
<mods:title id="173D3F4A354700D63CD95FBDC9DF8861">A taxonomic revision of Schoenus cuspidatus and allies (Cyperaceae, tribe Schoeneae) - Part 1 *</mods:title>
</mods:titleInfo>
<mods:name id="801594AD78D555F744AFE486E3E3C6CF" type="personal">
<mods:role id="60D7715E454A884D30CD72B374622B22">
<mods:roleTerm id="28464F7AE76BC2D07FC7DA44F0EA76C7">Author</mods:roleTerm>
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<mods:namePart id="0218BDC3B52C8C1F174BD6A5F6693A96">Elliott, T. L.</mods:namePart>
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<mods:namePart id="EBD0EBCCDD09280C6ADE66806E9AB573">Barrett, R. L.</mods:namePart>
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<mods:role id="31C7D089BF3505436846DFD5689B2C38">
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</mods:role>
<mods:namePart id="C9EF55E68022D17FD3D620A9620DFE76">Muasya, A. M.</mods:namePart>
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<mods:typeOfResource id="D65F4FBA1788F81B494133BD6358E263">text</mods:typeOfResource>
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<mods:title id="370256EBE2905805FEE1B00767CB1CAB">South African Journal of Botany</mods:title>
</mods:titleInfo>
<mods:part id="954AB32CA419F8CB6008503D46080652">
<mods:date id="EACEABBB9C1D94A62707834D779F1092">2019</mods:date>
<mods:detail id="EA67EC736466E2C20649136F491F0E69" type="pubDate">
<mods:number id="AC33C7400A23DA11A9C990D9BB55DE83">2019-03-31</mods:number>
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<mods:detail id="D09982D21D4F80B1585D82B645FF3448" type="volume">
<mods:number id="75386142469E5AFABAF2FA919F69293C">121</mods:number>
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<mods:start id="EBD188FD2842E633E527F58122A094B2">519</mods:start>
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<mods:url id="4F39295FC675F08B50F729298110A8BF">http://dx.doi.org/10.1016/j.sajb.2018.11.021</mods:url>
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<mods:classification id="1CED71149CCFC1581716D84BFD1C4204">journal article</mods:classification>
<mods:identifier id="E96F226B0BA0AD15B47E1772366D8410" type="DOI">10.1016/j.sajb.2018.11.021</mods:identifier>
<mods:identifier id="77D0F1A2D3B451E268D0B5F63157FB29" type="ISSN">1727-9321</mods:identifier>
<mods:identifier id="3D52E7375FD8E502EB2B3D584A4A63B8" type="Zenodo-Dep">10496596</mods:identifier>
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<treatment id="68465A3B6B781D4D0071D789F696FBBE" LSID="urn:lsid:plazi:treatment:68465A3B6B781D4D0071D789F696FBBE" httpUri="http://treatment.plazi.org/id/68465A3B6B781D4D0071D789F696FBBE" lastPageId="13" lastPageNumber="532" pageId="12" pageNumber="531">
<subSubSection id="A8F5B8A66B781D4C0071D789F13FFA34" box="[860,1377,1465,1486]" pageId="12" pageNumber="531" type="nomenclature">
<paragraph id="E050EB2D6B781D4C0071D789F13FFA34" blockId="12.[828,1499,155,1988]" box="[860,1377,1465,1486]" pageId="12" pageNumber="531">
<emphasis id="D29B373F6B781D4C0071D789F733FA34" bold="true" box="[860,877,1465,1485]" italics="true" pageId="12" pageNumber="531">8.</emphasis>
<taxonomicName id="27EF90AE6B781D4C005ED789F14FFA37" ID-CoL="8XW53" authority="T.L.Elliott &amp; Muasya" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" box="[883,1297,1465,1486]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="531" phylum="Tracheophyta" rank="species" species="quartziticus" status="sp. nov.">
<emphasis id="D29B373F6B781D4C005ED789F012FA34" bold="true" box="[883,1100,1465,1485]" italics="true" pageId="12" pageNumber="531">Schoenus quartziticus</emphasis>
T.L.Elliott &amp; Muasya
</taxonomicName>
,
<emphasis id="D29B373F6B781D4C0637D789F13FFA34" box="[1306,1377,1465,1485]" italics="true" pageId="12" pageNumber="531">
<taxonomicNameLabel id="C9A88A446B781D4C0637D789F13FFA34" box="[1306,1377,1465,1485]" pageId="12" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="A8F5B8A66B781D4C064BD78BF04CF9C4" pageId="12" pageNumber="531" type="materials_examined">
<paragraph id="E050EB2D6B781D4C064BD78BF04CF9C4" blockId="12.[828,1499,155,1988]" pageId="12" pageNumber="531">
<materialsCitation id="5087E1706B781D4C064BD78BF731F9C4" collectingDate="2016-10-01" collectionCode="BOL" collectorName="Elliott &amp; Muasya" country="South Africa" county="Bredasdorp" elevation="750" location="Overberg Renosterveld Nature Reserve" municipality="Haawegskloof Farm" pageId="12" pageNumber="531" specimenCount="1" stateProvince="Western Cape Province" typeStatus="holotype">
Type:
<collectingCountry id="98F8ABBD6B781D4C068CD78AF729FA10" name="South Africa" pageId="12" pageNumber="531">South Africa</collectingCountry>
,
<collectingRegion id="222B25CF6B781D4C00A9D7E7F02CFA10" box="[900,1138,1494,1514]" country="South Africa" name="Western Cape" pageId="12" pageNumber="531">Western Cape Province</collectingRegion>
, 3420 (
<collectingCounty id="093193A16B781D4C07EFD7E6F162FA10" box="[1218,1340,1494,1513]" pageId="12" pageNumber="531">Bredasdorp</collectingCounty>
):
<collectingMunicipality id="003471576B781D4C067DD7E6F733F9FF" pageId="12" pageNumber="531">Haawegskloof Farm</collectingMunicipality>
(
<location id="E530BDF66B781D4C005AD7C2F0B4F9FC" LSID="urn:lsid:plazi:treatment:68465A3B6B781D4D0071D789F696FBBE:E530BDF66B781D4C005AD7C2F0B4F9FC" box="[887,1258,1522,1541]" country="South Africa" county="Bredasdorp" municipality="Haawegskloof Farm" name="Overberg Renosterveld Nature Reserve" pageId="12" pageNumber="531" stateProvince="Western Cape Province">Overberg Renosterveld Nature Reserve</location>
),
<quantity id="271746C86B781D4C07D5D7C2F131F9FC" box="[1272,1391,1522,1541]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="12" pageNumber="531" unit="m" value="750.0" valueMax="1000.0" valueMin="500.0">
<elevation id="6BC20C1E6B781D4C07D5D7C2F131F9FC" box="[1272,1391,1522,1541]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="12" pageNumber="531" unit="m" value="750.0" valueMax="1000.0" valueMin="500.0">500–1000 m</elevation>
</quantity>
N.W. of research centre, (–AD),
<date id="9451CDED6B781D4C073CD43EF025F9D8" box="[1041,1147,1550,1569]" pageId="12" pageNumber="531" value="2016-10-01">
<collectingDate id="841534056B781D4C073CD43EF025F9D8" box="[1041,1147,1550,1569]" pageId="12" pageNumber="531" value="2016-10-01">1 Oct 2016</collectingDate>
</date>
,
<emphasis id="D29B373F6B781D4C07A8D43DF1C8F9D8" box="[1157,1430,1549,1569]" italics="true" pageId="12" pageNumber="531">
<collectorName id="4D1A8EFB6B781D4C07A8D43DF0E5F9D8" box="[1157,1211,1549,1569]" pageId="12" pageNumber="531">Elliott</collectorName>
&amp;
<collectorName id="4D1A8EFB6B781D4C07FBD43DF141F9D8" box="[1238,1311,1549,1569]" pageId="12" pageNumber="531">Muasya</collectorName>
TE2016_127
</emphasis>
(
<collectionCode id="86FE73E86B781D4C068FD43EF18DF9D8" box="[1442,1491,1550,1569]" collectionName="BOL" pageId="12" pageNumber="531">BOL</collectionCode>
!,
<typeStatus id="3F54558F6B781D4C0011D41AF737F9C4" box="[828,873,1578,1597]" pageId="12" pageNumber="531">holo</typeStatus>
.,
</materialsCitation>
<materialsCitation id="5087E1706B781D4C0058D41AF050F9C4" box="[885,1038,1578,1597]" collectionCode="MT, NBG" pageId="12" pageNumber="531" specimenCount="1" typeStatus="iso">
<collectionCode id="86FE73E86B781D4C0058D41AF7C5F9C4" box="[885,923,1578,1597]" pageId="12" pageNumber="531">MT</collectionCode>
!,
<collectionCode id="86FE73E86B781D4C008AD41AF787F9C4" box="[935,985,1578,1597]" collectionName="NBG" pageId="12" pageNumber="531">NBG</collectionCode>
!,
<typeStatus id="3F54558F6B781D4C00C8D41AF05AF9C4" box="[997,1028,1578,1597]" pageId="12" pageNumber="531">iso</typeStatus>
.)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A8F5B8A66B781D4D0071D476F6ECFE18" lastPageId="13" lastPageNumber="532" pageId="12" pageNumber="531" type="description">
<paragraph id="E050EB2D6B781D4D0071D476F6ECFE18" blockId="12.[828,1499,155,1988]" lastBlockId="13.[90,760,154,1402]" lastPageId="13" lastPageNumber="532" pageId="12" pageNumber="531">
Caespitose, phyllopodic perennial graminoid.
<emphasis id="D29B373F6B781D4C0625D475F11EF9A0" box="[1288,1344,1605,1625]" italics="true" pageId="12" pageNumber="531">Culms</emphasis>
terete, 310–678 ×
<quantity id="271746C86B781D4C007CD452F798F98C" box="[849,966,1634,1653]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="12" pageNumber="531" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 mm</quantity>
.
<emphasis id="D29B373F6B781D4C00E3D451F055F98C" box="[974,1035,1633,1653]" italics="true" pageId="12" pageNumber="531">Leaves</emphasis>
basal, 1–3(—6), 130–189(—273) × 0.2–1.0 mm, usually straight, proximally channelled, margin serrate above sheath.
<emphasis id="D29B373F6B781D4C0011D4A8F7DCF955" box="[828,898,1688,1708]" italics="true" pageId="12" pageNumber="531">Sheaths</emphasis>
firm, reddish-brown but paler above, longitudinally striate.
<emphasis id="D29B373F6B781D4C069AD4A8F706F931" italics="true" pageId="12" pageNumber="531">Ligule</emphasis>
usually firm but can be scarcely membranaceous,
<quantity id="271746C86B781D4C0618D486F1F8F930" box="[1333,1446,1717,1737]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="12" pageNumber="531" unit="mm" value="0.5" valueMax="0.8" valueMin="0.2">0.2–0.8 mm</quantity>
long.
<emphasis id="D29B373F6B781D4C0011D4E0F7EAF91D" box="[828,948,1744,1764]" italics="true" pageId="12" pageNumber="531">
In
<emphasis id="D29B373F6B781D4C0063D4E1F704F91D" bold="true" box="[846,858,1745,1764]" italics="true" pageId="12" pageNumber="531">fl</emphasis>
orescence
</emphasis>
a narrow pseudolateral panicle, 22–28(—47) ×
<quantity id="271746C86B781D4C06A9D4E1F188F91C" box="[1412,1494,1745,1765]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="12" pageNumber="531" unit="mm" value="5.5" valueMax="8.0" valueMin="3.0">3–8 mm</quantity>
, proximal rachis length varying in length,
<quantity id="271746C86B781D4C0797D4DEF148F8F8" box="[1210,1302,1773,1793]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.5" metricValueMin="0.7" pageId="12" pageNumber="531" unit="mm" value="16.0" valueMax="25.0" valueMin="7.0">7–25 mm</quantity>
.
<emphasis id="D29B373F6B781D4C0630D4DCF782F8E5" italics="true" pageId="12" pageNumber="531">
Proximal primary
<emphasis id="D29B373F6B781D4C0692D4DCF716F8E5" bold="true" italics="true" pageId="12" pageNumber="531">infl</emphasis>
orescence bracts
</emphasis>
sometimes channelled proximally, apex acute to acuminate, not widened at base, (26–)37–112(—134) mm long, exceeding length of inflorescence up to several times, longitudinal veins not notable. Proximal and subproximal primary inflorescence bracts without membranaceous extensions at base.
<emphasis id="D29B373F6B781D4C07BFD548F094F875" box="[1170,1226,1912,1932]" italics="true" pageId="12" pageNumber="531">Spikes</emphasis>
2–4, 7.0–15.0 mm long, usually dispersed along rachis.
<emphasis id="D29B373F6B781D4C0712D5A3F0D0F85E" box="[1087,1166,1939,1959]" italics="true" pageId="12" pageNumber="531">Spikelets</emphasis>
lanceolate, relatively short, 3.4–3.9 × 0.7–1.0 mm, pedicellate, 3–6 spikelets per spike, ranging from firm to membranaceous or chaffy in texture.
<emphasis id="D29B373F6B791D4D02E4D2AAF6E0FF57" box="[457,702,154,174]" italics="true" pageId="13" pageNumber="532">Proximal spikelet prophyll</emphasis>
1 per spikelet, with an inconspicuous vein extending to mucro,
<quantity id="271746C86B791D4D01A8D287F6A6FF33" box="[645,760,183,202]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" pageId="13" pageNumber="532" unit="mm" value="0.65" valueMax="0.9" valueMin="0.4">0.4–0.9 mm</quantity>
long, prophyll mucro (0.8–)
<quantity id="271746C86B791D4D0247D2E3F5BFFF1F" box="[362,481,211,230]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.4" metricValueMin="2.2" pageId="13" pageNumber="532" unit="mm" value="2.8" valueMax="3.4" valueMin="2.2">2.2–3.4 mm</quantity>
long.
<emphasis id="D29B373F6B791D4D0133D2E2F636FF1F" box="[542,616,210,230]" italics="true" pageId="13" pageNumber="532">Rachilla</emphasis>
0.6–2.4(—7.2) mm long.
<emphasis id="D29B373F6B791D4D0396D2DEF4A1FEFB" box="[187,255,238,258]" italics="true" pageId="13" pageNumber="532">Glumes</emphasis>
4–7 per spikelet, proximal glume (0.5–)1.0–
<quantity id="271746C86B791D4D0183D2DFF6A6FEFB" box="[686,760,239,258]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="13" pageNumber="532" unit="mm" value="1.8">1.8 mm</quantity>
long, subproximal glume 1.0–
<quantity id="271746C86B791D4D0254D33BF59DFEE7" box="[377,451,267,286]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="532" unit="mm" value="2.2">2.2 mm</quantity>
long, upper glumes longer than basal ones, lower glumes sometimes over half the length of spikelet, apex obtuse. Glume mucros absent or short, proximal mucro
<quantity id="271746C86B791D4D0377D36FF496FE8B" box="[90,200,351,370]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="17.0" metricValueMin="1.0" pageId="13" pageNumber="532" unit="mm" value="0.9" valueMax="1.7" valueMin="0.1">0.1–1.7 mm</quantity>
long, subproximal mucro
<quantity id="271746C86B791D4D0299D36FF67CFE8B" box="[436,546,351,370]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="16.0" metricValueMin="1.0" pageId="13" pageNumber="532" unit="mm" value="0.85" valueMax="1.6" valueMin="0.1">0.1–1.6 mm</quantity>
long.
<emphasis id="D29B373F6B791D4D017BD36DF6FCFE88" box="[598,674,349,369]" italics="true" pageId="13" pageNumber="532">Stamens</emphasis>
3 per floret, anthers not measured.
<emphasis id="D29B373F6B791D4D027AD349F5C0FE74" box="[343,414,377,397]" italics="true" pageId="13" pageNumber="532">Stigmas</emphasis>
3-branched, vestigial stigmas of second bisexual floret not observed.
<emphasis id="D29B373F6B791D4D02B5D3A5F675FE50" box="[408,555,405,425]" italics="true" pageId="13" pageNumber="532">Perianth bristles</emphasis>
about half the length of nutlet.
<emphasis id="D29B373F6B791D4D0398D381F4B0FE3C" box="[181,238,433,453]" italics="true" pageId="13" pageNumber="532">Nutlet</emphasis>
broad elliptic, trigonous, yellowish in colour, 1.9–2.0 ×
<quantity id="271746C86B791D4D0377D3FEF495FE1B" box="[90,203,462,482]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="13" pageNumber="532" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 mm</quantity>
.
<emphasis id="D29B373F6B791D4D03FED3FDF567FE18" box="[211,313,461,481]" italics="true" pageId="13" pageNumber="532">Nutlet beak</emphasis>
,
<quantity id="271746C86B791D4D026ED3FEF5ECFE1B" box="[323,434,462,482]" metricMagnitude="-4" metricUnit="m" metricValue="6" metricValueMax="8.0" metricValueMin="4.0" pageId="13" pageNumber="532" unit="mm" value="0.6" valueMax="0.8" valueMin="0.4">0.4–0.8 mm</quantity>
long, hispid (
<figureCitation id="78D4F7A86B791D4D011CD3FEF635FE18" box="[561,619,462,481]" captionStart="Fig" captionStartId="3.[90,115,602,616]" captionTargetBox="[328,1238,143,565]" captionTargetId="figure-246@3.[327,1238,142,566]" captionTargetPageId="3" captionText="Fig.2. Examples of nutlets from 11 species of the Schoenus cuspidatus and allies group found in southern Africa:A—S.auritus; B—S.graminifolius; C—S.ligulatus; D—S.exilis; E—S.bolusii; F— S.submarginalis; G—S.crassiculmis; H—S.quartziticus; I—S.limosus; J—S.purpurascens; and K—S.calceolus (immature).The black scale bar represents 1 mm.The mature fruits of S.calceolus and S.prophyllus are unknown." figureDoi="http://doi.org/10.5281/zenodo.10496600" httpUri="https://zenodo.org/record/10496600/files/figure.png" pageId="13" pageNumber="532">Figs. 2</figureCitation>
and
<figureCitation id="78D4F7A86B791D4D01B5D3FEF6FAFE18" box="[664,676,462,481]" captionStart="Fig" captionStartId="4.[113,138,983,997]" captionTargetBox="[275,1336,143,946]" captionTargetId="figure-478@4.[274,1336,142,947]" captionTargetPageId="4" captionText="Fig.4. Examples of plants from six species of the Schoenus cuspidatus andallies group found in southern Africa,showing the corresponding inflorescences and spikelets beneath each plant: A,G and M—S.crassiculmis; B,H and N—S.quartziticus, C,I andO—S.limosus; D,J,and P—S.purpurascens; E,K andQ—S.prophyllus; and F,L and R—S.calceolus. The black scale bar adjacent to the inflorescences in G—L represents 10 mm, while the scale bars adjacent to the spikelets in M—R represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10496604" httpUri="https://zenodo.org/record/10496604/files/figure.png" pageId="13" pageNumber="532">4</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="B490BBA56B781D4C03DCD584F6CFF83B" ID-DOI="http://doi.org/10.5281/zenodo.10496628" ID-Zenodo-Dep="10496628" box="[241,657,1971,1987]" httpUri="https://zenodo.org/record/10496628/files/figure.png" pageId="12" pageNumber="531" startId="12.[241,266,1972,1986]" targetBox="[147,749,1446,1932]" targetPageId="12" targetType="figure">
<paragraph id="E050EB2D6B781D4C03DCD584F6CFF83B" blockId="12.[241,657,1971,1987]" box="[241,657,1971,1987]" pageId="12" pageNumber="531">
<emphasis id="D29B373F6B781D4C03DCD584F577F83B" bold="true" box="[241,297,1972,1986]" pageId="12" pageNumber="531">Fig. 16.</emphasis>
Documented locations of
<taxonomicName id="27EF90AE6B781D4C02C0D583F6D0F83B" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" box="[493,654,1971,1987]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="531" phylum="Tracheophyta" rank="species" species="crassiculmis">Schoenus crassiculmis</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="A8F5B8A66B791D4D0357D3D9F57FFE04" box="[122,289,489,509]" pageId="13" pageNumber="532" type="biology_ecology">
<paragraph id="E050EB2D6B791D4D0357D3D9F57FFE04" blockId="13.[90,760,154,1402]" box="[122,289,489,509]" pageId="13" pageNumber="532">
<emphasis id="D29B373F6B791D4D0357D3D9F484FE04" box="[122,218,489,509]" italics="true" pageId="13" pageNumber="532">Flowering:</emphasis>
August
</paragraph>
</subSubSection>
<subSubSection id="A8F5B8A66B791D4D0357D035F615FDA8" pageId="13" pageNumber="532" type="distribution">
<paragraph id="E050EB2D6B791D4D0357D035F615FDA8" blockId="13.[90,760,154,1402]" pageId="13" pageNumber="532">
<emphasis id="D29B373F6B791D4D0357D035F662FDE0" box="[122,572,517,537]" italics="true" pageId="13" pageNumber="532">
Distribution and ecology:
<taxonomicName id="27EF90AE6B791D4D0242D035F662FDE0" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" box="[367,572,517,537]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="13" pageNumber="532" phylum="Tracheophyta" rank="species" species="quartziticus">Schoenus quartziticus</taxonomicName>
</emphasis>
has been collected from the Agulhas Plain region of
<collectingCountry id="98F8ABBD6B791D4D02B2D012F649FDCC" box="[415,535,546,565]" name="South Africa" pageId="13" pageNumber="532">South Africa</collectingCountry>
(
<figureCitation id="78D4F7A86B791D4D010BD012F638FDCC" box="[550,614,546,565]" captionStart="Fig" captionStartId="13.[219,244,1972,1986]" captionTargetBox="[122,1441,1441,1933]" captionTargetPageId="13" captionText="Fig. 17. Documented locations of Schoenus quartziticus." figureDoi="http://doi.org/10.5281/zenodo.10496630" httpUri="https://zenodo.org/record/10496630/files/figure.png" pageId="13" pageNumber="532">Fig. 17</figureCitation>
), appearing to be endemic to quartz patches on shale-derived soils.
</paragraph>
</subSubSection>
<subSubSection id="A8F5B8A66B791D4D0357D068F63FFD25" pageId="13" pageNumber="532" type="diagnosis">
<paragraph id="E050EB2D6B791D4D0357D068F63FFD25" blockId="13.[90,760,154,1402]" pageId="13" pageNumber="532">
<emphasis id="D29B373F6B791D4D0357D068F489FD95" box="[122,215,600,620]" italics="true" pageId="13" pageNumber="532">Diagnosis:</emphasis>
The lanceolate spikelets of
<taxonomicName id="27EF90AE6B791D4D02F0D068F602FD95" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" box="[477,604,600,620]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="13" pageNumber="532" phylum="Tracheophyta" rank="species" species="quartziticus">
<emphasis id="D29B373F6B791D4D02F0D068F602FD95" box="[477,604,600,620]" italics="true" pageId="13" pageNumber="532">S. quartziticus</emphasis>
</taxonomicName>
often have relatively thin, membranaceous glumes (
<figureCitation id="78D4F7A86B791D4D0290D045F65FFD70" box="[445,513,629,649]" captionStart="Fig" captionStartId="4.[113,138,983,997]" captionTargetBox="[275,1336,143,946]" captionTargetId="figure-478@4.[274,1336,142,947]" captionTargetPageId="4" captionText="Fig.4. Examples of plants from six species of the Schoenus cuspidatus andallies group found in southern Africa,showing the corresponding inflorescences and spikelets beneath each plant: A,G and M—S.crassiculmis; B,H and N—S.quartziticus, C,I andO—S.limosus; D,J,and P—S.purpurascens; E,K andQ—S.prophyllus; and F,L and R—S.calceolus. The black scale bar adjacent to the inflorescences in G—L represents 10 mm, while the scale bars adjacent to the spikelets in M—R represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10496604" httpUri="https://zenodo.org/record/10496604/files/figure.png" pageId="13" pageNumber="532">Fig. 4N</figureCitation>
) and somewhat resemble those of the more robust form of
<taxonomicName id="27EF90AE6B791D4D02B2D0A0F5BDFD5D" box="[415,483,656,676]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="13" pageNumber="532" phylum="Tracheophyta" rank="species" species="exilis">
<emphasis id="D29B373F6B791D4D02B2D0A0F5BDFD5D" box="[415,483,656,676]" italics="true" pageId="13" pageNumber="532">S. exilis</emphasis>
</taxonomicName>
in size and shape. However, the nutlets of the two species differ, with those of
<taxonomicName id="27EF90AE6B791D4D0111D09CF6E3FD39" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" box="[572,701,684,704]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="13" pageNumber="532" phylum="Tracheophyta" rank="species" species="quartziticus">
<emphasis id="D29B373F6B791D4D0111D09CF6E3FD39" box="[572,701,684,704]" italics="true" pageId="13" pageNumber="532">S. quartziticus</emphasis>
</taxonomicName>
being broad elliptic, while
<taxonomicName id="27EF90AE6B791D4D0236D0F8F500FD25" box="[283,350,712,732]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="13" pageNumber="532" phylum="Tracheophyta" rank="species" species="exilis">
<emphasis id="D29B373F6B791D4D0236D0F8F500FD25" box="[283,350,712,732]" italics="true" pageId="13" pageNumber="532">S. exilis</emphasis>
</taxonomicName>
has narrow elliptic nutlets.
</paragraph>
</subSubSection>
<subSubSection id="A8F5B8A66B791D4D0357D0D5F5F4FD01" box="[122,426,741,760]" pageId="13" pageNumber="532" type="multiple">
<paragraph id="E050EB2D6B791D4D0357D0D5F5F4FD01" blockId="13.[90,760,154,1402]" box="[122,426,741,760]" pageId="13" pageNumber="532">Additional collections examined</paragraph>
</subSubSection>
<subSubSection id="A8F5B8A66B791D4D0357D131F696FBBE" pageId="13" pageNumber="532" type="materials_examined">
<paragraph id="E050EB2D6B791D4D0357D131F696FBBE" blockId="13.[90,760,154,1402]" pageId="13" pageNumber="532">
<materialsCitation id="5087E1706B791D4D0357D131F6AAFCC9" collectingDate="2011-12-07" collectionCode="BOL" collectorName="Muasya &amp; Stirton &amp; Curtis" country="South Africa" county="Bredasdorp" location="Farm" municipality="Plaatjieskraal" pageId="13" pageNumber="532" specimenCount="1" stateProvince="WESTERN CAPE">
<collectingCountry id="98F8ABBD6B791D4D0357D131F4AAFCED" box="[122,244,769,788]" name="South Africa" pageId="13" pageNumber="532">South Africa</collectingCountry>
.
<collectingRegion id="222B25CF6B791D4D03D0D131F5C2FCED" box="[253,412,769,788]" country="South Africa" name="Western Cape" pageId="13" pageNumber="532">WESTERN CAPE</collectingRegion>
:
<emphasis id="D29B373F6B791D4D0288D131F634FCED" bold="true" box="[421,618,769,788]" pageId="13" pageNumber="532">
3420 (
<collectingCounty id="093193A16B791D4D02CFD131F601FCED" box="[482,607,769,788]" pageId="13" pageNumber="532">Bredasdorp</collectingCounty>
):
</emphasis>
<collectingMunicipality id="003471576B791D4D015CD131F6AAFCED" box="[625,756,769,788]" pageId="13" pageNumber="532">Plaatjieskraal</collectingMunicipality>
,
<location id="E530BDF66B791D4D0377D12DF4D2FCC9" LSID="urn:lsid:plazi:treatment:68465A3B6B781D4D0071D789F696FBBE:E530BDF66B791D4D0377D12DF4D2FCC9" box="[90,140,797,816]" country="South Africa" county="Bredasdorp" municipality="Plaatjieskraal" name="Farm" pageId="13" pageNumber="532" stateProvince="WESTERN CAPE">Farm</location>
(–AD),
<date id="9451CDED6B791D4D03C7D12DF53BFCC9" box="[234,357,797,816]" pageId="13" pageNumber="532" value="2011-12-07">
<collectingDate id="841534056B791D4D03C7D12DF53BFCC9" box="[234,357,797,816]" pageId="13" pageNumber="532" value="2011-12-07">7 Dec 2011</collectingDate>
</date>
,
<emphasis id="D29B373F6B791D4D0258D12CF5E1FCC9" box="[373,447,796,816]" italics="true" pageId="13" pageNumber="532">
<collectorName id="4D1A8EFB6B791D4D0258D12CF5E1FCC9" box="[373,447,796,816]" pageId="13" pageNumber="532">Muasya</collectorName>
</emphasis>
,
<emphasis id="D29B373F6B791D4D02FDD12CF6F4FCC9" box="[464,682,796,816]" italics="true" pageId="13" pageNumber="532">
<collectorName id="4D1A8EFB6B791D4D02FDD12CF651FCC9" box="[464,527,796,816]" pageId="13" pageNumber="532">Stirton</collectorName>
&amp;
<collectorName id="4D1A8EFB6B791D4D0115D12CF631FCC9" box="[568,623,796,816]" pageId="13" pageNumber="532">Curtis</collectorName>
6348
</emphasis>
(
<collectionCode id="86FE73E86B791D4D0190D12DF6B0FCC9" box="[701,750,797,816]" collectionName="BOL" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="532" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5087E1706B791D4D0377D109F6E9FC91" collectingDate="2016-10-01" collectionCode="BOL" collectorName="Elliott &amp; Muasya" country="South Africa" elevation="500" location="Haawegskloof Farm" municipality="Haawegskloof Farm" pageId="13" pageNumber="532" specimenCount="1" stateProvince="WESTERN CAPE">
<location id="E530BDF66B791D4D0377D109F544FCB5" LSID="urn:lsid:plazi:treatment:68465A3B6B781D4D0071D789F696FBBE:E530BDF66B791D4D0377D109F544FCB5" box="[90,282,825,844]" country="South Africa" municipality="Haawegskloof Farm" name="Haawegskloof Farm" pageId="13" pageNumber="532" stateProvince="WESTERN CAPE">Haawegskloof Farm</location>
,
<quantity id="271746C86B791D4D020ED109F53FFCB5" box="[291,353,825,844]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="532" unit="m" value="500.0">500 m</quantity>
along short-cut to reserve, W. of research centre, (–AD),
<date id="9451CDED6B791D4D03C0D165F506FC91" box="[237,344,853,872]" pageId="13" pageNumber="532" value="2016-10-01">
<collectingDate id="841534056B791D4D03C0D165F506FC91" box="[237,344,853,872]" pageId="13" pageNumber="532" value="2016-10-01">1 Oct 2016</collectingDate>
</date>
,
<emphasis id="D29B373F6B791D4D024FD164F629FC91" box="[354,631,852,872]" italics="true" pageId="13" pageNumber="532">
<collectorName id="4D1A8EFB6B791D4D024FD164F5C7FC91" box="[354,409,852,872]" pageId="13" pageNumber="532">Elliott</collectorName>
&amp;
<collectorName id="4D1A8EFB6B791D4D0298D164F5A0FC91" box="[437,510,852,872]" pageId="13" pageNumber="532">Muasya</collectorName>
TE2016_124
</emphasis>
(
<collectionCode id="86FE73E86B791D4D01AED165F6EFFC91" box="[643,689,853,872]" collectionName="BOL" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="532" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="5087E1706B791D4D01ECD164F521FC7D" collectionCode="BOL" collectorName="Elliott &amp; Muasya" country="South Africa" pageId="13" pageNumber="532" specimenCount="1" stateProvince="WESTERN CAPE">
<emphasis id="D29B373F6B791D4D01ECD164F565FC7A" italics="true" pageId="13" pageNumber="532">
<collectorName id="4D1A8EFB6B791D4D01ECD164F6A6FC91" box="[705,760,852,872]" pageId="13" pageNumber="532">Elliott</collectorName>
&amp;
<collectorName id="4D1A8EFB6B791D4D035ED15FF4E3FC7A" box="[115,189,879,899]" pageId="13" pageNumber="532">Muasya</collectorName>
TE2016_125
</emphasis>
(
<collectionCode id="86FE73E86B791D4D0267D141F527FC7D" box="[330,377,881,900]" collectionName="BOL" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="532" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="5087E1706B791D4D02A1D15FF6AAFC7D" box="[396,756,879,900]" collectionCode="BOL" collectorName="Elliott &amp; Muasya" country="South Africa" pageId="13" pageNumber="532" specimenCount="1" stateProvince="WESTERN CAPE">
<emphasis id="D29B373F6B791D4D02A1D15FF6F0FC7A" box="[396,686,879,899]" italics="true" pageId="13" pageNumber="532">
<collectorName id="4D1A8EFB6B791D4D02A1D15FF59DFC7A" box="[396,451,879,899]" pageId="13" pageNumber="532">Elliott</collectorName>
&amp;
<collectorName id="4D1A8EFB6B791D4D02CBD15FF66EFC7A" box="[486,560,879,899]" pageId="13" pageNumber="532">Muasya</collectorName>
TE2016_126
</emphasis>
(
<collectionCode id="86FE73E86B791D4D0190D141F6B0FC7D" box="[701,750,881,900]" collectionName="BOL" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="532" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5087E1706B791D4D0377D1BCF607FC42" collectingDate="2016-10-01" collectionCode="BOL" collectorName="Elliott &amp; Muasya" country="South Africa" elevation="750" location="Haawegskloof Farm" pageId="13" pageNumber="532" specimenCount="1" stateProvince="WESTERN CAPE">
<location id="E530BDF66B791D4D0377D1BCF540FC59" LSID="urn:lsid:plazi:treatment:68465A3B6B781D4D0071D789F696FBBE:E530BDF66B791D4D0377D1BCF540FC59" box="[90,286,908,928]" country="South Africa" name="Haawegskloof Farm" pageId="13" pageNumber="532" stateProvince="WESTERN CAPE">Haawegskloof Farm</location>
,
<quantity id="271746C86B791D4D0205D1BCF5FAFC59" box="[296,420,908,928]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="13" pageNumber="532" unit="m" value="750.0" valueMax="1000.0" valueMin="500.0">500–1000 m</quantity>
N.W. of research centre, (–AD),
<date id="9451CDED6B791D4D01C6D1BDF4E1FC42" pageId="13" pageNumber="532" value="2016-10-01">
<collectingDate id="841534056B791D4D01C6D1BDF4E1FC42" pageId="13" pageNumber="532" value="2016-10-01">1 Oct 2016</collectingDate>
</date>
,
<emphasis id="D29B373F6B791D4D03FED197F652FC42" box="[211,524,935,955]" italics="true" pageId="13" pageNumber="532">
<collectorName id="4D1A8EFB6B791D4D03FED197F555FC42" box="[211,267,935,955]" pageId="13" pageNumber="532">Elliott</collectorName>
&amp;
<collectorName id="4D1A8EFB6B791D4D0211D197F5D8FC42" box="[316,390,935,955]" pageId="13" pageNumber="532">Muasya</collectorName>
TE2016_127
</emphasis>
(
<collectionCode id="86FE73E86B791D4D010ED198F60DFC42" box="[547,595,936,955]" collectionName="BOL" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="532" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5087E1706B791D4D0143D198F6AAFC2E" collectingDate="2017-08-26" collectionCode="BOL" collectorName="Muasya" country="South Africa" location="Haawegskloof Renosterveld Reserve" municipality="Haawegskloof Renosterveld Reserve" pageId="13" pageNumber="532" specimenCount="1" stateProvince="WESTERN CAPE">
<location id="E530BDF66B791D4D0143D198F56DFC21" LSID="urn:lsid:plazi:treatment:68465A3B6B781D4D0071D789F696FBBE:E530BDF66B791D4D0143D198F56DFC21" country="South Africa" municipality="Haawegskloof Renosterveld Reserve" name="Haawegskloof Renosterveld Reserve" pageId="13" pageNumber="532" stateProvince="WESTERN CAPE">Haawegskloof Renosterveld Reserve</location>
, (–AD),
<date id="9451CDED6B791D4D02BFD1F4F647FC2E" box="[402,537,964,983]" pageId="13" pageNumber="532" value="2017-08-26">
<collectingDate id="841534056B791D4D02BFD1F4F647FC2E" box="[402,537,964,983]" pageId="13" pageNumber="532" value="2017-08-26">26 Aug 2017</collectingDate>
</date>
,
<emphasis id="D29B373F6B791D4D0105D1F3F6F5FC2E" box="[552,683,963,983]" italics="true" pageId="13" pageNumber="532">
<collectorName id="4D1A8EFB6B791D4D0105D1F3F62CFC2E" box="[552,626,963,983]" pageId="13" pageNumber="532">Muasya</collectorName>
7540
</emphasis>
(
<collectionCode id="86FE73E86B791D4D0190D1F4F6B0FC2E" box="[701,750,964,983]" collectionName="BOL" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="532" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5087E1706B791D4D0377D1D0F6AAFC0A" box="[90,756,991,1011]" collectingDate="2017-11-26" collectionCode="BOL" collectorName="Stirton" country="South Africa" location="Farm" municipality="Haawegskloof" pageId="13" pageNumber="532" specimenCount="1" stateProvince="WESTERN CAPE">
<collectingMunicipality id="003471576B791D4D0377D1D0F4BBFC0A" box="[90,229,992,1011]" pageId="13" pageNumber="532">Haawegskloof</collectingMunicipality>
,
<location id="E530BDF66B791D4D03D8D1D0F579FC0A" LSID="urn:lsid:plazi:treatment:68465A3B6B781D4D0071D789F696FBBE:E530BDF66B791D4D03D8D1D0F579FC0A" box="[245,295,992,1011]" country="South Africa" municipality="Haawegskloof" name="Farm" pageId="13" pageNumber="532" stateProvince="WESTERN CAPE">Farm</location>
(–AD),
<date id="9451CDED6B791D4D02A8D1D0F64EFC0A" box="[389,528,992,1011]" pageId="13" pageNumber="532" value="2017-11-26">
<collectingDate id="841534056B791D4D02A8D1D0F64EFC0A" box="[389,528,992,1011]" pageId="13" pageNumber="532" value="2017-11-26">26 Nov 2017</collectingDate>
</date>
,
<emphasis id="D29B373F6B791D4D0132D1EFF6F4FC0A" box="[543,682,991,1011]" italics="true" pageId="13" pageNumber="532">
<collectorName id="4D1A8EFB6B791D4D0132D1EFF601FC0A" box="[543,607,991,1011]" pageId="13" pageNumber="532">Stirton</collectorName>
14,441
</emphasis>
(
<collectionCode id="86FE73E86B791D4D0190D1D0F6B0FC0A" box="[701,750,992,1011]" collectionName="BOL" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="532" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5087E1706B791D4D0377D1CCF5DDFBBE" collectingDate="2017-11-30" collectionCode="BOL" collectorName="Elliott &amp; Smith" country="South Africa" elevation="200" location="Haarwegskloof Renosterveld Reserve" pageId="13" pageNumber="532" specimenCount="1" stateProvince="WESTERN CAPE">
<location id="E530BDF66B791D4D0377D1CCF592FBF6" LSID="urn:lsid:plazi:treatment:68465A3B6B781D4D0071D789F696FBBE:E530BDF66B791D4D0377D1CCF592FBF6" box="[90,460,1020,1039]" country="South Africa" name="Haarwegskloof Renosterveld Reserve" pageId="13" pageNumber="532" stateProvince="WESTERN CAPE">Haarwegskloof Renosterveld Reserve</location>
, c.
<quantity id="271746C86B791D4D02C2D1CCF671FBF6" box="[495,559,1020,1039]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="532" unit="m" value="200.0">200 m</quantity>
from entrance gate into renosterveld, c.
<quantity id="271746C86B791D4D0237D628F527FBD2" box="[282,377,1048,1067]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="13" pageNumber="532" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50–100 m</quantity>
from fenceline: (–AD),
<date id="9451CDED6B791D4D0171D628F688FBD2" box="[604,726,1048,1067]" pageId="13" pageNumber="532" value="2017-11-30">
<collectingDate id="841534056B791D4D0171D628F688FBD2" box="[604,726,1048,1067]" pageId="13" pageNumber="532" value="2017-11-30">30 Nov 2017</collectingDate>
</date>
,
<emphasis id="D29B373F6B791D4D01F3D627F51AFBBE" italics="true" pageId="13" pageNumber="532">
<collectorName id="4D1A8EFB6B791D4D01F3D627F420FBBE" pageId="13" pageNumber="532">Elliott</collectorName>
&amp;
<collectorName id="4D1A8EFB6B791D4D03B4D603F490FBBE" box="[153,206,1075,1095]" pageId="13" pageNumber="532">Smith</collectorName>
TE2016_331
</emphasis>
(
<collectionCode id="86FE73E86B791D4D0262D604F522FBBE" box="[335,380,1076,1095]" collectionName="BOL" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="532" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="5087E1706B791D4D02A6D603F696FBBE" box="[395,712,1075,1095]" collectionCode="BOL" collectorName="Elliott &amp; Smith" country="South Africa" pageId="13" pageNumber="532" specimenCount="1" stateProvince="WESTERN CAPE">
<emphasis id="D29B373F6B791D4D02A6D603F6D8FBBE" box="[395,646,1075,1095]" italics="true" pageId="13" pageNumber="532">
<collectorName id="4D1A8EFB6B791D4D02A6D603F59EFBBE" box="[395,448,1075,1095]" pageId="13" pageNumber="532">Elliott</collectorName>
&amp;
<collectorName id="4D1A8EFB6B791D4D02F6D603F64EFBBE" box="[475,528,1075,1095]" pageId="13" pageNumber="532">Smith</collectorName>
TE2016_332
</emphasis>
(
<collectionCode id="86FE73E86B791D4D01BCD604F6E1FBBE" box="[657,703,1076,1095]" collectionName="BOL" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="532" type="Herbarium">BOL</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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