treatments-xml/data/DD/A5/A2/DDA5A2567BF8A255FDA7DBB039B41AD5.xml
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<document id="7C5AD1CF2AF28297929253DEDB1FEFDC" ENCODING="UTF-8" ID-GBIF-Dataset="7b74dd89-3db5-41af-8499-cf216a4ab73d" ModsDocID="z01047p001" checkinTime="1247056604684" checkinUser="thomas" docAuthor="Heok Hee Ng &amp; David R. Edds" docDate="2005" docId="DDA5A2567BF8A255FDA7DBB039B41AD5" docLanguage="en" docName="2005_Ng_Edds_gg1.xml" docOrigin="Zootaxa 1047" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:E29D456B-DC4B-40EA-A11F-0A3F53380A68" docTitle="Pseudecheneis serracula Ng &amp; Edds, 2005, sp. nov." docType="treatment" docVersion="12" lastPageNumber="12" masterDocId="997C1BE742F8650C8FDD49DE36FCD2A1" masterDocTitle="Two new species of Pseudecheneis, rheophilic catfishes (Teleostei: Sisoridae) from Nepal." masterLastPageNumber="19" masterPageNumber="1" pageNumber="6" updateTime="1701314207313" updateUser="plazi">
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<mods:title id="3317A0E1769501E36E36C8398C43288F">Two new species of Pseudecheneis, rheophilic catfishes (Teleostei: Sisoridae) from Nepal.</mods:title>
</mods:titleInfo>
<mods:name id="E812BC42C6241CE426E31F9E2147DCB6" type="personal">
<mods:role id="4ACA51A2DE0426DBA84CCF5F0E981599">
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<mods:namePart id="4467919EC156E90D8C2ED95470E34508">Heok Hee Ng</mods:namePart>
</mods:name>
<mods:name id="CA05B57AC4051931F54DA702BE7B67C2" type="personal">
<mods:role id="D9B4EDEBF679AD025F9BF4F2BAB1445D">
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<mods:namePart id="AEA79A8CEC171D83AE60D06385C24C79">David R. Edds</mods:namePart>
</mods:name>
<mods:typeOfResource id="92D19B71580ED4BD7B4FCC92814E45FC">text</mods:typeOfResource>
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<mods:title id="AB859F5053AEB6A91EC0FDA93004F6B9">Zootaxa</mods:title>
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<mods:part id="2B3B7897CD381EB6686009DEFC061534">
<mods:date id="29F506AF7F139F0A53C6590906BF5B5D">2005</mods:date>
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<mods:number id="7FE16DC943FD8C5D1FA1F2ADEF23E036">1047</mods:number>
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<mods:classification id="10B067BEFED8485AF9F5BE355F4D8A42">journal article</mods:classification>
<mods:identifier id="75D094DEE42CE1B1FD344B42CEDA6E1D" type="Plazi-Custom">z01047p001</mods:identifier>
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<treatment id="DDA5A2567BF8A255FDA7DBB039B41AD5" ID-DOI="http://doi.org/10.5281/zenodo.6265982" ID-GBIF-Taxon="100124223" ID-Zenodo-Dep="6265982" LSID="urn:lsid:plazi:treatment:DDA5A2567BF8A255FDA7DBB039B41AD5" httpUri="http://treatment.plazi.org/id/DDA5A2567BF8A255FDA7DBB039B41AD5" lastPageNumber="12" pageNumber="6">
<subSubSection id="CA17504C38D06FE44453B9CDF2BD835C" type="nomenclature">
<paragraph id="4183AC4E74C1CCC87055CBCB142D75E5" pageNumber="6">
<taxonomicName id="66191CD1E5488A69D7803098BD758C71" ID-CoL="78CCS" LSID-ZBK="urn:lsid:zoobank.org:act:175A3E4F-DFFE-405A-A858-8920376B267D" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudecheneis serracula Ng &amp; Edds 2005" order="Siluriformes" pageNumber="6" phylum="Chordata" rank="species" species="serracula">Pseudecheneis serracula</taxonomicName>
<taxonomicNameLabel id="8BB6010C5756D72E0AB3AD6985EF1991" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="81DFEC614209DB15DC70F503C3536CFD" type="description">
<paragraph id="9727D3CAC511E28669479D8F8C46AAF2" pageNumber="6">(Fig. 3)</paragraph>
</subSubSection>
<subSubSection id="76EFB65043B4307363316DF2C8166F4D" type="reference_group">
<paragraph id="429289830D79D0BB20CAEBA0DC94B570" pageNumber="6">
<taxonomicName id="86F072F2C57803C7316F51FFCB14BDE6" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="6" phylum="Chordata" rank="species" species="sulcatus">Pseudecheneis sulcatus</taxonomicName>
(non MClelland, 1842) Edds, 1986a: 6; 1986b: 18; 1987: 11; 1993: 59; Shrestha, 1981: 197, Fig. 91 (in part); 1994: 62, Fig. 97 (in part).
</paragraph>
</subSubSection>
<subSubSection id="EF6297AF79491C7B77445C48CD94EAD1" type="nomenclature">
<paragraph id="D70C0ACD9899E95D081A8AEBC4FECC13" pageNumber="6">
Type material.
<materialsCitation id="AE67C708C503E710FA316E8774D22C90" ID-GBIF-Occurrence="923897731" country="Nepal" stateProvince="Mugu / Bajura">
<typeStatus id="0690FB7DBB5ACD9DDFDBFF43B489F97A">Holotype</typeStatus>
:
<collectionCode id="06017C6A420F3A7D1AD1AD9CB7F4FAF1">KU</collectionCode>
29554, 153.2 mm SL;
<collectingCountry id="AB930B8B8751FF557F87824AABC342A6">Nepal</collectingCountry>
:
<collectingRegion id="E2C2129680418168AE6EEA6BE404C14B">Mugu/Bajura</collectingRegion>
, Jhugala, Karnali River, purchased at Jhugala,
<geoCoordinate id="E707F1795383092A1080F5855172A456" direction="north" orientation="latitude" value="29.521666">29°31'18.0&quot;N</geoCoordinate>
<geoCoordinate id="723B8CF4BD600D8BBA0298901737C79F" direction="east" orientation="longitude" value="81.78001">81°46'48.0&quot;E</geoCoordinate>
; D. Edds,
<date id="6F4DB998951D5A44B28EC5AD5F7E6161" day="25" month="09" value="1996-09-25" year="1996">25 September 1996</date>
.
</materialsCitation>
</paragraph>
<paragraph id="1998A3EE3E6902085636CD77099AD448" pageNumber="6">
<materialsCitation id="F39695D8C8A2AB63F39D1AE5A4DCE9DD" ID-GBIF-Occurrence="923897733" country="Nepal" stateProvince="Chitawan">
<typeStatus id="6120DB8745517AA11625F17A93786122">Paratypes</typeStatus>
:
<collectionCode id="9A465A3B667268EC4E826FD358664DF4" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]">BMNH</collectionCode>
1985.9.16.50-51 (2), 48.0-48.2 mm SL;
<collectingCountry id="045E7FC7E452A3E2DCF2BAB1C4DBBA0B">Nepal</collectingCountry>
: Narayani River, Chitawan National Park; Edinburgh University,
<date id="6037CC1859594408EAF13EE4F50053AA" month="11" value="1984-11" year="1984">November 1984</date>
-
<date id="4D0D2FFF49F2964DFF0FBBA7DF225725" month="02" value="1985-02" year="1985">February 1985</date>
.
</materialsCitation>
<materialsCitation id="C772EE0CD5E6A85A232BA43F228FBCF0" ID-GBIF-Occurrence="923897748" country="Nepal" stateProvince="Kanchanpur">
<collectionCode id="F107965AE965912E2F2D7D393C06C863">KU</collectionCode>
28669 (5), 41.5-56.5 mm SL;
<collectingCountry id="4697B69CC18AEDEE1692FA2360B81D35">Nepal</collectingCountry>
:
<collectingRegion id="0B81226C6B1A28F48A84523462F9EAB4">Kanchanpur</collectingRegion>
, Brahamadev, Mahakali River at Brahamadev,
<geoCoordinate id="E32134ABFD27CD63CA1CCDF67E005DE1" direction="north" orientation="latitude" value="29.081696">29°4'54.1&quot;N</geoCoordinate>
<geoCoordinate id="DC4DCD996F3E59E36EA011C5E3E0A94A" direction="east" orientation="longitude" value="80.14169">80°8'30.1&quot;E</geoCoordinate>
; D. Edds,
<date id="B2A1369F259F5E2F012F811BD5BD5040" day="24" month="02" value="1996-02-24" year="1996">24 February 1996</date>
.
</materialsCitation>
<materialsCitation id="EF4130F441262843AAD849D2E629B8C3" ID-GBIF-Occurrence="923897732" country="Nepal" stateProvince="Gulmi / Syangja">
<collectionCode id="08A5FA4640715E8CB413A31E8F9598AA">KU</collectionCode>
29038 (1), 58.0 mm SL;
<collectingCountry id="00812FC0F051514A54110768BD1651AF">Nepal</collectingCountry>
:
<collectingRegion id="C96EA6C9526EA34FD49707510E7CE639">Gulmi/Syangja</collectingRegion>
, Kali Gandaki River at Ridi Bazar;
<geoCoordinate id="6A40969DBDE810B514AD773FB495F79E" direction="north" orientation="latitude" value="27.935">27°56'6.0&quot;N</geoCoordinate>
<geoCoordinate id="42AABF477ABFB67240C2067E1EC73626" direction="east" orientation="longitude" value="83.441696">83°26'30.1&quot;E</geoCoordinate>
; D. Edds,
<date id="6F152282421E220D7078C342862B7263" day="21" month="05" value="1996-05-21" year="1996">21 May 1996</date>
.
</materialsCitation>
<materialsCitation id="41B6BE731414545E62153D0B98BE9FB4" ID-GBIF-Occurrence="923897741" country="Nepal" stateProvince="Tanahun">
<collectionCode id="E68D3CA1A52836315BF009BF9C3318C0">KU</collectionCode>
35545 (2), 48.0-95.3 mm SL;
<collectingCountry id="7891A9E6347B3A192B5135018C31C110">Nepal</collectingCountry>
:
<collectingRegion id="315BFB135FF69316008A6557C960D90F">Tanahun</collectingRegion>
, Khairenitar, Seti River at Khairenitar,
<geoCoordinate id="DC4021DE8A1B8160736228B3A16C363E" direction="north" orientation="latitude" value="28.033333">28°2'0.0&quot;N</geoCoordinate>
<geoCoordinate id="AE909EA6B5C13907C9B60A9ABD77B4B4" direction="east" orientation="longitude" value="84.066666">84°4'0.0&quot;E</geoCoordinate>
; D. Edds,
<date id="564D3E8FB86C89B94E0489DF3A0BCDE7" day="15" month="06" value="1996-06-15" year="1996">15 June 1996</date>
.
</materialsCitation>
</paragraph>
<paragraph id="C453C6CA7CE991DF6208BA0C6E9E6F75" pageNumber="7">
<materialsCitation id="EF9C4655F9E63B19BA2C96199F1A8FCB" ID-GBIF-Occurrence="923897747" country="Nepal" stateProvince="Syangja">
<collectionCode id="7D2034A180F374E01D9F911725ACD432" collectionName="OSUS">OSUS</collectionCode>
15703 (4), 31.6-59.3 mm SL;
<collectingCountry id="643404E4DE2FABBDDFE4DD5CFC2E5170">Nepal</collectingCountry>
:
<collectingRegion id="AE061BB1C60DCC00C08F42A37A605D46">Syangja</collectingRegion>
, Kali Gandaki River at Nimaa; D. Edds,
<date id="0EA145A36AFD9B02F595B15B75ED7B0C" day="04" month="05" value="1985-05-04" year="1985">4 May 1985</date>
.
</materialsCitation>
<materialsCitation id="F2CD71C0E8B08851C9032E3628A60E20" ID-GBIF-Occurrence="923897730" country="Nepal" stateProvince="Gulmi / Syangja">
<collectionCode id="15E11063518B67A7571DEEB15960C9A1" collectionName="OSUS">OSUS</collectionCode>
15718 (9), 34.1-75.5 mm SL;
<collectingCountry id="606A3FA92F2FCC9E4DE0EFA2B875B660">Nepal</collectingCountry>
:
<collectingRegion id="10C72E658733012CCEE28ED0A0EAE821">Gulmi/Syangja</collectingRegion>
, Kali Gandaki River at Ridi Bazar;
<geoCoordinate id="6E995E92B4FACFE0D4A3F085AF393B1C" direction="north" orientation="latitude" value="27.935">27°56'6.0&quot;N</geoCoordinate>
<geoCoordinate id="8A21B04EA9AE9AE5BC323A31DEC1B2BE" direction="east" orientation="longitude" value="83.441696">83°26'30.1&quot;E</geoCoordinate>
; D. Edds,
<date id="C87F5E5A636178CBD444EB001ED652B5" day="22" month="05" value="1984-05-22" year="1984">22 May 1984</date>
.
</materialsCitation>
<materialsCitation id="3DD55518AC52500D12A6DB8790778B3B" ID-GBIF-Occurrence="923897749" country="Nepal" stateProvince="Baglung">
<collectionCode id="164E8D1273562A625117AD65DDAB91A7" collectionName="OSUS">OSUS</collectionCode>
15729 (3), 53.5-59.6 mm SL;
<collectingCountry id="8F0952034E5E38D2B285B3F25B7A3E41">Nepal</collectingCountry>
:
<collectingRegion id="3FE69F68F82F418911AE9EECE793485A">Baglung</collectingRegion>
, Kali Gandaki River at Sumsaa Ghat (Binamaare); D. Edds,
<date id="B73EFB8CDA3170719A25702EB293F9AC" day="25" month="05" value="1984-05-25" year="1984">25 May 1984</date>
.
</materialsCitation>
<materialsCitation id="A8757A20F368C395ECBA856367E9B1F6" ID-GBIF-Occurrence="923897737" country="Nepal" stateProvince="Myagdi">
<collectionCode id="2E78CF57AD928C3F704BBEC6646BD3E2" collectionName="OSUS">OSUS</collectionCode>
15736 (6), 19.8-54.4 mm SL;
<collectingCountry id="E31B9C7F7165BDE1AFF00BE1EC85F432">Nepal</collectingCountry>
:
<collectingRegion id="CC5D5F270AF303C0E793E74B930F6424">Myagdi</collectingRegion>
, Kali Gandaki River at Simaa; D. Edds,
<date id="608DABC12F33D65E1FE88A7741C5702A" day="27" month="05" value="1984-05-27" year="1984">27 May 1984</date>
.
</materialsCitation>
<materialsCitation id="22589CCEB3AE09B57ACBE27D946BEBDC" ID-GBIF-Occurrence="923897729" country="Nepal" stateProvince="Chitawan">
<collectionCode id="95B49F8B494C8D2691018EE57789E42F" collectionName="OSUS">OSUS</collectionCode>
16340 (1), 79.5 mm SL;
<collectingCountry id="8EB90722CB4B5635025A73843BD590F4">Nepal</collectingCountry>
:
<collectingRegion id="1B520F5D7357AF0C9940A6488F39DE98">Chitawan</collectingRegion>
, Narayani River at Narayanagarh, upstream from irrigation office; D. Edds,
<date id="88DB37E783C361B7A29EC21F0C862926" day="10" month="02" value="1985-02-10" year="1985">10 February 1985</date>
.
</materialsCitation>
<materialsCitation id="46A1210138EC79315B37E7CE52C30195" ID-GBIF-Occurrence="923897744" country="Nepal" stateProvince="Chitawan">
<collectionCode id="A64252CA12BD91289353F5D8D524164E" collectionName="OSUS">OSUS</collectionCode>
16609 (1), 22.2 mm SL;
<collectingCountry id="BAF89801C754ED93E9E2C067D4A39802">Nepal</collectingCountry>
:
<collectingRegion id="D8C0D54F6FD3D2FE3FF05234343FB3BC">Chitawan</collectingRegion>
, Narayani River at Amaltaari Ghat; D. Edds,
<date id="746B8617C04CD8A4458C7F7A18DEFB38" day="05" month="04" value="1985-04-05" year="1985">5 April 1985</date>
.
</materialsCitation>
<materialsCitation id="659347C35BD62D47410B53E685D3D484" ID-GBIF-Occurrence="923897743" country="Nepal" stateProvince="Chitawan">
<collectionCode id="CB6F1C9E95C607D471B12492670949DE" collectionName="OSUS">OSUS</collectionCode>
16637 (15), 63.0-130.5 mm SL;
<collectingCountry id="158E4F2C310CBCBDAEBE4F00B9CED2D6">Nepal</collectingCountry>
:
<collectingRegion id="F1781F07EEAE312A23DEEE16C549EFBC">Chitawan</collectingRegion>
, Narayani River at Narayangarh, upstream from irrigation office; D. Edds,
<date id="1E32A041FC74A4D84C020E26938EB049" day="19" month="04" value="1985-04-19" year="1985">19 April 1985</date>
.
</materialsCitation>
<materialsCitation id="66A0764994A90BD38B9428F41C577E8E" ID-GBIF-Occurrence="923897742" country="Nepal" stateProvince="Syangja">
<collectionCode id="43D75355600A70DF0B18F2C1D264C730" collectionName="OSUS">OSUS</collectionCode>
16695 (1), 62.3 mm SL;
<collectingCountry id="E84665526AD719B5B32AC15A847D57D5">Nepal</collectingCountry>
:
<collectingRegion id="B4CD194FBE0565FB6AE7162EC3178CED">Syangja</collectingRegion>
, Kali Gandaki River at Nimaa; D. Edds,
<date id="58B0DE7BED97A200F9B114633B9EA7A8" day="02" month="05" value="1985-05-02" year="1985">2 May 1985</date>
.
</materialsCitation>
<materialsCitation id="1D79D1982D1A46472D60EBF3BA65E543" ID-GBIF-Occurrence="923897746" country="Nepal" stateProvince="Syangja">
<collectionCode id="1FAE79FABB475CC125D282CBC4353088" collectionName="OSUS">OSUS</collectionCode>
17179 (1), 80.0 mm SL;
<collectingCountry id="757D010DB5C104E5E6DEE4EFF88686A4">Nepal</collectingCountry>
:
<collectingRegion id="843D8D53A6074F7391B244A1AD576831">Syangja</collectingRegion>
, Kali Gandaki River at Nimaa; D. Edds,
<date id="05D9A9F45875EB0FE68B938E75ADEF8A" day="06" month="01" value="1986-01-06" year="1986">6 January 1986</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="88AC64C09CF97907F5B5BB4A89DB748B" type="diagnosis">
<paragraph id="111BC1688253DCA32B97DDC6909E2A9C" pageNumber="7">
Diagnosis.
<taxonomicName id="996BC32FA5507F55C09026055AA75AE4" LSID-ZBK="urn:lsid:zoobank.org:act:175A3E4F-DFFE-405A-A858-8920376B267D" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudecheneis serracula Ng &amp; Edds 2005" order="Siluriformes" pageNumber="7" phylum="Chordata" rank="species" species="serracula">Pseudecheneis serracula</taxonomicName>
is distinguished from
<taxonomicName id="6B64D8EE53597B01F34E97F75C49288F" LSID-ZBK="urn:lsid:zoobank.org:act:31513E00-E57B-4618-A71C-41681FF7E329" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudecheneis paviei Vaillant 1892:126 [2]" order="Siluriformes" pageNumber="7" phylum="Chordata" rank="species" species="paviei">P. paviei</taxonomicName>
and
<taxonomicName id="20DFBD8B26FA34D7F16E58FEA8B1A742" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="7" phylum="Chordata" rank="species" species="sympelvica">P. sympelvica</taxonomicName>
in having an elongate body with 36-38 vertebrae (vs. short body with 33-35 vertebrae) and from
<taxonomicName id="CC39A29DE896A4A11380956C4D4A6968" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="7" phylum="Chordata" rank="species" species="sympelvica">P. sympelvica</taxonomicName>
in having separate (vs. fused) pelvic fins. It differs from other congeners in having strongly elevated neural spines of the last 2-3 preanal and first 6-7 postanal vertebrae (vs. neural spines of corresponding pre- and postanal vertebrae gradually increasing in height; Fig. 4) and, except for
<taxonomicName id="682E8B1248C0485068F61294974E7F4A" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="7" phylum="Chordata" rank="species" species="immaculata">P. immaculata</taxonomicName>
, in having a longer adipose-fin base (at least 2.0 times length of anal-fin base vs. 1.5-2.0 times; 26.8-30.4% SL vs. 17.8-24.3).
<taxonomicName id="3012DEE0253F5CD254EF965214709FE0" LSID-ZBK="urn:lsid:zoobank.org:act:175A3E4F-DFFE-405A-A858-8920376B267D" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudecheneis serracula Ng &amp; Edds 2005" order="Siluriformes" pageNumber="7" phylum="Chordata" rank="species" species="serracula">Pseudecheneis serracula</taxonomicName>
is further distinguished from
<taxonomicName id="C17980AB49A8352C3CEC8D205FB8746E" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="7" phylum="Chordata" rank="species" species="immaculata">P. immaculata</taxonomicName>
in the presence (vs. absence) of pale spots on the body, and from
<taxonomicName id="2836EE82DCA9C8D58A6997A90B713FE2" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="7" phylum="Chordata" rank="species" species="sulcata">P. sulcata</taxonomicName>
in having pelvic fins that reach (vs. do not reach) the base of the first anal-fin ray.
</paragraph>
</subSubSection>
<subSubSection id="35C63691EAF3D2588055E8E439505AF8" type="description">
<paragraph id="C51EC8DDC0C71736A5729DB3B1AA3E1F" pageNumber="7">Description. Morphometric data as in Table 2. Head and abdominal region narrow and strongly depressed. Dorsal profile rising gently from tip of snout to origin of dorsal fin, then almost horizontal or sloping very gently ventrally to end of caudal peduncle. Ventral profile horizontal to anal-fin base, then sloping very gently dorsally to end of caudal peduncle. Caudal peduncle long and moderately compressed. Anus and urogenital openings located at posteriormost extent of pelvic fin. Skin smooth, tuberculate in some areas. Lateral line complete and midlateral. Vertebrae 18+18=36 (14), 19+17=36 (1), 18+19=37 (14), 19+18=37* (19), 19+19=38 (3) or 20+18=38 (1).</paragraph>
<paragraph id="CBDA6F080326B213C49877028AE01222" pageNumber="7">Head acutely rounded when viewed from above. Gill openings moderate, extending from posttemporal region to base of first pectoral-fin element. Head covered with thick, tuberculate skin. Ventral surface of head with unculiferous collar on distal margin of branchiostegal membrane immediately anterior to thoracic adhesive apparatus.</paragraph>
<paragraph id="59D4F30DB9AD781BC3DE9F6CB1E2009C" pageNumber="9">Thoracic adhesive apparatus consisting of 13-18 transverse ridges (laminae) separated by grooves (sulcae); ridges frequently not meeting at midline of adhesive apparatus. Adhesive apparatus extending from immediately posterior to collar on distal margin of branchiostegal membrane to level of last pectoral-fin ray.</paragraph>
<paragraph id="E81E3E10BA07D2BCD3A16568E4DEBD24" lastPageNumber="10" pageNumber="9">Barbels flattened, and in four pairs. Maxillary barbel with ventral surface densely covered with papillae, and pointed tip; barbel extending about two-thirds of distance between its base and base of first pectoral-fin element. Distal half of barbel attached to snout via large, thin flap of skin. Nasal barbel with small flap of thin skin fringing posterior margin and extending midway to distance between posterior nares and anterior orbital margin. Inner mandibular-barbel densely covered with papillae; origin close to midline, extending to collar on distal margin of branchiostegal membrane. Outer mandibular barbel originates posterolateral of inner mandibular barbel, extending to level of anterior orbital margin. Eye small and almost rounded, subcutaneous and located on dorsal surface of head.</paragraph>
<paragraph id="985AF8212613A694B67812F5F10C0048" pageNumber="10">Mouth inferior, with moderately broad, thin papillate lips. Rictal lobe large and papillate. Premaxillary tooth band not exposed when mouth is closed. Premaxillary teeth short and conical, arranged in irregular rows on a moderately large quadrangular patch. Dentary teeth long, thin and somewhat rounded at tip; arranged in irregular rows on two separated, roughly triangular patches.</paragraph>
<paragraph id="4582F214ED9F20AB23044BC944AAED47" pageNumber="10">Dorsal-fin origin located at point through anterior third of body. First and second dorsal fin-ray elements not ossified, bearing i,5 (1), i,5,i (4) or i,6* (47) rays, and fin margin straight. Adipose fin with moderately long base, at least 2.0 times anal-fin base length; located in middle third of postdorsal region. Adipose fin margin gently convex; posterior end deeply incised. Caudal fin forked, with i,7,7,i (1), i,7,8,i* (47), i,8,7,i (1) or i,8,8,i (3) principal rays; procurrent rays symmetrical and extend only slightly anterior to fin base. Anal fin with short base extending less than half of adipose fin-base length and iv,7 (15), iv,8 (22), v,6 (1), v,6,i (7) or v,7* (7) rays. Anal fin margin almost straight.</paragraph>
<paragraph id="8707E24E84D682F180A59B0A260A9B8E" pageNumber="10">Pelvic-fin origin at vertical through second or third dorsal fin-ray base. Pelvic fin greatly enlarged, extending to base of first anal-fin ray. Anterior fin margin strongly convex, first element broadened and with regular striae on ventral surface; with i,5 (52) rays. Pectoral fin greatly enlarged and with convex anterior margin, reaching to just beyond pelvic-fin base. First element not ossified, broadened and with regular striae on ventral surface; fin with i,12 (2), i, 13* (40) or i, 14 (10) rays.</paragraph>
<paragraph id="075E3DA0404048FC801A198820D53CFA" pageNumber="10">Coloration. In 70% ethanol and 45% isopropanol: chestnut brown on dorsal and lateral surfaces of head and body, fading to very light brown on ventral region. Dorsal surfaces of head and body with distinctive series of small very light brown spots and bands: one ovate spot on base of first dorsal-fin ray, and another pair on each side of body immediately posterior to last dorsal-fin ray; one band on each side of body at adipose-fin origin, and another on caudal peduncle at base of caudal fin. Dorsal and anal fins hyaline, with brown base and brown subdistal band; brown coloration of base and subdistal band connected to each other at anterior third of fin. Adipose fin light brown, with lighter color around distal edge, especially at posterior end of fin. Caudal fin brown, with hyaline distal margin. Dorsal surfaces of pectoral and pelvic fins brown, ventral surfaces light yellow. Maxillary and nasal barbels brown dorsally and light yellow ventrally.</paragraph>
</subSubSection>
<subSubSection id="CEEC3522077B3E1150FF55757A6B99E2" type="distribution">
<paragraph id="645522104F835D66752CE4AFBE61DBE0" lastPageNumber="11" pageNumber="10">Distribution. Presently known only from rivers of the Middle Hills and Tarai (lowlands) of western (Karnali drainage) and central (Gandaki drainage) Nepal, in and at the base of foothills of the Himalayas (Fig. 2). The Karnali (= Ghaghara or Gogra in India) is a major tributary of the Ganges River in India. The Mahakali (= Sarda in India) forms the western border of Nepal with India, and is a tributary to the Ghaghara in India. The Seti and Kali Gandaki are tributaries to the Trisuli, which becomes the Narayani (= Sapta Gandaki) in Nepal. The Narayani (= Gandak in India) is a major tributary of the Ganges River in India.</paragraph>
</subSubSection>
<subSubSection id="E335E8B28DB095B472FF4A7DB432E05B" type="biology_ecology">
<paragraph id="6A351B41944BC908E78A19DF4F038BE2" pageNumber="11">Habitat and ecology. This species was captured exclusively in riffles, which were typically broad and shallow, well-oxygenated, and had swift water velocities over rocky substrate (Fig. 4). Habitat assessment for seven of these areas showed mean water depths of 10 cm (SE = 0.3), mean water velocities of 0.7 m/s (SE = 0.08), minimum dissolved oxygen of 8 mg/L, and pH from 8.0 to 8.75. Substrate was primarily cobble (ca. 64-256 mm) (68%, SE = 12.0) and pebble (ca. 32-64 mm) (17%, SE = 6.8).</paragraph>
<paragraph id="818BE5D289CEA1D1B99C7D586897D73C" pageNumber="11">
<taxonomicName id="5ADA0C476BC816416EB78D4158638F45" LSID-ZBK="urn:lsid:zoobank.org:act:175A3E4F-DFFE-405A-A858-8920376B267D" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudecheneis serracula Ng &amp; Edds 2005" order="Siluriformes" pageNumber="11" phylum="Chordata" rank="species" species="serracula">Pseudecheneis serracula</taxonomicName>
feeds on aquatic insects, including caddisflies (Edds, 1987). Species associates in our collections included
<taxonomicName id="0B7683414E5DB2E3E2E91DA0B7C5008D" family="Cyprinidae" genus="Crossocheilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="11" phylum="Chordata" rank="species" species="latius">Crossocheilus latius</taxonomicName>
(Cyprinidae),
<taxonomicName id="22116D1BC978057EE484EC7DE46107B9" LSID-ZBK="urn:lsid:zoobank.org:act:B5231D6F-6D10-4C4D-9860-C48BC4527D96" family="Cyprinidae" genus="Garra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Garra annandalei Hora 1921:657" order="Cypriniformes" pageNumber="11" phylum="Chordata" rank="species" species="annandalei">Garra annandalei</taxonomicName>
(Cyprinidae),
<taxonomicName id="38C183E04492C13C74BF59C311A13FAB" family="Cyprinidae" genus="Garra" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="11" phylum="Chordata" rank="species" species="gotyla">G. gotyla</taxonomicName>
(Cyprinidae),
<taxonomicName id="647839511229750EAE3711A3FA58466B" family="Cyprinidae" genus="Schizothoraichthys" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="11" phylum="Chordata" rank="species" species="progastus">Schizothoraichthys progastus</taxonomicName>
(Cyprinidae),
<taxonomicName id="A2D715E18BD5ED5B92585275838A51D8" family="Cyprinidae" genus="Schizothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="11" phylum="Chordata" rank="species" species="richardsonii">Schizothorax richardsonii</taxonomicName>
(Cyprinidae),
<taxonomicName id="A7631FC59FFE99F03EA7B726E15A8D17" LSID-ZBK="urn:lsid:zoobank.org:act:675C7B4C-7B57-48BA-A93E-6A68987F7FD4" family="Cobitidae" genus="Botia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Botia almorhae Gray 1831:8" order="Cypriniformes" pageNumber="11" phylum="Chordata" rank="species" species="almorhae">Botia almorhae</taxonomicName>
(Cobitidae), and
<taxonomicName id="A92A52F9824191374AF4AF230F19F7CF" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="11" phylum="Chordata" rank="species" species="pectinopterus">Glyptothorax cf. pectinopterus</taxonomicName>
(Sisoridae). In the Seti River,
<taxonomicName id="11BB73536F6C09AC51660CCAAB78C01C" LSID-ZBK="urn:lsid:zoobank.org:act:175A3E4F-DFFE-405A-A858-8920376B267D" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudecheneis serracula Ng &amp; Edds 2005" order="Siluriformes" pageNumber="11" phylum="Chordata" rank="species" species="serracula">P. serracula</taxonomicName>
was found syntopically with
<taxonomicName id="2E93C194B626629AE17B536289E65055" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="11" phylum="Chordata" rank="species" species="sulcata">P. sulcata</taxonomicName>
. Local fishermen report that
<taxonomicName id="45E05C82753D151E5EDBAD948A03898B" LSID-ZBK="urn:lsid:zoobank.org:act:175A3E4F-DFFE-405A-A858-8920376B267D" family="Sisoridae" genus="Pseudecheneis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudecheneis serracula Ng &amp; Edds 2005" order="Siluriformes" pageNumber="11" phylum="Chordata" rank="species" species="serracula">P. serracula</taxonomicName>
(local name = kabre) ascends the Kali Gandaki River to ca. 1000 m asl during monsoon (Edds, 1986b), suggesting these fish may breed in upper reaches of the river, then migrate downstream following spawning season.
</paragraph>
</subSubSection>
<subSubSection id="AFB35703C08D266A8D1FB5668649E3CE" type="etymology">
<paragraph id="D48ECC3D1D97F44CC96B76C16ACB6D3B" pageNumber="12">Etymology. From the Latin serraculum, meaning rudder, in allusion to the large adipose fin of this species. Used as a noun.</paragraph>
</subSubSection>
</treatment>
</document>