406 lines
49 KiB
XML
406 lines
49 KiB
XML
<document id="D042C43CF7489CE4D0AED5B97BBA5DC1" ID-CLB-Dataset="20422" ID-DOI="10.5852/ejt.2022.798.1669" ID-GBIF-Dataset="31fec661-5dac-4626-856d-4ba4680efee7" ID-ISSN="2118-9773" ID-Zenodo-Dep="6323002" ID-ZooBank="50692D26-A41C-4F85-B207-A6747FD07470" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1646225219873" checkinUser="felipe" docAuthor="Vagalinski, Boyan, Borissov, Simeon, Bobeva, Aneliya, Canciani, Giacomo & Antić, Dragan Ž." docDate="2022" docId="4A307579CF420D11FE6AFD30FABBFE33" docLanguage="en" docName="ejt-798_30-69.pdf" docOrigin="European Journal of Taxonomy 798" docSource="http://dx.doi.org/10.5852/ejt.2022.798.1669" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Stygiiulus illyricus Vagalinski & Borissov & Bobeva & Canciani & Antić 2022, comb. nov." docType="treatment" docVersion="11" lastPageNumber="46" masterDocId="B6090D01CF4D0D00FF95FFEAFFB8FF83" masterDocTitle="The mostly cavernicolous millipede genus Stygiiulus Verhoeff, 1929, stat. nov.: taxonomy, distribution and phylogenetic relationships (Diplopoda, Julida, Julidae)" masterLastPageNumber="69" masterPageNumber="30" pageNumber="45" updateTime="1699300459248" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1B9C5EFDF3F71DB720766811A866BEF1">The mostly cavernicolous millipede genus Stygiiulus Verhoeff, 1929, stat. nov.: taxonomy, distribution and phylogenetic relationships (Diplopoda, Julida, Julidae)</mods:title>
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<mods:namePart id="7387E7F53ED4FCF1244ADB43B1FBDE5F">Vagalinski, Boyan</mods:namePart>
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<mods:affiliation id="D2CC709384BD0C3F8F94510F4E4DDC8A">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113, Sofia, Bulgaria.</mods:affiliation>
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<mods:namePart id="7905A0AA9F4EB8C2B51512CC53D9B0BD">Borissov, Simeon</mods:namePart>
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<mods:affiliation id="F89511872013408CBD0E6521AE68568A">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113, Sofia, Bulgaria.</mods:affiliation>
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<mods:namePart id="092F7B1E123C7FD86AF2BAF7AA1BD1D0">Canciani, Giacomo</mods:namePart>
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<mods:affiliation id="394A5161E756407F92A014ED26DF0872">Jamarski Klub Kraški Krti / G. S. Talpe del Carso, loc. San Michele del Carso / Vrh, I- 34070 Savogna d’Isonzo (GO), Italy. & Associazione Speleologica Forum Julii Speleo, Via Carraria 99, I- 33043 Cividale del Friuli (UD), Italy.</mods:affiliation>
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<mods:affiliation id="0DD18C48A6DB78E86C7B8A4C85E80271">University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. & Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
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<mods:title id="9221C7EE6C5828E16D2A571AD479E2BB">European Journal of Taxonomy</mods:title>
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<treatment id="4A307579CF420D11FE6AFD30FABBFE33" ID-DOI="http://doi.org/10.5281/zenodo.8337989" ID-GBIF-Taxon="193854119" ID-Zenodo-Dep="8337989" LSID="urn:lsid:plazi:treatment:4A307579CF420D11FE6AFD30FABBFE33" httpUri="http://treatment.plazi.org/id/4A307579CF420D11FE6AFD30FABBFE33" lastPageId="17" lastPageNumber="46" pageId="15" pageNumber="45">
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<paragraph id="C226C46FCF420D0FFE6AFD30FB8CFD76" blockId="15.[511,1076,730,792]" box="[511,1076,730,757]" pageId="15" pageNumber="45">
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<taxonomicName id="0599BFECCF420D0FFE6AFD30FC13FD77" ID-CoL="6ZZNM" authority="(Verhoeff, 1929)" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Verhoeff" baseAuthorityYear="1929" box="[511,939,730,757]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="illyricus" status="comb. nov.">
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<emphasis id="F0ED187DCF420D0FFE6AFD30FD59FD76" bold="true" box="[511,737,730,757]" italics="true" pageId="15" pageNumber="45">Stygiiulus illyricus</emphasis>
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(
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<bibRefCitation id="A608B99ECF420D0FFD65FD30FC1BFD77" author="Verhoeff K." box="[752,931,730,756]" pageId="15" pageNumber="45" pagination="14 - 35" refId="ref22558" refString="Verhoeff K. 1929. Arthropoden aus sudostalpinen Hohlen (I). Gesammelt von Karl Strasser in Triest. Mitteilungen uber Hohlen- und Karstforschung 1929: 14 - 35." type="journal article" year="1929">Verhoeff, 1929</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="EBDEA506CF420D0FFC26FD30FB8CFD76" box="[947,1076,730,757]" pageId="15" pageNumber="45" rank="species">comb. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="8A8397E4CF420D0FFD25FD17FC3BFC9B" box="[688,899,765,792]" pageId="15" pageNumber="45" type="description">
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<paragraph id="C226C46FCF420D0FFD25FD17FC3BFC9B" blockId="15.[511,1076,730,792]" box="[688,899,765,792]" pageId="15" pageNumber="45">
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<figureCitation id="5AA2D8EACF420D0FFD25FD17FCA4FC9B" box="[688,796,765,792]" captionStart="Fig" captionStartId="26.[189,232,1822,1848]" captionTargetBox="[197,1384,282,1779]" captionTargetId="figure-12@26.[189,1398,263,1787]" captionTargetPageId="26" captionText="Fig. 10. Left gonopods of species of Stygiiulus stat. nov., mesal views. A. S. ausugi (Manfredi, 1953) comb.nov.,topotype♂(NHMD).B.S.illyricus(Verhoeff,1929)comb.nov.,redrawnfromVerhoeff (1929). C. S. insularis (Strasser, 1938) comb. nov., redrawn from Antić et al. (2018). D. S. maximus (Verhoeff, 1929) comb. nov., redrawn from Verhoeff (1930). E. S. seewaldi (Strasser, 1967) comb. nov., redrawn and modified from Strasser (1967). F. S. tobias (Berlese, 1886) comb. nov., ♂ from Grotta [Cave] del Subiolo (NHMD). Abbreviations: do = distal outgrowth of velum. Drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.6323028" httpUri="https://zenodo.org/record/6323028/files/figure.png" pageId="15" pageNumber="45">Figs 10B</figureCitation>
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,
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<figureCitation id="5AA2D8EACF420D0FFCBDFD14FCE2FC9B" box="[808,858,766,792]" captionStart="Fig" captionStartId="31.[189,231,1681,1707]" captionTargetBox="[345,1243,265,1635]" captionTargetId="figure-15@31.[325,1262,262,1645]" captionTargetPageId="31" captionText="Fig. 11. Left vulvae of species of Stygiiulus stat. nov. (A–D) and representatives of (other) Julini/ Leptoiulini (E) and Typhloiulini/blind Julini/Leptoiulini in caudal, somewhat mesal views. A. S. ausugi (Manfredi, 1953) comb. nov., ♀ from Grotta [Cave] del Calgeron (NHMD). B. S. illyricus (Verhoef, 1929) comb. nov., ♀ from Rabakova Cave (IZB). C. S. insularis (Strasser, 1938) comb. nov., topotype ♀ (CBSS). D. S. tobias (Berlese, 1886) comb. nov., ♀ from Grotta [Cave] del Subiolo (NHMD). E. Leptoiulus cf. trilineatus(C.L. Koch, 1847), ♀ from Belasitsa Mtn, Bulgaria. F. Serboiulus spelaeophilus Gulička, 1967, ♀ from Vodni Pech Cave (NMNHS). G. Typhloiulus (Inversotyphlus) lobifer Attems, 1951, ♀ from Minjera-Bauxite Mine (IBER). H. Typhloiulus (s. str.) strictus (Latzel, 1882), ♀ from Kunino (IBER). Abbreviations: at = anterior tube; la = lateral ampulla; lt = lateral tube; ma = median ampulla; mt = median tube; op = operculum; pa = posterior ampulla; pt = posterior tube. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6323030" httpUri="https://zenodo.org/record/6323030/files/figure.png" pageId="15" pageNumber="45">11B</figureCitation>
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,
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<figureCitation id="5AA2D8EACF420D0FFCF3FD14FC3BFC9B" box="[870,899,766,792]" captionStart="Fig" captionStartId="33.[189,232,1998,2024]" captionTargetBox="[189,1398,1099,1960]" captionTargetId="figure-332@33.[189,1398,1099,1960]" captionTargetPageId="33" captionText="Fig. 13. General distribution of the species of the genus Stygiiulus stat. nov." figureDoi="http://doi.org/10.5281/zenodo.6323034" httpUri="https://zenodo.org/record/6323034/files/figure.png" pageId="15" pageNumber="45">13</figureCitation>
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</paragraph>
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</subSubSection>
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<paragraph id="C226C46FCF420D10FF28FCD5FAFAFEA0" blockId="15.[189,915,831,858]" lastBlockId="16.[189,1346,265,433]" lastPageId="16" lastPageNumber="46" pageId="15" pageNumber="45">
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<treatmentCitationGroup id="E289E341CF420D10FF28FCD5FAFAFEA0" lastPageId="16" lastPageNumber="46" pageId="15" pageNumber="45">
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<treatmentCitation id="4338E27ECF420D0FFF28FCD5FCFDFCDA" author="Verhoeff K." box="[189,837,831,858]" page="23" pageId="15" pageNumber="45" year="1929">
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<taxonomicName id="0599BFECCF420D0FFF28FCD5FCFDFCDA" ID-CoL="7Z6NS" authority="Verhoeff, 1929: 20 - 23" authorityName="Verhoeff" authorityPageNumber="20 - 23" authorityYear="1929" box="[189,837,831,858]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="illyricus" subGenus="Stygiiulus">
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<emphasis id="F0ED187DCF420D0FFF28FCD5FEFCFCDA" box="[189,324,831,857]" italics="true" pageId="15" pageNumber="45">Typhloiulus</emphasis>
|
||
(
|
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<emphasis id="F0ED187DCF420D0FFEC0FCD5FE7FFCDA" box="[341,455,831,857]" italics="true" pageId="15" pageNumber="45">Stygiiulus</emphasis>
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)
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<emphasis id="F0ED187DCF420D0FFE4DFCD5FD8FFCDA" box="[472,567,831,857]" italics="true" pageId="15" pageNumber="45">illyricus</emphasis>
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<bibRefCitation id="A608B99ECF420D0FFDA8FCD5FCFDFCDA" author="Verhoeff K." box="[573,837,831,857]" pageId="15" pageNumber="45" pagination="14 - 35" refId="ref22558" refString="Verhoeff K. 1929. Arthropoden aus sudostalpinen Hohlen (I). Gesammelt von Karl Strasser in Triest. Mitteilungen uber Hohlen- und Karstforschung 1929: 14 - 35." type="journal article" year="1929">Verhoeff, 1929: 20–23</bibRefCitation>
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</taxonomicName>
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||
, fig. 1.
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||
<taxonomicName id="0599BFECCF5D0D10FF28FEE3FD8FFEA0" ID-CoL="7Z6NS" authorityName="Verhoeff" authorityYear="1929" box="[189,567,265,292]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="16" pageNumber="46" phylum="Arthropoda" rank="species" species="illyricus" subGenus="Stygiiulus">
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<emphasis id="F0ED187DCF5D0D10FF28FEE3FEFCFEA0" box="[189,324,265,291]" italics="true" pageId="16" pageNumber="46">Typhloiulus</emphasis>
|
||
(
|
||
<emphasis id="F0ED187DCF5D0D10FEC0FEE3FE7FFEA0" box="[341,455,265,291]" italics="true" pageId="16" pageNumber="46">Stygiiulus</emphasis>
|
||
)
|
||
<emphasis id="F0ED187DCF5D0D10FE4DFEE3FD8FFEA0" box="[472,567,265,291]" italics="true" pageId="16" pageNumber="46">illyricus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="4338E27ECF5D0D10FDC6FEE3FC88FEA0" author="Manfredi P." box="[595,816,265,292]" page="81" pageId="16" pageNumber="46" year="1932">
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||
<bibRefCitation id="A608B99ECF5D0D10FDC6FEE3FC88FEA0" author="Manfredi P." box="[595,816,265,292]" pageId="16" pageNumber="46" pagination="71 - 96" refId="ref21622" refString="Manfredi P. 1932. Contributo alla conoscenza della fauna cavernicola italiana. Miriapodi. Rivista di Scienze Naturali " Natura " 23: 71 - 96." type="journal article" year="1932">Manfredi 1932: 81</bibRefCitation>
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</treatmentCitation>
|
||
. —
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||
<treatmentCitation id="4338E27ECF5D0D10FCCAFEE3FB94FEA0" author="Strasser K." box="[863,1068,265,292]" page="20" pageId="16" pageNumber="46" year="1962">
|
||
<bibRefCitation id="A608B99ECF5D0D10FCCAFEE3FB94FEA0" author="Strasser K." box="[863,1068,265,292]" pageId="16" pageNumber="46" pagination="1 - 77" refId="ref22282" refString="Strasser K. 1962. Die Typhloiulini (Diplopoda Symphyognatha). Atti del Museo civico di Storia naturale di Trieste 23 (1): 1 - 77." type="journal article" year="1962">Strasser 1962: 20</bibRefCitation>
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</treatmentCitation>
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||
, 35–36, figs 11i, 36–37.
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||
</treatmentCitationGroup>
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||
</paragraph>
|
||
<caption id="96E694E7CF420D0FFF28F88EFB15F86A" ID-DOI="http://doi.org/10.5281/zenodo.6323020" ID-Zenodo-Dep="6323020" httpUri="https://zenodo.org/record/6323020/files/figure.png" pageId="15" pageNumber="45" startId="15.[189,231,1892,1918]" targetBox="[189,1398,949,1853]" targetPageId="15">
|
||
<paragraph id="C226C46FCF420D0FFF28F88EFB15F86A" blockId="15.[187,1399,1892,2025]" pageId="15" pageNumber="45">
|
||
<emphasis id="F0ED187DCF420D0FFF28F88EFEB1F8FC" bold="true" box="[189,265,1892,1919]" pageId="15" pageNumber="45">Fig. 6.</emphasis>
|
||
<taxonomicName id="0599BFECCF420D0FFE84F88FFD0FF8FC" ID-CoL="8YKQN" authority="(Strasser, 1971)" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[273,695,1893,1919]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="gentianae" status="comb. et stat. nov.">
|
||
<emphasis id="F0ED187DCF420D0FFE84F88FFE43F8FC" box="[273,507,1893,1919]" italics="true" pageId="15" pageNumber="45">Stygiiulus gentianae</emphasis>
|
||
(Strasser, 1971)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF420D0FFD2AF88FFC2CF8FC" box="[703,916,1893,1919]" pageId="15" pageNumber="45">comb. et stat. nov.</taxonomicNameLabel>
|
||
, ♂ from Grotta [cave] della Vecchia Diga (NHMD).
|
||
<emphasis id="F0ED187DCF420D0FFEA0F862FEF2F821" bold="true" box="[309,330,1928,1954]" pageId="15" pageNumber="45">A</emphasis>
|
||
. Right promere, caudal view.
|
||
<emphasis id="F0ED187DCF420D0FFD0EF862FD17F821" bold="true" box="[667,687,1928,1954]" pageId="15" pageNumber="45">B</emphasis>
|
||
. Right mesomere, oral view.
|
||
<emphasis id="F0ED187DCF420D0FFC6DF862FBB5F821" bold="true" box="[1016,1037,1928,1954]" pageId="15" pageNumber="45">C</emphasis>
|
||
. Distal part of left opisthomere, mesal view. Abbreviations: ab = anterior branch of solenomere; el = external lobe; il = internal lobe; pb = posterior branch of solenomere; ph = posterior hump; v = velum. Scale bars = 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C226C46FCF5D0D10FF28FEC6FBA0FEC5" blockId="16.[189,1346,265,433]" box="[189,1048,300,327]" pageId="16" pageNumber="46">
|
||
<treatmentCitationGroup id="E289E341CF5D0D10FF28FEC6FBA0FEC5" box="[189,1048,300,327]" pageId="16" pageNumber="46">
|
||
<taxonomicName id="0599BFECCF5D0D10FF28FEC6FE12FEC5" ID-CoL="7Z6NS" authorityName="Verhoeff" authorityYear="1929" box="[189,426,300,326]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="16" pageNumber="46" phylum="Arthropoda" rank="species" species="illyricus">
|
||
<emphasis id="F0ED187DCF5D0D10FF28FEC6FE12FEC5" box="[189,426,300,326]" italics="true" pageId="16" pageNumber="46">Typhloiulus illyricus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="4338E27ECF5D0D10FE53FEC6FD23FEC5" author="Verhoeff K." box="[454,667,300,327]" page="17" pageId="16" pageNumber="46" year="1930">
|
||
<bibRefCitation id="A608B99ECF5D0D10FE53FEC6FD23FEC5" author="Verhoeff K." box="[454,667,300,327]" pageId="16" pageNumber="46" pagination="97 - 113" refId="ref22589" refString="Verhoeff K. 1930. Arthropoden aus sudostalpinen Hohlen, gesammelt von Karl Strasser. 5. Aufsatz. Mitteilungen uber Hohlen- und Karstforschung 1930 (4): 97 - 113." type="journal article" year="1930">Verhoeff 1930: 17</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="4338E27ECF5D0D10FD5CFEC6FC71FEC5" ID-DOI="http://doi.org/10.5281/zenodo.5960745" ID-Zenodo-Dep="5960745" author="Antic D. Z. & Drazina T. & Rada T. & Lucic L. & Makarov S. E." box="[713,969,300,327]" httpUri="http://treatment.plazi.org/id/0383AF24200DFFEB14DFFE37FC48FB7E" page="263" pageId="16" pageNumber="46" year="2018">
|
||
<bibRefCitation id="A608B99ECF5D0D10FD5CFEC6FC71FEC5" author="Antic D. Z. & Drazina T. & Rada T. & Lucic L. & Makarov S. E." box="[713,969,300,327]" pageId="16" pageNumber="46" pagination="258 - 294" refId="ref20375" refString="Antic D. Z., Drazina T., Rada T., Lucic L. R & Makarov S. E. 2018. Review of the genus Typhloiulus Latzel, 1884 in the Dinaric region, with a description of four new species and the first description of the male of Typhloiulus insularis Strasser, 1938 (Diplopoda: Julida: Julidae). Zootaxa 4455 (2): 258 - 294. https: // doi. org / 10.11646 / zootaxa. 4455.2.2" type="journal article" year="2018">
|
||
Antić
|
||
<emphasis id="F0ED187DCF5D0D10FC84FEC7FCF3FEC5" box="[785,843,300,326]" italics="true" pageId="16" pageNumber="46">et al.</emphasis>
|
||
2018: 263
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 3.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF5D0D10FF28FEA5FBB7FEE9" blockId="16.[189,1346,265,433]" box="[189,1039,335,362]" pageId="16" pageNumber="46">
|
||
<treatmentCitationGroup id="E289E341CF5D0D10FF28FEA5FBB7FEE9" box="[189,1039,335,362]" pageId="16" pageNumber="46">
|
||
<taxonomicName id="0599BFECCF5D0D10FF28FEA5FE4BFEEA" authority="Verhoeff 1933" box="[189,499,335,361]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="16" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="illyricus" subSpecies="stygis">
|
||
<emphasis id="F0ED187DCF5D0D10FF28FEA5FE4BFEEA" box="[189,499,335,361]" italics="true" pageId="16" pageNumber="46">Typhloiulus illyricus stygis</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="4338E27ECF5D0D10FD9AFEA5FCA8FEE9" author="Verhoeff K. W." box="[527,784,335,362]" page="15" pageId="16" pageNumber="46" year="1933">
|
||
<bibRefCitation id="A608B99ECF5D0D10FD9AFEA5FCA8FEE9" author="Verhoeff K. W." box="[527,784,335,362]" pageId="16" pageNumber="46" pagination="1 - 21" refId="ref22622" refString="Verhoeff K. W. 1933. Arthropoden aus sudostalpinen Hohlen, gesammelt von Karl Strasser. 7. Aufsatz. Mitteilungen uber Hohlen- und Karstforschung 1933 (4): 1 - 21." type="journal article" year="1933">Verhoeff 1933: 15–17</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs 28–29,
|
||
<emphasis id="F0ED187DCF5D0D10FC31FEBAFBB7FEE9" bold="true" box="[932,1039,336,362]" pageId="16" pageNumber="46">syn. nov.</emphasis>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF5D0D10FF28FE98FCF9FE0E" blockId="16.[189,1346,265,433]" box="[189,833,370,397]" pageId="16" pageNumber="46">
|
||
<treatmentCitationGroup id="E289E341CF5D0D10FF28FE98FCF9FE0E" box="[189,833,370,397]" pageId="16" pageNumber="46">
|
||
<taxonomicName id="0599BFECCF5D0D10FF28FE98FDF4FE0F" ID-CoL="8YKQV" authority="Verhoeff, 1929" box="[189,588,370,397]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="16" pageNumber="46" phylum="Arthropoda" rank="variety" species="illyricus" variety="obscurus">
|
||
<emphasis id="F0ED187DCF5D0D10FF28FE98FE12FE0F" box="[189,426,370,396]" italics="true" pageId="16" pageNumber="46">Typhloiulus illyricus</emphasis>
|
||
var.
|
||
<emphasis id="F0ED187DCF5D0D10FE76FE98FDF4FE0F" box="[483,588,370,396]" italics="true" pageId="16" pageNumber="46">obscurus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="4338E27ECF5D0D10FDFDFE98FC85FE0E" ID-DOI="http://doi.org/10.5281/zenodo.6287499" ID-Zenodo-Dep="6287499" author="Verhoeff K. W." box="[616,829,370,397]" httpUri="http://treatment.plazi.org/id/D8D62D232BB5A19E469C094DF251B808" page="17" pageId="16" pageNumber="46" year="1933">
|
||
<bibRefCitation id="A608B99ECF5D0D10FDFDFE98FC85FE0E" author="Verhoeff K. W." box="[616,829,370,397]" pageId="16" pageNumber="46" pagination="1 - 21" refId="ref22622" refString="Verhoeff K. W. 1933. Arthropoden aus sudostalpinen Hohlen, gesammelt von Karl Strasser. 7. Aufsatz. Mitteilungen uber Hohlen- und Karstforschung 1933 (4): 1 - 21." type="journal article" year="1933">Verhoeff 1933: 17</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF5D0D10FF28FE7CFCFDFE32" blockId="16.[189,1346,265,433]" box="[189,837,406,433]" pageId="16" pageNumber="46">
|
||
<treatmentCitationGroup id="E289E341CF5D0D10FF28FE7CFCFDFE32" box="[189,837,406,433]" pageId="16" pageNumber="46">
|
||
<taxonomicName id="0599BFECCF5D0D10FF28FE7CFDF9FE33" authority="Verhoeff, 1929" box="[189,577,406,432]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="16" pageNumber="46" phylum="Arthropoda" rank="variety" species="illyricus" variety="obscura">
|
||
<emphasis id="F0ED187DCF5D0D10FF28FE7CFE12FE33" box="[189,426,406,432]" italics="true" pageId="16" pageNumber="46">Typhloiulus illyricus</emphasis>
|
||
var.
|
||
<emphasis id="F0ED187DCF5D0D10FE76FE7CFDF9FE33" box="[483,577,406,432]" italics="true" pageId="16" pageNumber="46">obscura</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="4338E27ECF5D0D10FDC9FE7CFCFDFE32" author="Wolf" box="[604,837,406,433]" page="515" pageId="16" pageNumber="46" year="1934-38">
|
||
<bibRefCitation id="A608B99ECF5D0D10FDC9FE7CFD6FFE33" author="Wolf B." box="[604,727,406,432]" pageId="16" pageNumber="46" refId="ref22657" refString="Wolf B. 1934 - 38. Animalium Cavernarus Catalogus. ' sGravenhage, W. Junk, Vol. 3." type="journal volume" year="1934">Wolf 1934</bibRefCitation>
|
||
–38: 515.
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8397E4CF5D0D10FF28FE3DFDDBFD2E" pageId="16" pageNumber="46" type="materials_examined">
|
||
<paragraph id="C226C46FCF5D0D10FF28FE3DFE10FE72" blockId="16.[189,424,471,497]" box="[189,424,471,497]" pageId="16" pageNumber="46">
|
||
<heading id="996E7303CF5D0D10FF28FE3DFE10FE72" bold="true" box="[189,424,471,497]" fontSize="11" level="3" pageId="16" pageNumber="46" reason="3">
|
||
<emphasis id="F0ED187DCF5D0D10FF28FE3DFE10FE72" bold="true" box="[189,424,471,497]" pageId="16" pageNumber="46">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF5D0D10FF28FDECFDDBFD2E" blockId="16.[189,1398,518,685]" pageId="16" pageNumber="46">
|
||
<materialsCitation id="72F1CE32CF5D0D10FF28FDECFDCBFDC0" ID-GBIF-Occurrence="3504034331" collectingDate="2012-04-19" collectionCode="2 topotype juvs, IZB" collectorName="T. Delic" country="Slovenia" county="Ocizla" determinerName="D. Antic" location="Besko-Ocizeljski System" municipality="Besko-Ocizeljski System" pageId="16" pageNumber="46" specimenCount="1">
|
||
<collectingCountry id="BA8E84FFCF5D0D10FF28FDECFEF5FDA3" box="[189,333,518,544]" name="Slovenia" pageId="16" pageNumber="46">SLOVENIA</collectingCountry>
|
||
•
|
||
<collectionCode id="A4885CAACF5D0D10FEF2FDECFDA7FDA3" box="[359,543,518,544]" pageId="16" pageNumber="46">2 topotype juvs</collectionCode>
|
||
;
|
||
<collectingCounty id="2B47BCE3CF5D0D10FDB9FDECFDC3FDA3" box="[556,635,518,544]" pageId="16" pageNumber="46">Ocizla</collectingCounty>
|
||
,
|
||
<location id="C74692B4CF5D0D10FD12FDECFC14FDA3" LSID="urn:lsid:plazi:treatment:4A307579CF420D11FE6AFD30FABBFE33:C74692B4CF5D0D10FD12FDECFC14FDA3" box="[647,940,518,544]" country="Slovenia" county="Ocizla" municipality="Besko-Ocizeljski System" name="Besko-Ocizeljski System" pageId="16" pageNumber="46">Beško-Ocizeljski System</location>
|
||
(=
|
||
<location id="C74692B4CF5D0D10FC43FDECFB79FDA2" LSID="urn:lsid:plazi:treatment:4A307579CF420D11FE6AFD30FABBFE33:C74692B4CF5D0D10FC43FDECFB79FDA2" box="[982,1217,518,545]" country="Slovenia" county="Ocizla" municipality="Besko-Ocizeljski System" name="Ocizeljska Jama Pit" pageId="16" pageNumber="46">Ocizeljska Jama Pit</location>
|
||
);
|
||
<date id="B627E2AFCF5D0D10FB43FDECFACAFDA3" box="[1238,1394,518,544]" pageId="16" pageNumber="46" value="2012-04-19">
|
||
<collectingDate id="A6631B47CF5D0D10FB43FDECFACAFDA3" box="[1238,1394,518,544]" pageId="16" pageNumber="46" value="2012-04-19">19 Apr. 2012</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="6F6CA1B9CF5D0D10FF2BFDC3FEA3FDC0" box="[190,283,553,579]" pageId="16" pageNumber="46">T. Delić</collectorName>
|
||
leg.;
|
||
<determinerName id="37FA28ABCF5D0D10FEC8FDC3FE78FDC0" box="[349,448,553,579]" pageId="16" pageNumber="46">D. Antić</determinerName>
|
||
det. 2018;
|
||
<collectionCode id="A4885CAACF5D0D10FDD1FDC3FDCBFDC0" box="[580,627,553,579]" pageId="16" pageNumber="46">IZB</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="72F1CE32CF5D0D10FD18FDC3FCCDFDE4" ID-GBIF-Occurrence="3504034302" collectingDate="2014-04-22" collectionCode="IZB" collectorName="M. Zagmajster" country="Slovenia" county="Dane" determinerName="D. Antic" location="Med jamah, Medjama" municipality="Mejame Pit" pageId="16" pageNumber="46" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Divaca">
|
||
<specimenCount id="D49F0FE6CF5D0D10FD18FDC3FD04FDC0" box="[653,700,553,579]" pageId="16" pageNumber="46" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="D49F0FE6CF5D0D10FD5DFDC3FD4EFDC0" box="[712,758,553,579]" pageId="16" pageNumber="46" type="female">1 ♀</specimenCount>
|
||
;
|
||
<collectingCounty id="2B47BCE3CF5D0D10FC90FDC3FCFDFDC0" box="[773,837,553,579]" pageId="16" pageNumber="46">Dane</collectingCounty>
|
||
,
|
||
<collectingRegion id="005D0A8DCF5D0D10FCC4FDC3FC1EFDC0" box="[849,934,553,579]" country="Slovenia" name="Divaca" pageId="16" pageNumber="46">Divača</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="22425E15CF5D0D10FC27FDC3FB82FDC0" box="[946,1082,553,579]" pageId="16" pageNumber="46">Mejame Pit</collectingMunicipality>
|
||
(=
|
||
<location id="C74692B4CF5D0D10FBF1FDC3FAD3FDC7" LSID="urn:lsid:plazi:treatment:4A307579CF420D11FE6AFD30FABBFE33:C74692B4CF5D0D10FBF1FDC3FAD3FDC7" box="[1124,1387,553,580]" country="Slovenia" county="Dane" municipality="Mejame Pit" name="Med jamah, Medjama" pageId="16" pageNumber="46" stateProvince="Divaca">Med jamah, Medjama</location>
|
||
);
|
||
<date id="B627E2AFCF5D0D10FF28FDA6FEE3FDE5" box="[189,347,588,614]" pageId="16" pageNumber="46" value="2014-04-22">
|
||
<collectingDate id="A6631B47CF5D0D10FF28FDA6FEE3FDE5" box="[189,347,588,614]" pageId="16" pageNumber="46" value="2014-04-22">22 Apr. 2014</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="6F6CA1B9CF5D0D10FEFDFDA7FDADFDE5" box="[360,533,588,615]" pageId="16" pageNumber="46">M. Zagmajster</collectorName>
|
||
leg.;
|
||
<determinerName id="37FA28ABCF5D0D10FDCDFDA7FD05FDE5" box="[600,701,588,615]" pageId="16" pageNumber="46">D. Antić</determinerName>
|
||
det. 2018;
|
||
<collectionCode id="A4885CAACF5D0D10FCD6FDA7FCCDFDE4" box="[835,885,589,615]" pageId="16" pageNumber="46">IZB</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="72F1CE32CF5D0D10FC17FDA6FB83FD09" ID-GBIF-Occurrence="3504034333" collectingDate="2013-06-30" collectionCode="IZB" collectorName="D. Superina" country="Croatia" county="Rocko polje" determinerName="D. Antic" location="Rabakova Cave" municipality="Roc" pageId="16" pageNumber="46" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Istria">
|
||
<collectingCountry id="BA8E84FFCF5D0D10FC17FDA6FBB8FDE5" box="[898,1024,588,614]" name="Croatia" pageId="16" pageNumber="46">CROATIA</collectingCountry>
|
||
•
|
||
<specimenCount id="D49F0FE6CF5D0D10FB8FFDA6FBF1FDE4" box="[1050,1097,588,615]" pageId="16" pageNumber="46" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="D49F0FE6CF5D0D10FBC2FDA6FB3EFDE4" box="[1111,1158,588,615]" pageId="16" pageNumber="46" type="female">1 ♀</specimenCount>
|
||
;
|
||
<collectingRegion id="005D0A8DCF5D0D10FB00FDA6FB6BFDE5" box="[1173,1235,588,614]" country="Croatia" name="Istarska" pageId="16" pageNumber="46">Istria</collectingRegion>
|
||
,
|
||
<collectingCounty id="2B47BCE3CF5D0D10FB74FDA6FACAFDE5" box="[1249,1394,588,614]" pageId="16" pageNumber="46">Ročko polje</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="22425E15CF5D0D10FF28FD9AFF57FD09" box="[189,239,624,650]" pageId="16" pageNumber="46">Roč</collectingMunicipality>
|
||
,
|
||
<location id="C74692B4CF5D0D10FF6CFD85FE0BFD0A" LSID="urn:lsid:plazi:treatment:4A307579CF420D11FE6AFD30FABBFE33:C74692B4CF5D0D10FF6CFD85FE0BFD0A" box="[249,435,623,649]" country="Croatia" county="Rocko polje" municipality="Roc" name="Rabakova Cave" pageId="16" pageNumber="46" stateProvince="Istria">Rabakova Cave</location>
|
||
;
|
||
<date id="B627E2AFCF5D0D10FE2BFD85FDEEFD0A" box="[446,598,623,650]" pageId="16" pageNumber="46" value="2013-06-30">
|
||
<collectingDate id="A6631B47CF5D0D10FE2BFD85FDEEFD0A" box="[446,598,623,650]" pageId="16" pageNumber="46" value="2013-06-30">30 Jun. 2013</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="6F6CA1B9CF5D0D10FDF4FD9AFD52FD0F" box="[609,746,624,652]" pageId="16" pageNumber="46">D. Šuperina</collectorName>
|
||
leg.;
|
||
<determinerName id="37FA28ABCF5D0D10FCBFFD9AFC34FD0A" box="[810,908,623,650]" pageId="16" pageNumber="46">D. Antić</determinerName>
|
||
det. 2018;
|
||
<collectionCode id="A4885CAACF5D0D10FB99FD9AFB83FD09" box="[1036,1083,624,650]" pageId="16" pageNumber="46">IZB</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="72F1CE32CF5D0D10FBC7FD85FDE7FD2E" ID-GBIF-Occurrence="3504034307" collectingDate="2013-04-16" collectionCode="IZB" collectorName="D. Superina" country="Croatia" county="Rocko" determinerName="D. Antic" location="Rabakova Cave" municipality="Roc" pageId="16" pageNumber="46" specimenCount="1" specimenCount-female="1" stateProvince="Istarska">
|
||
<specimenCount id="D49F0FE6CF5D0D10FBC7FD85FBC6FD09" box="[1106,1150,623,650]" pageId="16" pageNumber="46" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<date id="B627E2AFCF5D0D10FE12FD78FD99FD2E" box="[391,545,658,685]" pageId="16" pageNumber="46" value="2013-04-16">
|
||
<collectingDate id="A6631B47CF5D0D10FE12FD78FD99FD2E" box="[391,545,658,685]" pageId="16" pageNumber="46" value="2013-04-16">16 Apr. 2013</collectingDate>
|
||
</date>
|
||
;
|
||
<collectionCode id="A4885CAACF5D0D10FDB8FD79FDE7FD2E" box="[557,607,659,685]" pageId="16" pageNumber="46">IZB</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8397E4CF5D0D10FF28FD3EFBFCFA59" pageId="16" pageNumber="46" type="diagnosis">
|
||
<paragraph id="C226C46FCF5D0D10FF28FD3EFE8CFD6D" blockId="16.[189,308,724,750]" box="[189,308,724,750]" pageId="16" pageNumber="46">
|
||
<heading id="996E7303CF5D0D10FF28FD3EFE8CFD6D" bold="true" box="[189,308,724,750]" fontSize="11" level="3" pageId="16" pageNumber="46" reason="3">
|
||
<emphasis id="F0ED187DCF5D0D10FF28FD3EFE8CFD6D" bold="true" box="[189,308,724,750]" pageId="16" pageNumber="46">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF5D0D10FF28FCE9FB13FC05" blockId="16.[189,1399,770,903]" pageId="16" pageNumber="46">
|
||
A species of
|
||
<taxonomicName id="0599BFECCF5D0D10FEC3FCE8FE70FC9F" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" box="[342,456,770,796]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="16" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0ED187DCF5D0D10FEC3FCE8FE70FC9F" box="[342,456,770,796]" italics="true" pageId="16" pageNumber="46">Stygiiulus</emphasis>
|
||
</taxonomicName>
|
||
with normal mouthparts. Distinguishable from congeners by the combination of certain gonopodal characters (
|
||
<figureCitation id="5AA2D8EACF5D0D10FD89FCCCFD3EFCC3" box="[540,646,806,832]" captionStart="Fig" captionStartId="26.[189,232,1822,1848]" captionTargetBox="[197,1384,282,1779]" captionTargetId="figure-12@26.[189,1398,263,1787]" captionTargetPageId="26" captionText="Fig. 10. Left gonopods of species of Stygiiulus stat. nov., mesal views. A. S. ausugi (Manfredi, 1953) comb.nov.,topotype♂(NHMD).B.S.illyricus(Verhoeff,1929)comb.nov.,redrawnfromVerhoeff (1929). C. S. insularis (Strasser, 1938) comb. nov., redrawn from Antić et al. (2018). D. S. maximus (Verhoeff, 1929) comb. nov., redrawn from Verhoeff (1930). E. S. seewaldi (Strasser, 1967) comb. nov., redrawn and modified from Strasser (1967). F. S. tobias (Berlese, 1886) comb. nov., ♂ from Grotta [Cave] del Subiolo (NHMD). Abbreviations: do = distal outgrowth of velum. Drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.6323028" httpUri="https://zenodo.org/record/6323028/files/figure.png" pageId="16" pageNumber="46">Fig. 10B</figureCitation>
|
||
), viz., a mostly straight promere, a mesomere with the apex bent frontad, and an opisthomere with a small but distinct posterior hump, a broad and deeply serrated velum, and a vestigial posterior solenomeral branch, this being completely fused to the velum.
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF5D0D10FF28FC47FE29FC4B" blockId="16.[189,401,941,968]" box="[189,401,941,968]" pageId="16" pageNumber="46">
|
||
<heading id="996E7303CF5D0D10FF28FC47FE29FC4B" bold="true" box="[189,401,941,968]" fontSize="11" level="3" pageId="16" pageNumber="46" reason="3">
|
||
<emphasis id="F0ED187DCF5D0D10FF28FC47FE29FC4B" bold="true" box="[189,401,941,968]" pageId="16" pageNumber="46">Descriptive notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF5D0D10FF28FC36FD4CFBBF" blockId="16.[189,1399,988,1085]" pageId="16" pageNumber="46">ANTENNAE. 2.2 times as long as head and ca 1.8 times as long as H in males, and 1.7 times and ca 1.5– 1.55 times, respectively, in females; antennomere 5 ca 2.4 times as long as broad; antennomeres 2–5 subequal in length, ca 1.3–1.4 times as long as 6.</paragraph>
|
||
<paragraph id="C226C46FCF5D0D10FF28FB8EFD03FB21" blockId="16.[189,1398,1124,1186]" pageId="16" pageNumber="46">TARSUS OF MID- BODY LEGS. Ca 2.4 times as long as tibia and ca 2 times as long as apical claw. Mid-body legs ca 1.1 times as long as H in both sexes.</paragraph>
|
||
<paragraph id="C226C46FCF5D0D10FF28FB23FBFCFA59" blockId="16.[189,1400,1224,1498]" pageId="16" pageNumber="46">
|
||
FEMALE SEXUAL CHARACTERS. Leg-pairs 1 and 2 considerably shorter than, but as thick as, following legs. Vulva (
|
||
<figureCitation id="5AA2D8EACF5D0D10FE81FB06FEC3FA85" box="[276,379,1260,1286]" captionStart="Fig" captionStartId="31.[189,231,1681,1707]" captionTargetBox="[345,1243,265,1635]" captionTargetId="figure-15@31.[325,1262,262,1645]" captionTargetPageId="31" captionText="Fig. 11. Left vulvae of species of Stygiiulus stat. nov. (A–D) and representatives of (other) Julini/ Leptoiulini (E) and Typhloiulini/blind Julini/Leptoiulini in caudal, somewhat mesal views. A. S. ausugi (Manfredi, 1953) comb. nov., ♀ from Grotta [Cave] del Calgeron (NHMD). B. S. illyricus (Verhoef, 1929) comb. nov., ♀ from Rabakova Cave (IZB). C. S. insularis (Strasser, 1938) comb. nov., topotype ♀ (CBSS). D. S. tobias (Berlese, 1886) comb. nov., ♀ from Grotta [Cave] del Subiolo (NHMD). E. Leptoiulus cf. trilineatus(C.L. Koch, 1847), ♀ from Belasitsa Mtn, Bulgaria. F. Serboiulus spelaeophilus Gulička, 1967, ♀ from Vodni Pech Cave (NMNHS). G. Typhloiulus (Inversotyphlus) lobifer Attems, 1951, ♀ from Minjera-Bauxite Mine (IBER). H. Typhloiulus (s. str.) strictus (Latzel, 1882), ♀ from Kunino (IBER). Abbreviations: at = anterior tube; la = lateral ampulla; lt = lateral tube; ma = median ampulla; mt = median tube; op = operculum; pa = posterior ampulla; pt = posterior tube. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6323030" httpUri="https://zenodo.org/record/6323030/files/figure.png" pageId="16" pageNumber="46">Fig. 11B</figureCitation>
|
||
) slightly compressed in the sagittal plane, mostly symmetric; median cleft deeper than in most congeners; each valve of bursa with several, mostly vertically arranged setae, 1–2 setae on side sclerites; operculum (op) distally markedly thickened, with a concave apical margin, exceeding bursa by slightly more than ¼ of total height of vulva, medio-laterally with two longitudinal rows of setae each side, with a bunch of setae on each latero-apical corner. Receptaculum seminis consisting of a fine, slightly bent, mesal tube (mt) ending in a small piriform ampulla (ma), and a somewhat shorter lateral tube (lt) ending in an ampulla (la) of similar shape and size as the former one.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8397E4CF5D0D10FF28F9EAFDD8F9EE" pageId="16" pageNumber="46" type="distribution">
|
||
<paragraph id="C226C46FCF5D0D10FF28F9EAFEEDF999" blockId="16.[189,341,1536,1562]" box="[189,341,1536,1562]" pageId="16" pageNumber="46">
|
||
<heading id="996E7303CF5D0D10FF28F9EAFEEDF999" bold="true" box="[189,341,1536,1562]" fontSize="11" level="3" pageId="16" pageNumber="46" reason="3">
|
||
<emphasis id="F0ED187DCF5D0D10FF28F9EAFEEDF999" bold="true" box="[189,341,1536,1562]" pageId="16" pageNumber="46">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF5D0D10FF28F9C5FDD8F9EE" blockId="16.[188,1399,1583,1645]" pageId="16" pageNumber="46">
|
||
Known exclusively from caves and pits in the Istrian Peninsula in
|
||
<collectingCountry id="BA8E84FFCF5D0D10FC26F9C5FBB0F9CA" box="[947,1032,1583,1609]" name="Croatia" pageId="16" pageNumber="46">Croatia</collectingCountry>
|
||
and the northwestern Dinarides in
|
||
<collectingCountry id="BA8E84FFCF5D0D10FF4FF9B8FEF8F9EF" box="[218,320,1618,1644]" name="Slovenia" pageId="16" pageNumber="46">Slovenia</collectingCountry>
|
||
(
|
||
<figureCitation id="5AA2D8EACF5D0D10FEDAF9B8FE1CF9EF" box="[335,420,1618,1644]" captionStart="Fig" captionStartId="33.[189,232,1998,2024]" captionTargetBox="[189,1398,1099,1960]" captionTargetId="figure-332@33.[189,1398,1099,1960]" captionTargetPageId="33" captionText="Fig. 13. General distribution of the species of the genus Stygiiulus stat. nov." figureDoi="http://doi.org/10.5281/zenodo.6323034" httpUri="https://zenodo.org/record/6323034/files/figure.png" pageId="16" pageNumber="46">Fig. 13</figureCitation>
|
||
, green squares).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8397E4CF5D0D11FF28F97EFABBFE33" lastPageId="17" lastPageNumber="47" pageId="16" pageNumber="46" type="discussion">
|
||
<paragraph id="C226C46FCF5D0D10FF28F97EFE9BF92D" blockId="16.[189,291,1684,1710]" box="[189,291,1684,1710]" pageId="16" pageNumber="46">
|
||
<heading id="996E7303CF5D0D10FF28F97EFE9BF92D" bold="true" box="[189,291,1684,1710]" fontSize="11" level="3" pageId="16" pageNumber="46" reason="3">
|
||
<emphasis id="F0ED187DCF5D0D10FF28F97EFE9BF92D" bold="true" box="[189,291,1684,1710]" pageId="16" pageNumber="46">Remark</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF5D0D11FF28F929FABBFE33" blockId="16.[187,1400,1730,2003]" lastBlockId="17.[189,1399,265,432]" lastPageId="17" lastPageNumber="47" pageId="16" pageNumber="46">
|
||
From Rabakova Cave,
|
||
<bibRefCitation id="A608B99ECF5D0D10FE55F928FDC1F95E" author="Verhoeff K. W." box="[448,633,1730,1757]" pageId="16" pageNumber="46" pagination="1 - 21" refId="ref22622" refString="Verhoeff K. W. 1933. Arthropoden aus sudostalpinen Hohlen, gesammelt von Karl Strasser. 7. Aufsatz. Mitteilungen uber Hohlen- und Karstforschung 1933 (4): 1 - 21." type="journal article" year="1933">Verhoeff (1933)</bibRefCitation>
|
||
described the subspecies
|
||
<emphasis id="F0ED187DCF5D0D10FC0CF928FB87F95F" box="[921,1087,1730,1756]" italics="true" pageId="16" pageNumber="46">
|
||
<taxonomicName id="0599BFECCF5D0D10FC0CF928FC40F95F" authorityName="Verhoeff" authorityYear="1929" box="[921,1016,1730,1756]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="16" pageNumber="46" phylum="Arthropoda" rank="species" species="illyricus">illyricus</taxonomicName>
|
||
stygis
|
||
</emphasis>
|
||
, mentioning the possibility that the observed differences between the single male his description was based upon and the typical
|
||
<emphasis id="F0ED187DCF5D0D10FF28F8E2FEA8F8A1" box="[189,272,1800,1826]" italics="true" pageId="16" pageNumber="46">illyricu</emphasis>
|
||
s were due to periodomorphosis, but he nevertheless opted to give a formal name to the new morph. Indeed, the gonopods he drew seem to be underdeveloped (see
|
||
<bibRefCitation id="A608B99ECF5D0D10FC1DF8C6FBE3F8C5" author="Verhoeff K. W." box="[904,1115,1836,1862]" pageId="16" pageNumber="46" pagination="1 - 21" refId="ref22622" refString="Verhoeff K. W. 1933. Arthropoden aus sudostalpinen Hohlen, gesammelt von Karl Strasser. 7. Aufsatz. Mitteilungen uber Hohlen- und Karstforschung 1933 (4): 1 - 21." type="journal article" year="1933">Verhoeff 1933: 20</bibRefCitation>
|
||
, fig. 28). Fortunately, we examined here
|
||
<specimenCount id="D49F0FE6CF5D0D10FEE5F8A5FE65F8EA" box="[368,477,1871,1897]" pageId="16" pageNumber="46" type="adult">one adult</specimenCount>
|
||
male from Rabakova Cave, whose gonopods appeared completely identical to the gonopods of the typical
|
||
<taxonomicName id="0599BFECCF5D0D10FE6BF898FDE4F80F" authorityName="Verhoeff" authorityYear="1929" box="[510,604,1906,1932]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="16" pageNumber="46" phylum="Arthropoda" rank="species" species="illyricus">
|
||
<emphasis id="F0ED187DCF5D0D10FE6BF898FDE4F80F" box="[510,604,1906,1932]" italics="true" pageId="16" pageNumber="46">illyricus</emphasis>
|
||
</taxonomicName>
|
||
. We were even luckier to find that one of the juveniles from the
|
||
<typeStatus id="1D227ACDCF5D0D10FAD0F899FACFF80E" box="[1349,1399,1907,1933]" pageId="16" pageNumber="46">type</typeStatus>
|
||
locality of the typical
|
||
<taxonomicName id="0599BFECCF5D0D10FE54F87FFD98F82C" authorityName="Verhoeff" authorityYear="1929" box="[449,544,1941,1967]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="16" pageNumber="46" phylum="Arthropoda" rank="species" species="illyricus">
|
||
<emphasis id="F0ED187DCF5D0D10FE54F87FFD98F82C" box="[449,544,1941,1967]" italics="true" pageId="16" pageNumber="46">illyricus</emphasis>
|
||
</taxonomicName>
|
||
was a subadult male with unmodified first pair of legs. The gonopods of this subadult male are very similar to
|
||
<bibRefCitation id="A608B99ECF5D0D10FDF9F852FC8DF850" author="Verhoeff K. W." box="[620,821,1976,2003]" pageId="16" pageNumber="46" pagination="1 - 21" refId="ref22622" refString="Verhoeff K. W. 1933. Arthropoden aus sudostalpinen Hohlen, gesammelt von Karl Strasser. 7. Aufsatz. Mitteilungen uber Hohlen- und Karstforschung 1933 (4): 1 - 21." type="journal article" year="1933">Verhoeff's (1933)</bibRefCitation>
|
||
drawing of the gonopods of the subspecies
|
||
<emphasis id="F0ED187DCF5D0D10FAB8F853FAD7F851" box="[1325,1391,1977,2002]" italics="true" pageId="16" pageNumber="46">stygis</emphasis>
|
||
. Based on these observations, it is clear that Verhoeff's
|
||
<emphasis id="F0ED187DCF5C0D11FCD1FEE0FC3EFEA0" box="[836,902,266,291]" italics="true" pageId="17" pageNumber="47">stygis</emphasis>
|
||
was actually a non-adult male or “Junior- Männchen” [junior male] – a term coined by
|
||
<bibRefCitation id="A608B99ECF5C0D11FD71FEC6FC1FFEC4" author="Strasser K." box="[740,935,300,327]" pageId="17" pageNumber="47" pagination="313 - 345" refId="ref22368" refString="Strasser K. 1971 b. Uber Diplopoden Jugoslawiens. Senckenbergiana biologica 53: 313 - 345." type="journal article" year="1971">Strasser (1971b)</bibRefCitation>
|
||
, referring to typhloiulinine males with almost fully developed gonopods, but with non-modified first pair of legs and other, less conspicuous, underdeveloped external somatic structures (see also
|
||
<bibRefCitation id="A608B99ECF5C0D11FCD4FE98FBF4FE0E" author="Vagalinski B. & Stoev P. & Enghoff H." box="[833,1100,370,397]" pageId="17" pageNumber="47" pagination="334 - 362" refId="ref22493" refString="Vagalinski B., Stoev P. & Enghoff H. 2015. A review of the millipede genus Typhloiulus Latzel, 1884 (Diplopoda: Julida: Julidae), with a description of three new species from Bulgaria and Greece. Zootaxa 3999 (3): 334 - 362. https: // doi. org / 10.11646 / zootaxa. 3999.3.2" type="journal article" year="2015">
|
||
Vagalinski
|
||
<emphasis id="F0ED187DCF5C0D11FC52FE99FBBEFE0F" box="[967,1030,370,396]" italics="true" pageId="17" pageNumber="47">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
). We thus consider here
|
||
<taxonomicName id="0599BFECCF5C0D11FF28FE7CFE39FE33" box="[189,385,406,432]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="17" pageNumber="47" phylum="Arthropoda" rank="subSpecies" species="illyricus" subSpecies="stygis">
|
||
<emphasis id="F0ED187DCF5C0D11FF28FE7CFE39FE33" box="[189,385,406,432]" italics="true" pageId="17" pageNumber="47">S. illyricus stygis</emphasis>
|
||
</taxonomicName>
|
||
syn. nov. as a junior subjective synonym of the typical
|
||
<taxonomicName id="0599BFECCF5C0D11FB94FE7CFBC5FE33" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Verhoeff" baseAuthorityYear="1929" box="[1025,1149,406,432]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="17" pageNumber="47" phylum="Arthropoda" rank="species" species="illyricus" status="comb. nov.">
|
||
<emphasis id="F0ED187DCF5C0D11FB94FE7CFBC5FE33" box="[1025,1149,406,432]" italics="true" pageId="17" pageNumber="47">S. illyricus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF5C0D11FB16FE7CFABBFE33" box="[1155,1283,406,432]" pageId="17" pageNumber="47" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |