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<document id="53078A1D0E80DE2CD61930EA8CC6DF4E" ID-DOI="10.11646/zootaxa.4758.2.4" ID-GBIF-Dataset="a4c437e0-7983-41af-8f98-a3bd88cc2e69" ID-ISSN="1175-5326" ID-Zenodo-Dep="3734456" ID-ZooBank="2539ABCC-161E-44B2-BB2C-C949B1A7C94D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585658008729" checkinUser="plazi" docAuthor="Shear, William A." docDate="2020" docId="0F4F0D64253DFFA7FF69FD73FB46B8FE" docLanguage="en" docName="zootaxa.4758.2.4.pdf" docOrigin="Zootaxa 4758 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trisaria rex Shear 2020, new species" docType="treatment" docVersion="5" lastPageNumber="284" masterDocId="F376751C2534FFAEFFFEFFBBFFB9BE2B" masterDocTitle="The millipede family Striariidae Bollman, 1893: I. Introduction to the family, synonymy of Vaferaria Causey with Amplaria Chamberlin, the new subfamily Trisariinae, the new genus Trisaria, and three new species (Diplopoda, Chordeumatida, Striarioidea)" masterLastPageNumber="295" masterPageNumber="275" pageNumber="284" updateTime="1698733589358" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="60D16847864EAEEE77F07F8F180DC954">The millipede family Striariidae Bollman, 1893: I. Introduction to the family, synonymy of Vaferaria Causey with Amplaria Chamberlin, the new subfamily Trisariinae, the new genus Trisaria, and three new species (Diplopoda, Chordeumatida, Striarioidea)</mods:title>
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<mods:namePart id="3DB559D0A02267A54CE3D7D1F0C2B37D">Shear, William A.</mods:namePart>
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<mods:title id="5D75173A2E5D3B6D343CCA5C3864729B">Zootaxa</mods:title>
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<mods:date id="517438DF20C5726539B7F9958913F7A3">2020</mods:date>
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<mods:number id="C36E8363F48F4997B807F8C7A7BDD204">2020-03-30</mods:number>
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<treatment id="0F4F0D64253DFFA7FF69FD73FB46B8FE" ID-DOI="http://doi.org/10.5281/zenodo.3812351" ID-GBIF-Taxon="163417953" ID-Zenodo-Dep="3812351" LSID="urn:lsid:plazi:treatment:0F4F0D64253DFFA7FF69FD73FB46B8FE" httpUri="http://treatment.plazi.org/id/0F4F0D64253DFFA7FF69FD73FB46B8FE" lastPageNumber="284" pageId="9" pageNumber="284">
<subSubSection id="CFFCEFF9253DFFA7FF69FD73FE7CBCCA" box="[151,453,711,738]" pageId="9" pageNumber="284" type="nomenclature">
<paragraph id="8759BC72253DFFA7FF69FD73FE7CBCCA" blockId="9.[151,453,711,772]" box="[151,453,711,738]" pageId="9" pageNumber="284">
<heading id="DC110B1E253DFFA7FF69FD73FE7CBCCA" bold="true" box="[151,453,711,738]" fontSize="11" level="1" pageId="9" pageNumber="284" reason="1">
<emphasis id="B5926060253DFFA7FF69FD73FE7CBCCA" bold="true" box="[151,453,711,738]" pageId="9" pageNumber="284">
<taxonomicName id="40E6C7F1253DFFA7FF69FD73FE9FBCC9" ID-CoL="84DX9" authority="Shear, 2020" authorityName="Shear" authorityYear="2020" box="[151,294,712,738]" class="Diplopoda" family="Striariidae" genus="Trisaria" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="284" phylum="Arthropoda" rank="species" species="rex" status="sp. nov.">
<emphasis id="B5926060253DFFA7FF69FD73FE9FBCC9" bold="true" box="[151,294,712,738]" italics="true" pageId="9" pageNumber="284">Trisaria rex</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="AEA1DD1B253DFFA7FECBFD73FE7CBCCA" box="[309,453,711,738]" pageId="9" pageNumber="284" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFFCEFF9253DFFA7FF69FD51FC28BDA4" pageId="9" pageNumber="284" type="description">
<paragraph id="8759BC72253DFFA7FF69FD51FEA5BD2F" blockId="9.[151,453,711,772]" box="[151,284,745,772]" pageId="9" pageNumber="284">
<figureCitation id="1FDDA0F7253DFFA7FF69FD51FEA5BD2F" box="[151,284,745,772]" captionStart-0="FIGURES 1519" captionStart-1="FIGURES 2025" captionStart-2="FIGURES 2631" captionStartId-0="10.[151,264,1682,1707]" captionStartId-1="11.[151,264,1815,1840]" captionStartId-2="12.[151,264,1861,1886]" captionTargetBox-0="[151,1436,236,1657]" captionTargetBox-1="[225,1362,184,1787]" captionTargetBox-2="[197,1397,185,1830]" captionTargetId-0="figure@10.[151,1436,236,1657]" captionTargetId-1="figure@11.[225,1362,181,1789]" captionTargetId-2="figure@12.[190,1397,181,1836]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURES 1519. Trisaria rex. Fig. 15. Gnathochilarium of male, ventral view; isp, inner stipital palp, ll, laminae linguales, lp, lingual palp, m, promentum, osp, outer stipital palp, stg, stipes of gnathochilarium. Fig. 16. Terminal structures, ventral view; epi, epiproct, hypo, hypoproct, para, paraproct. Fig. 17. Specialized setae of legpair 1. Fig. 18. Legpair 3, anterior view; cx3, coxa, pf3, prefemur 3, s3, sternum 3. Fig. 19. Tip of coxal flask of left legpair 3, anterior view." captionText-1="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." captionText-2="FIGURES 2631. Trisaria rex and Trisaria washingtonensis. Figs. 2628. Trisaria rex. Fig 26. Anterior gonopods, anterior view; labels as in Figs. 21 and 22. Fig. 27. Gonopods, posterior view; labels as in Figs. 21 and 22. Fig. 28. Left leg 9 of male, anterior view; ap, apodeme; other labels as in Fig. 8. Figs. 2931. Trisaria washingtonensis. Fig. 29. Left leg 9 of male, anterior view; sp, spiracle; other labels as in Fig. 8. Fig. 30. Gonopods, anterior view; gst, gonopod sternum; other labels as in Figs. 21 &amp; 22. Fig. 31. Gonopods, posterior view; labels as in Fig. 27." figureDoi-0="http://doi.org/10.5281/zenodo.3734464" figureDoi-1="http://doi.org/10.5281/zenodo.3734466" figureDoi-2="http://doi.org/10.5281/zenodo.3734468" httpUri-0="https://zenodo.org/record/3734464/files/figure.png" httpUri-1="https://zenodo.org/record/3734466/files/figure.png" httpUri-2="https://zenodo.org/record/3734468/files/figure.png" pageId="9" pageNumber="284">Figs. 1528</figureCitation>
</paragraph>
<paragraph id="8759BC72253DFFA7FF69FC8BFC28BDA4" blockId="9.[151,1437,815,1750]" pageId="9" pageNumber="284">
<emphasis id="B5926060253DFFA7FF69FC8BFF5FBD61" bold="true" box="[151,230,816,842]" pageId="9" pageNumber="284">Types:</emphasis>
Male
<typeStatus id="585D02D0253DFFA7FECBFC94FE2EBD62" box="[309,407,815,841]" pageId="9" pageNumber="284" type="holotype">holotype</typeStatus>
and
<specimenCount id="91E077FB253DFFA7FE2AFC8BFD80BD62" box="[468,569,815,842]" pageId="9" pageNumber="284" type="female">3 female</specimenCount>
<typeStatus id="585D02D0253DFFA7FDBAFC8BFD16BD61" box="[580,687,816,842]" pageId="9" pageNumber="284" type="paratype">paratypes</typeStatus>
from. I-90 exit #37, 1300 asl,
<geoCoordinate id="E2D2DAB5253DFFA7FBD8FC94FB03BD62" box="[1062,1210,815,841]" degrees="47" direction="north" minutes="26.613" orientation="latitude" pageId="9" pageNumber="284" precision="1" value="47.44355">N47°26.613</geoCoordinate>
,
<geoCoordinate id="E2D2DAB5253DFFA7FB37FC94FAC8BD62" box="[1225,1393,815,841]" degrees="121" direction="west" minutes="40.061" orientation="longitude" pageId="9" pageNumber="284" precision="1" value="-121.66769">W121°40.061</geoCoordinate>
,
<date id="F3589AB2253DFFA7FA7EFC8BFE88BD47" pageId="9" pageNumber="284" value="2005-01-21">21 January 2005</date>
, W. Leonard,
<emphasis id="B5926060253DFFA7FE30FCEAFD83BD47" bold="true" box="[462,570,849,876]" pageId="9" pageNumber="284">King Co.</emphasis>
,
<collectingRegion id="45227290253DFFA7FDB7FCE9FD68BD47" box="[585,721,850,876]" country="United States of America" name="Washington" pageId="9" pageNumber="284">Washington</collectingRegion>
,
<materialsCitation id="378EB62F253DFFA7FD23FCEAFD19BDA4" ID-GBIF-Occurrence="2592925702" collectingDate="2004-02-25" collectorName="W. Leonard &amp; C. Richart &amp; Burke Memorial Museum &amp; University" location="Washington" pageId="9" pageNumber="284" specimenCount="1" stateProvince="Washington">
<date id="F3589AB2253DFFA7FD23FCEAFC11BD47" box="[733,936,849,876]" pageId="9" pageNumber="284" value="2004-02-25">
<collectingDate id="E31C635A253DFFA7FD23FCEAFC11BD47" box="[733,936,849,876]" pageId="9" pageNumber="284" value="2004-02-25">25 February 2004</collectingDate>
</date>
,
<collectorName id="2A13D9A4253DFFA7FC4DFCE8FB80BD47" box="[947,1081,850,877]" pageId="9" pageNumber="284">W. Leonard</collectorName>
,
<collectorName id="2A13D9A4253DFFA7FBBBFCE9FB02BD47" box="[1093,1211,850,876]" pageId="9" pageNumber="284">C. Richart</collectorName>
, deposited in
<collectorName id="2A13D9A4253DFFA7FAA6FCE9FECBBDA4" pageId="9" pageNumber="284">Burke Memorial Museum</collectorName>
,
<collectorName id="2A13D9A4253DFFA7FE83FCCEFE4ABDA4" box="[381,499,885,911]" pageId="9" pageNumber="284">University</collectorName>
of
<collectingRegion id="45227290253DFFA7FDE6FCCEFD19BDA4" box="[536,672,885,911]" country="United States of America" name="Washington" pageId="9" pageNumber="284">Washington</collectingRegion>
</materialsCitation>
, Seattle,
<collectingRegion id="45227290253DFFA7FCFAFCCEFC34BDA4" box="[772,909,885,911]" country="United States of America" name="Washington" pageId="9" pageNumber="284">Washington</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="CFFCEFF9253DFFA7FF39FC23FF42BDFF" pageId="9" pageNumber="284" type="etymology">
<paragraph id="8759BC72253DFFA7FF39FC23FF42BDFF" blockId="9.[151,1437,815,1750]" pageId="9" pageNumber="284">
<emphasis id="B5926060253DFFA7FF39FC23FEF7BD99" bold="true" box="[199,334,920,946]" pageId="9" pageNumber="284">Etymology:</emphasis>
The species epithet (Latin, “king”) refers to King County,
<collectingRegion id="45227290253DFFA7FC21FC2CFBDABD9A" box="[991,1123,919,945]" country="United States of America" name="Washington" pageId="9" pageNumber="284">Washington</collectingRegion>
, to which the species seems endemic.
</paragraph>
</subSubSection>
<subSubSection id="CFFCEFF9253DFFA7FF39FC67FA25BA17" pageId="9" pageNumber="284" type="diagnosis">
<paragraph id="8759BC72253DFFA7FF39FC67FA25BA17" blockId="9.[151,1437,815,1750]" pageId="9" pageNumber="284">
<emphasis id="B5926060253DFFA7FF39FC67FEFABDDC" bold="true" box="[199,323,988,1015]" pageId="9" pageNumber="284">Diagnosis:</emphasis>
Distinct in the form of the gonopods and ninth legs. The posterior branch of the median angiocoxite is very long (
<emphasis id="B5926060253DFFA7FED6FC44FED7BA32" bold="true" box="[296,366,1023,1049]" pageId="9" pageNumber="284">mac2,</emphasis>
<figureCitation id="1FDDA0F7253DFFA7FE8AFC44FE69BA31" box="[372,464,1023,1050]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Figs. 22</figureCitation>
,
<figureCitation id="1FDDA0F7253DFFA7FE24FBBBFE40BA31" box="[474,505,1024,1050]" captionStart="FIGURES 2631" captionStartId="12.[151,264,1861,1886]" captionTargetBox="[197,1397,185,1830]" captionTargetId="figure@12.[190,1397,181,1836]" captionTargetPageId="12" captionText="FIGURES 2631. Trisaria rex and Trisaria washingtonensis. Figs. 2628. Trisaria rex. Fig 26. Anterior gonopods, anterior view; labels as in Figs. 21 and 22. Fig. 27. Gonopods, posterior view; labels as in Figs. 21 and 22. Fig. 28. Left leg 9 of male, anterior view; ap, apodeme; other labels as in Fig. 8. Figs. 2931. Trisaria washingtonensis. Fig. 29. Left leg 9 of male, anterior view; sp, spiracle; other labels as in Fig. 8. Fig. 30. Gonopods, anterior view; gst, gonopod sternum; other labels as in Figs. 21 &amp; 22. Fig. 31. Gonopods, posterior view; labels as in Fig. 27." figureDoi="http://doi.org/10.5281/zenodo.3734468" httpUri="https://zenodo.org/record/3734468/files/figure.png" pageId="9" pageNumber="284">27</figureCitation>
), while it is short in both
<taxonomicName id="40E6C7F1253DFFA7FCF1FBBBFC3ABA31" box="[783,899,1024,1050]" class="Diplopoda" family="Striariidae" genus="Trisaria" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="284" phylum="Arthropoda" rank="species" species="olympia">
<emphasis id="B5926060253DFFA7FCF1FBBBFC3ABA31" box="[783,899,1024,1050]" italics="true" pageId="9" pageNumber="284">T. olympia</emphasis>
</taxonomicName>
and
<emphasis id="B5926060253DFFA7FC49FBBBFB2BBA31" box="[951,1170,1024,1050]" italics="true" pageId="9" pageNumber="284">
<taxonomicName id="40E6C7F1253DFFA7FC49FBBBFB37BA31" box="[951,1166,1024,1050]" class="Diplopoda" family="Striariidae" genus="Trisaria" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="284" phylum="Arthropoda" rank="species" species="washingtonensis">T. washingtonensis</taxonomicName>
.
</emphasis>
In contrast to the robust form of the ninth leg telopodite in the other two species, that of
<taxonomicName id="40E6C7F1253DFFA7FC9CFB98FC18BA17" authorityName="Shear" authorityYear="2020" box="[866,929,1059,1084]" class="Diplopoda" family="Striariidae" genus="Trisaria" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="284" phylum="Arthropoda" rank="species" species="rex">
<emphasis id="B5926060253DFFA7FC9CFB98FC18BA17" box="[866,929,1059,1084]" italics="true" pageId="9" pageNumber="284">T. rex</emphasis>
</taxonomicName>
is strongly compressed (
<emphasis id="B5926060253DFFA7FB46FB99FB6EBA17" bold="true" box="[1208,1239,1058,1084]" pageId="9" pageNumber="284">t9,</emphasis>
<figureCitation id="1FDDA0F7253DFFA7FB20FB99FADDBA17" box="[1246,1380,1058,1084]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Figs. 20, 23</figureCitation>
,
<figureCitation id="1FDDA0F7253DFFA7FA8EFB99FA29BA17" box="[1392,1424,1058,1084]" captionStart="FIGURES 2631" captionStartId="12.[151,264,1861,1886]" captionTargetBox="[197,1397,185,1830]" captionTargetId="figure@12.[190,1397,181,1836]" captionTargetPageId="12" captionText="FIGURES 2631. Trisaria rex and Trisaria washingtonensis. Figs. 2628. Trisaria rex. Fig 26. Anterior gonopods, anterior view; labels as in Figs. 21 and 22. Fig. 27. Gonopods, posterior view; labels as in Figs. 21 and 22. Fig. 28. Left leg 9 of male, anterior view; ap, apodeme; other labels as in Fig. 8. Figs. 2931. Trisaria washingtonensis. Fig. 29. Left leg 9 of male, anterior view; sp, spiracle; other labels as in Fig. 8. Fig. 30. Gonopods, anterior view; gst, gonopod sternum; other labels as in Figs. 21 &amp; 22. Fig. 31. Gonopods, posterior view; labels as in Fig. 27." figureDoi="http://doi.org/10.5281/zenodo.3734468" httpUri="https://zenodo.org/record/3734468/files/figure.png" pageId="9" pageNumber="284">28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CFFCEFF9253DFFA7FF39FBFFFF6DB80A" pageId="9" pageNumber="284" type="description">
<paragraph id="8759BC72253DFFA7FF39FBFFFCE7BBF7" blockId="9.[151,1437,815,1750]" pageId="9" pageNumber="284">
<emphasis id="B5926060253DFFA7FF39FBFFFCD3BA74" bold="true" box="[199,874,1092,1119]" pageId="9" pageNumber="284">Description: Male from Twin Falls/Iron Horse trailhead.</emphasis>
Length, 8.0 mm, width,
<quantity id="401E1197253DFFA7FB7EFBFEFB51BA74" box="[1152,1256,1093,1119]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="9" pageNumber="284" unit="mm" value="0.82">0.82 mm</quantity>
. Body form and secondary sexual modifications as described for genus. Gonopods (
<figureCitation id="1FDDA0F7253DFFA7FC71FBDCFBA8BAAA" box="[911,1041,1127,1153]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Figs.2022</figureCitation>
,
<figureCitation id="1FDDA0F7253DFFA7FBE2FBDCFBDDBAA9" box="[1052,1124,1127,1154]" captionStart="FIGURES 2631" captionStartId="12.[151,264,1861,1886]" captionTargetBox="[197,1397,185,1830]" captionTargetId="figure@12.[190,1397,181,1836]" captionTargetPageId="12" captionText="FIGURES 2631. Trisaria rex and Trisaria washingtonensis. Figs. 2628. Trisaria rex. Fig 26. Anterior gonopods, anterior view; labels as in Figs. 21 and 22. Fig. 27. Gonopods, posterior view; labels as in Figs. 21 and 22. Fig. 28. Left leg 9 of male, anterior view; ap, apodeme; other labels as in Fig. 8. Figs. 2931. Trisaria washingtonensis. Fig. 29. Left leg 9 of male, anterior view; sp, spiracle; other labels as in Fig. 8. Fig. 30. Gonopods, anterior view; gst, gonopod sternum; other labels as in Figs. 21 &amp; 22. Fig. 31. Gonopods, posterior view; labels as in Fig. 27." figureDoi="http://doi.org/10.5281/zenodo.3734468" httpUri="https://zenodo.org/record/3734468/files/figure.png" pageId="9" pageNumber="284">26, 27</figureCitation>
) robust, sternum (
<emphasis id="B5926060253DFFA7FACAFBD3FAEFBAA9" bold="true" box="[1332,1366,1128,1154]" pageId="9" pageNumber="284">gst</emphasis>
,
<figureCitation id="1FDDA0F7253DFFA7FA9BFBDCFF58BA8F" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Figs. 20, 21</figureCitation>
) broad, well-sclerotized. Median angiocoxite (
<emphasis id="B5926060253DFFA7FD02FB31FC97BA8F" bold="true" box="[764,814,1162,1188]" pageId="9" pageNumber="284">mac</emphasis>
,
<figureCitation id="1FDDA0F7253DFFA7FCC1FB31FC2BBA8F" box="[831,914,1162,1188]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Fig. 20</figureCitation>
) with two branches: the anteriormost (
<emphasis id="B5926060253DFFA7FAABFB31FA24BA8F" bold="true" box="[1365,1437,1162,1188]" pageId="9" pageNumber="284">mac1,</emphasis>
<figureCitation id="1FDDA0F7253DFFA7FF69FB16FF4DBAEC" box="[151,244,1197,1223]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Figs. 21</figureCitation>
,
<figureCitation id="1FDDA0F7253DFFA7FEFEFB16FEA4BAE3" box="[256,285,1197,1224]" captionStart="FIGURES 2631" captionStartId="12.[151,264,1861,1886]" captionTargetBox="[197,1397,185,1830]" captionTargetId="figure@12.[190,1397,181,1836]" captionTargetPageId="12" captionText="FIGURES 2631. Trisaria rex and Trisaria washingtonensis. Figs. 2628. Trisaria rex. Fig 26. Anterior gonopods, anterior view; labels as in Figs. 21 and 22. Fig. 27. Gonopods, posterior view; labels as in Figs. 21 and 22. Fig. 28. Left leg 9 of male, anterior view; ap, apodeme; other labels as in Fig. 8. Figs. 2931. Trisaria washingtonensis. Fig. 29. Left leg 9 of male, anterior view; sp, spiracle; other labels as in Fig. 8. Fig. 30. Gonopods, anterior view; gst, gonopod sternum; other labels as in Figs. 21 &amp; 22. Fig. 31. Gonopods, posterior view; labels as in Fig. 27." figureDoi="http://doi.org/10.5281/zenodo.3734468" httpUri="https://zenodo.org/record/3734468/files/figure.png" pageId="9" pageNumber="284">26</figureCitation>
) short, blunt; the posterior most (
<emphasis id="B5926060253DFFA7FD66FB16FD61BAEC" bold="true" box="[664,728,1197,1223]" pageId="9" pageNumber="284">mac2</emphasis>
,
<figureCitation id="1FDDA0F7253DFFA7FD18FB16FCFDBAEC" box="[742,836,1197,1223]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Figs. 22</figureCitation>
,
<figureCitation id="1FDDA0F7253DFFA7FCB1FB16FC2EBAEC" box="[847,919,1197,1223]" captionStart="FIGURES 2631" captionStartId="12.[151,264,1861,1886]" captionTargetBox="[197,1397,185,1830]" captionTargetId="figure@12.[190,1397,181,1836]" captionTargetPageId="12" captionText="FIGURES 2631. Trisaria rex and Trisaria washingtonensis. Figs. 2628. Trisaria rex. Fig 26. Anterior gonopods, anterior view; labels as in Figs. 21 and 22. Fig. 27. Gonopods, posterior view; labels as in Figs. 21 and 22. Fig. 28. Left leg 9 of male, anterior view; ap, apodeme; other labels as in Fig. 8. Figs. 2931. Trisaria washingtonensis. Fig. 29. Left leg 9 of male, anterior view; sp, spiracle; other labels as in Fig. 8. Fig. 30. Gonopods, anterior view; gst, gonopod sternum; other labels as in Figs. 21 &amp; 22. Fig. 31. Gonopods, posterior view; labels as in Fig. 27." figureDoi="http://doi.org/10.5281/zenodo.3734468" httpUri="https://zenodo.org/record/3734468/files/figure.png" pageId="9" pageNumber="284">26, 27</figureCitation>
) long, bent posteriorly at nearly a right angle, then dorsally at a slightly more obtuse angle. Lateral angiocoxite (
<emphasis id="B5926060253DFFA7FC70FB6BFC09BAC1" bold="true" box="[910,944,1232,1258]" pageId="9" pageNumber="284">lac</emphasis>
,
<figureCitation id="1FDDA0F7253DFFA7FC3EFB74FBABBAC1" box="[960,1042,1231,1258]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Fig. 20</figureCitation>
) with three branches: the anteriormost (
<emphasis id="B5926060253DFFA7FF21FB49FEA9BB27" bold="true" box="[223,272,1266,1292]" pageId="9" pageNumber="284">lac1</emphasis>
,
<figureCitation id="1FDDA0F7253DFFA7FEE0FB49FEC5BB27" box="[286,380,1266,1292]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Figs. 21</figureCitation>
,
<figureCitation id="1FDDA0F7253DFFA7FE77FB49FE68BB27" box="[393,465,1266,1292]" captionStart="FIGURES 2631" captionStartId="12.[151,264,1861,1886]" captionTargetBox="[197,1397,185,1830]" captionTargetId="figure@12.[190,1397,181,1836]" captionTargetPageId="12" captionText="FIGURES 2631. Trisaria rex and Trisaria washingtonensis. Figs. 2628. Trisaria rex. Fig 26. Anterior gonopods, anterior view; labels as in Figs. 21 and 22. Fig. 27. Gonopods, posterior view; labels as in Figs. 21 and 22. Fig. 28. Left leg 9 of male, anterior view; ap, apodeme; other labels as in Fig. 8. Figs. 2931. Trisaria washingtonensis. Fig. 29. Left leg 9 of male, anterior view; sp, spiracle; other labels as in Fig. 8. Fig. 30. Gonopods, anterior view; gst, gonopod sternum; other labels as in Figs. 21 &amp; 22. Fig. 31. Gonopods, posterior view; labels as in Fig. 27." figureDoi="http://doi.org/10.5281/zenodo.3734468" httpUri="https://zenodo.org/record/3734468/files/figure.png" pageId="9" pageNumber="284">26, 27</figureCitation>
) long, slightly curved, divided at tip into two processes, each of these further divided into three or four smaller, acute terminations, one set pointing anterior, the other posterior. Median branch (
<emphasis id="B5926060253DFFA7FA9BFAAEFA2CBB04" bold="true" box="[1381,1429,1301,1327]" pageId="9" pageNumber="284">lac2</emphasis>
,
<figureCitation id="1FDDA0F7253DFFA7FF69FA8CFEA6BB79" box="[151,287,1335,1362]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Figs. 21, 22</figureCitation>
,
<figureCitation id="1FDDA0F7253DFFA7FED5FA8CFECBBB79" box="[299,370,1335,1362]" captionStart="FIGURES 2631" captionStartId="12.[151,264,1861,1886]" captionTargetBox="[197,1397,185,1830]" captionTargetId="figure@12.[190,1397,181,1836]" captionTargetPageId="12" captionText="FIGURES 2631. Trisaria rex and Trisaria washingtonensis. Figs. 2628. Trisaria rex. Fig 26. Anterior gonopods, anterior view; labels as in Figs. 21 and 22. Fig. 27. Gonopods, posterior view; labels as in Figs. 21 and 22. Fig. 28. Left leg 9 of male, anterior view; ap, apodeme; other labels as in Fig. 8. Figs. 2931. Trisaria washingtonensis. Fig. 29. Left leg 9 of male, anterior view; sp, spiracle; other labels as in Fig. 8. Fig. 30. Gonopods, anterior view; gst, gonopod sternum; other labels as in Figs. 21 &amp; 22. Fig. 31. Gonopods, posterior view; labels as in Fig. 27." figureDoi="http://doi.org/10.5281/zenodo.3734468" httpUri="https://zenodo.org/record/3734468/files/figure.png" pageId="9" pageNumber="284">26, 27</figureCitation>
) set with dense array of fine spines, giving comb-like appearance. Posteriormost branch (
<emphasis id="B5926060253DFFA7FA9BFA8CFA2CBB7A" bold="true" box="[1381,1429,1335,1361]" pageId="9" pageNumber="284">lac3</emphasis>
,
<figureCitation id="1FDDA0F7253DFFA7FF69FAE1FF4CBB5F" box="[151,245,1370,1396]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Figs. 22</figureCitation>
,
<figureCitation id="1FDDA0F7253DFFA7FEFEFAE1FEA4BB5F" box="[256,285,1370,1396]" captionStart="FIGURES 2631" captionStartId="12.[151,264,1861,1886]" captionTargetBox="[197,1397,185,1830]" captionTargetId="figure@12.[190,1397,181,1836]" captionTargetPageId="12" captionText="FIGURES 2631. Trisaria rex and Trisaria washingtonensis. Figs. 2628. Trisaria rex. Fig 26. Anterior gonopods, anterior view; labels as in Figs. 21 and 22. Fig. 27. Gonopods, posterior view; labels as in Figs. 21 and 22. Fig. 28. Left leg 9 of male, anterior view; ap, apodeme; other labels as in Fig. 8. Figs. 2931. Trisaria washingtonensis. Fig. 29. Left leg 9 of male, anterior view; sp, spiracle; other labels as in Fig. 8. Fig. 30. Gonopods, anterior view; gst, gonopod sternum; other labels as in Figs. 21 &amp; 22. Fig. 31. Gonopods, posterior view; labels as in Fig. 27." figureDoi="http://doi.org/10.5281/zenodo.3734468" httpUri="https://zenodo.org/record/3734468/files/figure.png" pageId="9" pageNumber="284">27</figureCitation>
) short, simple, bluntly acuminate. Colpocoxites (
<emphasis id="B5926060253DFFA7FCB6FAE1FCD8BB5F" bold="true" box="[840,865,1370,1396]" pageId="9" pageNumber="284">cc</emphasis>
,
<figureCitation id="1FDDA0F7253DFFA7FC91FAE1FC75BB5F" box="[879,972,1370,1396]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Figs. 22</figureCitation>
,
<figureCitation id="1FDDA0F7253DFFA7FC29FAE1FBA6BB5F" box="[983,1055,1370,1396]" captionStart="FIGURES 2631" captionStartId="12.[151,264,1861,1886]" captionTargetBox="[197,1397,185,1830]" captionTargetId="figure@12.[190,1397,181,1836]" captionTargetPageId="12" captionText="FIGURES 2631. Trisaria rex and Trisaria washingtonensis. Figs. 2628. Trisaria rex. Fig 26. Anterior gonopods, anterior view; labels as in Figs. 21 and 22. Fig. 27. Gonopods, posterior view; labels as in Figs. 21 and 22. Fig. 28. Left leg 9 of male, anterior view; ap, apodeme; other labels as in Fig. 8. Figs. 2931. Trisaria washingtonensis. Fig. 29. Left leg 9 of male, anterior view; sp, spiracle; other labels as in Fig. 8. Fig. 30. Gonopods, anterior view; gst, gonopod sternum; other labels as in Figs. 21 &amp; 22. Fig. 31. Gonopods, posterior view; labels as in Fig. 27." figureDoi="http://doi.org/10.5281/zenodo.3734468" httpUri="https://zenodo.org/record/3734468/files/figure.png" pageId="9" pageNumber="284">26, 27</figureCitation>
) entirely membranous, with scaly cuticle. Ninth legs (
<figureCitation id="1FDDA0F7253DFFA7FE8DFAC6FE69BBBC" box="[371,464,1405,1431]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Figs. 23</figureCitation>
) much reduced, sternum (
<emphasis id="B5926060253DFFA7FD08FAC6FCAEBBBC" bold="true" box="[758,791,1405,1431]" pageId="9" pageNumber="284">st9</emphasis>
,
<figureCitation id="1FDDA0F7253DFFA7FCD8FAC6FC3ABBBC" box="[806,899,1405,1431]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Figs. 23</figureCitation>
), coxae (
<emphasis id="B5926060253DFFA7FC17FAC6FBBDBBBC" bold="true" box="[1001,1028,1405,1431]" pageId="9" pageNumber="284">c9</emphasis>
), telopodites (
<emphasis id="B5926060253DFFA7FB58FAC6FB07BBBC" bold="true" box="[1190,1214,1405,1431]" pageId="9" pageNumber="284">t9</emphasis>
) almost completely fused, telopodites much flattened, distally setose; coxae with minute anterior pore (
<emphasis id="B5926060253DFFA7FBCAFA1BFBE9BB91" bold="true" box="[1076,1104,1440,1466]" pageId="9" pageNumber="284">cp</emphasis>
,
<figureCitation id="1FDDA0F7253DFFA7FBA3FA24FB09BB91" box="[1117,1200,1439,1466]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Fig. 23</figureCitation>
). Tenth legs (
<figureCitation id="1FDDA0F7253DFFA7FABDFA24FA2DBB91" box="[1347,1428,1439,1466]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Fig. 24</figureCitation>
) with enlarged coxae (
<emphasis id="B5926060253DFFA7FE76FA79FE08BBF7" bold="true" box="[392,433,1474,1500]" pageId="9" pageNumber="284">c10</emphasis>
,
<figureCitation id="1FDDA0F7253DFFA7FE41FA79FDB6BBF7" box="[447,527,1474,1500]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Fig. 24</figureCitation>
) bearing glands (
<emphasis id="B5926060253DFFA7FD2DFA79FD48BBF7" bold="true" box="[723,753,1474,1500]" pageId="9" pageNumber="284">gp</emphasis>
,
<figureCitation id="1FDDA0F7253DFFA7FD01FA79FCEBBBF7" box="[767,850,1474,1500]" captionStart="FIGURES 2025" captionStartId="11.[151,264,1815,1840]" captionTargetBox="[225,1362,184,1787]" captionTargetId="figure@11.[225,1362,181,1789]" captionTargetPageId="11" captionText="FIGURES 2025. Trisaria rex. Fig. 20. Gonopod complex, ventral view; art7, articulating surface of paratergite 7, cx 10, cx11, coxae of legs 10 &amp; 11, gst, gonopod sternum, mac, median colpocoxite, lac, lateral colpocoxite, t9, telopodite of 9th leg. Fig. 21. Gonopods, anterior view; gst, gonopod sternum, lac1, lac2, first and second branches of lateral colpocoxite, mac1, mac 2, first and second branches of median colpocoxite. Fig. 22. Gonopods, posterior view; cc, colpocoxite, lac 3, third branch of lateral colpocoxite; other labels as in Fig. 21. Fig. 23. Legpair 9 of male; labels as in Fig. 13. Fig. 24. Legpair 10 of male; cx10, coxa, gp, coxal pore. Fig. 25. Part of prefemur of leg 10, showing modified setae." figureDoi="http://doi.org/10.5281/zenodo.3734466" httpUri="https://zenodo.org/record/3734466/files/figure.png" pageId="9" pageNumber="284">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="8759BC72253DFFA7FF39FA5EFF6DB80A" blockId="9.[151,1437,815,1750]" pageId="9" pageNumber="284">
<emphasis id="B5926060253DFFA7FF39FA5EFD50BBD4" bold="true" box="[199,745,1508,1535]" pageId="9" pageNumber="284">Female from Twin Falls/Iron Horse trailhead.</emphasis>
Length,
<quantity id="401E1197253DFFA7FCB1FA5EFC11BBD4" box="[847,936,1509,1535]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="9" pageNumber="284" unit="mm" value="9.2">9.2 mm</quantity>
, width
<quantity id="401E1197253DFFA7FC03FA5EFBDDBBD4" box="[1021,1124,1509,1535]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="284" unit="mm" value="0.9">0.90 mm</quantity>
. Nonsexual characters as in male.
</paragraph>
</subSubSection>
<subSubSection id="CFFCEFF9253DFFA7FF39F990FB46B8FE" pageId="9" pageNumber="284" type="distribution">
<paragraph id="8759BC72253DFFA7FF39F990FB4FB899" blockId="9.[151,1437,815,1750]" pageId="9" pageNumber="284">
<emphasis id="B5926060253DFFA7FF39F990FEDAB86D" bold="true" box="[199,355,1579,1606]" pageId="9" pageNumber="284">Distribution:</emphasis>
<collectingRegion id="45227290253DFFA7FE94F997FD90B86D" box="[362,553,1580,1606]" country="United States of America" name="Washington" pageId="9" pageNumber="284">WASHINGTON</collectingRegion>
:
<emphasis id="B5926060253DFFA7FDCAF990FD26B86D" bold="true" box="[564,671,1579,1606]" pageId="9" pageNumber="284">King Co.</emphasis>
: North of I90, Pratt Lake Trailhead,
<geoCoordinate id="E2D2DAB5253DFFA7FBC1F997FB73B86D" box="[1087,1226,1580,1606]" degrees="47" direction="north" minutes="23.829" orientation="latitude" pageId="9" pageNumber="284" precision="1" value="47.39715">N47°23.829</geoCoordinate>
,
<geoCoordinate id="E2D2DAB5253DFFA7FB2BF997FACDB86D" box="[1237,1396,1580,1606]" degrees="121" direction="west" minutes="29.109" orientation="longitude" pageId="9" pageNumber="284" precision="1" value="-121.48515">W121°29.109</geoCoordinate>
,
<date id="F3589AB2253DFFA7FA7EF990FE8DB841" pageId="9" pageNumber="284" value="2003-10-25">25 October 2003</date>
, W. Leonard, m, f. Same, but
<geoCoordinate id="E2D2DAB5253DFFA7FD7AF9EBFCA1B841" box="[644,792,1616,1642]" degrees="47" direction="north" minutes="23.821" orientation="latitude" pageId="9" pageNumber="284" precision="1" value="47.39702">N47°23.821</geoCoordinate>
,
<geoCoordinate id="E2D2DAB5253DFFA7FCDAF9EBFC75B841" box="[804,972,1616,1642]" degrees="121" direction="west" minutes="29.108" orientation="longitude" pageId="9" pageNumber="284" precision="1" value="-121.48513">W121°29.108</geoCoordinate>
, 1700 asl,
<date id="F3589AB2253DFFA7FBB3F9F4FAB4B841" box="[1101,1293,1615,1642]" pageId="9" pageNumber="284" value="2009-10-15">15 October 2009</date>
, mm. I-90 at Snoqualmie River, Twin Falls/Iron Horse Trailhead,
<geoCoordinate id="E2D2DAB5253DFFA7FD19F9C8FCC2B8A5" box="[743,891,1651,1678]" degrees="47" direction="north" minutes="26.651" orientation="latitude" pageId="9" pageNumber="284" precision="1" value="47.444183">N47°26.651</geoCoordinate>
,
<geoCoordinate id="E2D2DAB5253DFFA7FC79F9CFFB96B8A5" box="[903,1071,1652,1678]" degrees="121" direction="west" minutes="40.081" orientation="longitude" pageId="9" pageNumber="284" precision="1" value="-121.668015">W121°40.081</geoCoordinate>
,
<date id="F3589AB2253DFFA7FBC5F9C8FABEB8A5" box="[1083,1287,1651,1678]" pageId="9" pageNumber="284" value="2004-02-25">25 February 2004</date>
, W. Leonard, C. Richart, m, f; Tokul Creek, 520 asl, 47.556°N, 121.817°W,
<date id="F3589AB2253DFFA7FCA6F923FB8AB899" box="[856,1075,1687,1714]" pageId="9" pageNumber="284" value="1979-11-21">21 November 1979</date>
, R. Crawford, m.
</paragraph>
<paragraph id="8759BC72253DFFA7FF39F907FB46B8FE" blockId="9.[151,1437,815,1750]" box="[199,1279,1723,1750]" pageId="9" pageNumber="284">
<emphasis id="B5926060253DFFA7FF39F907FEAAB8FD" bold="true" box="[199,275,1724,1750]" pageId="9" pageNumber="284">Notes:</emphasis>
Presently this species is known only from central and western King County,
<collectingRegion id="45227290253DFFA7FB8CF900FB42B8FE" box="[1138,1275,1723,1749]" country="United States of America" name="Washington" pageId="9" pageNumber="284">Washington</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>