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<document id="369588257CE3F651264D4193D3E45A80" ID-DOI="10.5281/zenodo.188339" ID-GBIF-Dataset="877b8a20-6323-4bac-ab68-29b6d0f909fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="188339" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461089523673" checkinUser="plazi" docAuthor="Taylor, Christopher K. &amp; Hunt, Glenn S." docDate="2009" docId="F044879789433D4698E532AFFE74FEA4" docLanguage="en" docName="zt02130p059.pdf" docOrigin="Zootaxa 2130" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Neopantopsalis quasimodo Taylor &amp; Hunt, 2009, n. sp." docType="treatment" docVersion="9" lastPageNumber="49" masterDocId="0C7DFFEF89453D4F98723013FFE1FFE8" masterDocTitle="New genus of Megalopsalidinae (Arachnida: Opiliones: Monoscutidae) from north-eastern Australia" masterLastPageNumber="59" masterPageNumber="41" pageNumber="47" updateTime="1698588251978" updateUser="plazi">
<mods:mods id="A3B9B1C88697CD05C31D33295E55BA43" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="405A7CAF59B1EEF6866A4485E662FC99">
<mods:title id="3EEBCEF3FD14DD19458A3A3A24656E42">New genus of Megalopsalidinae (Arachnida: Opiliones: Monoscutidae) from north-eastern Australia</mods:title>
</mods:titleInfo>
<mods:name id="073F9A4FE7BE2D4F851C2025FCE8066F" type="personal">
<mods:role id="31A0A27A060DC786ECEB57770602887A">
<mods:roleTerm id="739E79B77FCF23278C4EE62F319B20EC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3BE095B1139498CCD908AA4EFE949ECB">Taylor, Christopher K.</mods:namePart>
</mods:name>
<mods:name id="3DBFC10330F5A867C9F15944FC4FBA99" type="personal">
<mods:role id="D886981EB8D3284D8DF40260BD17F47B">
<mods:roleTerm id="1982325F6FC3ADE0272F21348D4C7377">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="134C9277BC2624BAA1362B43666EF7C5">Hunt, Glenn S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="6800FB9A3E9A759BB0D1EA6D9E606292">text</mods:typeOfResource>
<mods:relatedItem id="2F3FD2FC30D552824AABE3E1C47B8DBA" type="host">
<mods:titleInfo id="D42547983C07B9A725D88193F74D2E5E">
<mods:title id="22B39447A55A85472E1AB27FBF1C4554">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7F3452AA44326A72A14E3C33228930D3">
<mods:date id="2DA7FF6BB3391D53F27B06673BEECADE">2009</mods:date>
<mods:detail id="A24682C79265ED9F94473FA30F4D2051" type="volume">
<mods:number id="0F984A867D7612177BF65837C1ED3493">2130</mods:number>
</mods:detail>
<mods:extent id="C9BD44AE8FA80665851777019DFFF647" unit="page">
<mods:start id="A1D2D4B2EA94D2A8DD9139CF7DE4018C">41</mods:start>
<mods:end id="0FA7AEE4BB6DD07548BA495453EB0C9D">59</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="DB286059A72F8541579686EF84AFE3E8">journal article</mods:classification>
<mods:identifier id="FA1A2BA25B527656AB7960DAB354AAE2" type="DOI">10.5281/zenodo.188339</mods:identifier>
<mods:identifier id="822FCCF03E276ED0F7AA2769705AAF3E" type="GBIF-Dataset">877b8a20-6323-4bac-ab68-29b6d0f909fa</mods:identifier>
<mods:identifier id="E27EAA99DEF48C58BDB6146FF4D27C40" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="12E1F10ED3608B7EAF991402B5090454" type="Zenodo-Dep">188339</mods:identifier>
<mods:identifier id="0AD11BA824DF38FC39F344201F10A72F" type="ZooBank">0FE195EE-ADD7-4161-8070-44027D3FC74C</mods:identifier>
</mods:mods>
<treatment id="F044879789433D4698E532AFFE74FEA4" ID-DOI="http://doi.org/10.5281/zenodo.6220693" ID-GBIF-Taxon="119619771" ID-Zenodo-Dep="6220693" LSID="urn:lsid:plazi:treatment:F044879789433D4698E532AFFE74FEA4" httpUri="http://treatment.plazi.org/id/F044879789433D4698E532AFFE74FEA4" lastPageId="9" lastPageNumber="49" pageId="6" pageNumber="47">
<subSubSection id="30F7650A89433D4998E532AFFEB8FD10" pageId="6" pageNumber="47" type="nomenclature">
<paragraph id="7852368189433D4998E532AFFDC4FD3D" blockId="6.[151,549,699,760]" box="[151,549,699,726]" pageId="6" pageNumber="47">
<heading id="231A81ED89433D4998E532AFFDC4FD3D" bold="true" box="[151,549,699,726]" fontSize="11" level="1" pageId="6" pageNumber="47" reason="1">
<emphasis id="4A99EA9389433D4998E532AFFDC4FD3D" bold="true" box="[151,549,699,726]" pageId="6" pageNumber="47">
<taxonomicName id="BFED4D0289433D4998E532AFFE3DFD3E" box="[151,476,700,726]" class="Arachnida" family="Monoscutidae" genus="Neopantopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="4A99EA9389433D4998E532AFFE3DFD3E" bold="true" box="[151,476,700,726]" italics="true" pageId="6" pageNumber="47">Neopantopsalis quasimodo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51AA57E889433D49999132A8FDC4FD3D" box="[483,549,699,725]" pageId="6" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="7852368189433D4998E532CDFEB8FD10" blockId="6.[151,549,699,760]" box="[151,345,734,760]" pageId="6" pageNumber="47">
(
<figureCitation id="E0D62A0489433D4998ED32CDFF0CFD10" box="[159,237,734,760]" captionStart="FIGURE 1" captionStartId="2.[151,260,1795,1819]" captionTargetBox="[219,1365,186,1777]" captionTargetId="figure@2.[188,1398,150,1820]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal view of Megalopsalidinae exemplars, showing division of propeltidium into three areas. A. Neopantopsalis quasimodo n. sp. (QM S 2481). B. Megalopsalis sp. (WAM T 72936). C. Megalopsalis? inconstans (QM S 74252). Scale bars = 1 mm." httpUri="https://zenodo.org/record/188340/files/figure.png" pageId="6" pageNumber="47">Figs. 1</figureCitation>
A, 3–12)
</paragraph>
</subSubSection>
<subSubSection id="30F7650A89433D4998E53338FD2EFA8B" pageId="6" pageNumber="47" type="materials_examined">
<paragraph id="7852368189433D4998E53338FD3DFC7B" blockId="6.[151,1438,811,1998]" pageId="6" pageNumber="47">
<emphasis id="4A99EA9389433D4998E53338FE74FCAD" bold="true" box="[151,405,811,837]" pageId="6" pageNumber="47">Material examined.</emphasis>
<emphasis id="4A99EA9389433D4999EF333FFDACFCAD" box="[413,589,812,837]" italics="true" pageId="6" pageNumber="47">
Male
<typeStatus id="A756882389433D499996333FFDACFCAD" box="[484,589,812,837]" pageId="6" pageNumber="47" type="holotype">holotype</typeStatus>
</emphasis>
. Mt. Finnigan,
<quantity id="BF159B6489433D499B7C333FFC48FCAE" box="[782,937,812,838]" metricMagnitude="2" metricUnit="m" metricValue="9.75" metricValueMax="11.0" metricValueMin="8.5" pageId="6" pageNumber="47" unit="m" value="975.0" valueMax="1100.0" valueMin="850.0">850–1100 m</quantity>
,
<quantity id="BF159B6489433D499BC5333FFBE4FCAE" box="[951,1029,812,838]" metricMagnitude="4" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="47" unit="km" value="37.0">37 km</quantity>
S. of Cooktown, N. Queensland,
<geoCoordinate id="1DD9504689433D4998E53341FF1BFC84" box="[151,250,850,876]" direction="south" orientation="latitude" pageId="6" pageNumber="47" precision="925" value="-15.816667">15°49’S</geoCoordinate>
<geoCoordinate id="1DD9504689433D4999703341FE99FC84" box="[258,376,850,876]" direction="east" orientation="longitude" pageId="6" pageNumber="47" precision="925" value="145.28334">145°17’E</geoCoordinate>
,
<date id="0C53104189433D4999F73341FDBEFC84" box="[389,607,850,876]" pageId="6" pageNumber="47" value="1982-04-19" valueMax="1982-04-22" valueMin="1982-04-19">19–22 April 1982</date>
, rainforest; Monteith, Yeates &amp; Cook (QM S2481, QM
<accessionNumber id="67BEAB6289433D499D773341FA85FC84" box="[1285,1380,850,876]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S39684" pageId="6" pageNumber="47" type="EnaNcbi">S39684</accessionNumber>
, this specimen has two accession numbers; measured).
</paragraph>
<paragraph id="7852368189433D4998B733B3FCBDFB20" blockId="6.[151,1438,811,1998]" pageId="6" pageNumber="47">
<emphasis id="4A99EA9389433D4998B733B3FEAAFC51" box="[197,331,928,953]" italics="true" pageId="6" pageNumber="47">
<typeStatus id="A756882389433D4998B733B3FEA6FC51" box="[197,327,928,953]" pageId="6" pageNumber="47" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="6EEBFD0889433D49992133B3FE4BFC52" box="[339,426,928,954]" pageId="6" pageNumber="47" type="male">1 male</specimenCount>
, as above (QM S2488);
<specimenCount id="6EEBFD0889433D499AA933B3FCD2FC52" box="[731,819,928,954]" pageId="6" pageNumber="47" type="male">1 male</specimenCount>
, Mt. Finnigan summit,
<quantity id="BF159B6489433D499C2B33B3FB51FC52" box="[1113,1200,928,954]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="47" unit="m" value="1100.0">1100m</quantity>
, via Helenville, N. Queensland,
<date id="0C53104189433D49994133D5FDFCFC08" box="[307,541,966,992]" pageId="6" pageNumber="47" value="1985-11-28">28 November 1985</date>
, G. Monteith &amp; D. Cook, pyrethrum knockdown/
<collectingCountry id="00FA761189433D499C1C33D5FB73FC08" box="[1134,1170,966,992]" name="Russia" pageId="6" pageNumber="47">RF</collectingCountry>
(QM S2491);
<specimenCount id="6EEBFD0889433D499D3133D5FA76FC08" box="[1347,1431,966,992]" pageId="6" pageNumber="47" type="male">1 male</specimenCount>
, Mt. Finnigan, ~
<quantity id="BF159B6489433D49992733FEFE0FFBEF" box="[341,494,1005,1031]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.05" metricValueMin="0.95" pageId="6" pageNumber="47" unit="m" value="1000.0" valueMax="1050.0" valueMin="950.0">950–1050 m</quantity>
(QM S2833);
<specimenCount id="6EEBFD0889433D499AD033FEFCE0FBEF" box="[674,769,1005,1031]" pageId="6" pageNumber="47" type="male">6 males</specimenCount>
(including
<specimenCount id="6EEBFD0889433D499BFC33FEFBE6FBEF" box="[910,1031,1005,1031]" pageId="6" pageNumber="47" type="juvenile">1 juvenile</specimenCount>
male), Mt. Finnigan summit, via Helensville,
<date id="0C53104189433D4999423407FDA8FBC6" box="[304,585,1044,1070]" pageId="6" pageNumber="47" value="1985-11-28" valueMax="1985-11-30" valueMin="1985-11-28">28–30 November 1985</date>
, Monteith, Cook &amp; Roberts (QM S3415;
<specimenCount id="6EEBFD0889433D499C363407FB40FBC6" box="[1092,1185,1044,1070]" pageId="6" pageNumber="47" type="male">5 males</specimenCount>
measured);
<specimenCount id="6EEBFD0889433D499D453407FA79FBC6" box="[1335,1432,1044,1070]" pageId="6" pageNumber="47" type="male">2 males</specimenCount>
, ditto,
<quantity id="BF159B6489433D4998AE3429FED8FBBC" box="[220,313,1082,1108]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="6" pageNumber="47" unit="m" value="1050.0">1050 m</quantity>
,
<date id="0C53104189433D4999373429FDD6FBBC" box="[325,567,1082,1108]" pageId="6" pageNumber="47" value="1990-12-03" valueMax="1990-12-05" valueMin="1990-12-03">3–5 December 1990</date>
, Monteith, Thompson, Cook, Sheridan &amp; Roberts (QM
<accessionNumber id="67BEAB6289433D499CA33429FACEFBBC" box="[1233,1327,1082,1108]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S35968" pageId="6" pageNumber="47" type="EnaNcbi">S35968</accessionNumber>
);
<specimenCount id="6EEBFD0889433D499D363429FA76FBBC" box="[1348,1431,1082,1108]" pageId="6" pageNumber="47" type="male">1 male</specimenCount>
, ditto, Monteith, Sheridan, Roberts &amp; Thompson, pyrethrum (QM S74269; measured), 1 minor male, ditto,
<quantity id="BF159B6489433D4998E5349BFEC5FB4A" box="[151,292,1160,1186]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="6" pageNumber="47" unit="m" value="900.0" valueMax="950.0" valueMin="850.0">850–950 m</quantity>
, Monteith, Thompson, Cook, Sheridan &amp; Roberts (QM
<accessionNumber id="67BEAB6289433D499BA4349BFBD9FB4A" box="[982,1080,1160,1186]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S74300" pageId="6" pageNumber="47" type="EnaNcbi">S74300</accessionNumber>
),
<specimenCount id="6EEBFD0889433D499C3C349BFB4CFB4A" box="[1102,1197,1160,1186]" pageId="6" pageNumber="47" type="male">5 males</specimenCount>
(including 2 minor males), ditto,
<quantity id="BF159B6489433D49994A34BDFE70FB20" box="[312,401,1198,1224]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="6" pageNumber="47" unit="m" value="1050.0">1050 m</quantity>
(QM
<accessionNumber id="67BEAB6289433D4999AA34BDFDD4FB20" box="[472,565,1198,1224]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S74301" pageId="6" pageNumber="47" type="EnaNcbi">S74301</accessionNumber>
; minor males measured).
</paragraph>
<paragraph id="7852368189433D4998B734C6FD2EFA8B" blockId="6.[151,1438,811,1998]" pageId="6" pageNumber="47">
<emphasis id="4A99EA9389433D4998B734C6FDC2FB07" bold="true" box="[197,547,1237,1263]" pageId="6" pageNumber="47">Other material examined.</emphasis>
<specimenCount id="6EEBFD0889433D499A5F34C6FD72FB07" box="[557,659,1237,1263]" pageId="6" pageNumber="47" type="male">5 males</specimenCount>
, Mt. Boolbun Sth. (Camp),
<quantity id="BF159B6489433D499B8834C6FBACFB07" box="[1018,1101,1237,1263]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="47" unit="m" value="850.0">850 m</quantity>
, NE Queensland,
<geoCoordinate id="1DD9504689433D499D4734C6FA7CFB07" box="[1333,1437,1237,1263]" direction="south" orientation="latitude" pageId="6" pageNumber="47" precision="925" value="-15.95">15°57’S</geoCoordinate>
<geoCoordinate id="1DD9504689433D4998E534EFFEEDFAFE" box="[151,268,1276,1302]" direction="east" orientation="longitude" pageId="6" pageNumber="47" precision="925" value="145.13333">145°08’E</geoCoordinate>
,
<date id="0C53104189433D49996B34EFFE13FAFE" box="[281,498,1276,1302]" pageId="6" pageNumber="47" value="1995-11-05">5 November 1995</date>
–
<date id="0C53104189433D499A6334EFFD3BFAFE" box="[529,730,1276,1302]" pageId="6" pageNumber="47" value="1996-01-11">11 January 1996</date>
, Monteith, Cook &amp; Roberts, intercept trap (QM
<accessionNumber id="67BEAB6289433D499D5034EFFA61FAFE" box="[1314,1408,1276,1302]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41346" pageId="6" pageNumber="47" type="EnaNcbi">S41346</accessionNumber>
; 1 measured);
<specimenCount id="6EEBFD0889433D4999533531FE9FFAD4" box="[289,382,1314,1340]" pageId="6" pageNumber="47" type="male">2 males</specimenCount>
, Mt. Hartley, SW slope,
<quantity id="BF159B6489433D499AD13531FD10FAD4" box="[675,753,1314,1340]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="47" unit="m" value="750.0">750 m</quantity>
,
<date id="0C53104189433D499A8F3531FC32FAD4" box="[765,979,1314,1340]" pageId="6" pageNumber="47" value="1995-11-08">8 November 1995</date>
–
<date id="0C53104189433D499B833531FB57FAD4" box="[1009,1206,1314,1340]" pageId="6" pageNumber="47" value="1996-01-16">16 January 1996</date>
, Monteith, Cook &amp; Roberts, intercept trap (QM
<accessionNumber id="67BEAB6289433D499991355AFDA0FA8B" box="[483,577,1353,1379]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41376" pageId="6" pageNumber="47" type="EnaNcbi">S41376</accessionNumber>
; measured).
</paragraph>
</subSubSection>
<subSubSection id="30F7650A89433D4998B7357CFEFCFA16" pageId="6" pageNumber="47" type="diagnosis">
<paragraph id="7852368189433D4998B7357CFEFCFA16" blockId="6.[151,1438,811,1998]" pageId="6" pageNumber="47">
<emphasis id="4A99EA9389433D4998B7357CFEA4FA61" bold="true" box="[197,325,1391,1417]" pageId="6" pageNumber="47">Diagnosis.</emphasis>
<taxonomicName id="BFED4D0289433D49993F3563FD6DFA61" box="[333,652,1392,1417]" class="Arachnida" family="Monoscutidae" genus="Neopantopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="quasimodo">
<emphasis id="4A99EA9389433D49993F3563FD6DFA61" box="[333,652,1392,1417]" italics="true" pageId="6" pageNumber="47">Neopantopsalis quasimodo</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="BFED4D0289433D499BE73563FC0EFA61" box="[917,1007,1392,1417]" class="Arachnida" family="Monoscutidae" genus="Neopantopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="psile">
<emphasis id="4A99EA9389433D499BE73563FC0EFA61" box="[917,1007,1392,1417]" italics="true" pageId="6" pageNumber="47">N. psile</emphasis>
</taxonomicName>
and
<taxonomicName id="BFED4D0289433D499C583563FB5DFA61" box="[1066,1212,1392,1417]" class="Arachnida" family="Monoscutidae" genus="Neopantopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="pentheter">
<emphasis id="4A99EA9389433D499C583563FB5DFA61" box="[1066,1212,1392,1417]" italics="true" pageId="6" pageNumber="47">N. pentheter</emphasis>
</taxonomicName>
by the presence of armature on the dorsal prosomal plate and ocularium. It is distinguished from
<taxonomicName id="BFED4D0289433D499CD93584FADCFA58" box="[1195,1341,1431,1456]" class="Arachnida" family="Monoscutidae" genus="Neopantopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="camelus">
<emphasis id="4A99EA9389433D499CD93584FADCFA58" box="[1195,1341,1431,1456]" italics="true" pageId="6" pageNumber="47">N. camelus</emphasis>
</taxonomicName>
and
<taxonomicName id="BFED4D0289433D499D0D3584FEA4FA3E" class="Arachnida" family="Monoscutidae" genus="Neopantopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="thaumatopoios">
<emphasis id="4A99EA9389433D499D0D3584FEA4FA3E" italics="true" pageId="6" pageNumber="47">N. thaumatopoios</emphasis>
</taxonomicName>
by having larger, fewer denticles on the dorsal prosomal plate and lacking pseudoarticulations in femur II.
</paragraph>
</subSubSection>
<subSubSection id="30F7650A89433D4798B73619FD57F8C6" lastPageId="8" lastPageNumber="49" pageId="6" pageNumber="47" type="description">
<paragraph id="7852368189433D4998B73619FE06F8B2" blockId="6.[151,1438,811,1998]" pageId="6" pageNumber="47">
<emphasis id="4A99EA9389433D4998B73619FEBCF9CC" bold="true" box="[197,349,1546,1572]" pageId="6" pageNumber="47">Description.</emphasis>
MAJOR MALE (N = 10). Prosoma length 2.76 (2.43–3.20), width 2.82 (2.45–3.20). Dorsal prosomal plate uniformly orange-brown, with bases of denticles silver. Anterior propeltidial area unarmed. Median propeltidial area denticulate, with pair of large denticles at apex of lateral mounds and further denticles anterior and lateral to mounds. Posterior propeltidial area mostly unarmed except for few small denticles lateral to lateral humps. Ocularium with row of few large denticles on each side. Lateral shelves mottled purple-brown, ozopore lobes white. Metapeltidium sclerotised medially to form distinctly round sclerotised area with dorsal prosomal plate. Dorsum of opisthosoma purple-brown, with large white spots, tan transverse stripes at segmental boundaries. Anal operculum white. Venter of opisthosoma tan, with some purple-brown areas laterally.
</paragraph>
<paragraph id="7852368189433D4998B73774FCA5F826" blockId="6.[151,1438,811,1998]" pageId="6" pageNumber="47">
<emphasis id="4A99EA9389433D4998B73774FEA7F868" box="[197,326,1895,1920]" italics="true" pageId="6" pageNumber="47">Chelicerae</emphasis>
. Segment I 10.74 (8.50–12.19), segment II 12.20 (9.69–13.74). Very long and slender; purplebrown, with both segments white distally. Both segments evenly denticulate; segment I with larger denticles dorsally than ventrally. Cheliceral fingers sinuous (
<figureCitation id="E0D62A0489433D499A9F37A7FCD4F826" box="[749,821,1972,1998]" captionStart="FIGURES 3 – 10" captionStartId="7.[151,272,1356,1380]" captionTargetBox="[159,1361,202,1317]" captionTargetId="figure@7.[151,1436,197,1332]" captionTargetPageId="7" captionText="FIGURES 3 – 10. Neopantopsalis quasimodo n. sp., male. 3. Lateral view of body showing chelicerae and pedipalps (QM S 2481, holotype). 4. Lateral view of body with appendages removed (QM S 2481, holotype). 5. Cheliceral fingers, frontal view (QM S 74301). 6. Femur to tibia of right leg I, dorsal view (QM S 2481, holotype). 7. Total penis, right lateral view (QM S 2488). 8. Glans, right lateral view (QM S 2488). 9. Glans of same male, ventral view (QM S 2488). 10. Glans of different male, ventral view (QM S 74301). Scale bars for Figs. 3 – 6 = 1 mm; Fig. 7 = 0.5 mm; Figs. 8 – 10 = 0.1 mm." httpUri="https://zenodo.org/record/188342/files/figure.png" pageId="6" pageNumber="47">Fig. 5</figureCitation>
).
</paragraph>
<caption id="2C92660989423D4898E5355FFA69FA07" httpUri="https://zenodo.org/record/188342/files/figure.png" pageId="7" pageNumber="48" targetBox="[159,1361,202,1317]" targetPageId="7">
<paragraph id="7852368189423D4898E5355FFA69FA07" blockId="7.[151,1436,1356,1519]" pageId="7" pageNumber="48">
<emphasis id="4A99EA9389423D4898E5355FFEB5FA8C" bold="true" box="[151,340,1356,1380]" pageId="7" pageNumber="48">FIGURES 3–10.</emphasis>
<taxonomicName id="BFED4D0289423D48992E355EFD67FA8C" box="[348,646,1357,1380]" class="Arachnida" family="Monoscutidae" genus="Neopantopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="4A99EA9389423D48992E355EFD67FA8C" box="[348,646,1357,1380]" italics="true" pageId="7" pageNumber="48">Neopantopsalis quasimodo</emphasis>
</taxonomicName>
<emphasis id="4A99EA9389423D489AFF355FFD2DFA8C" bold="true" box="[653,716,1356,1380]" pageId="7" pageNumber="48">
<taxonomicNameLabel id="51AA57E889423D489AFF355FFD2DFA8C" box="[653,716,1356,1380]" pageId="7" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male. 3. Lateral view of body showing chelicerae and pedipalps (QM S2481, holotype). 4. Lateral view of body with appendages removed (QM S2481, holotype). 5. Cheliceral fingers, frontal view (QM
<accessionNumber id="67BEAB6289423D48992A3581FE4FFA41" box="[344,430,1426,1449]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S74301" pageId="7" pageNumber="48" type="EnaNcbi">S74301</accessionNumber>
). 6. Femur to tibia of right leg I, dorsal view (QM S2481, holotype). 7. Total penis, right lateral view (QM S2488). 8. Glans, right lateral view (QM S2488). 9. Glans of same male, ventral view (QM S2488). 10. Glans of different male, ventral view (QM
<accessionNumber id="67BEAB6289423D489A6F35CBFD93FA07" box="[541,626,1496,1519]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S74301" pageId="7" pageNumber="48" type="EnaNcbi">S74301</accessionNumber>
). Scale bars for Figs. 3–6 = 1 mm; Fig. 7 = 0.5 mm; Figs. 8–10 = 0.1 mm.
</paragraph>
</caption>
<paragraph id="7852368189423D4898B7360EFD54F96C" blockId="7.[151,1438,1565,2016]" pageId="7" pageNumber="48">
<emphasis id="4A99EA9389423D4898B7360EFEA4F9DE" box="[197,325,1565,1590]" italics="true" pageId="7" pageNumber="48">Pedipalps</emphasis>
. Femur 1.56 (1.33–1.77), patella 0.63 (0.55–0.70), tibia 0.92 (0.81–1.00), tarsus 1.99 (1.76–2.17). Unarmed; white with purple-brown bands. No apophyses or hypersetose areas. No plumose setae (
<figureCitation id="E0D62A0489423D4898ED3679FF14F96C" box="[159,245,1642,1668]" captionStart="FIGURES 11 – 12" captionStartId="8.[151,272,973,997]" captionTargetBox="[175,1408,527,936]" captionTargetId="figure@8.[151,1436,506,948]" captionTargetPageId="8" captionText="FIGURES 11 – 12. Neopantopsalis quasimodo n. sp., male. 11. Pedipalp, patella / tibia junction, prolateral view (QM S 74269). 12. Close-up of glans to show sensory pore morphology (QM S 74300)." httpUri="https://zenodo.org/record/188343/files/figure.png" pageId="7" pageNumber="48">Fig. 11</figureCitation>
). Tarsal claw with many ventral teeth.
</paragraph>
<paragraph id="7852368189423D4798B73682FC94FECE" blockId="7.[151,1438,1565,2016]" lastBlockId="8.[151,1437,152,448]" lastPageId="8" lastPageNumber="49" pageId="7" pageNumber="48">
<emphasis id="4A99EA9389423D4898B73682FF1FF942" box="[197,254,1681,1706]" italics="true" pageId="7" pageNumber="48">Legs</emphasis>
. Femora 7.02 (6.15–7.73), 11.86 (10.54–13.08), 5.96 (5.31–6.22), 7.95 (7.12–8.72); patellae 1.25 (1.17–1.35), 1.37 (1.23–1.45), 1.28 (1.18–1.39), 1.37 (1.23–1.48); tibiae 6.27 (5.63–6.74), 12.58 (10.92–13.83), 5.82 (5.06–6.22), 7.99 (6.93–8.80). All segments light tan, with purple-brown bands except tan telotarsi. Coxae I–III with long spines on dorsal distal margins. Trochanter I with three long spines on dorsal prolateral margin, other trochanters unarmed. Femur I with proventral longitudinal row of long thin spines, longest at proximal and distal ends, dwindling to small denticles medially, turning along proventral margin at distal end (
<figureCitation id="E0D62A0489423D4899E9376AFE06F87B" box="[411,487,1913,1939]" captionStart="FIGURES 3 – 10" captionStartId="7.[151,272,1356,1380]" captionTargetBox="[159,1361,202,1317]" captionTargetId="figure@7.[151,1436,197,1332]" captionTargetPageId="7" captionText="FIGURES 3 – 10. Neopantopsalis quasimodo n. sp., male. 3. Lateral view of body showing chelicerae and pedipalps (QM S 2481, holotype). 4. Lateral view of body with appendages removed (QM S 2481, holotype). 5. Cheliceral fingers, frontal view (QM S 74301). 6. Femur to tibia of right leg I, dorsal view (QM S 2481, holotype). 7. Total penis, right lateral view (QM S 2488). 8. Glans, right lateral view (QM S 2488). 9. Glans of same male, ventral view (QM S 2488). 10. Glans of different male, ventral view (QM S 74301). Scale bars for Figs. 3 – 6 = 1 mm; Fig. 7 = 0.5 mm; Figs. 8 – 10 = 0.1 mm." httpUri="https://zenodo.org/record/188342/files/figure.png" pageId="7" pageNumber="48">Fig. 6</figureCitation>
). Dorsal longitudinal row of stouter large denticles, dwindling distally. Small scattered denticles on remainder of femur I. Patella I and tibia I with continuations of both longitudinal rows. Proventral spines long on patella I, turning along distal proventral margin; dwindling distally on tibia, ending ~1/3 of length from proximal end. Dorsal spines short on patella I, reduced to very few at proximal end of tibia. Remaining legs with small scattered denticles on femora, remaining segments unarmed except for distally-directed spines on dorsodistal edges of patellae. Tibia II with eight to eleven pseudosegments; tibia IV with three pseudosegments; femur II without pseudosegments.
</paragraph>
<paragraph id="78523681894D3D4798B73121FE93FE28" blockId="8.[151,1437,152,448]" pageId="8" pageNumber="49">
<emphasis id="4A99EA93894D3D4798B73121FEE7FEA3" box="[197,262,306,331]" italics="true" pageId="8" pageNumber="49">Penis</emphasis>
(
<figureCitation id="E0D62A04894D3D4799653121FE79FEA4" box="[279,408,306,332]" captionStart="FIGURES 3 – 10" captionStartId="7.[151,272,1356,1380]" captionTargetBox="[159,1361,202,1317]" captionTargetId="figure@7.[151,1436,197,1332]" captionTargetPageId="7" captionText="FIGURES 3 – 10. Neopantopsalis quasimodo n. sp., male. 3. Lateral view of body showing chelicerae and pedipalps (QM S 2481, holotype). 4. Lateral view of body with appendages removed (QM S 2481, holotype). 5. Cheliceral fingers, frontal view (QM S 74301). 6. Femur to tibia of right leg I, dorsal view (QM S 2481, holotype). 7. Total penis, right lateral view (QM S 2488). 8. Glans, right lateral view (QM S 2488). 9. Glans of same male, ventral view (QM S 2488). 10. Glans of different male, ventral view (QM S 74301). Scale bars for Figs. 3 – 6 = 1 mm; Fig. 7 = 0.5 mm; Figs. 8 – 10 = 0.1 mm." httpUri="https://zenodo.org/record/188342/files/figure.png" pageId="8" pageNumber="49">Figs. 7–10</figureCitation>
,
<figureCitation id="E0D62A04894D3D4799D73121FE27FEA4" box="[421,454,306,332]" captionStart="FIGURES 11 – 12" captionStartId="8.[151,272,973,997]" captionTargetBox="[175,1408,527,936]" captionTargetId="figure@8.[151,1436,506,948]" captionTargetPageId="8" captionText="FIGURES 11 – 12. Neopantopsalis quasimodo n. sp., male. 11. Pedipalp, patella / tibia junction, prolateral view (QM S 74269). 12. Close-up of glans to show sensory pore morphology (QM S 74300)." httpUri="https://zenodo.org/record/188343/files/figure.png" pageId="8" pageNumber="49">12</figureCitation>
). Bristle groups on left side of junction between shaft and glans absent or reduced; anterior right group also greatly reduced. Glans at low angle to shaft; long and dorsoventrally flattened; rectangular in profile. Pores on glans shallowly recessed, not raised on papillae (
<figureCitation id="E0D62A04894D3D479C193193FB30FE72" box="[1131,1233,384,410]" captionStart="FIGURES 11 – 12" captionStartId="8.[151,272,973,997]" captionTargetBox="[175,1408,527,936]" captionTargetId="figure@8.[151,1436,506,948]" captionTargetPageId="8" captionText="FIGURES 11 – 12. Neopantopsalis quasimodo n. sp., male. 11. Pedipalp, patella / tibia junction, prolateral view (QM S 74269). 12. Close-up of glans to show sensory pore morphology (QM S 74300)." httpUri="https://zenodo.org/record/188343/files/figure.png" pageId="8" pageNumber="49">Figs. 12</figureCitation>
). Stylus directed dorsad from glans.
</paragraph>
<caption id="2C926609894D3D4798E533DEFC16FBEF" httpUri="https://zenodo.org/record/188343/files/figure.png" pageId="8" pageNumber="49" targetBox="[175,1408,527,936]" targetPageId="8">
<paragraph id="78523681894D3D4798E533DEFC16FBEF" blockId="8.[151,1437,973,1031]" pageId="8" pageNumber="49">
<emphasis id="4A99EA93894D3D4798E533DEFE80FC0D" bold="true" box="[151,353,973,997]" pageId="8" pageNumber="49">FIGURES 11–12.</emphasis>
<taxonomicName id="BFED4D02894D3D47991B33DEFD75FC0C" box="[361,660,973,996]" class="Arachnida" family="Monoscutidae" genus="Neopantopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="4A99EA93894D3D47991B33DEFD75FC0C" box="[361,660,973,996]" italics="true" pageId="8" pageNumber="49">Neopantopsalis quasimodo</emphasis>
</taxonomicName>
<emphasis id="4A99EA93894D3D479AEE33DEFD3BFC0D" bold="true" box="[668,730,973,997]" pageId="8" pageNumber="49">
<taxonomicNameLabel id="51AA57E8894D3D479AEE33DEFD3BFC0D" box="[668,730,973,997]" pageId="8" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male. 11. Pedipalp, patella/tibia junction, prolateral view (QM S74269). 12. Close-up of glans to show sensory pore morphology (QM
<accessionNumber id="67BEAB62894D3D479BE733E3FC0AFBEF" box="[917,1003,1008,1031]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S74300" pageId="8" pageNumber="49" type="EnaNcbi">S74300</accessionNumber>
).
</paragraph>
</caption>
<paragraph id="78523681894D3D4798B73426FC4CFADF" blockId="8.[151,1437,1077,2031]" pageId="8" pageNumber="49">
MINOR MALE (N = 3). Prosoma length 1.37 (1.14–1.60), width 2.13 (2.03–2.18). Considerably smaller than major male, with lateral mounds not as raised and no postocularial hump. Armature on dorsal prosomal plate not as prominent.
<emphasis id="4A99EA93894D3D4799C83490FDDEFB74" box="[442,575,1155,1180]" italics="true" pageId="8" pageNumber="49">Chelicerae</emphasis>
. Segment I 4.23 (2.88–5.82), segment II 5.36 (3.96–6.89).
<emphasis id="4A99EA93894D3D479D693490FA74FB74" box="[1307,1429,1155,1180]" italics="true" pageId="8" pageNumber="49">Pedipalps</emphasis>
. Femur 1.29 (1.22–1.34), patella 0.55 (0.52–0.58), tibia 0.77 (0.72–0.81), tarsus 1.66 (1.61–1.75).
<emphasis id="4A99EA93894D3D479D2934BAFA74FB2A" box="[1371,1429,1193,1218]" italics="true" pageId="8" pageNumber="49">Legs</emphasis>
. Femora 5.90 (5.69–6.14), 10.19 (9.96–10.38), 5.23 (4.95–5.50), 7.03 (6.69–7.31); patellae 1.16 (1.14–1.21), 1.25 (1.21–1.28), 1.15 (1.06–1.23), 1.20 (1.16–1.28); tibiae 5.78 (5.44–6.15), 10.87 (10.46–11.46), 5.36 (4.97–5.75), 6.07 (6.70–7.35). Spine row on leg I not as developed.
</paragraph>
<paragraph id="78523681894D3D4798B73557FDD1F9F7" blockId="8.[151,1437,1077,2031]" pageId="8" pageNumber="49">
<emphasis id="4A99EA93894D3D4798B73557FEA2FAB6" bold="true" box="[197,323,1348,1374]" pageId="8" pageNumber="49">Variation.</emphasis>
Some specimens differ from the
<typeStatus id="A7568823894D3D479AA93557FCA5FAB6" box="[731,836,1348,1374]" pageId="8" pageNumber="49" type="holotype">holotype</typeStatus>
in that the mesopeltidium is not sclerotised. This probably indicates a younger age for these specimens, with less cuticular hardening. Cuticular hardening over time after the last moult can result in significant differences in appearance of individuals of Eupnoi at different ages (
<bibRefCitation id="1C7C4B70894D3D4798AF35ABFE95FA3A" author="Shultz" box="[221,372,1464,1490]" pageId="8" pageNumber="59" refString="Shultz, J. W. (2008) Leiobunum nigripes is a junior synonym of Leiobunum verrucosum (Opiliones, Sclerosomatidae). Journal of Arachnology, 36 (1), 184 - 186." type="journal article" year="2008">Shultz 2008</bibRefCitation>
) Some individual variation occurs in the amount of denticulation present on the dorsal prosomal plate. In particular, two specimens of QM S3415 show more denticles on the lateral humps, as well as denticles on the posterior hump.
</paragraph>
<paragraph id="78523681894D3D4798B7363FFD57F8C6" blockId="8.[151,1437,1077,2031]" pageId="8" pageNumber="49">
The most significant variation from a taxonomic point of view is that relating to the length of the left bristle groups on the penis. In some specimens (including the
<typeStatus id="A7568823894D3D479BF83641FC17F984" box="[906,1014,1618,1644]" pageId="8" pageNumber="49" type="holotype">holotype</typeStatus>
) the left bristle groups are entirely absent (
<figureCitation id="E0D62A04894D3D47988A366AFEA4F97B" box="[248,325,1657,1683]" captionStart="FIGURES 3 – 10" captionStartId="7.[151,272,1356,1380]" captionTargetBox="[159,1361,202,1317]" captionTargetId="figure@7.[151,1436,197,1332]" captionTargetPageId="7" captionText="FIGURES 3 – 10. Neopantopsalis quasimodo n. sp., male. 3. Lateral view of body showing chelicerae and pedipalps (QM S 2481, holotype). 4. Lateral view of body with appendages removed (QM S 2481, holotype). 5. Cheliceral fingers, frontal view (QM S 74301). 6. Femur to tibia of right leg I, dorsal view (QM S 2481, holotype). 7. Total penis, right lateral view (QM S 2488). 8. Glans, right lateral view (QM S 2488). 9. Glans of same male, ventral view (QM S 2488). 10. Glans of different male, ventral view (QM S 74301). Scale bars for Figs. 3 – 6 = 1 mm; Fig. 7 = 0.5 mm; Figs. 8 – 10 = 0.1 mm." httpUri="https://zenodo.org/record/188342/files/figure.png" pageId="8" pageNumber="49">Fig. 9</figureCitation>
), while in others small but distinct bristle groups are still present (
<figureCitation id="E0D62A04894D3D479CFA366AFB06F97B" box="[1160,1255,1657,1683]" captionStart="FIGURES 3 – 10" captionStartId="7.[151,272,1356,1380]" captionTargetBox="[159,1361,202,1317]" captionTargetId="figure@7.[151,1436,197,1332]" captionTargetPageId="7" captionText="FIGURES 3 – 10. Neopantopsalis quasimodo n. sp., male. 3. Lateral view of body showing chelicerae and pedipalps (QM S 2481, holotype). 4. Lateral view of body with appendages removed (QM S 2481, holotype). 5. Cheliceral fingers, frontal view (QM S 74301). 6. Femur to tibia of right leg I, dorsal view (QM S 2481, holotype). 7. Total penis, right lateral view (QM S 2488). 8. Glans, right lateral view (QM S 2488). 9. Glans of same male, ventral view (QM S 2488). 10. Glans of different male, ventral view (QM S 74301). Scale bars for Figs. 3 – 6 = 1 mm; Fig. 7 = 0.5 mm; Figs. 8 – 10 = 0.1 mm." httpUri="https://zenodo.org/record/188342/files/figure.png" pageId="8" pageNumber="49">Fig. 10</figureCitation>
). However, no external characters can be identified distinguishing specimens differing in this way. Genitalic polymorphism has been described only occasionally in arachnids, but cannot be ruled out
<emphasis id="4A99EA93894D3D479C6736D4FB93F908" box="[1045,1138,1735,1760]" italics="true" pageId="8" pageNumber="49">a priori</emphasis>
(Jocqué 2002). Until the
<taxonomicName id="BFED4D02894D3D4798E536FDFE37F8EF" box="[151,470,1774,1799]" class="Arachnida" family="Monoscutidae" genus="Neopantopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="quasimodo">
<emphasis id="4A99EA93894D3D4798E536FDFE37F8EF" box="[151,470,1774,1799]" italics="true" pageId="8" pageNumber="49">Neopantopsalis quasimodo</emphasis>
</taxonomicName>
species complex can be properly analysed (see comments below), I provisionally recognise only one species at the
<typeStatus id="A7568823894D3D479A6D3707FDB0F8C6" box="[543,593,1812,1838]" pageId="8" pageNumber="49">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="30F7650A894D3D4798B73729FCA0F893" pageId="8" pageNumber="49" type="etymology">
<paragraph id="78523681894D3D4798B73729FCA0F893" blockId="8.[151,1437,1077,2031]" pageId="8" pageNumber="49">
<emphasis id="4A99EA93894D3D4798B73729FEB5F8BC" bold="true" box="[197,340,1850,1876]" pageId="8" pageNumber="49">Etymology.</emphasis>
The name of this species was given by G. Hunt, and doubtlessly refers to the hunchbacked character from Victor Hugo’s novel
<emphasis id="4A99EA93894D3D479A4C3772FCD8F892" box="[574,825,1889,1914]" italics="true" pageId="8" pageNumber="49">Notre-Dame de Paris</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="30F7650A894D3D4698B7379BFE74FEA4" lastPageId="9" lastPageNumber="50" pageId="8" pageNumber="49" type="discussion">
<paragraph id="78523681894D3D4698B7379BFE74FEA4" blockId="8.[151,1437,1077,2031]" lastBlockId="9.[151,1436,152,332]" lastPageId="9" lastPageNumber="50" pageId="8" pageNumber="49">
<emphasis id="4A99EA93894D3D4798B7379BFEB4F84A" bold="true" box="[197,341,1928,1954]" pageId="8" pageNumber="49">Comments.</emphasis>
Specimens comparable to
<taxonomicName id="BFED4D02894D3D479AE8379BFCA0F849" box="[666,833,1928,1953]" class="Arachnida" family="Monoscutidae" genus="Neopantopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="quasimodo">
<emphasis id="4A99EA93894D3D479AE8379BFCA0F849" box="[666,833,1928,1953]" italics="true" pageId="8" pageNumber="49">N. quasimodo</emphasis>
</taxonomicName>
from many localities in north-eastern Queensland are held in the collection at QM. However, a wide range of variation can be seen within and between localities in size, development of the bristle groups on the penis and extent of the proventral spine row on leg I, and it seems likely that a complex of closely related species is involved rather than a single taxon. Unfortunately, establishing species boundaries within the group has proven difficult, and will require no small amount of study. Matters are complicated further by the uncertain nature of
<taxonomicName id="BFED4D02894C3D469BD030F6FBBFFF16" box="[930,1118,229,254]" class="Arachnida" family="Monoscutidae" genus="Neopantopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="continentalis">
<emphasis id="4A99EA93894C3D469BD030F6FBBFFF16" box="[930,1118,229,254]" italics="true" pageId="9" pageNumber="50">N. continentalis</emphasis>
</taxonomicName>
, as described below. Until such a study can be conducted, I recommend that specimens similar to
<taxonomicName id="BFED4D02894C3D469BA9311FFB9FFECD" box="[987,1150,268,293]" class="Arachnida" family="Monoscutidae" genus="Neopantopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="quasimodo">
<emphasis id="4A99EA93894C3D469BA9311FFB9FFECD" box="[987,1150,268,293]" italics="true" pageId="9" pageNumber="50">N. quasimodo</emphasis>
</taxonomicName>
be referred to as the ‘
<taxonomicName id="BFED4D02894C3D469DF3311FFEF9FEA3" class="Arachnida" family="Monoscutidae" genus="Neopantopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="quasimodo">
<emphasis id="4A99EA93894C3D469DF3311FFEF9FEA3" italics="true" pageId="9" pageNumber="50">N. quasimodo</emphasis>
</taxonomicName>
complex’.
</paragraph>
</subSubSection>
</treatment>
</document>
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