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<document id="4CC27C6B39038E5E8FECC907DA504F59" ID-DOI="10.11646/zootaxa.5323.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8212306" ID-ZooBank="DAAF3E31-A245-4421-A41B-E9BE1538ABF9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691078968945" checkinUser="plazi" docAuthor="Noguera, Felipe A. &amp; Santos-Silva, Antonio" docDate="2023" docId="FE18879D9B198316FF2BFDD6FAEBF962" docLanguage="en" docName="zootaxa.5323.4.1.pdf" docOrigin="Zootaxa 5323 (4)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phrynidius oaxacanensis Noguera &amp; Santos-Silva 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="469" masterDocId="0221FFE59B098304FFBCFFF4FFD1FFDF" masterDocTitle="New species and records in Phrynidius Lacordaire (Coleoptera, Cerambycidae, Lamiinae)" masterLastPageNumber="476" masterPageNumber="451" pageNumber="467" updateTime="1691079457083" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="8ED7CDD1F7C4F27CD1B1B32EA6653092" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E16B3A5ACE8A78ADA683047D6CBF6DD6">
<mods:title id="17803853FB8D68772C8C92C9A808FCF7">New species and records in Phrynidius Lacordaire (Coleoptera, Cerambycidae, Lamiinae)</mods:title>
</mods:titleInfo>
<mods:name id="B61199B761284CCC2C587A8078DB552D" type="personal">
<mods:role id="9D496EA3A6D066E57EBBC90155651A1C">
<mods:roleTerm id="CE65ED5159ED0F16B029EB04887A8E5C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BE9CAB2F01040FD4CC7620ADFE140D07">Noguera, Felipe A.</mods:namePart>
<mods:affiliation id="A69AA0C744C07F94F1150B26E07885F9">Estacioìn de Biologiìa Chamela, Instituto de Biologiìa, Universidad Nacional Autoìnoma de Meìxico, Apartado Postal 21, San Patricio, Jalisco 48980, Mexico.</mods:affiliation>
</mods:name>
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<mods:role id="DC07666466E4436FCB2FC04D88B455DE">
<mods:roleTerm id="A1714DB396B63A87CEB6E53BD04C54A7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8780C69240548299BD11B3399828834D">Santos-Silva, Antonio</mods:namePart>
<mods:affiliation id="B88A4D4665FD478C0BB8F696C561E92A">Museu de Zoologia, Universidade de São Paulo, São Paulo, SP, Brazil.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="1E5FC1FE54EB92CA2B5BFBF3F214DAA1">text</mods:typeOfResource>
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<mods:titleInfo id="EF4C7755DA6888B3C725E79DE139D17D">
<mods:title id="481A3CD2A80808D0BB32C4D5BE2D08B6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8FA8FDAE2841C87ABC71192B3E2D1883">
<mods:date id="9CCC7037AFF02D2B8FEC13C5C9F50AEF">2023</mods:date>
<mods:detail id="CEBCE492064EEE808D7E85FF0DBE458F" type="pubDate">
<mods:number id="E078F1243AF01A3E76C35E53893016BC">2023-08-03</mods:number>
</mods:detail>
<mods:detail id="03910AAB3743A95FC0210F65C470BF70" type="volume">
<mods:number id="6690F78CD1FCF6802ACF08DFBC2EBD40">5323</mods:number>
</mods:detail>
<mods:detail id="CA85BC14DC045E38C207B9F89C08542E" type="issue">
<mods:number id="F298D3AE37BD1239A8CCEFCDCB9EDC36">4</mods:number>
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<mods:start id="809D87B9937F7E151F33B9A3019B44DA">451</mods:start>
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<mods:location id="69ED5B997579A27049BB217F5D4A2BD7">
<mods:url id="B013EDB67F6D5B09B218F4814B8FC9FB">http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification id="5AFB3BE7B15F9072FD5A1B00CE14AB32">journal article</mods:classification>
<mods:identifier id="83AB378C1E421634B58E846EBC5332AB" type="DOI">10.11646/zootaxa.5323.4.1</mods:identifier>
<mods:identifier id="CF77F4FE4C4E05E8E9B1F083E364DF42" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="52F7CD08FE0C8943B4891B13C78A21AF" type="Zenodo-Dep">8212306</mods:identifier>
<mods:identifier id="09A09B33FB0B5CE0D4F2BCD43E0B67A4" type="ZooBank">DAAF3E31-A245-4421-A41B-E9BE1538ABF9</mods:identifier>
</mods:mods>
<treatment id="FE18879D9B198316FF2BFDD6FAEBF962" ID-DOI="http://doi.org/10.5281/zenodo.8212344" ID-Zenodo-Dep="8212344" LSID="urn:lsid:plazi:treatment:FE18879D9B198316FF2BFDD6FAEBF962" httpUri="http://treatment.plazi.org/id/FE18879D9B198316FF2BFDD6FAEBF962" lastPageId="18" lastPageNumber="469" pageId="16" pageNumber="467">
<subSubSection id="3EAB65009B198314FF2BFDD6FDFDFDE3" box="[151,556,546,573]" pageId="16" pageNumber="467" type="nomenclature">
<paragraph id="760E368B9B198314FF2BFDD6FDFDFDE3" blockId="16.[151,556,546,608]" box="[151,556,546,573]" pageId="16" pageNumber="467">
<heading id="2D4681E79B198314FF2BFDD6FDFDFDE3" bold="true" box="[151,556,546,573]" fontSize="11" level="1" pageId="16" pageNumber="467" reason="1">
<emphasis id="44C5EA999B198314FF2BFDD6FDFDFDE3" bold="true" box="[151,556,546,573]" pageId="16" pageNumber="467">
<taxonomicName id="B1B14D089B198314FF2BFDD6FE16FDE3" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[151,455,546,573]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="oaxacanensis" status="sp. nov.">
<emphasis id="44C5EA999B198314FF2BFDD6FE16FDE3" bold="true" box="[151,455,546,573]" italics="true" pageId="16" pageNumber="467">Phrynidius oaxacanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FF657E29B198314FE73FDD6FDFDFDE3" box="[463,556,546,572]" pageId="16" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3EAB65009B198314FF2BFDB2FEF6FDBF" box="[151,295,582,608]" pageId="16" pageNumber="467" type="description">
<paragraph id="760E368B9B198314FF2BFDB2FEF6FDBF" blockId="16.[151,556,546,608]" box="[151,295,582,608]" pageId="16" pageNumber="467">
(
<figureCitation id="EE8A2A0E9B198314FF23FDB2FECEFDBF" box="[159,287,582,608]" captionStart="FIGURES 36–41" captionStartId="17.[152,270,1966,1990]" captionTargetBox="[216,1371,309,1943]" captionTargetId="figure-88@17.[216,1371,309,1943]" captionTargetPageId="17" captionText="FIGURES 36–41. Phrynidius oaxacanensis sp. nov. 36–39) Holotype male: 36) Dorsal habitus; 37) Ventral habitus; 38) Lateral habitus; 39) Head, frontal view. 40–41) Paratype female: 40) Dorsal habitus; 41) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212332" httpUri="https://zenodo.org/record/8212332/files/figure.png" pageId="16" pageNumber="467">Figs 36–41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3EAB65009B198314FF2BFD7AFDCDFCCB" pageId="16" pageNumber="467" type="materials_examined">
<paragraph id="760E368B9B198314FF2BFD7AFDCDFCCB" blockId="16.[151,1437,654,2013]" pageId="16" pageNumber="467">
<emphasis id="44C5EA999B198314FF2BFD7AFE97FD77" bold="true" box="[151,326,654,680]" pageId="16" pageNumber="467">Type material.</emphasis>
<materialsCitation id="C6D93CD69B198314FEF1FD7AFDBDFD13" collectingDate="1982-08-29" collectionCode="FSCA" collectorName="C. W. &amp; L. O'Brien &amp; G. Wibmer" country="Mexico" location="Hwy" pageId="16" pageNumber="467" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
<typeStatus id="A90A88299B198314FEF1FD7AFE64FD77" box="[333,437,654,680]" pageId="16" pageNumber="467">Holotype</typeStatus>
male from
<collectingCountry id="0EA6761B9B198314FD85FD7AFD7BFD77" box="[569,682,654,680]" name="Mexico" pageId="16" pageNumber="467">MEXICO</collectingCountry>
,
<emphasis id="44C5EA999B198314FD0BFD7BFCDFFD76" box="[695,782,655,681]" italics="true" pageId="16" pageNumber="467">
<collectingRegion id="B475F8699B198314FD0BFD7BFCDFFD76" box="[695,782,655,681]" country="Mexico" name="Oaxaca" pageId="16" pageNumber="467">Oaxaca</collectingRegion>
</emphasis>
:
<location id="736E60509B198314FCA1FD7BFC82FD76" LSID="urn:lsid:plazi:treatment:FE18879D9B198316FF2BFDD6FAEBF962:736E60509B198314FCA1FD7BFC82FD76" box="[797,851,655,681]" country="Mexico" name="Hwy" pageId="16" pageNumber="467" stateProvince="Oaxaca">Hwy</location>
175,
<quantity id="B1499B6E9B198314FC28FD7AFC0DFD77" box="[916,988,654,680]" metricMagnitude="4" metricUnit="m" metricValue="7.6" pageId="16" pageNumber="467" unit="km" value="76.0">76 km</quantity>
NE
<collectingRegion id="B475F8699B198314FBACFD7AFBB6FD77" box="[1040,1127,654,680]" country="Mexico" name="Oaxaca" pageId="16" pageNumber="467">Oaxaca</collectingRegion>
, 9400’,
<date id="020F104B9B198314FB7FFD7AFA86FD77" box="[1219,1367,654,680]" pageId="16" pageNumber="467" value="1982-08-29">
<collectingDate id="124BE9A39B198314FB7FFD7AFA86FD77" box="[1219,1367,654,680]" pageId="16" pageNumber="467" value="1982-08-29">29.VIII.1982</collectingDate>
</date>
,
<collectorName id="DB44535D9B198314FAD8FD7AFEE3FD13" pageId="16" pageNumber="467">C.W. &amp; L. O’Brien</collectorName>
&amp;
<collectorName id="DB44535D9B198314FEE4FD46FE07FD13" box="[344,470,690,716]" pageId="16" pageNumber="467">G. Wibmer</collectorName>
leg. (
<collectionCode id="10A0AE4E9B198314FDA4FD46FDB5FD13" box="[536,612,690,716]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="16" pageNumber="467" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C6D93CD69B198314FDC4FD46FC85FD2E" collectingDate="1982-08-29" collectionCode="MZSP, FSCA" collectorName="C. W. &amp; L. O'Brien &amp; G. Wibmer" country="Mexico" location="Hwy" pageId="16" pageNumber="467" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca" typeStatus="paratype">
<typeStatus id="A90A88299B198314FDC4FD46FD37FD13" box="[632,742,690,716]" pageId="16" pageNumber="467">Paratypes</typeStatus>
—
<collectingCountry id="0EA6761B9B198314FCBFFD46FCA2FD13" box="[771,883,690,716]" name="Mexico" pageId="16" pageNumber="467">MEXICO</collectingCountry>
,
<emphasis id="44C5EA999B198314FCC3FD47FC07FD12" box="[895,982,691,717]" italics="true" pageId="16" pageNumber="467">
<collectingRegion id="B475F8699B198314FCC3FD47FC07FD12" box="[895,982,691,717]" country="Mexico" name="Oaxaca" pageId="16" pageNumber="467">Oaxaca</collectingRegion>
</emphasis>
:
<specimenCount id="60B7FD029B198314FC5AFD46FBE7FD13" box="[998,1078,690,716]" pageId="16" pageNumber="467" type="male">1 male</specimenCount>
, same data as holotype (
<collectionCode id="10A0AE4E9B198314FAF1FD46FA49FD13" box="[1357,1432,690,716]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="16" pageNumber="467" type="Museum">MZSP</collectionCode>
, formerly
<collectionCode id="10A0AE4E9B198314FEBEFD22FE9CFD2F" box="[258,333,726,752]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="16" pageNumber="467" type="Museum">FSCA</collectionCode>
);
<locationDeviation id="67BC2FA99B198314FEDFFD22FD87FD2F" box="[355,598,726,752]" location="Valle Nac'l" pageId="16" pageNumber="467">
<quantity id="B1499B6E9B198314FEDFFD22FE65FD2F" box="[355,436,726,752]" metricMagnitude="3" metricUnit="m" metricValue="5.9545728" pageId="16" pageNumber="467" unit="mi" value="3.7">3.7 mi.</quantity>
S Valle Nac’l
</locationDeviation>
[Nacional],
<location id="736E60509B198314FD54FD23FCCFFD2E" LSID="urn:lsid:plazi:treatment:FE18879D9B198316FF2BFDD6FAEBF962:736E60509B198314FD54FD23FCCFFD2E" box="[744,798,727,753]" country="Mexico" name="Hwy" pageId="16" pageNumber="467" stateProvince="Oaxaca">Hwy</location>
175
</materialsCitation>
,
<materialsCitation id="C6D93CD69B198314FCDEFD22FDC9FCCB" collectingDate="1983-05-27" collectionCode="MZSP, FSCA" collectorName="C. W. &amp; L. O'Brien" country="Mexico" pageId="16" pageNumber="467" specimenCount="2" specimenCount-female="2" stateProvince="Oaxaca" typeStatus="paratype">
<specimenCount id="60B7FD029B198314FCDEFD22FC02FD2F" box="[866,979,726,752]" pageId="16" pageNumber="467" type="female">2 females</specimenCount>
,
<date id="020F104B9B198314FC5DFD22FB87FD2F" box="[993,1110,726,752]" pageId="16" pageNumber="467" value="1983-05-27">
<collectingDate id="124BE9A39B198314FC5DFD22FB87FD2F" box="[993,1110,726,752]" pageId="16" pageNumber="467" value="1983-05-27">27.V.1983</collectingDate>
</date>
,
<collectorName id="DB44535D9B198314FBD8FD22FA93FD2F" box="[1124,1346,726,753]" pageId="16" pageNumber="467">C.W. &amp; L. O’Brien</collectorName>
leg. (1,
<collectionCode id="10A0AE4E9B198314FF2BFD0EFF31FCCB" box="[151,224,762,788]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="16" pageNumber="467" type="Museum">MZSP</collectionCode>
, formerly
<collectionCode id="10A0AE4E9B198314FEE8FD0EFE4FFCCB" box="[340,414,762,788]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="16" pageNumber="467" type="Museum">FSCA</collectionCode>
; 1,
<collectionCode id="10A0AE4E9B198314FE79FD0EFDC1FCCB" box="[453,528,762,788]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="16" pageNumber="467" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3EAB65009B198314FF7BFCEAFE28FBEB" pageId="16" pageNumber="467" type="diagnosis">
<paragraph id="760E368B9B198314FF7BFCEAFE28FBEB" blockId="16.[151,1437,654,2013]" pageId="16" pageNumber="467">
<emphasis id="44C5EA999B198314FF7BFCEAFE93FCE7" bold="true" box="[199,322,798,824]" pageId="16" pageNumber="467">Diagnosis.</emphasis>
<taxonomicName id="B1B14D089B198314FEF9FCEAFD8AFCE7" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[325,603,798,824]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="oaxacanensis" status="sp. nov.">
<emphasis id="44C5EA999B198314FEF9FCEAFD8AFCE7" box="[325,603,798,824]" italics="true" pageId="16" pageNumber="467">Phrynidius oaxacanensis</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B198314FDE3FCEAFD67FCE7" bold="true" box="[607,694,798,824]" pageId="16" pageNumber="467">
<taxonomicNameLabel id="5FF657E29B198314FDE3FCEAFD67FCE7" box="[607,694,798,824]" pageId="16" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="B1B14D089B198314FC80FCEBFC20FCE7" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[828,1009,798,824]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="pallifemoralis" status="sp. nov.">
<emphasis id="44C5EA999B198314FC80FCEBFC20FCE7" box="[828,1009,798,824]" italics="true" pageId="16" pageNumber="467">P.pallifemoralis</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B198314FC49FCEAFB9DFCE7" bold="true" box="[1013,1100,798,824]" pageId="16" pageNumber="467">
<taxonomicNameLabel id="5FF657E29B198314FC49FCEAFB9DFCE7" box="[1013,1100,798,824]" pageId="16" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but differs by the anterocentral tubercle of the pronotum irregular (regularly convex and straight in
<taxonomicName id="B1B14D089B198314FC22FCB7FB89FC83" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[926,1112,834,860]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="pallifemoralis" status="sp. nov.">
<emphasis id="44C5EA999B198314FC22FCB7FB89FC83" box="[926,1112,834,860]" italics="true" pageId="16" pageNumber="467">P. pallifemoralis</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B198314FBDDFCB6FB6DFC83" bold="true" box="[1121,1212,834,860]" pageId="16" pageNumber="467">
<taxonomicNameLabel id="5FF657E29B198314FBDDFCB6FB6DFC83" box="[1121,1212,834,860]" pageId="16" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), and by the elytral tubercles distinctly more elevated and acuminate apically (less elevated and mostly rounded in
<taxonomicName id="B1B14D089B198314FB00FC93FAA5FC5F" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[1212,1396,870,896]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="pallifemoralis" status="sp. nov.">
<emphasis id="44C5EA999B198314FB00FC93FAA5FC5F" box="[1212,1396,870,896]" italics="true" pageId="16" pageNumber="467">P. pallifemoralis</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B198314FAC7FC92FF19FC7B" bold="true" pageId="16" pageNumber="467">
<taxonomicNameLabel id="5FF657E29B198314FAC7FC92FF19FC7B" pageId="16" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). It differs from
<taxonomicName id="B1B14D089B198314FE35FC7FFD73FC7B" authority="Linsley, 1933" authorityName="Linsley" authorityYear="1933" box="[393,674,906,932]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="44C5EA999B198314FE35FC7FFDD0FC7B" box="[393,513,907,932]" italics="true" pageId="16" pageNumber="467">P. armatus</emphasis>
<bibRefCitation id="12204B7A9B198314FDB5FC7EFD73FC7B" author="Linsley, E. G." box="[521,674,906,932]" pageId="16" pageNumber="467" pagination="131 - 132" refId="ref17088" refString="Linsley, E. G. (1933) A new longicorn beetle from Central America. The Pan-Pacific Entomologist, 9 (3), 131 - 132." type="journal article" year="1933">Linsley, 1933</bibRefCitation>
</taxonomicName>
by the body slender (stout in
<taxonomicName id="B1B14D089B198314FC4BFC7FFBBEFC7B" authorityName="Linsley" authorityYear="1933" box="[1015,1135,907,932]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="44C5EA999B198314FC4BFC7FFBBEFC7B" box="[1015,1135,907,932]" italics="true" pageId="16" pageNumber="467">P. armatus</emphasis>
</taxonomicName>
), presence of anterolateral tubercle on the pronotum (absent in
<taxonomicName id="B1B14D089B198314FD84FC5BFD60FC17" authorityName="Linsley" authorityYear="1933" box="[568,689,943,968]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="44C5EA999B198314FD84FC5BFD60FC17" box="[568,689,943,968]" italics="true" pageId="16" pageNumber="467">P. armatus</emphasis>
</taxonomicName>
), and by elytral tubercles distinctly more elevated and acuminate apically (less elevated and mostly rounded in
<taxonomicName id="B1B14D089B198314FD21FC27FCC4FC33" authorityName="Linsley" authorityYear="1933" box="[669,789,979,1004]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="44C5EA999B198314FD21FC27FCC4FC33" box="[669,789,979,1004]" italics="true" pageId="16" pageNumber="467">P. armatus</emphasis>
</taxonomicName>
); from
<taxonomicName id="B1B14D089B198314FCD0FC27FA81FC32" authority="Franz, 1954" authorityName="Franz" authorityYear="1954" box="[876,1360,978,1005]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="subSpecies" species="salvadorensis" subSpecies="salvadorensis">
<emphasis id="44C5EA999B198314FCD0FC27FB13FC33" box="[876,1218,978,1004]" italics="true" pageId="16" pageNumber="467">P. salvadorensis salvadorensis</emphasis>
Franz, 1954
</taxonomicName>
and
<taxonomicName id="B1B14D089B198314FA34FC27FDA3FBCE" authority="Franz, 1954" authorityName="Franz" authorityYear="1954" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="subSpecies" species="salvadorensis" subSpecies="montecristensis">
<emphasis id="44C5EA999B198314FA34FC27FE35FBCF" italics="true" pageId="16" pageNumber="467">P. salvadorensis montecristensis</emphasis>
Franz, 1954
</taxonomicName>
, by the presence of anterolateral tubercle on the pronotum (absent in both subspecies of
<taxonomicName id="B1B14D089B198314FE88FBEFFE38FBEB" authorityName="Franz" authorityYear="1954" box="[308,489,1050,1076]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="salvadorensis">
<emphasis id="44C5EA999B198314FE88FBEFFE38FBEB" box="[308,489,1050,1076]" italics="true" pageId="16" pageNumber="467">P. salvadorensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="3EAB65009B198316FF7BFBCAFDB2F946" lastPageId="18" lastPageNumber="469" pageId="16" pageNumber="467" type="description">
<paragraph id="760E368B9B198314FF7BFBCAFDD4FB7E" blockId="16.[151,1437,654,2013]" pageId="16" pageNumber="467">
<emphasis id="44C5EA999B198314FF7BFBCAFDDEFB87" bold="true" box="[199,527,1086,1112]" pageId="16" pageNumber="467">
Description.
<typeStatus id="A90A88299B198314FEDCFBCAFE1CFB87" box="[352,461,1086,1112]" pageId="16" pageNumber="467">Holotype</typeStatus>
male
</emphasis>
(
<figureCitation id="EE8A2A0E9B198314FDA2FBCAFD70FB87" box="[542,673,1086,1112]" captionStart="FIGURES 36–41" captionStartId="17.[152,270,1966,1990]" captionTargetBox="[216,1371,309,1943]" captionTargetId="figure-88@17.[216,1371,309,1943]" captionTargetPageId="17" captionText="FIGURES 36–41. Phrynidius oaxacanensis sp. nov. 36–39) Holotype male: 36) Dorsal habitus; 37) Ventral habitus; 38) Lateral habitus; 39) Head, frontal view. 40–41) Paratype female: 40) Dorsal habitus; 41) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212332" httpUri="https://zenodo.org/record/8212332/files/figure.png" pageId="16" pageNumber="467">Figs 36–39</figureCitation>
). Integument mostly black; palpomeres dark brown with yellowish-brown apex; base of pedicel, and base and apex of antennomeres III–IV dark reddish brown; antennomeres V–XI brown with reddish-brown apex.
</paragraph>
<paragraph id="760E368B9B198315FF7BFB5EFAA5FEC2" blockId="16.[151,1437,654,2013]" lastBlockId="17.[151,1436,151,285]" lastPageId="17" lastPageNumber="468" pageId="16" pageNumber="467">
<emphasis id="44C5EA999B198314FF7BFB5EFEDFFB1B" bold="true" box="[199,270,1194,1220]" pageId="16" pageNumber="467">Head.</emphasis>
Frons abundantly, coarsely punctate; with abundant ochraceous pubescence obscuring integument on half close to clypeus, pubescence gradually slightly sparser, dark brown toward antennal tubercles; with short, decumbent, thick white setae interspersed.Antennal tubercles close to each other basally; coarsely punctate, punctures finer than on frons, except smooth apex; with dark-brown pubescence partially obscuring integument and short, decumbent, thick white setae interspersed, except glabrous smooth area. Vertex abundantly, coarsely punctate; with abundant, short brown pubescence not obscuring integument from area between antennal tubercles to anterior margin of upper eye lobes, with abundant yellowish-brown pubescence almost obscuring integument on remaining surface; with a few short, decumbent, thick white setae interspersed. Area behind eyes coarsely punctate, punctures slightly sparer toward inferior area of lower eye lobe; with dense yellowish-brown pubescence obscuring integument, and sparse, decumbent white setae interspersed. Genae coarsely punctate toward ventral surface, smooth toward clypeus; 1.9 times width of lower eye lobe; with abundant yellowish-brown pubescence obscuring integument, and short, bristly white setae interspersed. Wide central area of postclypeus with sparse white pubescence and a few long, erect yellowish-brown setae close to anteclypeus; sides of postclypeus almost glabrous. Labrum very finely punctate with a few coarser punctures interspersed; almost glabrous close to anteclypeus, with sparse yellowish-white pubescence centrally, pubescence longer and more abundant laterally, and dense fringe of yellow setae anteriorly. Eye lobes weakly divided; distance between upper eye lobes 0.52 times distance between outer margins of eyes; in frontal view, distance between lower eye lobes 0.73 times distance between outer margins of eyes; lower eye lobes longer than wide (1.45 times). Antennae 1.15 times elytral length, slightly surpassing base of posterior third of elytra. Scape somewhat abundantly and finely punctate; with abundant brown pubescence partially obscuring integument, except base and apex with abundant yellowish-white pubescence, and arched, thick, both dark brown and yellowish-white setae interspersed. Pedicel with yellowish-white pubescence, sparse on basal half, dense on apical half. Antennomere III sub-straight with pubescence and setae as on scape, but also with a few arched and thick yellowish-brown setae interspersed. Antennomere IV with yellowish-white pubescence basally and apically, pubescence dark brown on remaining surface, and arched, thick, both yellowish-white and yellowish-brown setae interspersed on basal 2/3. Antennomeres V–XI with yellowish-brown pubescence not obscuring integument, except apex with denser yellowish-white pubescence. Antennal formula (ratio) based on length of antennomere III: scape = 1.23; pedicel = 0.27; IV = 0.91; V = 0.54; VI = 0.50; VII = 0.48; VIII = 0.45; IX = 0.45; X = 0.45; XI = 0.59.
</paragraph>
<caption id="22CE66039B188315FF24F85AFB6DF834" ID-DOI="http://doi.org/10.5281/zenodo.8212332" ID-Zenodo-Dep="8212332" httpUri="https://zenodo.org/record/8212332/files/figure.png" pageId="17" pageNumber="468" startId="17.[152,270,1966,1990]" targetBox="[216,1371,309,1943]" targetPageId="17" targetType="figure">
<paragraph id="760E368B9B188315FF24F85AFB6DF834" blockId="17.[151,1436,1966,2027]" pageId="17" pageNumber="468">
<emphasis id="44C5EA999B188315FF24F85AFE8FF818" bold="true" box="[152,350,1966,1991]" pageId="17" pageNumber="468">FIGURES 36–41.</emphasis>
<taxonomicName id="B1B14D089B188315FED4F85BFDB8F819" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[360,617,1967,1991]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="oaxacanensis" status="sp. nov.">
<emphasis id="44C5EA999B188315FED4F85BFDB8F819" box="[360,617,1967,1991]" italics="true" pageId="17" pageNumber="468">Phrynidius oaxacanensis</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B188315FDCFF85BFCCBF819" bold="true" box="[627,794,1966,1991]" pageId="17" pageNumber="468">
<taxonomicNameLabel id="5FF657E29B188315FDCFF85BFD19F818" box="[627,712,1967,1991]" pageId="17" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
36–39)
</emphasis>
Holotype male:
<emphasis id="44C5EA999B188315FC72F85AFC21F819" bold="true" box="[974,1008,1966,1990]" pageId="17" pageNumber="468">36)</emphasis>
Dorsal habitus;
<emphasis id="44C5EA999B188315FB1EF85AFB12F819" bold="true" box="[1186,1219,1966,1990]" pageId="17" pageNumber="468">37)</emphasis>
Ventral habitus;
<emphasis id="44C5EA999B188315FAC6F85AFA4DF819" bold="true" box="[1402,1436,1966,1990]" pageId="17" pageNumber="468">38)</emphasis>
Lateral habitus;
<emphasis id="44C5EA999B188315FE80F826FE8CF835" bold="true" box="[316,349,2002,2026]" pageId="17" pageNumber="468">39)</emphasis>
Head, frontal view.
<emphasis id="44C5EA999B188315FD90F826FDA5F835" bold="true" box="[556,628,2002,2026]" pageId="17" pageNumber="468">40–41)</emphasis>
Paratype female:
<emphasis id="44C5EA999B188315FC97F826FC9CF835" bold="true" box="[811,845,2002,2026]" pageId="17" pageNumber="468">40)</emphasis>
Dorsal habitus;
<emphasis id="44C5EA999B188315FC48F826FBC4F835" bold="true" box="[1012,1045,2002,2026]" pageId="17" pageNumber="468">41)</emphasis>
Ventral habitus.
</paragraph>
</caption>
<paragraph id="760E368B9B1B8316FF7BFF63FA8EFB1A" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="469">
<emphasis id="44C5EA999B1B8316FF7BFF63FEF6FF6E" bold="true" box="[199,295,151,177]" pageId="18" pageNumber="469">Thorax.</emphasis>
Prothorax longer than wide; sides slightly rounded from anterolateral angles to posterior quarter, then narrowed and parallel-sided toward posterolateral angles. Pronotum moderately coarsely punctate and rugose; with five distinct gibbosities, one longitudinal, irregular, located centrally from anterior margin to apex of posterior quarter, one irregular, oblique, located on each side slightly before middle, and another more irregular located on each side slightly after middle; central area between gibbosities irregularly, longitudinally and slightly elevated; with short yellowish-brown pubescence not obscuring integument, pubescence denser and paler on posterocentral area and sides; with a few short, decumbent white setae interspersed, and decumbent yellowish-brown setae on gibbosities (yellowish-brown setae converging toward apex of gibbosities). Sides of prothorax with irregular surface, somewhat abundantly, coarsely punctate; with abundant yellowish-brown pubescence partially obscuring integument. Prosternum coarsely punctate; with abundant yellowish-white pubescence not obscuring integument, pubescence denser, yellower laterally and close to procoxal cavities. Prosternal process with sides slightly divergent from near base to wider apical area; abundantly, coarsely punctate; narrowest area 0.75 times procoxal width; with abundant whitish pubescence not obscuring integument, except glabrous sides. Mesoventrite smooth on wide central area, finely punctate laterally; with whitish pubescence not obscuring integument on wide central area, pubescence denser, yellower laterally. Mesanepisternum and mesepimeron somewhat abundantly and coarsely punctate; with abundant yellowish-brown pubescence partially obscuring integument, pubescence slightly sparser on mesepimeron. Metaventrite somewhat sparsely and finely punctate laterally, smooth centrally; with abundant yellowish-brown pubescence partially obscuring integument, except glabrous posterior half of central area, and short, decumbent, thick white setae interspersed. Mesoventral process slightly widened on posterior half; widest area 0.74 times mesocoxal width; pubescence as on prosternal process.
<emphasis id="44C5EA999B1B8316FC02FCB7FBC0FC82" bold="true" box="[958,1041,835,861]" pageId="18" pageNumber="469">Elytra.</emphasis>
Subelliptical; abundantly, coarsely punctate; with elevated conical tubercles, from near base to near apex, forming longitudinal row on dorsal posterior third of each elytron; with abundant yellowish-brown pubescence almost obscuring integument throughout, and short, decumbent, thick yellowish-white setae interspersed (yellowish-white setae converging toward apex on some tubercles).
<emphasis id="44C5EA999B1B8316FEA4FC27FE86FC32" bold="true" box="[280,343,979,1005]" pageId="18" pageNumber="469">Legs.</emphasis>
Femora with abundant yellowish-brown pubescence almost obscuring integument, and short, arched, thick, mostly white setae interspersed. Tibiae with dense yellowish pubescence dorsally and laterally on base, and dorsally on apex; sides of apical area of protibiae and sides and ventral apical area of meso- and metatibiae with white pubescence not obscuring integument; protibiae with dense, bristly yellowish-brown pubescence on apical quarter of ventral surface; dorsal sulcus of mesotibiae with dense, short, erect yellowish-brown setae; apex of all tibiae with thick yellowish-brown setae; remaining surface of tibiae with abundant brownish pubescence partially obscuring integument; all tibiae with short, arched, thick, both white and yellowish-brown setae interspersed.
</paragraph>
<paragraph id="760E368B9B1B8316FF7BFB3AFF2EFAA6" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="469">
<emphasis id="44C5EA999B1B8316FF7BFB3AFE92FB37" bold="true" box="[199,323,1230,1256]" pageId="18" pageNumber="469">Abdomen.</emphasis>
Ventrites with dense yellowish-brown pubescence laterally, especially on posterior area of ventrites 1 and 2, dense, pale longitudinal pubescent band centrally, especially on ventrites 2–4, and pale abundant pubescence not obscuring integument on remaining surface, pubescence sparser on entire posterior third of ventrite 5; with short, decumbent, thick white setae interspersed, except on posterior third of ventrite 5; ventrite 5 slightly depressed centrally.
</paragraph>
<paragraph id="760E368B9B1B8316FF7BFA77FEC3FA3A" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="469">
<emphasis id="44C5EA999B1B8316FF7BFA77FECFFA42" bold="true" box="[199,286,1411,1437]" pageId="18" pageNumber="469">Female</emphasis>
(
<figureCitation id="EE8A2A0E9B1B8316FE92FA77FE62FA42" box="[302,435,1411,1437]" captionStart="FIGURES 36–41" captionStartId="17.[152,270,1966,1990]" captionTargetBox="[216,1371,309,1943]" captionTargetId="figure-88@17.[216,1371,309,1943]" captionTargetPageId="17" captionText="FIGURES 36–41. Phrynidius oaxacanensis sp. nov. 36–39) Holotype male: 36) Dorsal habitus; 37) Ventral habitus; 38) Lateral habitus; 39) Head, frontal view. 40–41) Paratype female: 40) Dorsal habitus; 41) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212332" httpUri="https://zenodo.org/record/8212332/files/figure.png" pageId="18" pageNumber="469">Figs 40–41</figureCitation>
). Similar to male, differing especially by the shorter antennae, 1.05 times elytral length, reaching about middle of elytra; scape distinctly longer than antennomere III (1.7 times), and antennomere IV as long as III.
</paragraph>
<paragraph id="760E368B9B1B8316FF7BFA1AFC4DF9D6" blockId="18.[151,1437,151,1725]" box="[199,924,1518,1545]" pageId="18" pageNumber="469">
<emphasis id="44C5EA999B1B8316FF7BFA1AFE9CF9D7" bold="true" box="[199,333,1518,1544]" pageId="18" pageNumber="469">Variability.</emphasis>
Antennomeres V–XI light brown with reddish apex.
</paragraph>
<paragraph id="760E368B9B1B8316FF7BF9E6FDB2F946" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="469">
<emphasis id="44C5EA999B1B8316FF7BF9E6FE67F9F2" bold="true" box="[199,438,1554,1581]" pageId="18" pageNumber="469">Dimensions in mm</emphasis>
(
<typeStatus id="A90A88299B1B8316FE70F9E7FDE1F9F2" box="[460,560,1555,1581]" pageId="18" pageNumber="469">holotype</typeStatus>
male/
<typeStatus id="A90A88299B1B8316FDC0F9E7FD0DF9F2" box="[636,732,1555,1581]" pageId="18" pageNumber="469">paratype</typeStatus>
male/
<typeStatus id="A90A88299B1B8316FC96F9E7FC44F9F2" box="[810,917,1555,1581]" pageId="18" pageNumber="469">paratypes</typeStatus>
female). Total length, 8.30/7.90/9.35–9.55; prothoracic length, 2.30/2.10/2.60–2.80; anterior prothoracic width, 1.80/1.75/2.05–2.10; posterior prothoracic width, 1.75/1.70/2.05–2.10; maximum prothoracic width, 2.10/2.05/2.50–2.55; humeral width, 1.85/1.85/2.20– 2.30; elytral length, 5.75/4.95/6.10–6.20.
</paragraph>
</subSubSection>
<subSubSection id="3EAB65009B1B8316FF7BF957FAEBF962" box="[199,1338,1699,1725]" pageId="18" pageNumber="469" type="etymology">
<paragraph id="760E368B9B1B8316FF7BF957FAEBF962" blockId="18.[151,1437,151,1725]" box="[199,1338,1699,1725]" pageId="18" pageNumber="469">
<emphasis id="44C5EA999B1B8316FF7BF957FE9FF962" bold="true" box="[199,334,1699,1725]" pageId="18" pageNumber="469">Etymology.</emphasis>
The epithet “oaxacanensis” refers to the Mexican state of the type-locality of the species.
</paragraph>
</subSubSection>
</treatment>
</document>
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