215 lines
16 KiB
XML
215 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.391.7161" ID-GBIF-Dataset="2c378af4-8904-403d-b3ca-b16a2ee9951e" ID-PMC="PMC3974445" ID-Pensoft-Pub="1313-2970-391-75" ID-PubMed="24715781" ID-ZBK="D6A420ACB3BD4BE39E3C14766610838F" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-391-75" ModsDocOrigin="ZooKeys 391" ModsDocTitle="Four new spider species of the family Theridiosomatidae (Arachnida, Araneae) from caves in Laos" checkinTime="1451246140634" checkinUser="pensoft" docAuthor="Lin, Yucheng, Li, Shuqiang & Jaeger, Peter" docDate="2014" docId="0F3D94EF427443CB221676A89BF078CF" docLanguage="en" docName="ZooKeys 391: 75-102" docOrigin="ZooKeys 391" docSource="http://dx.doi.org/10.3897/zookeys.391.7161" docTitle="Alaria navicularis Lin, Li & Jaeger, 2014, sp. n." docType="treatment" docUuid="05825720-58CD-40F4-9FBE-4FBA4A97E5D1" docUuidSource="ZooBank" docVersion="4" lastPageNumber="83" masterDocId="FF9FFFFE412C3B2B1941FF90F72E8D0D" masterDocTitle="Four new spider species of the family Theridiosomatidae (Arachnida, Araneae) from caves in Laos" masterLastPageNumber="102" masterPageNumber="75" pageNumber="80" updateTime="1668158068389" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Four new spider species of the family Theridiosomatidae (Arachnida, Araneae) from caves in Laos</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lin, Yucheng</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Shuqiang</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jaeger, Peter</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>391</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>75</mods:start>
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<mods:end>102</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.391.7161</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.391.7161</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-391-75</mods:identifier>
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<mods:identifier type="ZBK">D6A420ACB3BD4BE39E3C14766610838F</mods:identifier>
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<mods:identifier type="ZooBank">D6A420ACB3BD4BE39E3C14766610838F</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152051758" LSID="urn:lsid:zoobank.org:act:05825720-58CD-40F4-9FBE-4FBA4A97E5D1" httpUri="http://treatment.plazi.org/id/0F3D94EF427443CB221676A89BF078CF" lastPageId="8" lastPageNumber="83" pageId="5" pageNumber="80">
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<subSubSection pageId="5" pageNumber="80" type="nomenclature">
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<paragraph pageId="5" pageNumber="80">
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<taxonomicName LSID="http://zoobank.org/05825720-58CD-40F4-9FBE-4FBA4A97E5D1" class="Gastropoda" family="Aporrhaidae" genus="Alaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alaria navicularis" order="Littorinimorpha" pageId="5" pageNumber="80" phylum="Mollusca" rank="species" species="navicularis">
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<pageBreakToken pageId="5" pageNumber="80" start="start">Alaria</pageBreakToken>
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navicularis
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</taxonomicName>
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<taxonomicNameLabel pageId="5" pageNumber="80">sp. n.</taxonomicNameLabel>
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Figs 5-10, 19
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="80" type="material examined">
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<paragraph pageId="5" pageNumber="80">Material examined.</paragraph>
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<paragraph pageId="5" pageNumber="80">
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Laos: Khammouan Province: Holotype: ♂ (SMF), 2.5 km WNW of Ban Tathot, entrance
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.631617">17°37.897'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="105.12503">105°07.502'E</geoCoordinate>
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, exit
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.633234">17°37.994'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="105.11992">105°07.195'E</geoCoordinate>
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, altitude 200 m, entrance area and in front of limestone cave, Tham Kamouk, leg. 30 April 2012. Paratypes: 2 ♀ (SMF), same data as holotype; 1 ♀ (SMF), 8.3 km NE of Thakek, Tham Noi,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.44425">17°26.655'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="104.862785">104°51.767'E</geoCoordinate>
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, altitude 158 m, in foot cave, leg. 26 November 2012; 1 ♀ (IZCAS), 2.5 km WNW of Ban Tathot, entrance
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.631617">17°37.897'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="105.12503">105°07.502'E</geoCoordinate>
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, exit
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.633234">17°37.994'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="105.11992">105°07.195'E</geoCoordinate>
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, altitude 200 m, entrance area and inner parts of limestone cave, Tham Kamouk, leg. 26 April 2012.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="80" type="etymology">
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<paragraph pageId="5" pageNumber="80">Etymology.</paragraph>
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<paragraph pageId="5" pageNumber="80">
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This specific name is derived from the Latin word
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<normalizedToken originalValue="“navicularis”">"navicularis"</normalizedToken>
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= "shaped like a boat", and refers to the shape of the median apophysis in the male palp; adjective.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="80" type="diagnosis">
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<paragraph pageId="5" pageNumber="80">Diagnosis.</paragraph>
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<paragraph pageId="5" pageNumber="80">
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The most significant difference of this new species to
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<taxonomicName class="Gastropoda" family="Aporrhaidae" genus="Alaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alaria chengguanensis" order="Littorinimorpha" pageId="5" pageNumber="80" phylum="Mollusca" rank="species" species="chengguanensis">Alaria chengguanensis</taxonomicName>
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(
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<bibRefCitation author="Zhao, Q" journalOrPublisher="Zookeys" pageId="13" pageNumber="88" pagination="1 - 48" title="Eleven new species of theridiosomatid spiders from southern China (Araneae, Theridiosomatidae)." url="10.3897/zookeys.255.3272" volume="255" year="2012">Zhao and Li 2012</bibRefCitation>
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: figs 1
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<normalizedToken originalValue="A–D">A-D</normalizedToken>
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, 2
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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, 3
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<normalizedToken originalValue="A–D">A-D</normalizedToken>
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, 4
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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, 5
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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) and
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<taxonomicName class="Gastropoda" family="Aporrhaidae" genus="Alaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alaria cavernicola" order="Littorinimorpha" pageId="5" pageNumber="80" phylum="Mollusca" rank="species" species="cavernicola">Alaria cavernicola</taxonomicName>
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sp. n. (Figs 1-4) is the navicular median apophysis in the male (Figs 5B, 6D, 8A, 9D), the triangular, weakly sclerotized scape and the nearly circular, juxtaposed spermathecae in the female (Figs 7
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<normalizedToken originalValue="B–D">B-D</normalizedToken>
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, 10
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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).
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</paragraph>
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<caption pageId="5" pageNumber="80">
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<paragraph pageId="5" pageNumber="80">
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Figure 5.
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<taxonomicName class="Gastropoda" family="Aporrhaidae" genus="Alaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alaria navicularis" order="Littorinimorpha" pageId="5" pageNumber="80" phylum="Mollusca" rank="species" species="navicularis">Alaria navicularis</taxonomicName>
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sp. n., holotype male. A Habitus, dorsal B Palp, prolateral C Ditto, retrolateral D Ditto, ventral E Cymbium, retrolateral F Palpal patella and tibia, retrolateral. Co = conductor; Em = embolus; MA = median apophysis; Pc = paracymbium; SCy = setae of cymbium; Te = tegulum.
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="80">
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<paragraph pageId="5" pageNumber="80">
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Figure 6.
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<taxonomicName class="Gastropoda" family="Aporrhaidae" genus="Alaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alaria navicularis" order="Littorinimorpha" pageId="5" pageNumber="80" phylum="Mollusca" rank="species" species="navicularis">Alaria navicularis</taxonomicName>
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sp. n., male holotype. A Bulb (median apophysis removed), distal B Ditto, prolateral C Ditto, retrolateral D Median apophysis, prolateral. Co = conductor; Em = embolus; MA = median apophysis; Te = tegulum.
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="80">
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<paragraph pageId="5" pageNumber="80">
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Figure 7.
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<taxonomicName class="Gastropoda" family="Aporrhaidae" genus="Alaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alaria navicularis" order="Littorinimorpha" pageId="5" pageNumber="80" phylum="Mollusca" rank="species" species="navicularis">Alaria navicularis</taxonomicName>
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sp. n., female paratype, from Tham Kamouk. A Habitus, dorsal B Epigyne, ventral C Vulva (lactic acid-treated), ventral D Ditto, dorsal (red line showing course of duct system). CD = copulatory duct; FD = fertilization duct; Sc = scape; Sp = spermathecae.
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="80">
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<paragraph pageId="5" pageNumber="80">
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Figure 8.
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<taxonomicName class="Gastropoda" family="Aporrhaidae" genus="Alaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alaria navicularis" order="Littorinimorpha" pageId="5" pageNumber="80" phylum="Mollusca" rank="species" species="navicularis">Alaria navicularis</taxonomicName>
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sp. n., holotype male. A Palp, prolateral B Ditto, retrolateral C Bulb (median apophysis removed), distal D Cymbium, retrolateral E Palpal patella and tibia, retrolateral. Co = conductor; Em = embolus; MA = median apophysis; Pc = paracymbium; SCy = setae of cymbium; Te = tegulum.
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="80">
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<paragraph pageId="5" pageNumber="80">
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Figure 9.
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<taxonomicName class="Gastropoda" family="Aporrhaidae" genus="Alaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alaria navicularis" order="Littorinimorpha" pageId="5" pageNumber="80" phylum="Mollusca" rank="species" species="navicularis">Alaria navicularis</taxonomicName>
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sp. n., holotype male. A Palp, ventral B Bulb (median apophysis removed), prolateral C Ditto, retrolateral D Median apophysis, prolateral. Co = conductor; Em = embolus; MA = median apophysis; Te = tegulum.
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="80">
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<paragraph pageId="5" pageNumber="80">
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Figure 10.
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<taxonomicName class="Gastropoda" family="Aporrhaidae" genus="Alaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alaria navicularis" order="Littorinimorpha" pageId="5" pageNumber="80" phylum="Mollusca" rank="species" species="navicularis">Alaria navicularis</taxonomicName>
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sp. n., female paratype, from Tham Kamouk. A Vulva (lactic acid-treated), ventral B Ditto, dorsal C Epigyne (untreated), ventral. CD = copulatory duct; FD = fertilization duct; Sc = scape; Sp = spermathecae.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="83" pageId="5" pageNumber="80" type="description">
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<paragraph pageId="5" pageNumber="80">Description.</paragraph>
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<paragraph pageId="5" pageNumber="80">Habitus see in Figs 5A, 7A. DS pear-shaped, yellow, eyes with black base, cervical groove distinct, sternum yellow. Legs yellow. Opisthosoma oval in male, pale yellow, dorsum with small slcerotized spots and a pair of black speckles; sub-spherical in female, pale, dorsum with white patches.</paragraph>
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<paragraph pageId="5" pageNumber="80">
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Male palp (Figs 5
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<normalizedToken originalValue="B–D">B-D</normalizedToken>
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, 8
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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, 9A): patella with strong macroseta (Figs 5C, 8B, 8E). Tibia with 3 trichobothria and lamellar process (Figs 5F, 8E). Paracymbium with basal hooked process and distal short spicula (Figs 5
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<normalizedToken originalValue="D–E">D-E</normalizedToken>
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, 8D, 9A). Tegulum medially smooth, marginally rugose (Figs 6A, 8C). Median apophysis especially large, navicular, surface with tiny aculei, proximal process short, distal process long and rugose (Figs 6D, 9D). Conductor chisel-shaped, distally slightly bent, longer than embolus, (Figs 6
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<normalizedToken originalValue="B–C">B-C</normalizedToken>
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, 9
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<normalizedToken originalValue="B–C">B-C</normalizedToken>
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). Embolus long, needle-shaped, sclerotized, distal part embedded in conductor, embolic tip not beyond distal end of conductor (Figs 6
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, 8C, 9
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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).
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</paragraph>
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<paragraph pageId="5" pageNumber="80">
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Female copulatory organ (Figs 7
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<normalizedToken originalValue="B–D">B-D</normalizedToken>
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, 10
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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): epigynal area covered with long setae (Figs 7B, 10A, 10C). Scape large, triangular, basally rugose and contracted, apically weakly sclerotized, with small distal pocket, protruding from beneath epigynal posterior margin (Figs 7
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<normalizedToken originalValue="B–D">B-D</normalizedToken>
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, 10
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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). Spermathecae sub-circular, juxtaposed; lateral copulatory ducts coils oval, bent (Figs 7D, 10B). Copulatory ducts narrow, ending laterally margin in spermathecae (Figs 7D, 10B). Fertilization ducts short, triangular, arising ventro-laterally from spermathecae (Figs 7D, 10B).
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</paragraph>
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<paragraph lastPageId="6" lastPageNumber="81" pageId="5" pageNumber="80">
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Male: total length 2.31, DS 1.10 long, 0.93 wide, clypeus 0.17, sternum 0.55 long, 0.45 wide, coxae IV separated by their width, opisthosoma 1.31 long, 1.20 wide. PME separated by less than half their diameter. Macrosetae: leg I: femur d2, p1, r1, patella d2, tibia d2, p3, r1, v2, metatarsus d1, p1, r1, v1; leg II: femur d2, r1, patella d2, tibia
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<pageBreakToken pageId="6" pageNumber="81" start="start">d</pageBreakToken>
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2, p1, r1, v1, metatarsus d2, r1, v1; leg III: femur d2, patella d2, tibia d2, p1, v1, metatarsus d1, p1, r1, v1; leg IV: femur d2, p1, patella d2, tibia d2, p1, r1, v1, metatarsus d1, p1, r1, v1. Metatarsal trichobothria: Tm I: 0.22; Tm II: 0.24; Tm III: 0.14. Leg measurements: I 3.34 (1.09, 0.45, 0.67, 0.69, 0.44); II 2.90 (0.93, 0.42, 0.57, 0.58, 0.40); III 2.16 (0.72, 0.32, 0.36, 0.43, 0.33); IV 2.65 (0.85, 0.37, 0.53, 0.55, 0.35).
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</paragraph>
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<paragraph lastPageId="8" lastPageNumber="83" pageId="6" pageNumber="81">
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Female (collected together with holotype): Total length 3.62, DS 1.24 long, 1.45 wide, clypeus 0.15, sternum 0.78 long, 0.68 wide, coxae IV separated by their width,
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<pageBreakToken pageId="7" pageNumber="82" start="start">opisthosoma</pageBreakToken>
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2.45 long, 2.35 wide. PME separated by less than half their diameter. Macrosetae: Leg I: femur d1, p1, r1, patella d2, tibia d2, p2, r1, v1, metatarsus p1, r1, v1; leg II: femur d2, r1, patella d2, tibia d2, p1, r1, v1, metatarsus p1, r1, v1; leg III: femur d1, v1, patella d2, tibia d2, p1, v1, metatarsus d1, p1, r1, v1; leg IV: femur d2, p1, patella d2, tibia d2, p1, r1, v1, metatarsus d1, p1, r1, v1. Metatarsal trichobothria: Tm I: 0.30; Tm II: 0.25; Tm III: 0.15. Leg measurements: I 4.46 (1.45, 0.61, 0.93,
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<pageBreakToken pageId="8" pageNumber="83" start="start">0.88</pageBreakToken>
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, 0.59); II 3.85 (1.20, 0.56, 0.78, 0.77, 0.54); III 2.83 (0.92, 0.42, 0.49, 0.54, 0.46); IV 3.54 (1.16, 0.52, 0.70, 0.69, 0.47).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="83" type="variation">
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<paragraph pageId="8" pageNumber="83">Variation.</paragraph>
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<paragraph pageId="8" pageNumber="83">The total length ranges from 3.42 to 3.70 in females (n = 4).</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="83" type="distribution">
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<paragraph pageId="8" pageNumber="83">Distribution.</paragraph>
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<paragraph pageId="8" pageNumber="83">See in Fig. 19.</paragraph>
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</subSubSection>
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</treatment>
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</document> |