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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.181.69740" ID-GBIF-Dataset="dca39c99-b65a-42c7-accd-4f7d8e8b421a" ID-PMC="PMC8421323" ID-Pensoft-Pub="1314-2003-181-29" ID-Pensoft-UUID="25B16C1AE4375702BC69E64E28291B0A" ID-PubMed="34557054" ModsDocID="1314-2003-181-29" checkinTime="1630349217782" checkinUser="pensoft" docAuthor="Moroni, Pablo, Martinez, Agustina &amp; Simpson, Michael G." docDate="2021" docId="A3EBB54A2AFA52419A077F3FF6822682" docLanguage="en" docName="PhytoKeys 181: 29-47" docOrigin="PhytoKeys 181" docPubDate="2021-08-30" docSource="http://dx.doi.org/10.3897/phytokeys.181.69740" docTitle="Cryptantha chaetocalyx (Phil.) I. M. Johnst., Contr. Gray Herb. 78: 43. 1927." docType="treatment" docVersion="4" id="25B16C1AE4375702BC69E64E28291B0A" lastPageNumber="29" masterDocId="25B16C1AE4375702BC69E64E28291B0A" masterDocTitle="Nomenclatural revision of Cryptantha (Boraginaceae s. str.) names linked to South American taxa" masterLastPageNumber="47" masterPageNumber="29" pageNumber="29" updateTime="1668140413250" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Nomenclatural revision of Cryptantha (Boraginaceae s. str.) names linked to South American taxa</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moroni, Pablo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5306-476X</mods:nameIdentifier>
<mods:affiliation>Instituto de Botanica Darwinion (ANCEFN-CONICET), Labarden 200, CC 22, B 1642 HYD, San Isidro, Buenos Aires, Argentina</mods:affiliation>
<mods:nameIdentifier type="email">pmoroni@darwin.edu.ar</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Martinez, Agustina</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4768-664X</mods:nameIdentifier>
<mods:affiliation>Instituto de Botanica Darwinion (ANCEFN-CONICET), Labarden 200, CC 22, B 1642 HYD, San Isidro, Buenos Aires, Argentina</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Simpson, Michael G.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6197-2132</mods:nameIdentifier>
<mods:affiliation>Department of Biology, San Diego State University, San Diego, California 92182, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-08-30</mods:number>
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<mods:detail type="volume">
<mods:number>181</mods:number>
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<mods:start>29</mods:start>
<mods:end>47</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.181.69740</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.181.69740</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-181-29</mods:identifier>
<mods:identifier type="Pensoft-UUID">25B16C1AE4375702BC69E64E28291B0A</mods:identifier>
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<treatment ID-GBIF-Taxon="185371626" LSID="urn:lsid:plazi:treatment:A3EBB54A2AFA52419A077F3FF6822682" httpUri="http://treatment.plazi.org/id/A3EBB54A2AFA52419A077F3FF6822682" lastPageNumber="29" pageId="0" pageNumber="29">
<subSubSection pageId="0" pageNumber="29" type="nomenclature">
<paragraph pageId="0" pageNumber="29">
6.
<taxonomicName LSID="A3EBB54A-2AFA-5241-9A07-7F3FF6822682" authority="(Phil.) I. M. Johnst., Contr. Gray Herb. 78: 43. 1927." authorityName="(Phil.) I. M. Johnst., Contr. Gray Herb. 78: 43. 1927." class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cryptantha chaetocalyx" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="chaetocalyx">Cryptantha chaetocalyx (Phil.) I.M. Johnst., Contr. Gray Herb. 78: 43. 1927.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29" type="reference_group">
<paragraph pageId="0" pageNumber="29">
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cryptantha chaetocalyx" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="chaetocalyx">Cryptantha chaetocalyx</taxonomicName>
<taxonomicName authorityName="Philippi" authorityYear="1860" class="Magnoliopsida" family="Boraginaceae" genus="Eritrichium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eritrichium chaetocalyx" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="chaetocalyx">Eritrichium chaetocalyx</taxonomicName>
Phil., Fl. Atacam. 39. 1860. Type: Chile.
<normalizedToken originalValue="Región">Region</normalizedToken>
de Atacama: Caldera, Pan de
<normalizedToken originalValue="Azúcar">Azucar</normalizedToken>
, Dec. 1853,
<emphasis italics="true" pageId="0" pageNumber="29">R.A. Philippi s.n.</emphasis>
(first-step lectotype, designated by
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">Johnston 1927</bibRefCitation>
, pg. 43: SGO; second-step lectotype, designated here: SGO [SGO000004073 digital image!]; isolectotypes: GH [GH00096375 digital image!], SGO [SGO000004050 digital image!]).
</paragraph>
<paragraph pageId="0" pageNumber="29">
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cryptantha chaetocalyx" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="chaetocalyx">Cryptantha chaetocalyx</taxonomicName>
=
<taxonomicName authorityName="Philippi" authorityYear="1895" class="Magnoliopsida" family="Boraginaceae" genus="Eritrichium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eritrichium divaricatum" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="divaricatum">Eritrichium divaricatum</taxonomicName>
Phil., Anales Univ. Chile 90: 534. 1895.
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cryptantha divaricata" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="divaricata">Cryptantha divaricata</taxonomicName>
(Phil.) Reiche, Anales Univ. Nac. Tec. Altiplano 121: 827. 1907. Type: Chile.
<normalizedToken originalValue="Región">Region</normalizedToken>
de Atacama: Caldera, Sep. 1885,
<emphasis italics="true" pageId="0" pageNumber="29">F. Philippi s.n</emphasis>
. (first-step lectotype, designated by
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">Johnston 1927</bibRefCitation>
, pg. 43: SGO; second-step lectotype, designated here: SGO [SGO000004070 digital image!]; isolectotypes: GH [GH00096373 digital image!], SGO [SGO000004071 digital image!]).
</paragraph>
<paragraph pageId="0" pageNumber="29">
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cryptantha chaetocalyx" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="chaetocalyx">Cryptantha chaetocalyx</taxonomicName>
=
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Eritrichium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eritrichium pustulosum" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="pustulosum">Eritrichium pustulosum</taxonomicName>
Phil., Anales Univ. Chile 90: 537. 1895. Type: Chile.
<normalizedToken originalValue="Región">Region</normalizedToken>
de Atacama: Caldera, Sep. 1879,
<emphasis italics="true" pageId="0" pageNumber="29">R.. Philippi s.n</emphasis>
. (first-step lectotype, designated by
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">Johnston 1927</bibRefCitation>
, pg. 43: SGO; second-step lectotype, designated here: SGO [SGO000004125 digital image!]; isolectotypes: GH [GH00096374 digital image!], SGO [SGO000004126 digital image!], SGO [SGO000004127 digital image!]).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29" type="notes">
<paragraph pageId="0" pageNumber="29">Notes.</paragraph>
<paragraph pageId="0" pageNumber="29">
In describing
<taxonomicName authorityName="Philippi" authorityYear="1860" class="Magnoliopsida" family="Boraginaceae" genus="Eritrichium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eritrichium chaetocalyx" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="chaetocalyx">
<emphasis italics="true" pageId="0" pageNumber="29">Eritrichium chaetocalyx</emphasis>
</taxonomicName>
,
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.162865" author="Philippi, RA" journalOrPublisher="INTA, Buenos Aires" pageId="0" pageNumber="29" refId="B35" refString="Philippi, RA, 1860. Florula Atacamensis seu enumeratio plantarum, quas in itinere per desertum Atacamense. Sumptibus E. Anton. https://doi.org/10.5962/bhl.title.162865" title="Florula Atacamensis seu enumeratio plantarum, quas in itinere per desertum Atacamense. Sumptibus E. Anton." url="https://doi.org/10.5962/bhl.title.162865" year="1860">Rudolph A. Philippi (1860)</bibRefCitation>
cited a gathering he made near Pan de
<normalizedToken originalValue="Azúcar">Azucar</normalizedToken>
, Chile. Two duplicates of the type collection in agreement with the protologue and annotated by him as &quot;
<taxonomicName authorityName="Philippi" authorityYear="1860" class="Magnoliopsida" family="Boraginaceae" genus="Eritrichium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eritrichium chaetocalyx" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="chaetocalyx">Eritrichium chaetocalyx</taxonomicName>
&quot; were located at SGO.
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">Johnston (1927)</bibRefCitation>
discussed this name and its original material, referring to a sheet lodged at SGO as the
<normalizedToken originalValue="“type”">&quot;type&quot;</normalizedToken>
of the name. In this context,
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">
<normalizedToken originalValue="Johnstons">Johnston's</normalizedToken>
(1927)
</bibRefCitation>
statement must thus be considered as a first-step typification. Thus, in order to narrow this designation, the sheet SGO000004073 is here selected as a second-step lectotype.
</paragraph>
<paragraph pageId="0" pageNumber="29">
In the protologue of
<taxonomicName authorityName="Philippi" authorityYear="1895" class="Magnoliopsida" family="Boraginaceae" genus="Eritrichium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eritrichium divaricatum" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="divaricatum">
<emphasis italics="true" pageId="0" pageNumber="29">Eritrichium divaricatum</emphasis>
</taxonomicName>
,
<bibRefCitation DOI="https://doi.org/10.5354/0717-8883.2012.20872" author="Philippi, RA" journalOrPublisher="Anales de la Universidad de Chile" pageId="0" pageNumber="29" pagination="511 - 566" refId="B39" refString="Philippi, RA, 1895. Plantas nuevas chilenas de las familias que corresponden al tomo IV de la obra de Gay: (continuacion). Anales de la Universidad de Chile 90: 511 - 566, DOI: https://doi.org/10.5354/0717-8883.2012.20872" title="Plantas nuevas chilenas de las familias que corresponden al tomo IV de la obra de Gay: (continuacion)." url="https://doi.org/10.5354/0717-8883.2012.20872" volume="90" year="1895">Rudolph A. Philippi (1895)</bibRefCitation>
cited as type material a collection made by F. Philippi in Caldera, Chile.
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">Johnston (1927)</bibRefCitation>
referred to a specimen kept at SGO as the
<normalizedToken originalValue="“type”">&quot;type&quot;</normalizedToken>
of the name, although two sheets annotated, in R. A.
<normalizedToken originalValue="Philippis">Philippi's</normalizedToken>
hand, as &quot;
<taxonomicName authorityName="Philippi" authorityYear="1895" class="Magnoliopsida" family="Boraginaceae" genus="Eritrichium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eritrichium divaricatum" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="divaricatum">Eritrichium divaricatum</taxonomicName>
&quot; are found there. This situation determines that
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">
<normalizedToken originalValue="Johnstons">Johnston's</normalizedToken>
(1927)
</bibRefCitation>
statement must be considered as a first-step lectotypification. In this context, the duplicate SGO000004070 is here selected as a second-step lectotype (Art. 9.17 of the ICN;
<bibRefCitation DOI="https://doi.org/10.12705/Code.2018" author="Turland, NJ" journalOrPublisher="Annals of the Missouri Botanical Garden" pageId="0" pageNumber="29" refId="B51" refString="Turland, NJ, Wiersema, JH, Barrie, FR, Greuter, W, Hawksworth, DL, Herendeen, PS, Knapp, S, Kusber, WH, Li, DZ, Marhold, K, May, TW, McNeill, J, Monro, AM, Prad, J, Price, MJ, Smith, GF, 2018. International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten. https://doi.org/10.12705/Code.2018" title="International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten." url="https://doi.org/10.12705/Code.2018" year="2018">Turland et al. 2018</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="29">
<bibRefCitation DOI="https://doi.org/10.5354/0717-8883.2012.20872" author="Philippi, RA" journalOrPublisher="Anales de la Universidad de Chile" pageId="0" pageNumber="29" pagination="511 - 566" refId="B39" refString="Philippi, RA, 1895. Plantas nuevas chilenas de las familias que corresponden al tomo IV de la obra de Gay: (continuacion). Anales de la Universidad de Chile 90: 511 - 566, DOI: https://doi.org/10.5354/0717-8883.2012.20872" title="Plantas nuevas chilenas de las familias que corresponden al tomo IV de la obra de Gay: (continuacion)." url="https://doi.org/10.5354/0717-8883.2012.20872" volume="90" year="1895">Rudolph A. Philippi (1895)</bibRefCitation>
described
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Eritrichium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eritrichium pustulosum" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="pustulosum">
<emphasis italics="true" pageId="0" pageNumber="29">Eritrichium pustulosum</emphasis>
</taxonomicName>
based on a specimen he collected in Caldera, Chile. According to
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">Johnston (1927)</bibRefCitation>
, the type material of the name is lodged at SGO. However, three sheets linked to
<taxonomicName lsidName="E. pustulosum" pageId="0" pageNumber="29" rank="species" species="pustulosum">
<emphasis italics="true" pageId="0" pageNumber="29">E. pustulosum</emphasis>
</taxonomicName>
, in agreement with the protologue, were located at SGO, which can certainly be considered as original material. In this context,
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">
<normalizedToken originalValue="Johnstons">Johnston's</normalizedToken>
(1927)
</bibRefCitation>
statement must be considered as a first-step lectotypification. In order to narrow this earlier designation, the sheet SGO000004125 is here selected as a second-step lectotype.
</paragraph>
</subSubSection>
</treatment>
</document>