treatments-xml/data/84/78/3E/84783E54FFBFFFC0AFDA788EFD247543.xml
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<document id="0EB5E29D663F9A08E32F390791ED4D19" ID-CLB-Dataset="9628" ID-DOI="10.1093/zoolinnean/zly024" ID-GBIF-Dataset="c93d9a7f-28c5-4282-a677-5f28410fc66c" ID-Zenodo-Dep="4253044" ID-ZooBank="65FFC8DB-4738-49FF-99F5-FC8532CC9795" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1604692953150" checkinUser="valdenar" docAuthor="Taylor, Peter J., Macdonald, Angus, Goodman, Steven M., Kearney, Teresa, Cotterill, Fenton P. D., Stoffberg, Sam, Monadjem, Ara, Schoeman, M. Corrie, Guyton, Jennifer, Naskrecki, Pitor &amp; Richards, Leigh R." docDate="2018" docId="84783E54FFBFFFC0AFDA788EFD247543" docLanguage="en" docName="ZoolJLinnSoc.184.12491276.pdf.imf" docOrigin="Zoological Journal of the Linnean Society 184" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Rhinolophus gorongosae GORONGOSAE 2018, SP. NOV." docType="treatment" docVersion="13" lastPageNumber="1265" masterDocId="7841462CFFB2FFD0AF2C7C04FFB67043" masterDocTitle="Integrative taxonomy resolves three new cryptic species of small southern African horseshoe bats (Rhinolophus)" masterLastPageNumber="1276" masterPageNumber="1249" pageNumber="1262" updateTime="1698846174469" updateUser="ExternalLinkService">
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<mods:title id="D652D828A9CE156AC4013623BA60F65F">Integrative taxonomy resolves three new cryptic species of small southern African horseshoe bats (Rhinolophus)</mods:title>
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<mods:namePart id="4B34B0D6D74B8CDCC9C6AB6368CE01DE">Taylor, Peter J.</mods:namePart>
<mods:affiliation id="1DA04CE8024B67A800B62314583C0AF2">SARChI Chair on Biodiversity Value &amp; Change and Core Team Member of the Centre for Invasion Biology, School of Mathematical &amp; Natural Sciences, University of Venda, Private Bag X 5050, Thohoyandou 0950, South Africa &amp; School of Life Sciences, University of KwaZulu-Natal, Biological Sciences Building, South Ring Road, Westville Campus, Durban, KwaZulu-Natal 3630, South Africa</mods:affiliation>
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<mods:namePart id="4D16FCDDA7BE6369C4328A28D9A30830">Macdonald, Angus</mods:namePart>
<mods:affiliation id="B67CF44812B7AC0B5AE8F561343A6BF7">School of Life Sciences, University of KwaZulu-Natal, Biological Sciences Building, South Ring Road, Westville Campus, Durban, KwaZulu-Natal 3630, South Africa</mods:affiliation>
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<mods:namePart id="9A5F825FB108111F30AD1D30978C46CB">Goodman, Steven M.</mods:namePart>
<mods:affiliation id="A969655B5DFED33BF1EE866E644199E8">Field Museumof Natural History, 1400 South Lake Shore Drive, Chicago, IL 60605, USA &amp; Association Vahatra, BP 3972, Antananarivo 101, Madagascar</mods:affiliation>
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<mods:namePart id="368687E5E6FAF1D2CA1897CB8E21121E">Kearney, Teresa</mods:namePart>
<mods:affiliation id="B49180A46CB3344988FBB61622ECDEBB">Ditsong National Museumof Natural History, POBox 413, Pretoria 0001, South Africa &amp; School of Animal, Plant and Environmental Sciences, University of the Witwatersrand, Private Bag 3, Wits 2050, South Africa</mods:affiliation>
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<mods:namePart id="F107EBAF9FDB0CF10E92BE193BBF40CA">Cotterill, Fenton P. D.</mods:namePart>
<mods:affiliation id="E6B6F0AEBFE847E6DBE9CB365E5A24E3">Geoecodynamics Research Hub, c / o Department of Botany and Zoology, University of Stellenbosch, Private Bag X 1 Matieland, 7602, Stellenbosch, South Africa</mods:affiliation>
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<mods:namePart id="DC3BA7F861194833B9B1FE0CB2812FD9">Stoffberg, Sam</mods:namePart>
<mods:affiliation id="4BEE0C76FE0B5A0FAA44D89B10DF44E4">Department of Botany and Zoology, University of Stellenbosch, Private Bag X 1 Matieland, 7602, Stellenbosch, South Africa</mods:affiliation>
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<mods:namePart id="943ED99B64EA681421BD90B6C552BB69">Monadjem, Ara</mods:namePart>
<mods:affiliation id="48824A08A2D29615CF627E19D49D4AAD">All Out Africa Research Unit, Department of Biological Sciences, University of Swaziland, Private Bag 4, Kwaluseni, Swaziland</mods:affiliation>
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<mods:namePart id="F078753CBAF8D7767939536C26B75446">Schoeman, M. Corrie</mods:namePart>
<mods:affiliation id="2719252D954296BFB709AE27FC9940DB">School of Life Sciences, University of KwaZulu-Natal, Biological Sciences Building, South Ring Road, Westville Campus, Durban, KwaZulu-Natal 3630, South Africa</mods:affiliation>
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<mods:namePart id="5765194BDC64FD65E26E129E44B73DD9">Guyton, Jennifer</mods:namePart>
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<mods:namePart id="E06F43DD13345941AAEA0D377ADE6EF3">Naskrecki, Pitor</mods:namePart>
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<mods:namePart id="6D0238B2B46F5D5FDA2C4F7A9E0E35A0">Richards, Leigh R.</mods:namePart>
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<mods:title id="44E02FF3C1A252353AD47CC506A91666">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="87E65B957035A1DD878C79A089E24217">2018</mods:date>
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<mods:identifier id="6897BE5B549524DE49261FD37710B000" type="DOI">10.1093/zoolinnean/zly024</mods:identifier>
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<treatment id="84783E54FFBFFFC0AFDA788EFD247543" ID-DOI="http://doi.org/10.5281/zenodo.4328253" ID-GBIF-Taxon="168821772" ID-Zenodo-Dep="4328253" LSID="urn:lsid:plazi:treatment:84783E54FFBFFFC0AFDA788EFD247543" httpUri="http://treatment.plazi.org/id/84783E54FFBFFFC0AFDA788EFD247543" lastPageId="16" lastPageNumber="1265" pageId="13" pageNumber="1262">
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<taxonomicName id="CBD1F4C1FFBFFFDDAFDA788EFDFB74E2" ID-CoL="7NDGV" authority="GORONGOSAE" authorityName="GORONGOSAE" authorityYear="2018" box="[246,589,1162,1185]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="1262" phylum="Chordata" rank="species" species="gorongosae" status="sp. nov.">RHINOLOPHUS GORONGOSAE</taxonomicName>
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<taxonomicNameLabel id="2596EE2BFFBFFFDDAD797889FD0E74E1" box="[597,696,1162,1186]" pageId="13" pageNumber="1262" rank="species">SP. NOV.</taxonomicNameLabel>
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<subSubSection id="44CBDCC9FFBFFFDDAE7F78B5FDEC748B" box="[339,602,1201,1226]" pageId="13" pageNumber="1262" type="description">
<paragraph id="0C6E8F42FFBFFFDDAE7F78B5FDEC748B" blockId="13.[339,602,1201,1226]" box="[339,602,1201,1226]" pageId="13" pageNumber="1262">LEAST HORSESHOE BAT</paragraph>
</subSubSection>
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<materialsCitation id="BCB9851FFFBFFFDDAF8F78DDFEDD7660" ID-GBIF-Occurrence="2960754303" accessionNumber="MN025922.1" collectingDate="2015-04-25" collectionCode="JAG" country="Mozambique" latitude="-18.599" location="Sofala Province" longLatPrecision="76" longitude="34.343" pageId="13" pageNumber="1262" specimenCode="JAG196" specimenCount="1" stateProvince="Sofala Province" typeStatus="holotype">
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<typeStatus id="D36A31E0FFBFFFDDAF8F78DDFEA674AD" box="[163,272,1241,1262]" pageId="13" pageNumber="1262" type="holotype">Holotype</typeStatus>
:
</emphasis>
Durban Natural Science Museum (DM) No. 14820 (field number
<specimenCode id="5C772739FFBFFFDDAEE078FCFD98754D" box="[460,558,1272,1294]" pageId="13" pageNumber="1262">JAG196</specimenCode>
), is an adult male, preserved in 70% ethyl alcohol, originally part of series of specimens collected by J. A. Guyton on
<date id="786FA982FFBFFFDDAD847931FF697529" pageId="13" pageNumber="1262" value="2015-04-25">
<collectingDate id="682B506AFFBFFFDDAD847931FF697529" pageId="13" pageNumber="1262" value="2015-04-25">25 April 2015</collectingDate>
</date>
. The cranium and baculum has been extracted and examined for this study. The specimen has been included in both morphometric analyses.
</paragraph>
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<emphasis id="3EA55350FFBFFFDDAF8F79D4FEF775A6" box="[163,321,1488,1509]" italics="true" pageId="13" pageNumber="1262">Type locality:</emphasis>
Bunga Inselberg, Gorongosa National Park,
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,
<collectingCountry id="74C6CFD2FFBFFFDDAEE279EAFDD97647" box="[462,623,1518,1540]" name="Mozambique" pageId="13" pageNumber="1262">Mozambique</collectingCountry>
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,
<geoCoordinate id="69E5E985FFBFFFDDAF8F7A09FEA47660" box="[163,274,1549,1571]" degrees="34.343" direction="east" orientation="longitude" pageId="13" pageNumber="1262" precision="55" value="34.343">34.343° E</geoCoordinate>
,
<quantity id="CB2922A7FFBFFFDDAE317A09FED07660" box="[285,358,1549,1571]" metricMagnitude="2" metricUnit="m" metricValue="2.12" pageId="13" pageNumber="1262" unit="m" value="212.0">212 m</quantity>
.
</paragraph>
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<materialsCitation id="BCB9851FFFBFFFDDAF8F7A48FEC176C3" ID-GBIF-Occurrence="3126007302" collectingDate="2015-04-24" collectingDateMax="2015-04-25" collectingDateMin="2015-04-24" collectionCode="DM" pageId="13" pageNumber="1262" specimenCode="DM 14815-14819" specimenCount="1" typeStatus="paratype">
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<typeStatus id="D36A31E0FFBFFFDDAF8F7A48FEA47622" box="[163,274,1612,1633]" pageId="13" pageNumber="1262" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="1AD744CBFFBFFFDDAE337A48FE6F7621" box="[287,473,1612,1634]" pageId="13" pageNumber="1262" type="generic">Eight specimens</specimenCount>
collected
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<collectingDate id="682B506AFFBFFFDDAD657A48FCBC7621" box="[585,778,1612,1634]" pageId="13" pageNumber="1262" value="2015-04-24" valueMax="2015-04-25" valueMin="2015-04-24">2425 April 2015</collectingDate>
</date>
(
<specimenCode id="5C772739FFBFFFDDAF867A6FFEDA76C3" box="[170,364,1642,1664]" pageId="13" pageNumber="1262">DM 1481514819</specimenCode>
),
</materialsCitation>
<materialsCitation id="BCB9851FFFBFFFDDAE507A6FFD3476C2" ID-GBIF-Occurrence="3126007335" box="[380,642,1642,1665]" collectingDate="2015-05-02" collectionCode="DM" pageId="13" pageNumber="1262" specimenCode="DM 14828" specimenCount="1" typeStatus="paratype">
<date id="786FA982FFBFFFDDAE507A6FFE4E76C3" box="[380,504,1642,1664]" pageId="13" pageNumber="1262" value="2015-05-02">
<collectingDate id="682B506AFFBFFFDDAE507A6FFE4E76C3" box="[380,504,1642,1664]" pageId="13" pageNumber="1262" value="2015-05-02">2 May 2015</collectingDate>
</date>
(
<specimenCode id="5C772739FFBFFFDDAD287A6FFDC176C2" box="[516,631,1643,1665]" pageId="13" pageNumber="1262">DM 14828</specimenCode>
),
</materialsCitation>
<materialsCitation id="BCB9851FFFBFFFDDADAB7A6EFEE976DC" ID-GBIF-Occurrence="3126007321" collectingDate="2015-11-05" collectionCode="DM" pageId="13" pageNumber="1262" specimenCode="DM 14843" specimenCount="1" typeStatus="paratype">
<date id="786FA982FFBFFFDDADAB7A6EFF6E76DC" pageId="13" pageNumber="1262" value="2015-11-05">
<collectingDate id="682B506AFFBFFFDDADAB7A6EFF6E76DC" pageId="13" pageNumber="1262" value="2015-11-05">5 November 2015</collectingDate>
</date>
(
<specimenCode id="5C772739FFBFFFDDAFC97A8DFEEF76DC" box="[229,345,1673,1695]" pageId="13" pageNumber="1262">DM 14843</specimenCode>
)
</materialsCitation>
and
<materialsCitation id="BCB9851FFFBFFFDDAEBA7A8DFD0676DC" ID-GBIF-Occurrence="3126007309" box="[406,688,1673,1695]" collectingDate="2015-07-22" collectionCode="DM" pageId="13" pageNumber="1262" specimenCode="DM 14865" specimenCount="1" typeStatus="paratype">
<date id="786FA982FFBFFFDDAEBA7A8DFD9776DC" box="[406,545,1673,1695]" pageId="13" pageNumber="1262" value="2015-07-22">
<collectingDate id="682B506AFFBFFFDDAEBA7A8DFD9776DC" box="[406,545,1673,1695]" pageId="13" pageNumber="1262" value="2015-07-22">22 July 2015</collectingDate>
</date>
(
<specimenCode id="5C772739FFBFFFDDAD027A8DFD1276DC" box="[558,676,1673,1695]" pageId="13" pageNumber="1262">DM 14865</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="0C6E8F42FFBFFFDDAF8F7ACCFBB5715B" blockId="13.[163,779,1736,1881]" lastBlockId="13.[827,1443,197,280]" pageId="13" pageNumber="1262">
<emphasis id="3EA55350FFBFFFDDAF8F7ACCFCBD769E" box="[163,779,1736,1757]" italics="true" pageId="13" pageNumber="1262">Referred specimens having molecular identification:</emphasis>
<materialsCitation id="BCB9851FFFBFFFDDAF8F7AE3FE20771B" ID-GBIF-Occurrence="3126007322" collectionCode="DM, JAG" collectorName=". A. Guyton" country="Mozambique" elevation="212" latitude="-18.599" location="Bunga Inselberg" longLatPrecision="76" longitude="34.343" municipality="Gorongosa National Park" pageId="13" pageNumber="1262" specimenCode="DM 14815, JAG 188" specimenCount="1" specimenCount-female="1" stateProvince="Sofala Province">
<specimenCode id="5C772739FFBFFFDDAF8F7AE3FE9476BF" box="[163,290,1766,1788]" pageId="13" pageNumber="1262">DM 14815</specimenCode>
(
<specimenCode id="5C772739FFBFFFDDAE1F7AE3FE1676BE" box="[307,416,1767,1789]" pageId="13" pageNumber="1262">JAG 188</specimenCode>
) a
<specimenCount id="1AD744CBFFBFFFDDAEE67AE2FDAA76BF" box="[458,540,1766,1788]" pageId="13" pageNumber="1262" type="female">female</specimenCount>
specimen collected by J
<collectorName id="A124EA94FFBFFFDDAFFA7B02FEED7759" box="[214,347,1797,1819]" pageId="13" pageNumber="1262">. A. Guyton</collectorName>
from
<collectingCountry id="74C6CFD2FFBFFFDDAE887B01FD887758" box="[420,574,1797,1819]" name="Mozambique" pageId="13" pageNumber="1262">Mozambique</collectingCountry>
,
<collectingRegion id="CE1541A0FFBFFFDDAD667B01FCB07758" box="[586,774,1797,1819]" country="Mozambique" name="Sofala" pageId="13" pageNumber="1262">Sofala Province</collectingRegion>
,
<collectingMunicipality id="EC0A1538FFBFFFDDAF8F7B20FE797779" box="[163,463,1828,1850]" pageId="13" pageNumber="1262">Gorongosa National Park</collectingMunicipality>
,
<location id="090ED999FFBFFFDDAEF77B20FD287779" LSID="urn:lsid:plazi:treatment:84783E54FFBFFFC0AFDA788EFD247543:090ED999FFBFFFDDAEF77B20FD287779" box="[475,670,1828,1850]" country="Mozambique" latitude="-18.599" longLatPrecision="76" longitude="34.343" municipality="Gorongosa National Park" name="Bunga Inselberg" pageId="13" pageNumber="1262" stateProvince="Sofala Province">Bunga Inselberg</location>
, -
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,
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,
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<elevation id="87FC6871FFBFFFDDAE697B47FE27771B" box="[325,401,1859,1880]" metricMagnitude="2" metricUnit="m" metricValue="2.12" pageId="13" pageNumber="1262" unit="m" value="212.0">212 m</elevation>
</quantity>
;
</materialsCitation>
<materialsCitation id="BCB9851FFFBFFFDDAEB27B47FBB5715B" ID-GBIF-Occurrence="3126007312" collectionCode="DM, JAG" collectorName="J. A. Guyton" country="Mozambique" elevation="308" latitude="-18.694" location="Gorongosa National Park" longLatPrecision="76" longitude="34.208" municipality="Gorongosa National Park" pageId="13" pageNumber="1262" specimenCode="DM 14843, JAG 228" specimenCount="1" specimenCount-female="1" stateProvince="Sofala Province">
<specimenCode id="5C772739FFBFFFDDAEB27B47FDAC771B" box="[414,538,1859,1880]" pageId="13" pageNumber="1262">DM 14843</specimenCode>
(
<specimenCode id="5C772739FFBFFFDDAD067B47FD22771A" box="[554,660,1859,1881]" pageId="13" pageNumber="1262">JAG 228</specimenCode>
) a
<specimenCount id="1AD744CBFFBFFFDDAD977B46FCBC771B" box="[699,778,1858,1880]" pageId="13" pageNumber="1262" type="female">female</specimenCount>
specimen collected by
<collectorName id="A124EA94FFBFFFDDAB167CC2FB707099" box="[1082,1222,197,219]" pageId="13" pageNumber="1262">J. A. Guyton</collectorName>
from
<collectingCountry id="74C6CFD2FFBFFFDDAA277CC1FA297098" box="[1291,1439,197,219]" name="Mozambique" pageId="13" pageNumber="1262">Mozambique</collectingCountry>
,
<collectingRegion id="CE1541A0FFBFFFDDAC177CE0FC5870B9" box="[827,1006,228,250]" country="Mozambique" name="Sofala" pageId="13" pageNumber="1262">Sofala Province</collectingRegion>
,
<collectingMunicipality id="EC0A1538FFBFFFDDACD57CE0FAA870B9" box="[1017,1310,228,250]" pageId="13" pageNumber="1262">Gorongosa National Park</collectingMunicipality>
, -
<geoCoordinate id="69E5E985FFBFFFDDAA1C7CE0FA2870BA" box="[1328,1438,228,250]" degrees="18.694" direction="south" orientation="latitude" pageId="13" pageNumber="1262" precision="55" value="-18.694">18.694° S</geoCoordinate>
,
<geoCoordinate id="69E5E985FFBFFFDDAC177D07FC1C715B" box="[827,938,259,280]" degrees="34.208" direction="east" orientation="longitude" pageId="13" pageNumber="1262" precision="55" value="34.208">34.208° E</geoCoordinate>
,
<quantity id="CB2922A7FFBFFFDDAC997D07FC48715B" box="[949,1022,259,280]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="13" pageNumber="1262" unit="m" value="308.0">
<elevation id="87FC6871FFBFFFDDAC997D07FC48715B" box="[949,1022,259,280]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="13" pageNumber="1262" unit="m" value="308.0">308 m</elevation>
</quantity>
.
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</paragraph>
<paragraph id="0C6E8F42FFBFFFDDAC177D45FB6C72EB" blockId="13.[827,1443,321,680]" pageId="13" pageNumber="1262">
<materialsCitation id="BCB9851FFFBFFFDDAC177D45FB9C71B3" ID-GBIF-Occurrence="3126007304" collectingDate="2013-04-21" collectionCode="TMSA, JAG" collectorName="J. A. Guyton" country="Mozambique" elevation="213" latitude="-18.635" location="Cheringoma Plateau" longLatPrecision="76" longitude="34.808" municipality="Gorongosa National Park" pageId="13" pageNumber="1262" specimenCode="TMSA 49116, JAG 31" specimenCount="1" specimenCount-male="1" stateProvince="Sofala Province">
<emphasis id="3EA55350FFBFFFDDAC177D45FC5E7136" italics="true" pageId="13" pageNumber="1262">Referred specimens having only morphological identification:</emphasis>
<specimenCode id="5C772739FFBFFFDDACDC7D64FB397135" box="[1008,1167,352,374]" pageId="13" pageNumber="1262">TMSA 49116</specimenCode>
(
<specimenCode id="5C772739FFBFFFDDAB8D7D64FAB67135" box="[1185,1280,352,374]" pageId="13" pageNumber="1262">JAG 31</specimenCode>
), adult
<specimenCount id="1AD744CBFFBFFFDDAA4E7D64FA297135" box="[1378,1439,352,374]" pageId="13" pageNumber="1262" type="male">male</specimenCount>
, collected on
<date id="786FA982FFBFFFDDACF07D7BFB3A71D7" box="[988,1164,382,404]" pageId="13" pageNumber="1262" value="2013-04-21">
<collectingDate id="682B506AFFBFFFDDACF07D7BFB3A71D7" box="[988,1164,382,404]" pageId="13" pageNumber="1262" value="2013-04-21">21 April 2013</collectingDate>
</date>
by
<collectorName id="A124EA94FFBFFFDDAB937D7BFAE971D7" box="[1215,1375,383,404]" pageId="13" pageNumber="1262">J. A. Guyton</collectorName>
from
<collectingCountry id="74C6CFD2FFBFFFDDAC177D99FC6071F0" box="[827,982,413,435]" name="Mozambique" pageId="13" pageNumber="1262">Mozambique</collectingCountry>
,
<collectingRegion id="CE1541A0FFBFFFDDACCF7D99FB1571F0" box="[995,1187,413,435]" country="Mozambique" name="Sofala" pageId="13" pageNumber="1262">Sofala Province</collectingRegion>
,
<collectingMunicipality id="EC0A1538FFBFFFDDAB9D7D99FCC27191" pageId="13" pageNumber="1262">Gorongosa National Park</collectingMunicipality>
,
<location id="090ED999FFBFFFDDAC517DB8FBDF7191" LSID="urn:lsid:plazi:treatment:84783E54FFBFFFC0AFDA788EFD247543:090ED999FFBFFFDDAC517DB8FBDF7191" box="[893,1129,444,466]" country="Mozambique" latitude="-18.635" longLatPrecision="76" longitude="34.808" municipality="Gorongosa National Park" name="Cheringoma Plateau" pageId="13" pageNumber="1262" stateProvince="Sofala Province">Cheringoma Plateau</location>
,
<location id="090ED999FFBFFFDDAB587DB8FB597191" LSID="urn:lsid:plazi:treatment:84783E54FFBFFFC0AFDA788EFD247543:090ED999FFBFFFDDAB587DB8FB597191" box="[1140,1263,444,466]" country="Mozambique" latitude="-18.635" longLatPrecision="76" longitude="34.808" municipality="Gorongosa National Park" name="Gorge Rim" pageId="13" pageNumber="1262" stateProvince="Sofala Province">Gorge Rim</location>
,
<location id="090ED999FFBFFFDDABD57DB8FA907191" LSID="urn:lsid:plazi:treatment:84783E54FFBFFFC0AFDA788EFD247543:090ED999FFBFFFDDABD57DB8FA907191" box="[1273,1318,444,466]" country="Mozambique" latitude="-18.635" longLatPrecision="76" longitude="34.808" municipality="Gorongosa National Park" name="Site" pageId="13" pageNumber="1262" stateProvince="Sofala Province">Site</location>
1, -
<geoCoordinate id="69E5E985FFBFFFDDAA617DB8FCFD71B3" degrees="18.635" direction="south" orientation="latitude" pageId="13" pageNumber="1262" precision="55" value="-18.635">18.635° S</geoCoordinate>
,
<geoCoordinate id="69E5E985FFBFFFDDAC747DDFFC7B71B3" box="[856,973,475,496]" degrees="34.808" direction="east" orientation="longitude" pageId="13" pageNumber="1262" precision="55" value="34.808">34.808° E</geoCoordinate>
,
<quantity id="CB2922A7FFBFFFDDACF67DDFFB9171B3" box="[986,1063,475,496]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="13" pageNumber="1262" unit="m" value="213.0">
<elevation id="87FC6871FFBFFFDDACF67DDFFB9171B3" box="[986,1063,475,496]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="13" pageNumber="1262" unit="m" value="213.0">213 m</elevation>
</quantity>
.
</materialsCitation>
<materialsCitation id="BCB9851FFFBFFFDDAB1D7DDFFB60720F" ID-GBIF-Occurrence="3126007315" collectingDate="2015-07-21" collectionCode="DM" collectorName="J. A. Guyton" country="Mozambique" elevation="1150" latitude="-18.465" location="Gorongosa National Park" longLatPrecision="76" longitude="34.052" municipality="Gorongosa National Park" pageId="13" pageNumber="1262" specimenCode="DM 14864" specimenCount="1" specimenCount-female="1" stateProvince="Sofala Province">
<emphasis id="3EA55350FFBFFFDDAB1D7DDFFB6571AC" box="[1073,1235,474,496]" italics="true" pageId="13" pageNumber="1262">
<taxonomicName id="CBD1F4C1FFBFFFDDAB1D7DDFFB2771B3" authorityName="Brito &amp; Gallo" authorityYear="2002" box="[1073,1169,475,496]" class="Actinopterygii" higherTaxonomySource="GBIF" kingdom="Animalia" order="Incertae" pageId="13" pageNumber="1262" phylum="Chordata" rank="order">Incertae</taxonomicName>
sedis
</emphasis>
:
<specimenCode id="5C772739FFBFFFDDABCE7DDFFAEB71B3" box="[1250,1373,474,496]" pageId="13" pageNumber="1262">DM 14864</specimenCode>
adult
<specimenCount id="1AD744CBFFBFFFDDAC177DFDFC3E724C" box="[827,904,505,527]" pageId="13" pageNumber="1262" type="female">female</specimenCount>
collected by
<collectorName id="A124EA94FFBFFFDDAB307DFEFB1E724D" box="[1052,1192,505,527]" pageId="13" pageNumber="1262">J. A. Guyton</collectorName>
on
<date id="786FA982FFBFFFDDABFF7DFDFAD3724C" box="[1235,1381,505,527]" pageId="13" pageNumber="1262" value="2015-07-21">
<collectingDate id="682B506AFFBFFFDDABFF7DFDFAD3724C" box="[1235,1381,505,527]" pageId="13" pageNumber="1262" value="2015-07-21">21 July 2015</collectingDate>
</date>
from
<collectingCountry id="74C6CFD2FFBFFFDDAC177E1CFC60726D" box="[827,982,536,558]" name="Mozambique" pageId="13" pageNumber="1262">Mozambique</collectingCountry>
,
<collectingRegion id="CE1541A0FFBFFFDDACCF7E1CFB15726D" box="[995,1187,536,558]" country="Mozambique" name="Sofala" pageId="13" pageNumber="1262">Sofala Province</collectingRegion>
,
<collectingMunicipality id="EC0A1538FFBFFFDDAB9D7E1CFCC2720F" pageId="13" pageNumber="1262">Gorongosa National Park</collectingMunicipality>
, -
<geoCoordinate id="69E5E985FFBFFFDDACAB7E32FC40720F" box="[903,1014,566,588]" degrees="18.465" direction="south" orientation="latitude" pageId="13" pageNumber="1262" precision="55" value="-18.465">18.465° S</geoCoordinate>
,
<geoCoordinate id="69E5E985FFBFFFDDAB2E7E32FBC5720F" box="[1026,1139,566,588]" degrees="34.052" direction="east" orientation="longitude" pageId="13" pageNumber="1262" precision="55" value="34.052">34.052° E</geoCoordinate>
,
<quantity id="CB2922A7FFBFFFDDAB537E32FB60720F" box="[1151,1238,566,588]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="13" pageNumber="1262" unit="m" value="1150.0">
<elevation id="87FC6871FFBFFFDDAB537E32FB60720F" box="[1151,1238,566,588]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="13" pageNumber="1262" unit="m" value="1150.0">1150 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="BCB9851FFFBFFFDDABCD7E33FB6C72EB" ID-GBIF-Occurrence="3126007310" collectingDate="2009-05-01" collectionCode="DM" collectorName="J. Bayliss" country="Mozambique" elevation="1478" latitude="-15.045" location="Mount Inago Forest Camp" longLatPrecision="76" longitude="37.396" pageId="13" pageNumber="1262" specimenCode="DM 11482" specimenCount="1" specimenCount-female="1" stateProvince="Nampula Province">
<specimenCode id="5C772739FFBFFFDDABCD7E33FAEC720F" box="[1249,1370,567,588]" pageId="13" pageNumber="1262">DM 11482</specimenCode>
, adult
<specimenCount id="1AD744CBFFBFFFDDAC177E51FC3A7228" box="[827,908,597,619]" pageId="13" pageNumber="1262" type="female">female</specimenCount>
collected on
<date id="786FA982FFBFFFDDAB027E51FB0B7229" box="[1070,1213,597,619]" pageId="13" pageNumber="1262" value="2009-05-01">
<collectingDate id="682B506AFFBFFFDDAB027E51FB0B7229" box="[1070,1213,597,619]" pageId="13" pageNumber="1262" value="2009-05-01">1 May 2009</collectingDate>
</date>
by
<collectorName id="A124EA94FFBFFFDDABC07E52FAD47228" box="[1260,1378,597,619]" pageId="13" pageNumber="1262">J. Bayliss</collectorName>
from
<collectingCountry id="74C6CFD2FFBFFFDDAC177E70FC6672C9" box="[827,976,628,650]" name="Mozambique" pageId="13" pageNumber="1262">Mozambique</collectingCountry>
,
<collectingRegion id="CE1541A0FFBFFFDDACF77E70FB0572C9" box="[987,1203,628,650]" country="Mozambique" name="Nampula" pageId="13" pageNumber="1262">Nampula Province</collectingRegion>
,
<location id="090ED999FFBFFFDDAB927E70FC3672EB" LSID="urn:lsid:plazi:treatment:84783E54FFBFFFC0AFDA788EFD247543:090ED999FFBFFFDDAB927E70FC3672EB" country="Mozambique" latitude="-15.045" longLatPrecision="76" longitude="37.396" name="Mount Inago Forest Camp" pageId="13" pageNumber="1262" stateProvince="Nampula Province">Mount Inago Forest Camp</location>
, -
<geoCoordinate id="69E5E985FFBFFFDDACBE7E96FC4972EB" box="[914,1023,658,680]" degrees="15.045" direction="south" orientation="latitude" pageId="13" pageNumber="1262" precision="55" value="-15.045">15.045° S</geoCoordinate>
,
<geoCoordinate id="69E5E985FFBFFFDDAB267E96FBCF72EB" box="[1034,1145,658,680]" degrees="37.396" direction="east" orientation="longitude" pageId="13" pageNumber="1262" precision="55" value="37.396">37.396° E</geoCoordinate>
<quantity id="CB2922A7FFBFFFDDAB537E97FB6072EB" box="[1151,1238,659,680]" metricMagnitude="3" metricUnit="m" metricValue="1.478" pageId="13" pageNumber="1262" unit="m" value="1478.0">
<elevation id="87FC6871FFBFFFDDAB537E97FB6072EB" box="[1151,1238,659,680]" metricMagnitude="3" metricUnit="m" metricValue="1.478" pageId="13" pageNumber="1262" unit="m" value="1478.0">1478 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="44CBDCC9FFBFFFDDAC177ED5FBB97300" pageId="13" pageNumber="1262" type="etymology">
<paragraph id="0C6E8F42FFBFFFDDAC177ED5FBB97300" blockId="13.[827,1443,721,835]" pageId="13" pageNumber="1262">
<emphasis id="3EA55350FFBFFFDDAC177ED5FC7772A5" box="[827,961,721,742]" italics="true" pageId="13" pageNumber="1262">Etymology:</emphasis>
The species derives its name from the Gorongosa district of Mozambique, in particular Gorongosa National Park, a biologically diverse region of southern Africa.
</paragraph>
</subSubSection>
<subSubSection id="44CBDCC9FFBFFFDDAC177F68FA28752F" pageId="13" pageNumber="1262" type="diagnosis">
<paragraph id="0C6E8F42FFBFFFDDAC177F68FA28752F" blockId="13.[827,1443,876,1388]" pageId="13" pageNumber="1262">
<emphasis id="3EA55350FFBFFFDDAC177F68FC0073C2" box="[827,950,876,897]" italics="true" pageId="13" pageNumber="1262">Diagnosis:</emphasis>
The species can be clearly distinguished from both
<emphasis id="3EA55350FFBFFFDDAC827F8FFBFD73E3" box="[942,1099,907,928]" italics="true" pageId="13" pageNumber="1262">
<taxonomicName id="CBD1F4C1FFBFFFDDAC827F8FFB9273E3" authorityName="Gough" authorityYear="1908" box="[942,1060,907,928]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="1262" phylum="Chordata" rank="species" sensu="stricto" species="saeinnyi">R. swinnyi</taxonomicName>
<taxonomicNameLabel id="2596EE2BFFBFFFDDAB077F8FFBFD73E3" box="[1067,1099,907,928]" pageId="13" pageNumber="1262" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="CBD1F4C1FFBFFFDDABAF7F8FFABD73DC" baseAuthorityName="TMSA" baseAuthorityYear="1325" box="[1155,1291,906,928]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="1262" phylum="Chordata" rank="species" species="rhodesiae">
<emphasis id="3EA55350FFBFFFDDABAF7F8FFABD73DC" box="[1155,1291,906,928]" italics="true" pageId="13" pageNumber="1262">R. rhodesiae</emphasis>
</taxonomicName>
on molecular grounds (
<figureCitation id="94EA93C7FFBFFFDDAC887FADFC5E73FD" box="[932,1000,937,959]" captionStart="Figure 2" captionStartId="6.[145,225,1835,1857]" captionTargetBox="[398,1168,195,1786]" captionTargetId="figure-27@6.[397,1172,195,1795]" captionTargetPageId="6" captionText="Figure 2. Bayesian tree of partial Cytb sequences, with nodal support values based on Bayesian probabilities and bootstrap values for Maximum Likelihood and Maximum Parsimony. Samples included in all analyses (molecular, morphometric and bacular) are underlined." figureDoi="http://doi.org/10.5281/zenodo.4253048" httpUri="https://zenodo.org/record/4253048/files/figure.png" pageId="13" pageNumber="1262">Figs 2</figureCitation>
,
<figureCitation id="94EA93C7FFBFFFDDACD97FADFBB473FD" box="[1013,1026,937,958]" captionStart="Figure 3" captionStartId="14.[145,223,1313,1335]" captionTargetBox="[306,1265,196,1272]" captionTargetId="figure-27@14.[305,1265,195,1273]" captionTargetPageId="14" captionText="Figure 3. Haplotype network using minimum-spanning inference method based on 667 base pairs of Cytb." figureDoi="http://doi.org/10.5281/zenodo.4253053" httpUri="https://zenodo.org/record/4253053/files/figure.png" pageId="13" pageNumber="1262">3</figureCitation>
) as well as by its smaller size (
<figureCitation id="94EA93C7FFBFFFDDAA737FADFA2A73FD" box="[1375,1436,937,959]" captionStart="Figure 4" captionStartId="14.[145,223,1848,1870]" captionTargetBox="[354,1217,1410,1808]" captionTargetId="figure-45@14.[353,1217,1409,1809]" captionTargetPageId="14" captionText="Figure 4. PCA variation for five log-transformed cranial variables for six proposed small southern Africa Rhinolophus spp. defined on molecular and morphological grounds." figureDoi="http://doi.org/10.5281/zenodo.4253055" httpUri="https://zenodo.org/record/4253055/files/figure.png" pageId="13" pageNumber="1262">Fig. 4</figureCitation>
;
<tableCitation id="4153BAF9FFBFFFDDAC177FCCFC38739E" box="[827,910,968,990]" captionStart="Table 2" captionStartId="8.[141,205,424,446]" captionTargetPageId="8" captionText="Table 2. Summary statistics for six external and three cranial variables (in mm), mass (in g) and echolocation peak frequency for museum-type specimens and series of taxa sampled in this study in the saeinnyi, simulator and landeri groups. Except in the case of type specimens and certain series where indicated, all measurements were taken by the authors of this paper" httpUri="http://table.plazi.org/id/58AEDFCAFFBAFFD8AFA17DACFD7D71BA" pageId="13" pageNumber="1262" tableUuid="58AEDFCAFFBAFFD8AFA17DACFD7D71BA">Table 2</tableCitation>
), distinct cranial shape (
<figureCitation id="94EA93C7FFBFFFDDAB897FCCFB5C739D" box="[1189,1258,968,990]" captionStart="Figure 5" captionStartId="15.[163,242,944,966]" captionTargetBox="[373,1234,196,903]" captionTargetId="figure-81@15.[371,1235,195,904]" captionTargetPageId="15" captionText="Figure 5. Biplot showing the first and second principal components from a PCA of 2D landmark data for small southern Africa Rhinolophus spp., including data from type specimens of R. saeinnyi and R. s. rhodesiae. Deformation grids illustrate cranial shape changes associated with each PC." figureDoi="http://doi.org/10.5281/zenodo.4253057" httpUri="https://zenodo.org/record/4253057/files/figure.png" pageId="13" pageNumber="1262">Figs 5</figureCitation>
,
<figureCitation id="94EA93C7FFBFFFDDABDA7FCCFAB2739D" box="[1270,1284,968,990]" captionStart="Figure 6" captionStartId="15.[163,243,1848,1870]" captionTargetBox="[164,1442,1208,1808]" captionTargetId="figure-128@15.[163,1443,1207,1809]" captionTargetPageId="15" captionText="Figure 6. Overall lateral cranial morphology of small southern African Rhinolophus taxa as illustrated by deformation grids (exaggerated 3×)." figureDoi="http://doi.org/10.5281/zenodo.4253059" httpUri="https://zenodo.org/record/4253059/files/figure.png" pageId="13" pageNumber="1262">6</figureCitation>
), echolocation call peakfrequency (
<tableCitation id="4153BAF9FFBFFFDDAB067FE2FBC873BF" box="[1066,1150,998,1020]" captionStart="Table 2" captionStartId="8.[141,205,424,446]" captionTargetPageId="8" captionText="Table 2. Summary statistics for six external and three cranial variables (in mm), mass (in g) and echolocation peak frequency for museum-type specimens and series of taxa sampled in this study in the saeinnyi, simulator and landeri groups. Except in the case of type specimens and certain series where indicated, all measurements were taken by the authors of this paper" httpUri="http://table.plazi.org/id/58AEDFCAFFBAFFD8AFA17DACFD7D71BA" pageId="13" pageNumber="1262" tableUuid="58AEDFCAFFBAFFD8AFA17DACFD7D71BA">Table 2</tableCitation>
), baculum (
<figureCitation id="94EA93C7FFBFFFDDAA2A7FE2FAFB73BF" box="[1286,1357,998,1020]" captionStart="Figure 7" captionStartId="16.[145,224,715,737]" captionTargetBox="[477,1093,196,674]" captionTargetId="figure-514@16.[476,1094,195,675]" captionTargetPageId="16" captionText="Figure 7. Biplot showing the first and second principal components from a PCA of eight bacular measurements for small southern Africa Rhinolophus taxa. Numbers indicate the position of taxa referred to in Table 5." figureDoi="http://doi.org/10.5281/zenodo.4253061" httpUri="https://zenodo.org/record/4253061/files/figure.png" pageId="13" pageNumber="1262">Figs 7</figureCitation>
,
<figureCitation id="94EA93C7FFBFFFDDAA767FE3FADE73BF" box="[1370,1384,999,1020]" captionStart="Figure 8" captionStartId="17.[163,243,1805,1827]" captionTargetBox="[326,1282,201,1765]" captionTargetId="figure-26@17.[323,1283,195,1766]" captionTargetPageId="17" captionText="Figure 8. Dorsal, lateral and ventral views (from left to right) of bacula of nine of the ten taxa as identified by PCA of bacular measurements. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4253063" httpUri="https://zenodo.org/record/4253063/files/figure.png" pageId="13" pageNumber="1262">8</figureCitation>
) and noseleaf (
<figureCitation id="94EA93C7FFBFFFDDAC817801FC417459" box="[941,1015,1029,1051]" captionStart="Figure 9" captionStartId="18.[145,223,1693,1715]" captionTargetBox="[309,1263,199,1652]" captionTargetId="figure-103@18.[305,1265,195,1653]" captionTargetPageId="18" captionText="Figure 9. Noseleaf structure of representatives of various Rhinolophus taxa (not drawn to scale)." figureDoi="http://doi.org/10.5281/zenodo.4253067" httpUri="https://zenodo.org/record/4253067/files/figure.png" pageId="13" pageNumber="1262">Figs 9</figureCitation>
,
<figureCitation id="94EA93C7FFBFFFDDAB2A7801FB947459" box="[1030,1058,1029,1050]" captionStart="Figure 12" captionStartId="21.[163,241,1848,1870]" captionTargetBox="[404,1203,864,1809]" captionTargetId="figure-355@21.[403,1203,863,1809]" captionTargetPageId="21" captionText="Figure 12. Portraits of: A, the lateral facial profile showing the connecting process (marked by arrow); B, general noseleaf morphology; and, C, in flight behaviour of Rhinolophus gorongosae sp. nov. (Photographs by P. Naskrecki.)" figureDoi="http://doi.org/10.5281/zenodo.4253073" httpUri="https://zenodo.org/record/4253073/files/figure.png" pageId="13" pageNumber="1262">12</figureCitation>
) characteristics. Although some measurements overlap, there is minimal overlap in condylocanine skull length and zygomatic skull width between this species (14.815.2 mm; 8.138.56 mm) and
<taxonomicName id="CBD1F4C1FFBFFFDDAC5C7884FC5874D6" authorityName="Gough" authorityYear="1908" box="[880,1006,1152,1173]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="1262" phylum="Chordata" rank="species" species="saeinnyi">
<emphasis id="3EA55350FFBFFFDDAC5C7884FC3474D6" box="[880,898,1152,1173]" italics="true" pageId="13" pageNumber="1262">R</emphasis>
.
<emphasis id="3EA55350FFBFFFDDACBE7884FC5874D6" box="[914,1006,1152,1173]" italics="true" pageId="13" pageNumber="1262">swinnyi</emphasis>
</taxonomicName>
(14.316.0 mm; 8.79.2 mm) and
<taxonomicName id="CBD1F4C1FFBFFFDDAAA67884FC1174F0" baseAuthorityName="TMSA" baseAuthorityYear="1325" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="1262" phylum="Chordata" rank="species" species="rhodesiae">
<emphasis id="3EA55350FFBFFFDDAAA67884FA2A74D6" box="[1418,1436,1152,1173]" italics="true" pageId="13" pageNumber="1262">R</emphasis>
.
<emphasis id="3EA55350FFBFFFDDAC17789AFC1174F0" box="[827,935,1182,1203]" italics="true" pageId="13" pageNumber="1262">rhodesiae</emphasis>
</taxonomicName>
(15.115.7; 8.468.96 mm). The small size of this form makes it even smaller than
<taxonomicName id="CBD1F4C1FFBFFFDDABDE78B9FA9A7491" baseAuthorityName="Monadjem et al." baseAuthorityYear="2010" box="[1266,1324,1213,1234]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="1262" phylum="Chordata" rank="species" species="denti">
<emphasis id="3EA55350FFBFFFDDABDE78B9FA9A7491" box="[1266,1324,1213,1234]" italics="true" pageId="13" pageNumber="1262">denti</emphasis>
</taxonomicName>
(regarded by
<bibRefCitation id="6840F2B3FFBFFFDDAC7378D8FB8A74B2" author="Csorba G &amp; Ujhelyip P &amp; Thomas N" box="[863,1084,1244,1266]" journalOrPublisher="Shropshire: Alana books" pageId="13" pageNumber="1262" refId="ref17499" refString="Csorba G, Ujhelyip P, Thomas N. 2003. Horseshoe bats of the aeorld (Chiroptera: Rhinolophidae). Shropshire: Alana books." title="Horseshoe bats of the aeorld (Chiroptera: Rhinolophidae)" type="book" year="2003">
Csorba
<emphasis id="3EA55350FFBFFFDDAC9678D8FC4374B2" box="[954,1013,1244,1265]" italics="true" pageId="13" pageNumber="1262">et al.</emphasis>
, 2003
</bibRefCitation>
as the smallest species in the Ethiopian region), therefore making this new species Africas smallest horseshoe bat. Comparing means for
<taxonomicName id="CBD1F4C1FFBFFFDDAAA7791EFC02750E" authority="Taylor &amp; Macdonald &amp; Goodman &amp; Kearney &amp; Cotterill &amp; Stoffberg &amp; Monadjem &amp; Schoeman &amp; Guyton &amp; Richards, 2018" authorityName="Taylor &amp; Macdonald &amp; Goodman &amp; Kearney &amp; Cotterill &amp; Stoffberg &amp; Monadjem &amp; Schoeman &amp; Guyton &amp; Richards" authorityYear="2018" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="1262" phylum="Chordata" rank="species" species="gorongosae" status="sp. nov.">
<emphasis id="3EA55350FFBFFFDDAAA7791EFA2B756C" box="[1419,1437,1306,1327]" italics="true" pageId="13" pageNumber="1262">R</emphasis>
.
<emphasis id="3EA55350FFBFFFDDAC17793CFC02750E" box="[827,948,1336,1357]" italics="true" pageId="13" pageNumber="1262">gorongosae</emphasis>
</taxonomicName>
<emphasis id="3EA55350FFBFFFDDAC95793CFBA4750E" bold="true" box="[953,1042,1336,1357]" pageId="13" pageNumber="1262">
<taxonomicNameLabel id="2596EE2BFFBFFFDDAC95793CFBA4750E" box="[953,1042,1336,1357]" pageId="13" pageNumber="1262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<tableCitation id="4153BAF9FFBFFFDDAB33793CFBD8750E" box="[1055,1134,1336,1358]" captionStart="Table 2" captionStartId="8.[141,205,424,446]" captionTargetPageId="8" captionText="Table 2. Summary statistics for six external and three cranial variables (in mm), mass (in g) and echolocation peak frequency for museum-type specimens and series of taxa sampled in this study in the saeinnyi, simulator and landeri groups. Except in the case of type specimens and certain series where indicated, all measurements were taken by the authors of this paper" httpUri="http://table.plazi.org/id/58AEDFCAFFBAFFD8AFA17DACFD7D71BA" pageId="13" pageNumber="1262" tableUuid="58AEDFCAFFBAFFD8AFA17DACFD7D71BA">Table 2</tableCitation>
) and
<taxonomicName id="CBD1F4C1FFBFFFDDAB86793CFCCF752F" authority="(Monadjem et al., 2010)" baseAuthorityName="Monadjem et al." baseAuthorityYear="2010" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="1262" phylum="Chordata" rank="species" species="denti">
<emphasis id="3EA55350FFBFFFDDAB86793CFB54750E" box="[1194,1250,1336,1357]" italics="true" pageId="13" pageNumber="1262">denti</emphasis>
(
<bibRefCitation id="6840F2B3FFBFFFDDABC3793CFCC67528" author="Monadjem A &amp; Taylor PJ &amp; Cotterill FPD &amp; Schoeman MC" journalOrPublisher="Johannesburg: Wits University Press" pageId="13" pageNumber="1262" refId="ref18903" refString="Monadjem A, Taylor PJ, Cotterill FPD, Schoeman MC. 2010. Bats of Southern and South-central Africa: a biogeographic and taxonomic synthesis. Johannesburg: Wits University Press." title="Bats of Southern and South-central Africa: a biogeographic and taxonomic synthesis" type="book" year="2010">
Monadjem
<emphasis id="3EA55350FFBFFFDDAA45793CFA2B750E" box="[1385,1437,1336,1357]" italics="true" pageId="13" pageNumber="1262">et al.</emphasis>
, 2010
</bibRefCitation>
)
</taxonomicName>
: forearm 41.3 mm cf. 43.1 mm; mass 5.6 g cf. 7.0 g.
</paragraph>
</subSubSection>
<subSubSection id="44CBDCC9FFBFFFC0AC177992FE4D73F0" lastPageId="16" lastPageNumber="1265" pageId="13" pageNumber="1262" type="description">
<paragraph id="0C6E8F42FFBFFFC0AC177992FE4D73F0" blockId="13.[827,1443,1429,1880]" lastBlockId="16.[145,761,802,947]" lastPageId="16" lastPageNumber="1265" pageId="13" pageNumber="1262">
<emphasis id="3EA55350FFBFFFDDAC177992FC7575E8" box="[827,963,1430,1451]" italics="true" pageId="13" pageNumber="1262">Description:</emphasis>
The genetically-distinct
<taxonomicName id="CBD1F4C1FFBFFFDDABF07992FACE75E8" authority="Taylor &amp; Macdonald &amp; Goodman &amp; Kearney &amp; Cotterill &amp; Stoffberg &amp; Monadjem &amp; Schoeman &amp; Guyton &amp; Richards, 2018" authorityName="Taylor &amp; Macdonald &amp; Goodman &amp; Kearney &amp; Cotterill &amp; Stoffberg &amp; Monadjem &amp; Schoeman &amp; Guyton &amp; Richards" authorityYear="2018" box="[1244,1400,1430,1451]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="1262" phylum="Chordata" rank="species" species="gorongosae" status="sp. nov.">
<emphasis id="3EA55350FFBFFFDDABF07992FACE75E8" box="[1244,1400,1430,1451]" italics="true" pageId="13" pageNumber="1262">R. gorongosae</emphasis>
</taxonomicName>
<emphasis id="3EA55350FFBFFFDDAAAC7991FCD9758A" bold="true" pageId="13" pageNumber="1262">
<taxonomicNameLabel id="2596EE2BFFBFFFDDAAAC7991FCD9758A" pageId="13" pageNumber="1262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar in pelage colour but distinctly smaller in external (mean total length 68 mm, mean forearm length = 41 mm) and cranial (mean condylocanine length 15.1 mm) measurements (
<tableCitation id="4153BAF9FFBFFFDDABEA7A14FAA87666" box="[1222,1310,1552,1574]" captionStart="Table 2" captionStartId="8.[141,205,424,446]" captionTargetPageId="8" captionText="Table 2. Summary statistics for six external and three cranial variables (in mm), mass (in g) and echolocation peak frequency for museum-type specimens and series of taxa sampled in this study in the saeinnyi, simulator and landeri groups. Except in the case of type specimens and certain series where indicated, all measurements were taken by the authors of this paper" httpUri="http://table.plazi.org/id/58AEDFCAFFBAFFD8AFA17DACFD7D71BA" pageId="13" pageNumber="1262" tableUuid="58AEDFCAFFBAFFD8AFA17DACFD7D71BA">Table 2</tableCitation>
) than both
<taxonomicName id="CBD1F4C1FFBFFFDDAC177A2AFC1C7600" baseAuthorityName="TMSA" baseAuthorityYear="1325" box="[827,938,1582,1603]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="1262" phylum="Chordata" rank="species" species="rhodesiae">
<emphasis id="3EA55350FFBFFFDDAC177A2AFC1C7600" box="[827,938,1582,1603]" italics="true" pageId="13" pageNumber="1262">rhodesiae</emphasis>
</taxonomicName>
(mean totallength 75 mm, forearmlength 44 mm, condyolocanine length 15.4 mm) and
<emphasis id="3EA55350FFBFFFDDAA0A7A4AFA157620" box="[1318,1443,1614,1635]" italics="true" pageId="13" pageNumber="1262">
<taxonomicName id="CBD1F4C1FFBFFFDDAA0A7A4AFACB7620" authorityName="Gough" authorityYear="1908" box="[1318,1405,1614,1635]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="1262" phylum="Chordata" rank="species" sensu="stricto" species="saeinnyi">swinnyi</taxonomicName>
<taxonomicNameLabel id="2596EE2BFFBFFFDDAAAE7A4AFA157620" box="[1410,1443,1614,1635]" pageId="13" pageNumber="1262" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
from South Africa (mean totallength 73 mm, forearm length 44 mm, condyolocanine length 15.4 mm). Based on geometric morphometric results, the diminutive
<taxonomicName id="CBD1F4C1FFBFFFDDAC177ACCFC65769E" authority="Taylor &amp; Macdonald &amp; Goodman &amp; Kearney &amp; Cotterill &amp; Stoffberg &amp; Monadjem &amp; Schoeman &amp; Guyton &amp; Richards, 2018" authorityName="Taylor &amp; Macdonald &amp; Goodman &amp; Kearney &amp; Cotterill &amp; Stoffberg &amp; Monadjem &amp; Schoeman &amp; Guyton &amp; Richards" authorityYear="2018" box="[827,979,1736,1757]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="1262" phylum="Chordata" rank="species" species="gorongosae" status="sp. nov.">
<emphasis id="3EA55350FFBFFFDDAC177ACCFC65769E" box="[827,979,1736,1757]" italics="true" pageId="13" pageNumber="1262">R. gorongosae</emphasis>
</taxonomicName>
<emphasis id="3EA55350FFBFFFDDACF57ACCFB85769E" bold="true" box="[985,1075,1736,1757]" pageId="13" pageNumber="1262">
<taxonomicNameLabel id="2596EE2BFFBFFFDDACF57ACCFB85769E" box="[985,1075,1736,1757]" pageId="13" pageNumber="1262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a reduced foramen magnum, a noticeable depression along the parietal region, low set bullae, a narrow braincase and a narrow and high set nasal inflation with a sharp slope from the nasal inflation to the maxillae (
<figureCitation id="94EA93C7FFBFFFDDAB7C7B46FB3D771B" box="[1104,1163,1858,1880]" captionStart="Figure 6" captionStartId="15.[163,243,1848,1870]" captionTargetBox="[164,1442,1208,1808]" captionTargetId="figure-128@15.[163,1443,1207,1809]" captionTargetPageId="15" captionText="Figure 6. Overall lateral cranial morphology of small southern African Rhinolophus taxa as illustrated by deformation grids (exaggerated 3×)." figureDoi="http://doi.org/10.5281/zenodo.4253059" httpUri="https://zenodo.org/record/4253059/files/figure.png" pageId="13" pageNumber="1262">Fig. 6</figureCitation>
). Lancet is subtriangular with straight to slightly concave sides, and a high, rounded connecting process; the sella has a diminutive, pointed tip (
<figureCitation id="94EA93C7FFBDFFDFAE007884FEC174D6" box="[300,375,1152,1174]" captionStart="Figure 12" captionStartId="21.[163,241,1848,1870]" captionTargetBox="[404,1203,864,1809]" captionTargetId="figure-355@21.[403,1203,863,1809]" captionTargetPageId="21" captionText="Figure 12. Portraits of: A, the lateral facial profile showing the connecting process (marked by arrow); B, general noseleaf morphology; and, C, in flight behaviour of Rhinolophus gorongosae sp. nov. (Photographs by P. Naskrecki.)" figureDoi="http://doi.org/10.5281/zenodo.4253073" httpUri="https://zenodo.org/record/4253073/files/figure.png" pageId="15" pageNumber="1264">Fig. 12</figureCitation>
). Baculum short with reduced base, with a characteristic notch on one side of the shaft tip (position variable). The slightly narrower baculum shaft with narrower tip distinguishes
<taxonomicName id="CBD1F4C1FFBDFFDFAB897884FA8974D6" authority="Taylor &amp; Macdonald &amp; Goodman &amp; Kearney &amp; Cotterill &amp; Stoffberg &amp; Monadjem &amp; Schoeman &amp; Guyton &amp; Richards, 2018" authorityName="Taylor &amp; Macdonald &amp; Goodman &amp; Kearney &amp; Cotterill &amp; Stoffberg &amp; Monadjem &amp; Schoeman &amp; Guyton &amp; Richards" authorityYear="2018" box="[1189,1343,1152,1173]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="1264" phylum="Chordata" rank="species" species="gorongosae" status="sp. nov.">
<emphasis id="3EA55350FFBDFFDFAB897884FA8974D6" box="[1189,1343,1152,1173]" italics="true" pageId="15" pageNumber="1264">R. gorongosae</emphasis>
</taxonomicName>
<emphasis id="3EA55350FFBDFFDFAA6A7884FA1474D6" bold="true" box="[1350,1442,1152,1173]" pageId="15" pageNumber="1264">
<taxonomicNameLabel id="2596EE2BFFBDFFDFAA6A7884FA1474D6" box="[1350,1442,1152,1173]" pageId="15" pageNumber="1264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="CBD1F4C1FFA2FFC0AFE07F27FEF7737B" authorityName="Gough" authorityYear="1908" box="[204,321,803,824]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="1265" phylum="Chordata" rank="species" species="saeinnyi">
<emphasis id="3EA55350FFA2FFC0AFE07F27FF68737B" box="[204,222,803,824]" italics="true" pageId="16" pageNumber="1265">R</emphasis>
.
<emphasis id="3EA55350FFA2FFC0AFC67F27FEF7737B" box="[234,321,803,824]" italics="true" pageId="16" pageNumber="1265">swinnyi</emphasis>
</taxonomicName>
(
<tableCitation id="4153BAF9FFA2FFC0AE7D7F26FE14737B" box="[337,418,802,824]" captionStart="Table 5" captionStartId="11.[163,227,196,217]" captionTargetPageId="11" captionText="Table 5. Bacula measurements for 33 individuals of three small African Rhinolophus taxa. Mean, standard deviation, range and sample sizes are provided. All measurements recorded in mm. Numbers in superscript refer to the position of taxa in Fig. 7" httpUri="http://table.plazi.org/id/58AEDFCAFFB9FFDBAF8F7CC0FE9A7157" pageId="16" pageNumber="1265" tableUuid="58AEDFCAFFB9FFDBAF8F7CC0FE9A7157">Table 5</tableCitation>
;
<figureCitation id="94EA93C7FFA2FFC0AE837F26FE44737B" box="[431,498,802,824]" captionStart="Figure 7" captionStartId="16.[145,224,715,737]" captionTargetBox="[477,1093,196,674]" captionTargetId="figure-514@16.[476,1094,195,675]" captionTargetPageId="16" captionText="Figure 7. Biplot showing the first and second principal components from a PCA of eight bacular measurements for small southern Africa Rhinolophus taxa. Numbers indicate the position of taxa referred to in Table 5." figureDoi="http://doi.org/10.5281/zenodo.4253061" httpUri="https://zenodo.org/record/4253061/files/figure.png" pageId="16" pageNumber="1265">Figs 7</figureCitation>
,
<figureCitation id="94EA93C7FFA2FFC0AED37F27FDBA737B" box="[511,524,803,824]" captionStart="Figure 8" captionStartId="17.[163,243,1805,1827]" captionTargetBox="[326,1282,201,1765]" captionTargetId="figure-26@17.[323,1283,195,1766]" captionTargetPageId="17" captionText="Figure 8. Dorsal, lateral and ventral views (from left to right) of bacula of nine of the ten taxa as identified by PCA of bacular measurements. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4253063" httpUri="https://zenodo.org/record/4253063/files/figure.png" pageId="16" pageNumber="1265">8</figureCitation>
). Maxillary toothrow with minute and laterally displaced PM
<superScript id="FBA4220AFFA2FFC0AD667F44FDE4730F" attach="left" box="[586,594,832,844]" fontSize="5" pageId="16" pageNumber="1265">1</superScript>
with a distinct gap between the alveolar borders of canine and PM
<superScript id="FBA4220AFFA2FFC0ADDD7F5BFD4F7328" attach="left" box="[753,761,863,875]" fontSize="5" pageId="16" pageNumber="1265">2</superScript>
(
<figureCitation id="94EA93C7FFA2FFC0AFB67F7AFF5873D7" box="[154,238,894,916]" captionStart="Figure 10" captionStartId="19.[163,243,1715,1737]" captionTargetBox="[324,1279,196,1672]" captionTargetId="figure-76@19.[323,1283,195,1675]" captionTargetPageId="19" captionText="Figure 10. Occlusional views of the maxillary toothrows of the various Rhinolophus taxa detailed in this study. Arrows indicate the position of the small anterior premolar." figureDoi="http://doi.org/10.5281/zenodo.4253069" httpUri="https://zenodo.org/record/4253069/files/figure.png" pageId="16" pageNumber="1265">Fig. 10</figureCitation>
). Mean peak echolocation CF frequency is 106 kHz (104108 kHz;
<tableCitation id="4153BAF9FFA2FFC0AEB77F99FE5A73F1" box="[411,492,925,947]" captionStart="Table 2" captionStartId="8.[141,205,424,446]" captionTargetPageId="8" captionText="Table 2. Summary statistics for six external and three cranial variables (in mm), mass (in g) and echolocation peak frequency for museum-type specimens and series of taxa sampled in this study in the saeinnyi, simulator and landeri groups. Except in the case of type specimens and certain series where indicated, all measurements were taken by the authors of this paper" httpUri="http://table.plazi.org/id/58AEDFCAFFBAFFD8AFA17DACFD7D71BA" pageId="16" pageNumber="1265" tableUuid="58AEDFCAFFBAFFD8AFA17DACFD7D71BA">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
<caption id="58AEDFCAFFBCFFDEAFBD7925FB447574" ID-DOI="http://doi.org/10.5281/zenodo.4253053" ID-Zenodo-Dep="4253053" box="[145,1266,1313,1335]" httpUri="https://zenodo.org/record/4253053/files/figure.png" pageId="14" pageNumber="1263" startId="14.[145,223,1313,1335]" targetBox="[306,1265,196,1272]" targetPageId="14">
<paragraph id="0C6E8F42FFBCFFDEAFBD7925FB447574" blockId="14.[145,1266,1313,1335]" box="[145,1266,1313,1335]" pageId="14" pageNumber="1263">
<emphasis id="3EA55350FFBCFFDEAFBD7925FF4F7575" bold="true" box="[145,249,1313,1335]" pageId="14" pageNumber="1263">Figure 3.</emphasis>
Haplotype network using minimum-spanning inference method based on 667 base pairs of
<emphasis id="3EA55350FFBCFFDEAB907925FB5A7574" box="[1212,1260,1313,1335]" italics="true" pageId="14" pageNumber="1263">Cytb</emphasis>
.
</paragraph>
</caption>
<caption id="58AEDFCAFFBCFFDEAFBD7B3CFD22772F" ID-DOI="http://doi.org/10.5281/zenodo.4253055" ID-Zenodo-Dep="4253055" httpUri="https://zenodo.org/record/4253055/files/figure.png" pageId="14" pageNumber="1263" startId="14.[145,223,1848,1870]" targetBox="[354,1217,1410,1808]" targetPageId="14">
<paragraph id="0C6E8F42FFBCFFDEAFBD7B3CFD22772F" blockId="14.[145,1425,1848,1900]" pageId="14" pageNumber="1263">
<emphasis id="3EA55350FFBCFFDEAFBD7B3CFF4E770E" bold="true" box="[145,248,1848,1870]" pageId="14" pageNumber="1263">Figure 4.</emphasis>
PCA variation for five log-transformed cranial variables for six proposed small southern Africa
<taxonomicName id="CBD1F4C1FFBCFFDEABF17B3DFAD4770C" authorityName="GORONGOSAE" authorityYear="2018" box="[1245,1378,1849,1871]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="1263" phylum="Chordata" rank="genus">
<emphasis id="3EA55350FFBCFFDEABF17B3DFAD4770C" box="[1245,1378,1849,1871]" italics="true" pageId="14" pageNumber="1263">Rhinolophus</emphasis>
</taxonomicName>
spp. defined on molecular and morphological grounds.
</paragraph>
</caption>
<caption id="58AEDFCAFFBDFFDFAF8F7FB4FD227442" ID-DOI="http://doi.org/10.5281/zenodo.4253057" ID-Zenodo-Dep="4253057" httpUri="https://zenodo.org/record/4253057/files/figure.png" pageId="15" pageNumber="1264" startId="15.[163,242,944,966]" targetBox="[373,1234,196,903]" targetPageId="15">
<paragraph id="0C6E8F42FFBDFFDFAF8F7FB4FD227442" blockId="15.[163,1443,944,1025]" pageId="15" pageNumber="1264">
<emphasis id="3EA55350FFBDFFDFAF8F7FB4FEBA7386" bold="true" box="[163,268,944,966]" pageId="15" pageNumber="1264">Figure 5.</emphasis>
Biplot showing the first and second principal components from a PCA of 2D landmark data for small southern Africa
<taxonomicName id="CBD1F4C1FFBDFFDFAFC47FC9FEDB73A0" authorityName="GORONGOSAE" authorityYear="2018" box="[232,365,973,995]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="1264" phylum="Chordata" rank="genus">
<emphasis id="3EA55350FFBDFFDFAFC47FC9FEDB73A0" box="[232,365,973,995]" italics="true" pageId="15" pageNumber="1264">Rhinolophus</emphasis>
</taxonomicName>
spp., including data from type specimens of
<taxonomicName id="CBD1F4C1FFBDFFDFAC177FCAFC1C73A0" authorityName="Gough" authorityYear="1908" box="[827,938,973,995]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="1264" phylum="Chordata" rank="species" species="saeinnyi">
<emphasis id="3EA55350FFBDFFDFAC177FCAFC1C73A0" box="[827,938,973,995]" italics="true" pageId="15" pageNumber="1264">R. swinnyi</emphasis>
</taxonomicName>
and
<emphasis id="3EA55350FFBDFFDFACF17FCAFBC773A0" box="[989,1137,973,995]" italics="true" pageId="15" pageNumber="1264">
R. s.
<taxonomicName id="CBD1F4C1FFBDFFDFAB217FC9FBC773A0" baseAuthorityName="TMSA" baseAuthorityYear="1325" box="[1037,1137,973,995]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="1264" phylum="Chordata" rank="species" species="rhodesiae">rhodesiae</taxonomicName>
</emphasis>
. Deformation grids illustrate cranial shape changes associated with each PC.
</paragraph>
</caption>
<caption id="58AEDFCAFFBDFFDFAF8F7B3CFE22772F" ID-DOI="http://doi.org/10.5281/zenodo.4253059" ID-Zenodo-Dep="4253059" httpUri="https://zenodo.org/record/4253059/files/figure.png" pageId="15" pageNumber="1264" startId="15.[163,243,1848,1870]" targetBox="[164,1442,1208,1808]" targetPageId="15">
<paragraph id="0C6E8F42FFBDFFDFAF8F7B3CFE22772F" blockId="15.[163,1443,1848,1900]" pageId="15" pageNumber="1264">
<emphasis id="3EA55350FFBDFFDFAF8F7B3CFEBB770E" bold="true" box="[163,269,1848,1870]" pageId="15" pageNumber="1264">Figure 6.</emphasis>
Overall lateral cranial morphology of small southern African
<taxonomicName id="CBD1F4C1FFBDFFDFAC847B3DFB99770C" authorityName="GORONGOSAE" authorityYear="2018" box="[936,1071,1849,1871]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="1264" phylum="Chordata" rank="genus">
<emphasis id="3EA55350FFBDFFDFAC847B3DFB99770C" box="[936,1071,1849,1871]" italics="true" pageId="15" pageNumber="1264">Rhinolophus</emphasis>
</taxonomicName>
taxa as illustrated by deformation grids (exaggerated 3×).
</paragraph>
</caption>
<caption id="58AEDFCAFFA2FFC0AFBD7ECFFBC572BD" ID-DOI="http://doi.org/10.5281/zenodo.4253061" ID-Zenodo-Dep="4253061" httpUri="https://zenodo.org/record/4253061/files/figure.png" pageId="16" pageNumber="1265" startId="16.[145,224,715,737]" targetBox="[477,1093,196,674]" targetPageId="16">
<paragraph id="0C6E8F42FFA2FFC0AFBD7ECFFBC572BD" blockId="16.[145,1425,715,766]" pageId="16" pageNumber="1265">
<emphasis id="3EA55350FFA2FFC0AFBD7ECFFF4F72A3" bold="true" box="[145,249,715,737]" pageId="16" pageNumber="1265">Figure 7.</emphasis>
Biplot showing the first and second principal components from a PCA of eight bacular measurements for small southern Africa
<emphasis id="3EA55350FFA2FFC0AE167EECFE0372BD" box="[314,437,744,766]" italics="true" pageId="16" pageNumber="1265">Rhinolophu</emphasis>
s taxa. Numbers indicate the position of taxa referred to in Table 5.
</paragraph>
</caption>
<subSubSection id="44CBDCC9FFA2FFC0AFBD7FD2FD247543" pageId="16" pageNumber="1265" type="distribution">
<paragraph id="0C6E8F42FFA2FFC0AFBD7FD2FD247543" blockId="16.[145,761,982,1280]" pageId="16" pageNumber="1265">
<emphasis id="3EA55350FFA2FFC0AFBD7FD2FE0E73A8" box="[145,440,982,1003]" italics="true" pageId="16" pageNumber="1265">Distribution and biology:</emphasis>
So far, it appears that this tiny species is restricted in its distribution to Gorongosa National Park in
<collectingCountry id="74C6CFD2FFA2FFC0AE727810FE447469" box="[350,498,1044,1066]" name="Mozambique" pageId="16" pageNumber="1265">Mozambique</collectingCountry>
(
<figureCitation id="94EA93C7FFA2FFC0AD2F7810FDE4746A" box="[515,594,1044,1066]" captionStart="Figure 11" captionStartId="20.[145,223,1121,1143]" captionTargetBox="[386,1185,196,1081]" captionTargetId="figure-438@20.[385,1185,195,1081]" captionTargetPageId="20" captionText="Figure 11. Final distribution of small Rhinolophus species classified according to the present study." figureDoi="http://doi.org/10.5281/zenodo.4253071" httpUri="https://zenodo.org/record/4253071/files/figure.png" pageId="16" pageNumber="1265">Fig. 11</figureCitation>
), although we provisionally refer a very small adult individual from Mount Inago in Mozambique to this taxon. Molecular sequences are required from a wider range of localities to determine the range of this taxon. Given that the two individuals sequenced from Gorongosa National Park were distinct from each other, the possibility exists that more than one cryptic species may be present.
</paragraph>
</subSubSection>
</treatment>
</document>