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<document id="E4FCB27FD8803BA70C33BC6DC2254373" ID-DOI="10.11646/zootaxa.4742.1.12" ID-GBIF-Dataset="dfca4816-9e5a-4629-a619-fe33b2324079" ID-ISSN="1175-5326" ID-Zenodo-Dep="3674407" ID-ZooBank="B4C86DC6-6213-4405-9F5B-47A2E0EC9C32" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582096829951" checkinUser="plazi" docAuthor="Miessen, Geoffrey, Uliana, Marco &amp; Keith, Denis" docDate="2020" docId="585687E3FFA1FFC7FB9FFA0A1DB2F9A0" docLanguage="en" docName="zootaxa.4742.1.12.pdf" docOrigin="Zootaxa 4742 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chilotrogus matejiceki Miessen, Uliana &amp; Keith 2020, new species" docType="treatment" docUuid="46973C58-9546-4BD6-872E-228027A68496" docUuidSource="ZooBank" docVersion="6" lastPageNumber="188" masterDocId="A46FFF9BFFA5FFC0FB08FF991F70FFDC" masterDocTitle="Review of the genus Chilotrogus Reitter (Coleoptera: Scarabaeidae: Melolonthinae Rhizotrogini) in Iran with description of two new species" masterLastPageNumber="193" masterPageNumber="181" pageNumber="185" updateTime="1698719186324" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="67DDBDB0B2FE0600B2F1C029A5D00F5B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3DBCDEBE3CACAFE6795DDBEE30A1FAEF">Review of the genus Chilotrogus Reitter (Coleoptera: Scarabaeidae: Melolonthinae Rhizotrogini) in Iran with description of two new species</mods:title>
</mods:titleInfo>
<mods:name id="4C00181E4371C26BC1DA2715A970AA83" type="personal">
<mods:role id="DE171EDB97ABCC50286EA9D378EC5D14">
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<mods:namePart id="67E0652AD39929772D2EE7A8A8B849AF">Miessen, Geoffrey</mods:namePart>
</mods:name>
<mods:name id="E44D0A3278B0F36B3E348B2CE3A98FC1" type="personal">
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<mods:roleTerm id="F588605572D087D4CC42C8ADDBAE94F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="06B8A5B8D57DF4CF6107BCCB74D7E6FC">Uliana, Marco</mods:namePart>
</mods:name>
<mods:name id="B16873E2CCD928DD332179E96B14A199" type="personal">
<mods:role id="A95A8DAF31BB8BD1984CF203E0F04182">
<mods:roleTerm id="D2B19FAA99AD917F727FCC768B7F3218">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="32A6D4F70F600EDBA23749717E986F64">Keith, Denis</mods:namePart>
</mods:name>
<mods:typeOfResource id="2440C1E3C1FC27A5691005C69D7D870A">text</mods:typeOfResource>
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<mods:title id="AF7A4B4557B987D7F7870F7BD5C6708E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7F04BE539FE6AFCEC268382E9A1618C8">
<mods:date id="80EF301883114F26FA227E9F3BB73EE3">2020</mods:date>
<mods:detail id="1416BFD980AC59954B7ED68FF1925CF9" type="pubDate">
<mods:number id="3A9A88A7419DEF03A80944971D2D9E0C">2020-02-19</mods:number>
</mods:detail>
<mods:detail id="FE8D6EDF949FEC7A51916FBDAF959E4E" type="volume">
<mods:number id="635C568479D4E924B046E7CB45A7D3FC">4742</mods:number>
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<mods:number id="362AD1B4F5037F522EDB1B3D6C77D72E">1</mods:number>
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<mods:classification id="313BE571ABF68D20DBF5B4B5270A7DD0">journal article</mods:classification>
<mods:identifier id="A80FF0849F958E03EA2AC752F54951F2" type="DOI">10.11646/zootaxa.4742.1.12</mods:identifier>
<mods:identifier id="1FFED6B57175D85EB277B2B08FC6ACAE" type="GBIF-Dataset">dfca4816-9e5a-4629-a619-fe33b2324079</mods:identifier>
<mods:identifier id="DED0501893786ED5B86EF32AC12C7C80" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CCF14C3654D4B4E1A1F53C1647DB9E33" type="Zenodo-Dep">3674407</mods:identifier>
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<treatment id="585687E3FFA1FFC7FB9FFA0A1DB2F9A0" ID-DOI="http://doi.org/10.5281/zenodo.3681155" ID-GBIF-Taxon="162163935" ID-Zenodo-Dep="3681155" LSID="urn:lsid:zoobank.org:act:46973C58-9546-4BD6-872E-228027A68496" httpUri="http://treatment.plazi.org/id/585687E3FFA1FFC7FB9FFA0A1DB2F9A0" lastPageId="7" lastPageNumber="188" pageId="4" pageNumber="185">
<subSubSection id="98E5657EFFA1FFC4FB9FFA0A1CF4FA72" box="[151,900,1427,1454]" pageId="4" pageNumber="185" type="nomenclature">
<paragraph id="D04036F5FFA1FFC4FB9FFA0A1CF4FA72" blockId="4.[151,900,1427,1490]" box="[151,900,1427,1454]" pageId="4" pageNumber="185">
<heading id="8B088199FFA1FFC4FB9FFA0A1CF4FA72" bold="true" box="[151,900,1427,1454]" fontSize="11" level="1" pageId="4" pageNumber="185" reason="1">
<emphasis id="E28BEAE7FFA1FFC4FB9FFA0A1CF4FA72" bold="true" box="[151,900,1427,1454]" pageId="4" pageNumber="185">
<taxonomicName id="17FF4D76FFA1FFC4FB9FFA0A1D97FA72" authority="Miessen, Uliana &amp; Keith" authorityName="Miessen, Uliana &amp; Keith" authorityYear="2020" box="[151,743,1427,1454]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="185" phylum="Arthropoda" rank="species" species="matejiceki" status="sp. nov.">
<emphasis id="E28BEAE7FFA1FFC4FB9FFA0A1EDAFA72" bold="true" box="[151,426,1427,1454]" italics="true" pageId="4" pageNumber="185">Chilotrogus matejiceki</emphasis>
Miessen, Uliana &amp; Keith
</taxonomicName>
,
<taxonomicNameLabel id="F9B8579CFFA1FFC4F9FCFA0D1CF4FA72" box="[756,900,1428,1454]" pageId="4" pageNumber="185" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="98E5657EFFA1FFC4FB9FFA211DC6FA0E" box="[151,694,1463,1490]" pageId="4" pageNumber="185" type="description">
<paragraph id="D04036F5FFA1FFC4FB9FFA211DC6FA0E" blockId="4.[151,900,1427,1490]" box="[151,694,1463,1490]" pageId="4" pageNumber="185">
(
<figureCitation id="48C42A70FFA1FFC4FB97FA211F9FFA0E" box="[159,239,1464,1490]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1933,1958]" captionTargetBox="[296,1284,278,1895]" captionTargetId="figure@2.[291,1295,270,1905]" captionTargetPageId="2" captionText="FIGURES 1–4. Male habitus of Iranian Chilotrogus Reitter, 1905. 1, Ch. panotrogoides Reitter, 1905, specimen from Iran: Sarbijan (Shinagara); 2, Ch. matejiceki Miessen, Uliana &amp; Keith, new species, paratype from Iran: Mijam; 3, Ch. farsensis Keith, 2003, specimen from 40 Km NW Shiraz, Sangar; 4, Ch. schoolmeestersi Miessen, Uliana &amp; Keith, new species, holotype." figureDoi="http://doi.org/10.5281/zenodo.3674409" httpUri="https://zenodo.org/record/3674409/files/figure.png" pageId="4" pageNumber="185">Figs. 2</figureCitation>
,
<figureCitation id="48C42A70FFA1FFC4FBF3FA2E1E7BFA0E" box="[251,267,1463,1490]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1266,1291]" captionTargetBox="[151,1436,194,1241]" captionTargetId="figure@3.[151,1436,181,1241]" captionTargetPageId="3" captionText="FIGURES 5–8. Type specimens of Chilotrogus Reitter, 1905. 5, Ch. matejiceki Miessen, Uliana &amp; Keith, new species, holotype and labels; 6, Ch. panotrogoides Reitter, 1905, holotype and labels; 7, Ch. panotrogoides, labels of false allotype in MHNG; 8, Ch. panotrogoides, labels of false paratype in MHNG." figureDoi="http://doi.org/10.5281/zenodo.3674411" httpUri="https://zenodo.org/record/3674411/files/figure.png" pageId="4" pageNumber="185">5</figureCitation>
,
<figureCitation id="48C42A70FFA1FFC4FA1FFA211E10FA0E" box="[279,352,1464,1490]" captionStart="FIGURES 14–18" captionStartId="5.[151,264,937,962]" captionTargetBox="[159,1431,191,899]" captionTargetId="figure@5.[151,1436,181,911]" captionTargetPageId="5" captionText="FIGURES 14–18. Male genitalia of Ch. matejiceki Miessen, Uliana &amp; Keith, new species, holotype. 14, aedeagus (endophallic sclerite dissected), right lateral view; 15, same, dorsal view; 16, same, apical view; 17: endophallic sclerite, dorsal view; 18, same, right lateral view." figureDoi="http://doi.org/10.5281/zenodo.3674415" httpUri="https://zenodo.org/record/3674415/files/figure.png" pageId="4" pageNumber="185">14–18</figureCitation>
,
<figureCitation id="48C42A70FFA1FFC4FA63FA211EC4FA0E" box="[363,436,1464,1490]" captionStart="FIGURES 19–22" captionStartId="5.[151,264,1730,1755]" captionTargetBox="[157,1393,1070,1669]" captionTargetId="figure@5.[157,1430,1053,1705]" captionTargetPageId="5" captionText="FIGURES 19–22. Paramera of Ch. panotrogoides Reitter, 1905 (19, 20) and Ch. matejiceki Miessen, Uliana &amp; Keith, new species (21, 22), extremes of variability, left inclined lateral view. 19, specimen from Iran: Sarbijan (Shinagara); 20, specimen from Iran: Kerman (holotype); 21, specimen from Iran: Deh Bakhri (holotype); 22, specimen from Iran: Mijam (paratype)." figureDoi="http://doi.org/10.5281/zenodo.3674417" httpUri="https://zenodo.org/record/3674417/files/figure.png" pageId="4" pageNumber="185">21, 22</figureCitation>
,
<figureCitation id="48C42A70FFA1FFC4FAB7FA2E1D77FA0E" box="[447,519,1463,1490]" captionStart="FIGURES 23–26" captionStartId="6.[151,264,1866,1891]" captionTargetBox="[177,1410,189,1828]" captionTargetId="figure@6.[177,1410,181,1840]" captionTargetPageId="6" captionText="FIGURES 23–26. Endophallic sclerite of Ch. panotrogoides Reitter, 1905 (23, 24) and Ch. matejiceki Miessen, Uliana &amp; Keith, new species (25, 26), extremes of variability. 23, specimen from Iran: Kerman (holotype); 24, specimen from Iran: Sarbijan (Shinagara); 25, specimen from Iran: Deh Bakhri (holotype); 26, specimen from Iran: Mijam (paratype). a, left lateral view; b, inclined lateral view; c, frontal view, apex of internal tubular structure in evidence. Red lines indicate the boundary between sclerotized and soft tissue on the apical meatus." figureDoi="http://doi.org/10.5281/zenodo.3674419" httpUri="https://zenodo.org/record/3674419/files/figure.png" pageId="4" pageNumber="185">25, 26</figureCitation>
,
<figureCitation id="48C42A70FFA1FFC4F91BFA211D2BFA0E" box="[531,603,1464,1490]" captionStart="FIGURES 27–30" captionStartId="7.[151,264,627,652]" captionTargetBox="[152,1436,181,601]" captionTargetId="figure@7.[152,1436,181,601]" captionTargetPageId="7" captionText="FIGURES 27–30. Female genitalia of Ch. panotrogoides Reitter, 1905 (27, 28) and Ch. matejiceki Miessen, Uliana &amp; Keith, new species (29, 30). 27, 28, specimens from Iran: Sirjan; 29, specimen from Iran: Deh Bakhri (allotype); 30, specimen from Iran: Deh Bakhri (paratype)." figureDoi="http://doi.org/10.5281/zenodo.3674421" httpUri="https://zenodo.org/record/3674421/files/figure.png" pageId="4" pageNumber="185">29, 30</figureCitation>
,
<figureCitation id="48C42A70FFA1FFC4F96FFA2E1DF5FA0E" box="[615,645,1463,1490]" captionStart="FIGURES 45, 46" captionStartId="10.[151,264,1848,1873]" captionTargetBox="[151,1436,1332,1822]" captionTargetId="figure@10.[151,1436,1332,1822]" captionTargetPageId="10" captionText="FIGURES 45, 46. Detail of pronotal punctures. 45, Ch. matejiceki Miessen, Uliana &amp; Keith, new species, paratype from Iran: Mijam; 46, Ch. farsensis Keith, 2003, specimen from Iran: 15 Km N Vazag, Kuh-e-Dinar." figureDoi="http://doi.org/10.5281/zenodo.3674429" httpUri="https://zenodo.org/record/3674429/files/figure.png" pageId="4" pageNumber="185">45</figureCitation>
,
<figureCitation id="48C42A70FFA1FFC4F999FA211DDEFA0E" box="[657,686,1464,1490]" captionStart="FIGURE 47" captionStartId="11.[151,250,1655,1680]" captionTargetBox="[175,1412,669,1631]" captionTargetId="figure@11.[175,1412,669,1631]" captionTargetPageId="11" captionText="FIGURE 47. Known distribution of Iranian Chilotrogus Reitter, 1905, based on verified records." figureDoi="http://doi.org/10.5281/zenodo.3674431" httpUri="https://zenodo.org/record/3674431/files/figure.png" pageId="4" pageNumber="185">47</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="98E5657EFFA1FFC4FB9FF9991DF4F92E" pageId="4" pageNumber="185" type="materials_examined">
<paragraph id="D04036F5FFA1FFC4FB9FF9991DF4F92E" blockId="4.[151,1437,1535,2030]" pageId="4" pageNumber="185">
<emphasis id="E28BEAE7FFA1FFC4FB9FF9991E31F9C6" bold="true" box="[151,321,1535,1562]" pageId="4" pageNumber="185">Type material</emphasis>
(
<specimenCount id="C6F9FD7CFFA1FFC4FA5CF9991EFDF9C6" box="[340,397,1536,1562]" pageId="4" pageNumber="185" type="male">9♂♂</specimenCount>
and
<specimenCount id="C6F9FD7CFFA1FFC4FAC1F9991D72F9C6" box="[457,514,1536,1562]" pageId="4" pageNumber="185" type="female">3♀♀</specimenCount>
).
<typeStatus id="0F448857FFA1FFC4F914F9991DF7F9C6" box="[540,647,1536,1562]" pageId="4" pageNumber="185" type="holotype">Holotype</typeStatus>
,
<materialsCitation id="60973CA8FFA1FFC4F99DF9991C25F9BE" ID-GBIF-Occurrence="2566071128" collectingDate="1973-05-03" collectionCode="NMPC" collectorName="E. Iran" determinerName="D Keith" elevation="1725" pageId="4" pageNumber="185" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="C6F9FD7CFFA1FFC4F99DF9991DDAF9C6" box="[661,682,1536,1562]" pageId="4" pageNumber="185" type="male">♂</specimenCount>
(
<collectionCode id="B6EEAE30FFA1FFC4F9B4F9991C7FF9C6" box="[700,783,1536,1562]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="185">NMPC</collectionCode>
) labelled: “
<collectorName id="7D0A5323FFA1FFC4F89FF9991C9AF9C6" box="[919,1002,1536,1562]" pageId="4" pageNumber="185">E. Iran</collectorName>
, Deh Bakhri |
<quantity id="17079B10FFA1FFC4FF9FFA661A33F9C6" box="[1175,1347,1535,1562]" metricMagnitude="3" metricUnit="m" metricValue="1.725" metricValueMax="1.75" metricValueMin="1.7" pageId="4" pageNumber="185" unit="m" value="1725.0" valueMax="1750.0" valueMin="1700.0">
<elevation id="5BD2D1C6FFA1FFC4FF9FFA661A33F9C6" box="[1175,1347,1535,1562]" metricMagnitude="3" metricUnit="m" metricValue="1.725" metricValueMax="1.75" metricValueMin="1.7" pageId="4" pageNumber="185" unit="m" value="1725.0" valueMax="1750.0" valueMin="1700.0">1700—1750 m</elevation>
</quantity>
| 30.4–
<date id="A4411035FFA1FFC4FB9FF9BA1F89F9E2" box="[151,249,1571,1598]" pageId="4" pageNumber="185" value="1973-05-03">
<collectingDate id="B405E9DDFFA1FFC4FB9FF9BA1F89F9E2" box="[151,249,1571,1598]" pageId="4" pageNumber="185" value="1973-05-03">3.5.1973</collectingDate>
</date>
[printed] || Loc. no. 186 | Exp. Nat. Mus. | Praha [printed] ||
<taxonomicName id="17FF4D76FFA1FFC4F8B7F9BD1B33F9E2" authorityName="Reitter" authorityYear="1905" box="[959,1091,1572,1598]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="185" phylum="Arthropoda" rank="genus">Chilotrogus</taxonomicName>
[handwritten] | panotrogoides [handwritten] | Rtt. [handwritten]
<determinerName id="259CDA31FFA1FFC4F905F9D11D15F9BE" box="[525,613,1608,1634]" pageId="4" pageNumber="185">D Keith</determinerName>
det [20]07 [printed]”
</materialsCitation>
.
<materialsCitation id="60973CA8FFA1FFC4F856F9D11AE8F9BE" ID-GBIF-Occurrence="2566071064" box="[862,1432,1608,1634]" collectingDate="1973-05-03" collectionCode="CRS, CGM" collectorName="E. Iran" determinerName="D Keith" elevation="1725" pageId="4" pageNumber="185" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="0F448857FFA1FFC4F856F9D11CBFF9BE" box="[862,975,1608,1634]" pageId="4" pageNumber="185" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="C6F9FD7CFFA1FFC4F8D1F9D11C8CF9BE" box="[985,1020,1608,1634]" pageId="4" pageNumber="185" type="male">1♂</specimenCount>
(
<collectionCode id="B6EEAE30FFA1FFC4FF01F9D11B32F9BE" box="[1033,1090,1608,1634]" pageId="4" pageNumber="185">CRS</collectionCode>
,
<collectionCode id="B6EEAE30FFA1FFC4FF44F9D11BE0F9BE" box="[1100,1168,1608,1634]" pageId="4" pageNumber="185">CGM</collectionCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="60973CA8FFA1FFC4FB9FF9F51D29F976" ID-GBIF-Occurrence="2566071113" collectingDate="2014-05-28" collectionCode="CMU, CJM" collectorName="Cerny Fr." elevation="1650" latitude="28.874722" location="Jiroft" longLatPrecision="20" longitude="57.883614" municipality="Mijam" pageId="4" pageNumber="185" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Kerman" typeStatus="paratype">
<specimenCount id="C6F9FD7CFFA1FFC4FB9FF9F51FBFF95A" box="[151,207,1644,1670]" pageId="4" pageNumber="185" type="male">4♂♂</specimenCount>
and
<specimenCount id="C6F9FD7CFFA1FFC4FA0EF9F51E59F95A" box="[262,297,1644,1670]" pageId="4" pageNumber="185" type="female">1♀</specimenCount>
(
<collectionCode id="B6EEAE30FFA1FFC4FA30F9F51E0CF95A" box="[312,380,1644,1670]" country="Thailand" lsid="urn:lsid:biocol.org:col:12797" name="Chiang Mai University" pageId="4" pageNumber="185" type="Herbarium">CMU</collectionCode>
,
<collectionCode id="B6EEAE30FFA1FFC4FA80F9F51EB4F95A" box="[392,452,1644,1670]" pageId="4" pageNumber="185">CJM</collectionCode>
): “dat. 27-
<date id="A4411035FFA1FFC4F93EF9F21DD9F95A" box="[566,681,1643,1670]" pageId="4" pageNumber="185" value="2014-05-28">
<collectingDate id="B405E9DDFFA1FFC4F93EF9F21DD9F95A" box="[566,681,1643,1670]" pageId="4" pageNumber="185" value="2014-05-28">28.5.2014</collectingDate>
</date>
Irán IR |
<collectingRegion id="123BF817FFA1FFC4F81DF9F51CDFF95A" box="[789,943,1644,1670]" country="Iran" name="Kerman" pageId="4" pageNumber="185">Kerman prov.</collectingRegion>
|
<collectingMunicipality id="3024AC8FFFA1FFC4F8CBF9F51B7EF95A" box="[963,1038,1644,1670]" pageId="4" pageNumber="185">Mijam</collectingMunicipality>
(
<location id="D520602EFFA1FFC4FF15F9F51B2CF95B" LSID="urn:lsid:plazi:treatment:585687E3FFA1FFC7FB9FFA0A1DB2F9A0:D520602EFFA1FFC4FF15F9F51B2CF95B" box="[1053,1116,1644,1671]" latitude="28.874722" longLatPrecision="20" longitude="57.883614" municipality="Mijam" name="Jiroft" pageId="4" pageNumber="185" stateProvince="Kerman">Jiroft</location>
) |
<geoCoordinate id="B5CB5032FFA1FFC4FF7FF9F51A78F95A" box="[1143,1288,1643,1670]" degrees="28" direction="north" minutes="52" orientation="latitude" pageId="4" pageNumber="185" precision="15" seconds="29" value="28.874722">N 28°52’29”</geoCoordinate>
<geoCoordinate id="B5CB5032FFA1FFC4FE07F9F51AECF95A" box="[1295,1436,1643,1670]" degrees="57" direction="east" minutes="53" orientation="longitude" pageId="4" pageNumber="185" precision="15" seconds="01" value="57.883614">E 57°53’01”</geoCoordinate>
| leg.
<collectorName id="7D0A5323FFA1FFC4FBDEF9161E34F976" box="[214,324,1679,1706]" pageId="4" pageNumber="185">Černý Fr.</collectorName>
,
<quantity id="17079B10FFA1FFC4FA58F9161EDAF976" box="[336,426,1679,1706]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="4" pageNumber="185" unit="m" value="1650.0">
<elevation id="5BD2D1C6FFA1FFC4FA58F9161EDAF976" box="[336,426,1679,1706]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="4" pageNumber="185" unit="m" value="1650.0">1650 m</elevation>
</quantity>
.n.m. [printed]”
</materialsCitation>
;
<materialsCitation id="60973CA8FFA1FFC4F96DF9091C33F912" ID-GBIF-Occurrence="2566071098" collectingDate="1997-05-31" collectionCode="CRS, CGM, V" country="Iran" elevation="2300" latitude="28.983334" location="Deh Bakhri" longLatPrecision="1229" longitude="57.916668" pageId="4" pageNumber="185" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Kerman" typeStatus="paratype">
<specimenCount id="C6F9FD7CFFA1FFC4F96DF9091DEEF976" box="[613,670,1680,1706]" pageId="4" pageNumber="185" type="male">2♂♂</specimenCount>
and
<specimenCount id="C6F9FD7CFFA1FFC4F9DEF9091D89F976" box="[726,761,1680,1706]" pageId="4" pageNumber="185" type="female">1♀</specimenCount>
(
<collectionCode id="B6EEAE30FFA1FFC4F801F9091C32F976" box="[777,834,1680,1706]" pageId="4" pageNumber="185">CRS</collectionCode>
,
<collectionCode id="B6EEAE30FFA1FFC4F847F9091CE4F976" box="[847,916,1680,1706]" pageId="4" pageNumber="185">CGM</collectionCode>
): “
<collectingCountry id="A8E87665FFA1FFC4F8BAF9091C8CF976" box="[946,1020,1680,1706]" name="Iran" pageId="4" pageNumber="185">IRAN</collectingCountry>
,
<collectingRegion id="123BF817FFA1FFC4FF00F9091BD3F976" box="[1032,1187,1680,1706]" country="Iran" name="Kerman" pageId="4" pageNumber="185">Kerman prov.</collectingRegion>
|
<quantity id="17079B10FFA1FFC4FFB0F9161B83F976" box="[1208,1267,1679,1706]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="185" unit="km" value="5.0">5 km</quantity>
S.
<location id="D520602EFFA1FFC4FE11F9091AECF976" LSID="urn:lsid:plazi:treatment:585687E3FFA1FFC7FB9FFA0A1DB2F9A0:D520602EFFA1FFC4FE11F9091AECF976" box="[1305,1436,1680,1706]" country="Iran" latitude="28.983334" longLatPrecision="1229" longitude="57.916668" name="Deh Bakhri" pageId="4" pageNumber="185" stateProvince="Kerman">Deh Bakhri</location>
|
<geoCoordinate id="B5CB5032FFA1FFC4FBAFF92A1E78F912" box="[167,264,1715,1742]" degrees="28" direction="north" minutes="59" orientation="latitude" pageId="4" pageNumber="185" precision="925" value="28.983334">28°59’N</geoCoordinate>
<geoCoordinate id="B5CB5032FFA1FFC4FA1AF92A1E02F912" box="[274,370,1715,1742]" degrees="57" direction="east" minutes="55" orientation="longitude" pageId="4" pageNumber="185" precision="925" value="57.916668">57°55’E</geoCoordinate>
,
<quantity id="17079B10FFA1FFC4FA89F92D1EA9F912" box="[385,473,1716,1742]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="185" unit="m" value="2300.0">
<elevation id="5BD2D1C6FFA1FFC4FA89F92D1EA9F912" box="[385,473,1716,1742]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="185" unit="m" value="2300.0">2300 m</elevation>
</quantity>
|
<date id="A4411035FFA1FFC4FAFBF92D1D18F912" box="[499,616,1716,1742]" pageId="4" pageNumber="185" value="1997-05-31">
<collectingDate id="B405E9DDFFA1FFC4FAFBF92D1D18F912" box="[499,616,1716,1742]" pageId="4" pageNumber="185" value="1997-05-31">
31.
<collectionCode id="B6EEAE30FFA1FFC4F91AF92D1D58F912" box="[530,552,1716,1742]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="185" type="Museum">V</collectionCode>
.1997
</collectingDate>
</date>
, A. Hofmann lgt.”
</materialsCitation>
;
<materialsCitation id="60973CA8FFA1FFC4F85AF92D1DF0F92E" ID-GBIF-Occurrence="2566071066" collectingDate="1997-05-31" collectingDateMax="1997-06-01" collectingDateMin="1997-05-31" collectionCode="CRS, VI" collectorName="A. Hofmann &amp; P. Kautt" country="Iran" elevation="2300" latitude="28.983334" location="Deh Bakhri" longLatPrecision="1229" longitude="57.916668" pageId="4" pageNumber="185" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kerman" typeStatus="allotype">
<specimenCount id="C6F9FD7CFFA1FFC4F85AF92D1C05F912" box="[850,885,1716,1742]" pageId="4" pageNumber="185" type="male">1♂</specimenCount>
and
<specimenCount id="C6F9FD7CFFA1FFC4F8BAF92D1CA4F912" box="[946,980,1716,1742]" pageId="4" pageNumber="185" type="female">1♀</specimenCount>
(
<typeStatus id="0F448857FFA1FFC4F8EEF92D1B20F913" box="[998,1104,1716,1743]" pageId="4" pageNumber="185" type="allotype">allotypus</typeStatus>
) (
<collectionCode id="B6EEAE30FFA1FFC4FF63F92D1BD5F912" box="[1131,1189,1716,1742]" pageId="4" pageNumber="185">CRS</collectionCode>
), same data, but “
<date id="A4411035FFA1FFC4FE71F92D1E5EF92E" pageId="4" pageNumber="185" value="1997-05-31" valueMax="1997-06-01" valueMin="1997-05-31">
<collectingDate id="B405E9DDFFA1FFC4FE71F92D1E5EF92E" pageId="4" pageNumber="185" value="1997-05-31" valueMax="1997-06-01" valueMin="1997-05-31">
31. V–1.
<collectionCode id="B6EEAE30FFA1FFC4FBC4F9411F9DF92E" box="[204,237,1752,1778]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="4" pageNumber="185">VI</collectionCode>
.1997
</collectingDate>
</date>
, leg.
<collectorName id="7D0A5323FFA1FFC4FA60F9411E84F92E" box="[360,500,1752,1778]" pageId="4" pageNumber="185">A. Hofmann</collectorName>
&amp;
<collectorName id="7D0A5323FFA1FFC4F91FF9411D06F92E" box="[535,630,1752,1778]" pageId="4" pageNumber="185">P. Kautt</collectorName>
”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="98E5657EFFA1FFC7FBCFF9621A53FC28" lastPageId="7" lastPageNumber="188" pageId="4" pageNumber="185" type="description">
<paragraph id="D04036F5FFA1FFC4FBCFF9621EDBF85E" blockId="4.[151,1437,1535,2030]" pageId="4" pageNumber="185">
<emphasis id="E28BEAE7FFA1FFC4FBCFF9621D76F8CA" bold="true" box="[199,518,1787,1814]" pageId="4" pageNumber="185">
Description of the
<typeStatus id="0F448857FFA1FFC4FA97F9651D76F8CA" box="[415,518,1788,1814]" pageId="4" pageNumber="185" type="holotype">holotype</typeStatus>
</emphasis>
(
<specimenCount id="C6F9FD7CFFA1FFC4F91DF9651D5BF8CA" box="[533,555,1788,1814]" pageId="4" pageNumber="185" type="male">♂</specimenCount>
). Length 16.6 mm. Habitus dark brown and matt (
<figureCitation id="48C42A70FFA1FFC4FF5CF9651BE7F8CA" box="[1108,1175,1787,1814]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1266,1291]" captionTargetBox="[151,1436,194,1241]" captionTargetId="figure@3.[151,1436,181,1241]" captionTargetPageId="3" captionText="FIGURES 5–8. Type specimens of Chilotrogus Reitter, 1905. 5, Ch. matejiceki Miessen, Uliana &amp; Keith, new species, holotype and labels; 6, Ch. panotrogoides Reitter, 1905, holotype and labels; 7, Ch. panotrogoides, labels of false allotype in MHNG; 8, Ch. panotrogoides, labels of false paratype in MHNG." figureDoi="http://doi.org/10.5281/zenodo.3674411" httpUri="https://zenodo.org/record/3674411/files/figure.png" pageId="4" pageNumber="185">Fig. 5</figureCitation>
). Clypeus and forehead with large, strong and dense punctuation and traces of microscopic pilosity on the edges. Clypeus slightly larger than the distance between the eyes. Antennae with ten antennomeres, trimerous antennal club short, as long as funicle and scape together.
</paragraph>
<paragraph id="D04036F5FFA1FFC4FBCFF8151E06F832" blockId="4.[151,1437,1535,2030]" pageId="4" pageNumber="185">Pronotum glabrous with large, strong, dense and deep punctuation, the punctures slightly more spaced than those on clypeus and forehead. Lateral margins crenulate with thick setae, slightly shorter than those of epipleuron in the humeral area.</paragraph>
<caption id="8480667DFFA0FFC5FB9FFC301EFBFBD6" ID-DOI="http://doi.org/10.5281/zenodo.3674415" ID-Zenodo-Dep="3674415" httpUri="https://zenodo.org/record/3674415/files/figure.png" pageId="5" pageNumber="186" startId="5.[151,264,937,962]" targetBox="[159,1431,191,899]" targetPageId="5">
<paragraph id="D04036F5FFA0FFC5FB9FFC301EFBFBD6" blockId="5.[151,1437,937,1034]" pageId="5" pageNumber="186">
<emphasis id="E28BEAE7FFA0FFC5FB9FFC301E24FC1E" bold="true" box="[151,340,937,962]" pageId="5" pageNumber="186">FIGURES 14–18.</emphasis>
Male genitalia of
<taxonomicName id="17FF4D76FFA0FFC5F903FC301CD2FC1E" authority="Miessen, Uliana &amp; Keith" authorityName="Miessen, Uliana &amp; Keith" authorityYear="2020" box="[523,930,937,962]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="186" phylum="Arthropoda" rank="species" species="matejiceki" status="sp. nov.">
<emphasis id="E28BEAE7FFA0FFC5F903FC301DEDFC1D" box="[523,669,937,961]" italics="true" pageId="5" pageNumber="186">Ch. matejiceki</emphasis>
Miessen, Uliana &amp; Keith
</taxonomicName>
,
<taxonomicNameLabel id="F9B8579CFFA0FFC5F8A3FC331B57FC1E" box="[939,1063,937,962]" pageId="5" pageNumber="186" rank="species">new species</taxonomicNameLabel>
, holotype. 14, aedeagus (endophallic sclerite dissected), right lateral view; 15, same, dorsal view; 16, same, apical view; 17: endophallic sclerite, dorsal view; 18, same, right lateral view.
</paragraph>
</caption>
<caption id="8480667DFFA0FFC5FB9FF95B1A4CF8FE" ID-DOI="http://doi.org/10.5281/zenodo.3674417" ID-Zenodo-Dep="3674417" httpUri="https://zenodo.org/record/3674417/files/figure.png" pageId="5" pageNumber="186" startId="5.[151,264,1730,1755]" targetBox="[157,1393,1070,1669]" targetPageId="5">
<paragraph id="D04036F5FFA0FFC5FB9FF95B1A4CF8FE" blockId="5.[151,1436,1730,1827]" pageId="5" pageNumber="186">
<emphasis id="E28BEAE7FFA0FFC5FB9FF95B1E24F907" bold="true" box="[151,340,1730,1755]" pageId="5" pageNumber="186">FIGURES 19–22.</emphasis>
Paramera of
<taxonomicName id="17FF4D76FFA0FFC5FAD1F95B1C6DF907" authority="Reitter, 1905" authorityName="Reitter" authorityYear="1905" box="[473,797,1730,1755]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="186" phylum="Arthropoda" rank="species" species="panotrogoides">
<emphasis id="E28BEAE7FFA0FFC5FAD1F95B1DE4F906" box="[473,660,1730,1754]" italics="true" pageId="5" pageNumber="186">Ch. panotrogoides</emphasis>
<bibRefCitation id="B46E4B04FFA0FFC5F992F95B1C6DF907" author="Reitter, E." box="[666,797,1730,1755]" pageId="5" pageNumber="186" pagination="201 - 206" refId="ref7163" refString="Reitter, E. (1905) Neun neue Coleopteren aus der palaearktischen Fauna. Wiener Entomologische Zeitung, 24, 201 - 206. https: // doi. org / 10.5962 / bhl. part. 5395" type="journal article" year="1905">Reitter, 1905</bibRefCitation>
</taxonomicName>
(19, 20) and
<taxonomicName id="17FF4D76FFA0FFC5F8ABF95B1A49F907" authority="Miessen, Uliana &amp; Keith" authorityName="Miessen, Uliana &amp; Keith" authorityYear="2020" box="[931,1337,1730,1755]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="186" phylum="Arthropoda" rank="species" species="matejiceki" status="sp. nov.">
<emphasis id="E28BEAE7FFA0FFC5F8ABF95B1B44F906" box="[931,1076,1730,1754]" italics="true" pageId="5" pageNumber="186">Ch. matejiceki</emphasis>
Miessen, Uliana &amp; Keith
</taxonomicName>
,
<taxonomicNameLabel id="F9B8579CFFA0FFC5FE4AF95A1FCEF923" pageId="5" pageNumber="186" rank="species">new species</taxonomicNameLabel>
(21, 22), extremes of variability, left inclined lateral view. 19, specimen from Iran: Sarbijan (Shinagara); 20, specimen from Iran: Kerman (holotype); 21, specimen from Iran: Deh Bakhri (holotype); 22, specimen from Iran: Mijam (paratype).
</paragraph>
</caption>
<paragraph id="D04036F5FFA0FFC5FBCFF8C81A54F853" blockId="5.[151,1436,1873,2043]" pageId="5" pageNumber="186">Elytra matt, pruinose with coarse punctuation, sparser on the interstriae. Pilosity very short and sparse, distributed over the entire surface. Epipleuron with long and thick pilosity, gradually shortening towards apex.</paragraph>
<paragraph id="D04036F5FFA0FFC5FBCFF8001C21F827" blockId="5.[151,1436,1873,2043]" pageId="5" pageNumber="186">Protibiae tridentate, the basal tooth at the middle, the median tooth slightly closer to the apical. Inner spur inserted slightly behind the median tooth. Metatarsi elongate, 1.8 times longer than metatibiae which are dented on upper face. At the base of each dent is one thick and long seta.</paragraph>
<caption id="8480667DFFA3FFC6FB9FF8D31D09F82E" ID-DOI="http://doi.org/10.5281/zenodo.3674419" ID-Zenodo-Dep="3674419" httpUri="https://zenodo.org/record/3674419/files/figure.png" pageId="6" pageNumber="187" startId="6.[151,264,1866,1891]" targetBox="[177,1410,189,1828]" targetPageId="6">
<paragraph id="D04036F5FFA3FFC6FB9FF8D31D09F82E" blockId="6.[151,1436,1866,2035]" pageId="6" pageNumber="187">
<emphasis id="E28BEAE7FFA3FFC6FB9FF8D31E23F8BF" bold="true" box="[151,339,1866,1891]" pageId="6" pageNumber="187">FIGURES 23–26.</emphasis>
Endophallic sclerite of
<taxonomicName id="17FF4D76FFA3FFC6F948F8D21CF2F8BF" authority="Reitter, 1905" authorityName="Reitter" authorityYear="1905" box="[576,898,1866,1891]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="187" phylum="Arthropoda" rank="species" species="panotrogoides">
<emphasis id="E28BEAE7FFA3FFC6F948F8D21D8AF8BF" box="[576,762,1867,1891]" italics="true" pageId="6" pageNumber="187">Ch. panotrogoides</emphasis>
<bibRefCitation id="B46E4B04FFA3FFC6F808F8D31CF2F8BF" author="Reitter, E." box="[768,898,1866,1891]" pageId="6" pageNumber="187" pagination="201 - 206" refId="ref7163" refString="Reitter, E. (1905) Neun neue Coleopteren aus der palaearktischen Fauna. Wiener Entomologische Zeitung, 24, 201 - 206. https: // doi. org / 10.5962 / bhl. part. 5395" type="journal article" year="1905">Reitter, 1905</bibRefCitation>
</taxonomicName>
(23, 24) and
<taxonomicName id="17FF4D76FFA3FFC6FF0DF8D21AE9F8BF" authority="Miessen, Uliana &amp; Keith" authorityName="Miessen, Uliana &amp; Keith" authorityYear="2020" box="[1029,1433,1866,1891]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="187" phylum="Arthropoda" rank="species" species="matejiceki" status="sp. nov.">
<emphasis id="E28BEAE7FFA3FFC6FF0DF8D21BE7F8BF" box="[1029,1175,1867,1891]" italics="true" pageId="6" pageNumber="187">Ch. matejiceki</emphasis>
Miessen, Uliana &amp; Keith
</taxonomicName>
,
<taxonomicNameLabel id="F9B8579CFFA3FFC6FB9FF8F61E63F85B" box="[151,275,1902,1927]" pageId="6" pageNumber="187" rank="species">new species</taxonomicNameLabel>
(25, 26), extremes of variability. 23, specimen from Iran: Kerman (holotype); 24, specimen from Iran: Sarbijan (Shinagara); 25, specimen from Iran: Deh Bakhri (holotype); 26, specimen from Iran: Mijam (paratype). a, left lateral view; b, inclined lateral view; c, frontal view, apex of internal tubular structure in evidence. Red lines indicate the boundary between sclerotized and soft tissue on the apical meatus.
</paragraph>
</caption>
<caption id="8480667DFFA2FFC7FB9FFDEA1EC9FD0F" ID-DOI="http://doi.org/10.5281/zenodo.3674421" ID-Zenodo-Dep="3674421" httpUri="https://zenodo.org/record/3674421/files/figure.png" pageId="7" pageNumber="188" startId="7.[151,264,627,652]" targetBox="[152,1436,181,601]" targetPageId="7">
<paragraph id="D04036F5FFA2FFC7FB9FFDEA1EC9FD0F" blockId="7.[151,1436,627,724]" pageId="7" pageNumber="188">
<emphasis id="E28BEAE7FFA2FFC7FB9FFDEA1E25FD50" bold="true" box="[151,341,627,652]" pageId="7" pageNumber="188">FIGURES 27–30.</emphasis>
Female genitalia of
<taxonomicName id="17FF4D76FFA2FFC7F920FDEA1C01FD50" authority="Reitter, 1905" authorityName="Reitter" authorityYear="1905" box="[552,881,627,652]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="panotrogoides">
<emphasis id="E28BEAE7FFA2FFC7F920FDEA1D3DFD57" box="[552,589,627,651]" italics="true" pageId="7" pageNumber="188">Ch.</emphasis>
<emphasis id="E28BEAE7FFA2FFC7F95CFDEA1D95FD57" box="[596,741,627,651]" italics="true" pageId="7" pageNumber="188">panotrogoides</emphasis>
<bibRefCitation id="B46E4B04FFA2FFC7F9E4FDEA1C01FD50" author="Reitter, E." box="[748,881,627,652]" pageId="7" pageNumber="188" pagination="201 - 206" refId="ref7163" refString="Reitter, E. (1905) Neun neue Coleopteren aus der palaearktischen Fauna. Wiener Entomologische Zeitung, 24, 201 - 206. https: // doi. org / 10.5962 / bhl. part. 5395" type="journal article" year="1905">Reitter, 1905</bibRefCitation>
</taxonomicName>
(27, 28) and
<taxonomicName id="17FF4D76FFA2FFC7F8F4FDEA1AE9FD50" authority="Miessen, Uliana &amp; Keith" authorityName="Miessen, Uliana &amp; Keith" authorityYear="2020" box="[1020,1433,627,652]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="matejiceki" status="sp. nov.">
<emphasis id="E28BEAE7FFA2FFC7F8F4FDEA1B50FD57" box="[1020,1056,627,651]" italics="true" pageId="7" pageNumber="188">Ch.</emphasis>
<emphasis id="E28BEAE7FFA2FFC7FF2FFDEA1BFFFD57" box="[1063,1167,627,651]" italics="true" pageId="7" pageNumber="188">matejiceki</emphasis>
Miessen, Uliana &amp; Keith
</taxonomicName>
,
<taxonomicNameLabel id="F9B8579CFFA2FFC7FB9FFD011E62FD6C" box="[151,274,663,688]" pageId="7" pageNumber="188" rank="species">new species</taxonomicNameLabel>
(29, 30). 27, 28, specimens from Iran: Sirjan; 29, specimen from Iran: Deh Bakhri (allotype); 30, specimen from Iran: Deh Bakhri (paratype).
</paragraph>
</caption>
<paragraph id="D04036F5FFA2FFC7FBCFFC9B1DD3FCC0" blockId="7.[151,1437,770,1660]" box="[199,675,770,796]" pageId="7" pageNumber="188">Pygidium with raised setae in lateral view.</paragraph>
<paragraph id="D04036F5FFA2FFC7FBCFFCBF1DEFFC54" blockId="7.[151,1437,770,1660]" pageId="7" pageNumber="188">
Aedeagus of characteristic shape; parameres (
<figureCitation id="48C42A70FFA2FFC7F9CAFCBF1C37FC9D" box="[706,839,806,833]" captionStart="FIGURES 14–18" captionStartId="5.[151,264,937,962]" captionTargetBox="[159,1431,191,899]" captionTargetId="figure@5.[151,1436,181,911]" captionTargetPageId="5" captionText="FIGURES 14–18. Male genitalia of Ch. matejiceki Miessen, Uliana &amp; Keith, new species, holotype. 14, aedeagus (endophallic sclerite dissected), right lateral view; 15, same, dorsal view; 16, same, apical view; 17: endophallic sclerite, dorsal view; 18, same, right lateral view." figureDoi="http://doi.org/10.5281/zenodo.3674415" httpUri="https://zenodo.org/record/3674415/files/figure.png" pageId="7" pageNumber="188">Figs. 14–16</figureCitation>
) in dorsal view with a marked angle at the level of the central opening. Internal piece broadly open dorsally, exposing an internal tubular structure that bears a backwards protruding spine at the apex (
<figureCitation id="48C42A70FFA2FFC7FAD7FCF71D16FC54" box="[479,614,878,904]" captionStart="FIGURES 14–18" captionStartId="5.[151,264,937,962]" captionTargetBox="[159,1431,191,899]" captionTargetId="figure@5.[151,1436,181,911]" captionTargetPageId="5" captionText="FIGURES 14–18. Male genitalia of Ch. matejiceki Miessen, Uliana &amp; Keith, new species, holotype. 14, aedeagus (endophallic sclerite dissected), right lateral view; 15, same, dorsal view; 16, same, apical view; 17: endophallic sclerite, dorsal view; 18, same, right lateral view." figureDoi="http://doi.org/10.5281/zenodo.3674415" httpUri="https://zenodo.org/record/3674415/files/figure.png" pageId="7" pageNumber="188">Figs. 17, 18</figureCitation>
,
<figureCitation id="48C42A70FFA2FFC7F97AFCF41DE2FC54" box="[626,658,877,904]" captionStart="FIGURES 23–26" captionStartId="6.[151,264,1866,1891]" captionTargetBox="[177,1410,189,1828]" captionTargetId="figure@6.[177,1410,181,1840]" captionTargetPageId="6" captionText="FIGURES 23–26. Endophallic sclerite of Ch. panotrogoides Reitter, 1905 (23, 24) and Ch. matejiceki Miessen, Uliana &amp; Keith, new species (25, 26), extremes of variability. 23, specimen from Iran: Kerman (holotype); 24, specimen from Iran: Sarbijan (Shinagara); 25, specimen from Iran: Deh Bakhri (holotype); 26, specimen from Iran: Mijam (paratype). a, left lateral view; b, inclined lateral view; c, frontal view, apex of internal tubular structure in evidence. Red lines indicate the boundary between sclerotized and soft tissue on the apical meatus." figureDoi="http://doi.org/10.5281/zenodo.3674419" httpUri="https://zenodo.org/record/3674419/files/figure.png" pageId="7" pageNumber="188">25</figureCitation>
).
</paragraph>
<paragraph id="D04036F5FFA2FFC7FBCFFC081B86FC0D" blockId="7.[151,1437,770,1660]" pageId="7" pageNumber="188">
<emphasis id="E28BEAE7FFA2FFC7FBCFFC081ECAFC70" bold="true" box="[199,442,913,940]" pageId="7" pageNumber="188">Variability of males.</emphasis>
Length: 16.1–17.5 mm. Parameres in dorsal view with the marked angle at the level of the central opening more or less developed (
<figureCitation id="48C42A70FFA2FFC7F98FFC2F1C7FFC0C" box="[647,783,950,976]" captionStart="FIGURES 19–22" captionStartId="5.[151,264,1730,1755]" captionTargetBox="[157,1393,1070,1669]" captionTargetId="figure@5.[157,1430,1053,1705]" captionTargetPageId="5" captionText="FIGURES 19–22. Paramera of Ch. panotrogoides Reitter, 1905 (19, 20) and Ch. matejiceki Miessen, Uliana &amp; Keith, new species (21, 22), extremes of variability, left inclined lateral view. 19, specimen from Iran: Sarbijan (Shinagara); 20, specimen from Iran: Kerman (holotype); 21, specimen from Iran: Deh Bakhri (holotype); 22, specimen from Iran: Mijam (paratype)." figureDoi="http://doi.org/10.5281/zenodo.3674417" httpUri="https://zenodo.org/record/3674417/files/figure.png" pageId="7" pageNumber="188">Figs. 21, 22</figureCitation>
), internal piece also variable (
<figureCitation id="48C42A70FFA2FFC7FF68FC2F1B99FC0D" box="[1120,1257,949,977]" captionStart="FIGURES 23–26" captionStartId="6.[151,264,1866,1891]" captionTargetBox="[177,1410,189,1828]" captionTargetId="figure@6.[177,1410,181,1840]" captionTargetPageId="6" captionText="FIGURES 23–26. Endophallic sclerite of Ch. panotrogoides Reitter, 1905 (23, 24) and Ch. matejiceki Miessen, Uliana &amp; Keith, new species (25, 26), extremes of variability. 23, specimen from Iran: Kerman (holotype); 24, specimen from Iran: Sarbijan (Shinagara); 25, specimen from Iran: Deh Bakhri (holotype); 26, specimen from Iran: Mijam (paratype). a, left lateral view; b, inclined lateral view; c, frontal view, apex of internal tubular structure in evidence. Red lines indicate the boundary between sclerotized and soft tissue on the apical meatus." figureDoi="http://doi.org/10.5281/zenodo.3674419" httpUri="https://zenodo.org/record/3674419/files/figure.png" pageId="7" pageNumber="188">Figs. 25, 26</figureCitation>
).
</paragraph>
<paragraph id="D04036F5FFA2FFC7FBCFFC431A53FC28" blockId="7.[151,1437,770,1660]" box="[199,1315,985,1012]" pageId="7" pageNumber="188">
<emphasis id="E28BEAE7FFA2FFC7FBCFFC431E55FC28" bold="true" box="[199,293,986,1012]" pageId="7" pageNumber="188">Female.</emphasis>
Length: 15.7–16.1 mm. Similar to male, but antennal club shorter, as well as the metatarsi.
</paragraph>
</subSubSection>
<subSubSection id="98E5657EFFA2FFC7FBCFFC671A75FBE0" pageId="7" pageNumber="188" type="etymology">
<paragraph id="D04036F5FFA2FFC7FBCFFC671A75FBE0" blockId="7.[151,1437,770,1660]" pageId="7" pageNumber="188">
<emphasis id="E28BEAE7FFA2FFC7FBCFFC671E3EFBC4" bold="true" box="[199,334,1022,1048]" pageId="7" pageNumber="188">Etymology.</emphasis>
Dedicated to Jan Matějíček (
<collectingRegion id="123BF817FFA2FFC7F992FC671C21FBC4" box="[666,849,1022,1048]" country="Czech Republic" name="Kralovehradecky" pageId="7" pageNumber="188">Hradec Králové</collectingRegion>
,
<collectingCountry id="A8E87665FFA2FFC7F856FC671B66FBC5" box="[862,1046,1022,1049]" name="Czech Republic" pageId="7" pageNumber="188">Czech Republic</collectingCountry>
), a specialist on
<taxonomicName id="17FF4D76FFA2FFC7FFDAFC671A1CFBC4" box="[1234,1388,1022,1048]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="188" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
and keen collector of Scarabaeoidea, who entrusted one of us (MU) with the first specimen that we study.
</paragraph>
</subSubSection>
<subSubSection id="98E5657EFFA2FFC7FBCFFBDC1D8BFB58" pageId="7" pageNumber="188" type="distribution">
<paragraph id="D04036F5FFA2FFC7FBCFFBDC1D8BFB58" blockId="7.[151,1437,770,1660]" pageId="7" pageNumber="188">
<emphasis id="E28BEAE7FFA2FFC7FBCFFBDC1E11FBBC" bold="true" box="[199,353,1093,1120]" pageId="7" pageNumber="188">Distribution.</emphasis>
Currently only known from three nearby collecting sites on Jebal Barez mountain range, in southeastern
<collectingCountry id="A8E87665FFA2FFC7FBE5FBF31E6DFB58" box="[237,285,1130,1156]" name="Iran" pageId="7" pageNumber="188">Iran</collectingCountry>
, between
<date id="A4411035FFA2FFC7FA85FBF01D43FB58" box="[397,563,1129,1156]" pageId="7" pageNumber="188" value="1650" valueMax="2300">
1650 and
<quantity id="17079B10FFA2FFC7FAF3FBF31D43FB58" box="[507,563,1130,1156]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="188" unit="m" value="2300.0">2300</quantity>
</date>
m a.s.l. (
<figureCitation id="48C42A70FFA2FFC7F992FBF31D9FFB58" box="[666,751,1130,1156]" captionStart="FIGURE 47" captionStartId="11.[151,250,1655,1680]" captionTargetBox="[175,1412,669,1631]" captionTargetId="figure@11.[175,1412,669,1631]" captionTargetPageId="11" captionText="FIGURE 47. Known distribution of Iranian Chilotrogus Reitter, 1905, based on verified records." figureDoi="http://doi.org/10.5281/zenodo.3674431" httpUri="https://zenodo.org/record/3674431/files/figure.png" pageId="7" pageNumber="188">Fig. 47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="98E5657EFFA2FFC7FBCFFB141DB2F9A0" pageId="7" pageNumber="188" type="diagnosis">
<paragraph id="D04036F5FFA2FFC7FBCFFB141DF4F9E9" blockId="7.[151,1437,770,1660]" pageId="7" pageNumber="188">
<emphasis id="E28BEAE7FFA2FFC7FBCFFB141EA1FB74" bold="true" box="[199,465,1165,1192]" pageId="7" pageNumber="188">Differential diagnosis.</emphasis>
<taxonomicName id="17FF4D76FFA2FFC7FAD3FB171DAEFB74" authority="Miessen &amp; Uliana &amp; Keith, 2020" authorityName="Miessen &amp; Uliana &amp; Keith" authorityYear="2020" box="[475,734,1166,1192]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="matejiceki" status="sp. nov.">
<emphasis id="E28BEAE7FFA2FFC7FAD3FB171DAEFB74" box="[475,734,1166,1192]" italics="true" pageId="7" pageNumber="188">Chilotrogus matejiceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F9B8579CFFA2FFC7F9E0FB171C02FB74" box="[744,882,1166,1192]" pageId="7" pageNumber="188" rank="species">new species</taxonomicNameLabel>
appears to be externally indistinguishable from
<taxonomicName id="17FF4D76FFA2FFC7FB9FFB2B1E17FB10" authorityName="Reitter" authorityYear="1905" box="[151,359,1202,1228]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="panotrogoides">
<emphasis id="E28BEAE7FFA2FFC7FB9FFB2B1E17FB10" box="[151,359,1202,1228]" italics="true" pageId="7" pageNumber="188">Ch. panotrogoides</emphasis>
</taxonomicName>
(
<emphasis id="E28BEAE7FFA2FFC7FA7EFB2B1EE3FB10" box="[374,403,1202,1228]" italics="true" pageId="7" pageNumber="188">cf.</emphasis>
habitus in
<figureCitation id="48C42A70FFA2FFC7F905FB2B1D2BFB10" box="[525,603,1202,1228]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1933,1958]" captionTargetBox="[296,1284,278,1895]" captionTargetId="figure@2.[291,1295,270,1905]" captionTargetPageId="2" captionText="FIGURES 1–4. Male habitus of Iranian Chilotrogus Reitter, 1905. 1, Ch. panotrogoides Reitter, 1905, specimen from Iran: Sarbijan (Shinagara); 2, Ch. matejiceki Miessen, Uliana &amp; Keith, new species, paratype from Iran: Mijam; 3, Ch. farsensis Keith, 2003, specimen from 40 Km NW Shiraz, Sangar; 4, Ch. schoolmeestersi Miessen, Uliana &amp; Keith, new species, holotype." figureDoi="http://doi.org/10.5281/zenodo.3674409" httpUri="https://zenodo.org/record/3674409/files/figure.png" pageId="7" pageNumber="188">Figs. 2</figureCitation>
,
<figureCitation id="48C42A70FFA2FFC7F96EFB281D04FB10" box="[614,628,1201,1228]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1266,1291]" captionTargetBox="[151,1436,194,1241]" captionTargetId="figure@3.[151,1436,181,1241]" captionTargetPageId="3" captionText="FIGURES 5–8. Type specimens of Chilotrogus Reitter, 1905. 5, Ch. matejiceki Miessen, Uliana &amp; Keith, new species, holotype and labels; 6, Ch. panotrogoides Reitter, 1905, holotype and labels; 7, Ch. panotrogoides, labels of false allotype in MHNG; 8, Ch. panotrogoides, labels of false paratype in MHNG." figureDoi="http://doi.org/10.5281/zenodo.3674411" httpUri="https://zenodo.org/record/3674411/files/figure.png" pageId="7" pageNumber="188">5</figureCitation>
and
<figureCitation id="48C42A70FFA2FFC7F9A2FB2B1DC8FB10" box="[682,696,1202,1228]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1933,1958]" captionTargetBox="[296,1284,278,1895]" captionTargetId="figure@2.[291,1295,270,1905]" captionTargetPageId="2" captionText="FIGURES 1–4. Male habitus of Iranian Chilotrogus Reitter, 1905. 1, Ch. panotrogoides Reitter, 1905, specimen from Iran: Sarbijan (Shinagara); 2, Ch. matejiceki Miessen, Uliana &amp; Keith, new species, paratype from Iran: Mijam; 3, Ch. farsensis Keith, 2003, specimen from 40 Km NW Shiraz, Sangar; 4, Ch. schoolmeestersi Miessen, Uliana &amp; Keith, new species, holotype." figureDoi="http://doi.org/10.5281/zenodo.3674409" httpUri="https://zenodo.org/record/3674409/files/figure.png" pageId="7" pageNumber="188">1</figureCitation>
,
<figureCitation id="48C42A70FFA2FFC7F9CEFB2B1DA6FB11" box="[710,726,1202,1229]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1266,1291]" captionTargetBox="[151,1436,194,1241]" captionTargetId="figure@3.[151,1436,181,1241]" captionTargetPageId="3" captionText="FIGURES 5–8. Type specimens of Chilotrogus Reitter, 1905. 5, Ch. matejiceki Miessen, Uliana &amp; Keith, new species, holotype and labels; 6, Ch. panotrogoides Reitter, 1905, holotype and labels; 7, Ch. panotrogoides, labels of false allotype in MHNG; 8, Ch. panotrogoides, labels of false paratype in MHNG." figureDoi="http://doi.org/10.5281/zenodo.3674411" httpUri="https://zenodo.org/record/3674411/files/figure.png" pageId="7" pageNumber="188">6</figureCitation>
), and identification can be achieved only by examination of the male genitalia (
<emphasis id="E28BEAE7FFA2FFC7FA4EFB4F1E12FB2C" box="[326,354,1238,1264]" italics="true" pageId="7" pageNumber="188">cf.</emphasis>
<figureCitation id="48C42A70FFA2FFC7FA61FB4F1EAFFB2C" box="[361,479,1238,1264]" captionStart="FIGURES 9–13" captionStartId="4.[151,264,1285,1310]" captionTargetBox="[159,1430,534,1249]" captionTargetId="figure@4.[151,1436,525,1260]" captionTargetPageId="4" captionText="FIGURES 9–13. Male genitalia of Ch. panotrogoides Reitter, 1905, holotype. 9, aedeagus (endophallic sclerite dissected), right lateral view; 10, same, dorsal view; 11, same, apical view; 12: endophallic sclerite, dorsal view; 13, same, right lateral view." figureDoi="http://doi.org/10.5281/zenodo.3674413" httpUri="https://zenodo.org/record/3674413/files/figure.png" pageId="7" pageNumber="188">Figs. 9–13</figureCitation>
and
<figureCitation id="48C42A70FFA2FFC7F91FFB4F1D11FB2C" box="[535,609,1238,1264]" captionStart="FIGURES 14–18" captionStartId="5.[151,264,937,962]" captionTargetBox="[159,1431,191,899]" captionTargetId="figure@5.[151,1436,181,911]" captionTargetPageId="5" captionText="FIGURES 14–18. Male genitalia of Ch. matejiceki Miessen, Uliana &amp; Keith, new species, holotype. 14, aedeagus (endophallic sclerite dissected), right lateral view; 15, same, dorsal view; 16, same, apical view; 17: endophallic sclerite, dorsal view; 18, same, right lateral view." figureDoi="http://doi.org/10.5281/zenodo.3674415" httpUri="https://zenodo.org/record/3674415/files/figure.png" pageId="7" pageNumber="188">14–18</figureCitation>
). Parameres of
<taxonomicName id="17FF4D76FFA2FFC7F807FB4F1CC2FB2C" authority="Miessen &amp; Uliana &amp; Keith, 2020" authorityName="Miessen &amp; Uliana &amp; Keith" authorityYear="2020" box="[783,946,1238,1264]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="matejiceki" status="sp. nov.">
<emphasis id="E28BEAE7FFA2FFC7F807FB4F1CC2FB2C" box="[783,946,1238,1264]" italics="true" pageId="7" pageNumber="188">Ch. matejiceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F9B8579CFFA2FFC7F8B1FB4F1B30FB2C" box="[953,1088,1238,1264]" pageId="7" pageNumber="188" rank="species">new species</taxonomicNameLabel>
are characterised by an angled to tooth-like projection of the internal margin, which marks the distal part of a broad basal concavity. The
<typeStatus id="0F448857FFA2FFC7FE2EFB631A26FAC8" box="[1318,1366,1274,1300]" pageId="7" pageNumber="188">type</typeStatus>
series shows some degree of variability (including between syntopic specimens) in this trait (
<emphasis id="E28BEAE7FFA2FFC7FF55FA871B01FAE4" box="[1117,1137,1310,1336]" italics="true" pageId="7" pageNumber="188">cf</emphasis>
.
<figureCitation id="48C42A70FFA2FFC7FF77FA871A75FAE4" box="[1151,1285,1310,1336]" captionStart="FIGURES 19–22" captionStartId="5.[151,264,1730,1755]" captionTargetBox="[157,1393,1070,1669]" captionTargetId="figure@5.[157,1430,1053,1705]" captionTargetPageId="5" captionText="FIGURES 19–22. Paramera of Ch. panotrogoides Reitter, 1905 (19, 20) and Ch. matejiceki Miessen, Uliana &amp; Keith, new species (21, 22), extremes of variability, left inclined lateral view. 19, specimen from Iran: Sarbijan (Shinagara); 20, specimen from Iran: Kerman (holotype); 21, specimen from Iran: Deh Bakhri (holotype); 22, specimen from Iran: Mijam (paratype)." figureDoi="http://doi.org/10.5281/zenodo.3674417" httpUri="https://zenodo.org/record/3674417/files/figure.png" pageId="7" pageNumber="188">Figs. 21, 22</figureCitation>
), that is however well distinct from the gently curved morphology of
<taxonomicName id="17FF4D76FFA2FFC7F828FADB1C81FA80" authorityName="Reitter" authorityYear="1905" box="[800,1009,1346,1372]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="panotrogoides">
<emphasis id="E28BEAE7FFA2FFC7F828FADB1C81FA80" box="[800,1009,1346,1372]" italics="true" pageId="7" pageNumber="188">Ch. panotrogoides</emphasis>
</taxonomicName>
(
<figureCitation id="48C42A70FFA2FFC7FF09FADB1BFDFA80" box="[1025,1165,1346,1372]" captionStart="FIGURES 19–22" captionStartId="5.[151,264,1730,1755]" captionTargetBox="[157,1393,1070,1669]" captionTargetId="figure@5.[157,1430,1053,1705]" captionTargetPageId="5" captionText="FIGURES 19–22. Paramera of Ch. panotrogoides Reitter, 1905 (19, 20) and Ch. matejiceki Miessen, Uliana &amp; Keith, new species (21, 22), extremes of variability, left inclined lateral view. 19, specimen from Iran: Sarbijan (Shinagara); 20, specimen from Iran: Kerman (holotype); 21, specimen from Iran: Deh Bakhri (holotype); 22, specimen from Iran: Mijam (paratype)." figureDoi="http://doi.org/10.5281/zenodo.3674417" httpUri="https://zenodo.org/record/3674417/files/figure.png" pageId="7" pageNumber="188">Figs. 19, 20</figureCitation>
). Additional differences are found in the distal part of the internal sclerite. Its dorsal spine is variable in elongation in both species, but in
<taxonomicName id="17FF4D76FFA2FFC7FE7DFAFF1E7AFA78" authority="Miessen &amp; Uliana &amp; Keith, 2020" authorityName="Miessen &amp; Uliana &amp; Keith" authorityYear="2020" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="matejiceki" status="sp. nov.">
<emphasis id="E28BEAE7FFA2FFC7FE7DFAFF1E7AFA78" italics="true" pageId="7" pageNumber="188">Ch. matejiceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F9B8579CFFA2FFC7FA18FA131EE5FA78" box="[272,405,1418,1444]" pageId="7" pageNumber="188" rank="species">new species</taxonomicNameLabel>
it is broad at the base and flattened in its distal part (
<figureCitation id="48C42A70FFA2FFC7F8DEFA131B2BFA79" box="[982,1115,1417,1445]" captionStart="FIGURES 23–26" captionStartId="6.[151,264,1866,1891]" captionTargetBox="[177,1410,189,1828]" captionTargetId="figure@6.[177,1410,181,1840]" captionTargetPageId="6" captionText="FIGURES 23–26. Endophallic sclerite of Ch. panotrogoides Reitter, 1905 (23, 24) and Ch. matejiceki Miessen, Uliana &amp; Keith, new species (25, 26), extremes of variability. 23, specimen from Iran: Kerman (holotype); 24, specimen from Iran: Sarbijan (Shinagara); 25, specimen from Iran: Deh Bakhri (holotype); 26, specimen from Iran: Mijam (paratype). a, left lateral view; b, inclined lateral view; c, frontal view, apex of internal tubular structure in evidence. Red lines indicate the boundary between sclerotized and soft tissue on the apical meatus." figureDoi="http://doi.org/10.5281/zenodo.3674419" httpUri="https://zenodo.org/record/3674419/files/figure.png" pageId="7" pageNumber="188">Figs. 25, 26</figureCitation>
), while in
<taxonomicName id="17FF4D76FFA2FFC7FFC6FA131AECFA78" authorityName="Reitter" authorityYear="1905" box="[1230,1436,1418,1444]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="panotrogoides">
<emphasis id="E28BEAE7FFA2FFC7FFC6FA131AECFA78" box="[1230,1436,1418,1444]" italics="true" pageId="7" pageNumber="188">Ch. panotrogoides</emphasis>
</taxonomicName>
the same spine is narrow at the base and is cylindrical in its distal part (
<figureCitation id="48C42A70FFA2FFC7F8A8FA371B57FA14" box="[928,1063,1454,1480]" captionStart="FIGURES 23–26" captionStartId="6.[151,264,1866,1891]" captionTargetBox="[177,1410,189,1828]" captionTargetId="figure@6.[177,1410,181,1840]" captionTargetPageId="6" captionText="FIGURES 23–26. Endophallic sclerite of Ch. panotrogoides Reitter, 1905 (23, 24) and Ch. matejiceki Miessen, Uliana &amp; Keith, new species (25, 26), extremes of variability. 23, specimen from Iran: Kerman (holotype); 24, specimen from Iran: Sarbijan (Shinagara); 25, specimen from Iran: Deh Bakhri (holotype); 26, specimen from Iran: Mijam (paratype). a, left lateral view; b, inclined lateral view; c, frontal view, apex of internal tubular structure in evidence. Red lines indicate the boundary between sclerotized and soft tissue on the apical meatus." figureDoi="http://doi.org/10.5281/zenodo.3674419" httpUri="https://zenodo.org/record/3674419/files/figure.png" pageId="7" pageNumber="188">Figs. 23, 24</figureCitation>
). Also, the subcircular apex of the tube-like structure that bears the spine is almost completely occupied by soft tissue (evertible part of endophallus) in
<taxonomicName id="17FF4D76FFA2FFC7FBBCFA6F1EF5F9CC" authorityName="Reitter" authorityYear="1905" box="[180,389,1526,1552]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="panotrogoides">
<emphasis id="E28BEAE7FFA2FFC7FBBCFA6F1EF5F9CC" box="[180,389,1526,1552]" italics="true" pageId="7" pageNumber="188">Ch. panotrogoides</emphasis>
</taxonomicName>
(
<figureCitation id="48C42A70FFA2FFC7FA9CFA6F1D47F9CC" box="[404,567,1526,1552]" captionStart="FIGURES 23–26" captionStartId="6.[151,264,1866,1891]" captionTargetBox="[177,1410,189,1828]" captionTargetId="figure@6.[177,1410,181,1840]" captionTargetPageId="6" captionText="FIGURES 23–26. Endophallic sclerite of Ch. panotrogoides Reitter, 1905 (23, 24) and Ch. matejiceki Miessen, Uliana &amp; Keith, new species (25, 26), extremes of variability. 23, specimen from Iran: Kerman (holotype); 24, specimen from Iran: Sarbijan (Shinagara); 25, specimen from Iran: Deh Bakhri (holotype); 26, specimen from Iran: Mijam (paratype). a, left lateral view; b, inclined lateral view; c, frontal view, apex of internal tubular structure in evidence. Red lines indicate the boundary between sclerotized and soft tissue on the apical meatus." figureDoi="http://doi.org/10.5281/zenodo.3674419" httpUri="https://zenodo.org/record/3674419/files/figure.png" pageId="7" pageNumber="188">Figs. 23c, 24c</figureCitation>
), while the same soft tissue occupies only about the lower half of the apex in
<taxonomicName id="17FF4D76FFA2FFC7FB9FF9831E49F9E8" authority="Miessen &amp; Uliana &amp; Keith, 2020" authorityName="Miessen &amp; Uliana &amp; Keith" authorityYear="2020" box="[151,313,1562,1588]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="matejiceki" status="sp. nov.">
<emphasis id="E28BEAE7FFA2FFC7FB9FF9831E49F9E8" box="[151,313,1562,1588]" italics="true" pageId="7" pageNumber="188">Ch. matejiceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F9B8579CFFA2FFC7FA48F9831EB7F9E8" box="[320,455,1562,1588]" pageId="7" pageNumber="188" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="48C42A70FFA2FFC7FADEF9831D08F9E9" box="[470,632,1561,1589]" captionStart="FIGURES 23–26" captionStartId="6.[151,264,1866,1891]" captionTargetBox="[177,1410,189,1828]" captionTargetId="figure@6.[177,1410,181,1840]" captionTargetPageId="6" captionText="FIGURES 23–26. Endophallic sclerite of Ch. panotrogoides Reitter, 1905 (23, 24) and Ch. matejiceki Miessen, Uliana &amp; Keith, new species (25, 26), extremes of variability. 23, specimen from Iran: Kerman (holotype); 24, specimen from Iran: Sarbijan (Shinagara); 25, specimen from Iran: Deh Bakhri (holotype); 26, specimen from Iran: Mijam (paratype). a, left lateral view; b, inclined lateral view; c, frontal view, apex of internal tubular structure in evidence. Red lines indicate the boundary between sclerotized and soft tissue on the apical meatus." figureDoi="http://doi.org/10.5281/zenodo.3674419" httpUri="https://zenodo.org/record/3674419/files/figure.png" pageId="7" pageNumber="188">Figs. 25c, 26c</figureCitation>
).
</paragraph>
<paragraph id="D04036F5FFA2FFC7FBCFF9A71DB2F9A0" blockId="7.[151,1437,770,1660]" pageId="7" pageNumber="188">
Females of the two species show very moderate differences in the genital sclerites (
<figureCitation id="48C42A70FFA2FFC7FF7EF9A71B8FF984" box="[1142,1279,1598,1624]" captionStart="FIGURES 27–30" captionStartId="7.[151,264,627,652]" captionTargetBox="[152,1436,181,601]" captionTargetId="figure@7.[152,1436,181,601]" captionTargetPageId="7" captionText="FIGURES 27–30. Female genitalia of Ch. panotrogoides Reitter, 1905 (27, 28) and Ch. matejiceki Miessen, Uliana &amp; Keith, new species (29, 30). 27, 28, specimens from Iran: Sirjan; 29, specimen from Iran: Deh Bakhri (allotype); 30, specimen from Iran: Deh Bakhri (paratype)." figureDoi="http://doi.org/10.5281/zenodo.3674421" httpUri="https://zenodo.org/record/3674421/files/figure.png" pageId="7" pageNumber="188">Figs. 27–30</figureCitation>
), whose diagnostic value should be verified on a longer series.
</paragraph>
</subSubSection>
</treatment>
</document>
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