212 lines
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212 lines
30 KiB
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<document id="864324CB21B9951D8E3D7D723DCBB3D6" ID-DOI="10.11646/zootaxa.4758.2.4" ID-GBIF-Dataset="a4c437e0-7983-41af-8f98-a3bd88cc2e69" ID-ISSN="1175-5326" ID-Zenodo-Dep="3734456" ID-ZooBank="2539ABCC-161E-44B2-BB2C-C949B1A7C94D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585658008729" checkinUser="plazi" docAuthor="Shear, William A." docDate="2020" docId="0F4F0D642533FFA9FF69FF64FB80BB06" docLanguage="en" docName="zootaxa.4758.2.4.pdf" docOrigin="Zootaxa 4758 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vaferaria Causey 1958" docType="treatment" docVersion="5" lastPageNumber="282" masterDocId="F376751C2534FFAEFFFEFFBBFFB9BE2B" masterDocTitle="The millipede family Striariidae Bollman, 1893: I. Introduction to the family, synonymy of Vaferaria Causey with Amplaria Chamberlin, the new subfamily Trisariinae, the new genus Trisaria, and three new species (Diplopoda, Chordeumatida, Striarioidea)" masterLastPageNumber="295" masterPageNumber="275" pageNumber="282" updateTime="1698733589358" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7734B4E1863DA4D9BEDA7E4960ABC50E">The millipede family Striariidae Bollman, 1893: I. Introduction to the family, synonymy of Vaferaria Causey with Amplaria Chamberlin, the new subfamily Trisariinae, the new genus Trisaria, and three new species (Diplopoda, Chordeumatida, Striarioidea)</mods:title>
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<mods:namePart id="9E311BB3A1EE21F2434F50580E858E19">Shear, William A.</mods:namePart>
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<treatment id="0F4F0D642533FFA9FF69FF64FB80BB06" ID-DOI="http://doi.org/10.5281/zenodo.3812335" ID-GBIF-Taxon="163417941" ID-Zenodo-Dep="3812335" LSID="urn:lsid:plazi:treatment:0F4F0D642533FFA9FF69FF64FB80BB06" httpUri="http://treatment.plazi.org/id/0F4F0D642533FFA9FF69FF64FB80BB06" lastPageNumber="282" pageId="7" pageNumber="282">
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<subSubSection id="CFFCEFF92533FFA9FF69FF64FDE6BED1" box="[151,607,223,250]" pageId="7" pageNumber="282" type="nomenclature">
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<paragraph id="8759BC722533FFA9FF69FF64FDE6BED1" blockId="7.[151,607,223,250]" box="[151,607,223,250]" pageId="7" pageNumber="282">
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<heading id="DC110B1E2533FFA9FF69FF64FDE6BED1" bold="true" box="[151,607,223,250]" fontSize="11" level="1" pageId="7" pageNumber="282" reason="1">
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<emphasis id="B59260602533FFA9FF69FF64FDE6BED1" bold="true" box="[151,607,223,250]" pageId="7" pageNumber="282">
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Synonymy of
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<taxonomicName id="40E6C7F12533FFA9FEBCFF64FDE6BED1" ID-CoL="d1f8bbb6-e81a-461a-96bb-903f4f1194ce" authority="Causey, 1958" authorityName="Causey" authorityYear="1958" box="[322,607,223,250]" class="Diplopoda" family="Striariidae" genus="Vaferaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">
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<emphasis id="B59260602533FFA9FEBCFF64FE0DBED2" bold="true" box="[322,436,223,249]" italics="true" pageId="7" pageNumber="282">Vaferaria</emphasis>
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<bibRefCitation id="E377C1832533FFA9FE42FF64FDE6BED1" author="Causey, N. B." box="[444,607,223,250]" pageId="7" pageNumber="282" pagination="179 - 184" refId="ref10808" refString="Causey, N. B. (1958) New records and descriptions of a new genus and a new species of millipeds of the family Striariidae (Chordeumida). Proceedings of the Biological Society of Washington, 71, 179 - 184." type="journal article" year="1958">Causey, 1958</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CFFCEFF92533FFA9FF69FE9EFB80BB06" pageId="7" pageNumber="282" type="description">
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<paragraph id="8759BC722533FFA9FF69FE9EFD63BCB2" blockId="7.[151,1437,293,1325]" pageId="7" pageNumber="282">
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<bibRefCitation id="E377C1832533FFA9FF69FE9EFEF8BF6B" author="Loomis, H. F." box="[151,321,293,320]" pageId="7" pageNumber="282" pagination="404 - 409" refId="ref11257" refString="Loomis, H. F. (1936) New millipeds of the American family Striariidae. Journal of the Washington Academy of Science, 26, 404 - 409." type="journal article" year="1936">Loomis (1936)</bibRefCitation>
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surveyed the known striariids, and added two new species,
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<taxonomicName id="40E6C7F12533FFA9FC03FE9EFB2FBF14" authorityName="Loomis" authorityYear="1936" box="[1021,1174,293,319]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="species" species="nana">
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<emphasis id="B59260602533FFA9FC03FE9EFB2FBF14" box="[1021,1174,293,319]" italics="true" pageId="7" pageNumber="282">Striaria nana</emphasis>
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</taxonomicName>
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and
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<emphasis id="B59260602533FFA9FB2CFE9EFA24BF14" box="[1234,1437,293,319]" italics="true" pageId="7" pageNumber="282">
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<taxonomicName id="40E6C7F12533FFA9FB2CFE9EFA20BF14" authorityName="Loomis" authorityYear="1936" box="[1234,1433,293,319]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="species" species="imberbis">Striaria imberbis</taxonomicName>
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.
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</emphasis>
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The latter species, according to Loomis, had characters that were not found in other
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<taxonomicName id="40E6C7F12533FFA9FBB5FEF3FB1BBF49" authorityName="Bollman" authorityYear="1888" box="[1099,1186,328,354]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">
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<emphasis id="B59260602533FFA9FBB5FEF3FB1BBF49" box="[1099,1186,328,354]" italics="true" pageId="7" pageNumber="282">Striaria</emphasis>
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</taxonomicName>
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species, including the dorsum of the rings, which was said to be “depressed throughout, but [on] the anterior ones more strikingly so.” Additionally, the single male
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<typeStatus id="585D02D02533FFA9FE42FE36FDA7BF8C" box="[444,542,397,423]" pageId="7" pageNumber="282" type="holotype">holotype</typeStatus>
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appeared to lack the sinuate spines on the angles of the labrum, had prominently expanded lateralmost crests (the sixth) on the second ring, and prominent lobes on the trochanters of the second legs. Loomis only provided drawings of a second leg. Much later,
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<bibRefCitation id="E377C1832533FFA9FC6BFE69FB82BFC7" author="Causey, N. B." box="[917,1083,465,492]" pageId="7" pageNumber="282" pagination="179 - 184" refId="ref10808" refString="Causey, N. B. (1958) New records and descriptions of a new genus and a new species of millipeds of the family Striariidae (Chordeumida). Proceedings of the Biological Society of Washington, 71, 179 - 184." type="journal article" year="1958">Causey (1958)</bibRefCitation>
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cited some of these characters (still unillustrated) and named a new genus,
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<emphasis id="B59260602533FFA9FD6EFE4EFCB8BC24" box="[656,769,501,527]" italics="true" pageId="7" pageNumber="282">
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<taxonomicName id="40E6C7F12533FFA9FD6EFE4EFD44BC24" authorityName="Causey" authorityYear="1958" box="[656,765,501,527]" class="Diplopoda" family="Striariidae" genus="Vaferaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">Vaferaria</taxonomicName>
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,
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</emphasis>
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for
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<emphasis id="B59260602533FFA9FCCCFE4EFC01BC24" box="[818,952,501,527]" italics="true" pageId="7" pageNumber="282">
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<taxonomicName id="40E6C7F12533FFA9FCCCFE4EFC0DBC24" authorityName="Loomis" authorityYear="1936" box="[818,948,501,527]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="species" species="imberbis">S. imberbis</taxonomicName>
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,
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</emphasis>
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emphasizing especially the “flattening” of the anterior rings. It is not clear from her description that she actually studied the
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<typeStatus id="585D02D02533FFA9FBCDFDACFB2CBC1A" box="[1075,1173,535,561]" pageId="7" pageNumber="282" type="holotype">holotype</typeStatus>
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of
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<emphasis id="B59260602533FFA9FB42FDA3FAFBBC19" box="[1212,1346,536,562]" italics="true" pageId="7" pageNumber="282">
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<taxonomicName id="40E6C7F12533FFA9FB42FDA3FA87BC19" authorityName="Loomis" authorityYear="1936" box="[1212,1342,536,562]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="species" species="imberbis">S. imberbis</taxonomicName>
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.
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</emphasis>
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Indeed, it appears that no mature male specimens of any of the species discussed (except possibly for
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<emphasis id="B59260602533FFA9FB44FD80FAF6BC7F" box="[1210,1359,570,597]" italics="true" pageId="7" pageNumber="282">
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<taxonomicName id="40E6C7F12533FFA9FB44FD80FAF3BC7F" authorityName="Bollman" authorityYear="1888" box="[1210,1354,570,597]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="species" species="granulosa">S. granulosa</taxonomicName>
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,
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</emphasis>
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and in that case not the types) were examined.
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<bibRefCitation id="E377C1832533FFA9FDADFDE6FCBBBC5C" author="Hoffman, R. L." box="[595,770,605,631]" pageId="7" pageNumber="282" refId="ref11201" refString="Hoffman, R. L. (1979) Classification of the Diplopoda. Museum d'Histoire Naturelle, Geneve, 237 pp." type="book" year="1979">Hoffman (1979</bibRefCitation>
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,
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<bibRefCitation id="E377C1832533FFA9FCF3FDE6FCFFBC5C" author="Hoffman, R. L." box="[781,838,605,631]" pageId="7" pageNumber="282" pagination="1 - 584" refId="ref11224" refString="Hoffman, R. L. (1999) Checklist of the millipeds of North and Middle America. Virginia Museum of Natural History Special Publications, 8, 1 - 584." type="journal article" year="1999">1999</bibRefCitation>
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) persistently misspelled the genus name as “
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<taxonomicName id="40E6C7F12533FFA9FAC6FDE6FA30BC5C" authorityName="Causey" authorityYear="1958" box="[1336,1417,605,631]" class="Diplopoda" family="Striariidae" genus="Vaferia" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">
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<emphasis id="B59260602533FFA9FAC6FDE6FA30BC5C" box="[1336,1417,605,631]" italics="true" pageId="7" pageNumber="282">Vaferia</emphasis>
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</taxonomicName>
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,” and
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<bibRefCitation id="E377C1832533FFA9FF39FDC4FE7EBCB1" author="Shear, W. A. & Krejca, J. K." box="[199,455,639,666]" pageId="7" pageNumber="282" pagination="23 - 39" refId="ref11524" refString="Shear, W. A. & Krejca, J. K. (2007) Revalidation of the millipede genus Amplaria Chamberlin 1941 (Diplopoda, Chordeumatida, Striariidae) and description of two new species from caves in Sequoia and Kings Canyon National Parks, California. Zootaxa, 1532 (1), 23 - 39. https: // doi. org / 10.11646 / zootaxa. 1532.1.2" type="journal article" year="2007">Shear & Krejca (2007)</bibRefCitation>
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made the same mistake.
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</paragraph>
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<paragraph id="8759BC722533FFA9FF39FD19FB74BDF9" blockId="7.[151,1437,293,1325]" pageId="7" pageNumber="282">
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As a part of a survey of the described species of
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<emphasis id="B59260602533FFA9FD1EFD18FC87BC96" box="[736,830,675,701]" italics="true" pageId="7" pageNumber="282">
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<taxonomicName id="40E6C7F12533FFA9FD1EFD18FC83BC96" authorityName="Bollman" authorityYear="1888" box="[736,826,675,701]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">Striaria</taxonomicName>
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,
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</emphasis>
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I studied the
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<typeStatus id="585D02D02533FFA9FC2AFD19FB8FBC97" box="[980,1078,674,700]" pageId="7" pageNumber="282" type="holotype">holotype</typeStatus>
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and only known male specimen of
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<taxonomicName id="40E6C7F12533FFA9FF4BFD7EFE8ABCF4" authorityName="Loomis" authorityYear="1936" box="[181,307,709,735]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="species" species="imberbis">
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<emphasis id="B59260602533FFA9FF4BFD7EFE8ABCF4" box="[181,307,709,735]" italics="true" pageId="7" pageNumber="282">S. imberbis</emphasis>
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</taxonomicName>
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(Smithsonian National Museum of Natural History); collected south of Atascadero, San Luis Obispo Co.,
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<collectingRegion id="452272902533FFA9FF31FD5CFEFABD2A" box="[207,323,743,769]" country="United States of America" name="California" pageId="7" pageNumber="282">California</collectingRegion>
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, by O. F. Cook,
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<date id="F3589AB22533FFA9FDFBFD53FD0EBD29" box="[517,695,744,770]" pageId="7" pageNumber="282" value="1928-01-01">1 January 1928</date>
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, along with a female and “a number of young”, unfortunately in rather poor condition. The gonopods had not been dissected. While the shelf-like sixth crest of the second segment, the lack of a labral spine, and the presence of large lobes on the second leg trochanters all proved to be repeatable observations, the “flattening” of the body, even anteriorly, was not. The structure of the rings was typical of other striariids, and it remains unclear what exactly Loomis meant in his description by “depressed,” though he may have been referring to the median sulcus, deeply infolded in species of
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<taxonomicName id="40E6C7F12533FFA9FC75FC2EFC5BBD84" authorityName="Bollman" authorityYear="1888" box="[907,994,917,943]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">
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<emphasis id="B59260602533FFA9FC75FC2EFC5BBD84" box="[907,994,917,943]" italics="true" pageId="7" pageNumber="282">Striaria</emphasis>
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</taxonomicName>
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. I dissected and studied the gonopods, ninth legs and second legs at high magnification under the compound microscope (
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<figureCitation id="1FDDA0F72533FFA9FBC6FC0CFB79BDF9" box="[1080,1216,951,978]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1872,1897]" captionTargetBox="[154,1323,153,1811]" captionTargetId="figure@6.[151,1373,151,1855]" captionTargetPageId="6" captionText="FIGURES 9–14. Striaria columbiana and Amplaria imberbis. Figs. 9, 10. Striaria columbiana. Fig. 9. Left gonopod, mesal view; pcc, posterior lobe of colpocoxite; other labels as in Fig. 6. Fig. 10. Right gonopod, lateral view; labels as in Fig. 9. Figs. 11–14. Amplaria imberbis male holotype. Fig. 11. Right gonopod, lateral view; labels as in Fig. 6. Fig. 12. Angiocoxite tips of right gonopod, mesal view; labels as in Fig. 6. Fig. 13. Legpair 9 of male, anterior view; cp, coxal pore; other labels as in Fig. 8." figureDoi="http://doi.org/10.5281/zenodo.3734462" httpUri="https://zenodo.org/record/3734462/files/figure.png" pageId="7" pageNumber="282">Figs. 11–14</figureCitation>
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).
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</paragraph>
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<paragraph id="8759BC722533FFA9FF39FC60FB80BB06" blockId="7.[151,1437,293,1325]" pageId="7" pageNumber="282">
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I found that the gonopods (
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<figureCitation id="1FDDA0F72533FFA9FE05FC61FD3BBDDF" box="[507,642,986,1012]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1872,1897]" captionTargetBox="[154,1323,153,1811]" captionTargetId="figure@6.[151,1373,151,1855]" captionTargetPageId="6" captionText="FIGURES 9–14. Striaria columbiana and Amplaria imberbis. Figs. 9, 10. Striaria columbiana. Fig. 9. Left gonopod, mesal view; pcc, posterior lobe of colpocoxite; other labels as in Fig. 6. Fig. 10. Right gonopod, lateral view; labels as in Fig. 9. Figs. 11–14. Amplaria imberbis male holotype. Fig. 11. Right gonopod, lateral view; labels as in Fig. 6. Fig. 12. Angiocoxite tips of right gonopod, mesal view; labels as in Fig. 6. Fig. 13. Legpair 9 of male, anterior view; cp, coxal pore; other labels as in Fig. 8." figureDoi="http://doi.org/10.5281/zenodo.3734462" httpUri="https://zenodo.org/record/3734462/files/figure.png" pageId="7" pageNumber="282">Figs. 11, 12</figureCitation>
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) and ninth legs (
|
||
<figureCitation id="1FDDA0F72533FFA9FCBCFC61FC2CBDDF" box="[834,917,986,1012]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1872,1897]" captionTargetBox="[154,1323,153,1811]" captionTargetId="figure@6.[151,1373,151,1855]" captionTargetPageId="6" captionText="FIGURES 9–14. Striaria columbiana and Amplaria imberbis. Figs. 9, 10. Striaria columbiana. Fig. 9. Left gonopod, mesal view; pcc, posterior lobe of colpocoxite; other labels as in Fig. 6. Fig. 10. Right gonopod, lateral view; labels as in Fig. 9. Figs. 11–14. Amplaria imberbis male holotype. Fig. 11. Right gonopod, lateral view; labels as in Fig. 6. Fig. 12. Angiocoxite tips of right gonopod, mesal view; labels as in Fig. 6. Fig. 13. Legpair 9 of male, anterior view; cp, coxal pore; other labels as in Fig. 8." figureDoi="http://doi.org/10.5281/zenodo.3734462" httpUri="https://zenodo.org/record/3734462/files/figure.png" pageId="7" pageNumber="282">Fig. 13</figureCitation>
|
||
), while in some ways simple, were consistent with the striariid species now assigned to a revalidated
|
||
<taxonomicName id="40E6C7F12533FFA9FD05FC46FB8DBA3C" authority="Chamberlin, 1941" authorityName="Chamberlin" authorityYear="1941" box="[763,1076,1021,1047]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B59260602533FFA9FD05FC46FCDCBA3C" box="[763,869,1021,1047]" italics="true" pageId="7" pageNumber="282">Amplaria</emphasis>
|
||
<bibRefCitation id="E377C1832533FFA9FC95FC46FB8DBA3C" author="Chamberlin, R. V." box="[875,1076,1021,1047]" pageId="7" pageNumber="282" pagination="3 - 23" refId="ref10976" refString="Chamberlin, R. V. (1941) New western millipeds. Bulletin of the University of Utah, Biological Series, 31 (2), 6, 3 - 23." type="journal article" year="1941">Chamberlin, 1941</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="E377C1832533FFA9FBBCFC46FA8BBA3C" author="Shear, W. A. & Krejca, J. K." box="[1090,1330,1021,1047]" pageId="7" pageNumber="282" pagination="23 - 39" refId="ref11524" refString="Shear, W. A. & Krejca, J. K. (2007) Revalidation of the millipede genus Amplaria Chamberlin 1941 (Diplopoda, Chordeumatida, Striariidae) and description of two new species from caves in Sequoia and Kings Canyon National Parks, California. Zootaxa, 1532 (1), 23 - 39. https: // doi. org / 10.11646 / zootaxa. 1532.1.2" type="journal article" year="2007">Shear & Krejca 2007</bibRefCitation>
|
||
, Shear,
|
||
<emphasis id="B59260602533FFA9FA76FC45FF0DBA11" italics="true" pageId="7" pageNumber="282">et al.</emphasis>
|
||
2017). The second legpair (
|
||
<figureCitation id="1FDDA0F72533FFA9FE0CFBA4FDFDBA11" box="[498,580,1055,1082]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1872,1897]" captionTargetBox="[154,1323,153,1811]" captionTargetId="figure@6.[151,1373,151,1855]" captionTargetPageId="6" captionText="FIGURES 9–14. Striaria columbiana and Amplaria imberbis. Figs. 9, 10. Striaria columbiana. Fig. 9. Left gonopod, mesal view; pcc, posterior lobe of colpocoxite; other labels as in Fig. 6. Fig. 10. Right gonopod, lateral view; labels as in Fig. 9. Figs. 11–14. Amplaria imberbis male holotype. Fig. 11. Right gonopod, lateral view; labels as in Fig. 6. Fig. 12. Angiocoxite tips of right gonopod, mesal view; labels as in Fig. 6. Fig. 13. Legpair 9 of male, anterior view; cp, coxal pore; other labels as in Fig. 8." figureDoi="http://doi.org/10.5281/zenodo.3734462" httpUri="https://zenodo.org/record/3734462/files/figure.png" pageId="7" pageNumber="282">Fig. 14</figureCitation>
|
||
) and other secondary sexual characters including those cited in the description of the genus also conformed to the other species of
|
||
<taxonomicName id="40E6C7F12533FFA9FCEAFBF9FCC7BA77" authorityName="Chamberlin" authorityYear="1941" box="[788,894,1090,1116]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B59260602533FFA9FCEAFBF9FCC7BA77" box="[788,894,1090,1116]" italics="true" pageId="7" pageNumber="282">Amplaria</emphasis>
|
||
</taxonomicName>
|
||
. In redescribing
|
||
<taxonomicName id="40E6C7F12533FFA9FBC3FBF8FA21BA77" authority="(Chamberlin, 1910)" baseAuthorityName="Chamberlin" baseAuthorityYear="1910" box="[1085,1432,1090,1117]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="species" species="nazinta">
|
||
<emphasis id="B59260602533FFA9FBC3FBF8FB16BA77" box="[1085,1199,1091,1117]" italics="true" pageId="7" pageNumber="282">A. nazinta</emphasis>
|
||
(
|
||
<bibRefCitation id="E377C1832533FFA9FB41FBF9FA29BA77" author="Chamberlin, R. V." box="[1215,1424,1090,1116]" pageId="7" pageNumber="282" pagination="233 - 262" refId="ref10890" refString="Chamberlin, R. V. (1910) Diplopoda from the western states. Annals of the Entomological Society of America, 3, 233 - 262, pls. XXX-XLIII. https: // doi. org / 10.1093 / aesa / 3.4.233" type="journal article" year="1910">Chamberlin, 1910</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<bibRefCitation id="E377C1832533FFA9FF69FBDEFED3BA54" author="Shear, W. A. & Nosler, P. & Marek, P. E." box="[151,362,1125,1151]" pageId="7" pageNumber="282" pagination="233 - 240" refId="ref11657" refString="Shear, W. A., Nosler, P. & Marek, P. E. (2017) The identity of Amplaria nazinta (Chamberlin, 1910): a century-old millipede mystery resolved (Diplopoda, Chordeumatida, Striariidae). Zootaxa, 4311 (2), 233 - 240. https: // doi. org / 10.11646 / zootaxa. 4311.2.4" type="journal article" year="2017">
|
||
Shear
|
||
<emphasis id="B59260602533FFA9FF21FBDDFEA1BA54" box="[223,280,1125,1151]" italics="true" pageId="7" pageNumber="282">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
noted that the labral spines of this species were strongly reduced. In all species of
|
||
<taxonomicName id="40E6C7F12533FFA9FAECFBDEFAC5BA54" authorityName="Chamberlin" authorityYear="1941" box="[1298,1404,1125,1151]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B59260602533FFA9FAECFBDEFAC5BA54" box="[1298,1404,1125,1151]" italics="true" pageId="7" pageNumber="282">Amplaria</emphasis>
|
||
</taxonomicName>
|
||
so far studied, including several awaiting description, the second leg’s (
|
||
<figureCitation id="1FDDA0F72533FFA9FC65FB3CFC54BA89" box="[923,1005,1159,1186]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1872,1897]" captionTargetBox="[154,1323,153,1811]" captionTargetId="figure@6.[151,1373,151,1855]" captionTargetPageId="6" captionText="FIGURES 9–14. Striaria columbiana and Amplaria imberbis. Figs. 9, 10. Striaria columbiana. Fig. 9. Left gonopod, mesal view; pcc, posterior lobe of colpocoxite; other labels as in Fig. 6. Fig. 10. Right gonopod, lateral view; labels as in Fig. 9. Figs. 11–14. Amplaria imberbis male holotype. Fig. 11. Right gonopod, lateral view; labels as in Fig. 6. Fig. 12. Angiocoxite tips of right gonopod, mesal view; labels as in Fig. 6. Fig. 13. Legpair 9 of male, anterior view; cp, coxal pore; other labels as in Fig. 8." figureDoi="http://doi.org/10.5281/zenodo.3734462" httpUri="https://zenodo.org/record/3734462/files/figure.png" pageId="7" pageNumber="282">Fig. 14</figureCitation>
|
||
) trochanters bear setae-crowned lobes of varying lengths, and some showed strong development of the sixth crest of the second ring. Since the gonopods (
|
||
<figureCitation id="1FDDA0F72533FFA9FF61FB76FE9DBACC" box="[159,292,1229,1255]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1872,1897]" captionTargetBox="[154,1323,153,1811]" captionTargetId="figure@6.[151,1373,151,1855]" captionTargetPageId="6" captionText="FIGURES 9–14. Striaria columbiana and Amplaria imberbis. Figs. 9, 10. Striaria columbiana. Fig. 9. Left gonopod, mesal view; pcc, posterior lobe of colpocoxite; other labels as in Fig. 6. Fig. 10. Right gonopod, lateral view; labels as in Fig. 9. Figs. 11–14. Amplaria imberbis male holotype. Fig. 11. Right gonopod, lateral view; labels as in Fig. 6. Fig. 12. Angiocoxite tips of right gonopod, mesal view; labels as in Fig. 6. Fig. 13. Legpair 9 of male, anterior view; cp, coxal pore; other labels as in Fig. 8." figureDoi="http://doi.org/10.5281/zenodo.3734462" httpUri="https://zenodo.org/record/3734462/files/figure.png" pageId="7" pageNumber="282">Figs. 11, 12</figureCitation>
|
||
) follow the
|
||
<taxonomicName id="40E6C7F12533FFA9FE52FB76FDAFBACC" authorityName="Chamberlin" authorityYear="1941" box="[428,534,1229,1255]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B59260602533FFA9FE52FB76FDAFBACC" box="[428,534,1229,1255]" italics="true" pageId="7" pageNumber="282">Amplaria</emphasis>
|
||
</taxonomicName>
|
||
plan and the other characters cited by
|
||
<bibRefCitation id="E377C1832533FFA9FC3AFB76FBD1BACC" author="Causey, N. B." box="[964,1128,1228,1255]" pageId="7" pageNumber="282" pagination="179 - 184" refId="ref10808" refString="Causey, N. B. (1958) New records and descriptions of a new genus and a new species of millipeds of the family Striariidae (Chordeumida). Proceedings of the Biological Society of Washington, 71, 179 - 184." type="journal article" year="1958">Causey (1958)</bibRefCitation>
|
||
as distinguishing
|
||
<taxonomicName id="40E6C7F12533FFA9FACCFB76FA25BACC" authorityName="Causey" authorityYear="1958" box="[1330,1436,1229,1255]" class="Diplopoda" family="Striariidae" genus="Vaferaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B59260602533FFA9FACCFB76FA25BACC" box="[1330,1436,1229,1255]" italics="true" pageId="7" pageNumber="282">Vaferaria</emphasis>
|
||
</taxonomicName>
|
||
are well within the ranges shown by other
|
||
<taxonomicName id="40E6C7F12533FFA9FD91FB4BFD60BB21" authorityName="Chamberlin" authorityYear="1941" box="[623,729,1264,1290]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B59260602533FFA9FD91FB4BFD60BB21" box="[623,729,1264,1290]" italics="true" pageId="7" pageNumber="282">Amplaria</emphasis>
|
||
</taxonomicName>
|
||
species, I consider
|
||
<taxonomicName id="40E6C7F12533FFA9FC4DFB4BFBA4BB21" authorityName="Causey" authorityYear="1958" box="[947,1053,1264,1290]" class="Diplopoda" family="Striariidae" genus="Vaferaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B59260602533FFA9FC4DFB4BFBA4BB21" box="[947,1053,1264,1290]" italics="true" pageId="7" pageNumber="282">Vaferaria</emphasis>
|
||
</taxonomicName>
|
||
to be a junior subjective synonym of
|
||
<emphasis id="B59260602533FFA9FF48FAA9FE9EBB07" box="[182,295,1298,1324]" italics="true" pageId="7" pageNumber="282">
|
||
<taxonomicName id="40E6C7F12533FFA9FF48FAA9FE9ABB07" authorityName="Chamberlin" authorityYear="1941" box="[182,291,1298,1324]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="genus">Amplaria</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
This gives us
|
||
<emphasis id="B59260602533FFA9FE36FAAAFB8BBB07" bold="true" box="[456,1074,1297,1324]" pageId="7" pageNumber="282">
|
||
<taxonomicName id="40E6C7F12533FFA9FE36FAAAFCE0BB07" authority="(Loomis, 1936)" authorityName="Shear" authorityYear="2020" baseAuthorityName="Loomis" baseAuthorityYear="1936" box="[456,857,1297,1324]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="282" phylum="Arthropoda" rank="species" species="imberbis" status="comb. nov.">
|
||
<emphasis id="B59260602533FFA9FE36FAAAFD26BB07" bold="true" box="[456,671,1297,1324]" italics="true" pageId="7" pageNumber="282">Amplaria imberbis</emphasis>
|
||
(
|
||
<bibRefCitation id="E377C1832533FFA9FD50FAAAFCE8BB07" author="Loomis, H. F." box="[686,849,1297,1324]" pageId="7" pageNumber="282" pagination="404 - 409" refId="ref11257" refString="Loomis, H. F. (1936) New millipeds of the American family Striariidae. Journal of the Washington Academy of Science, 26, 404 - 409." type="journal article" year="1936">Loomis, 1936</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="AEA1DD1B2533FFA9FC9BFAA9FB8BBB07" box="[869,1074,1297,1324]" pageId="7" pageNumber="282" rank="species">new combination</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |