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<document ID-DOI="http://dx.doi.org/10.3897/jhr.94.94263" ID-Pensoft-Pub="1314-2607-94-13" ID-Pensoft-UUID="ADCFB8021287566FB2D7E8A8711D5CAE" ID-ZooBank="6CB807239A47403FABEC9AF8AE7F417F" ModsDocID="1314-2607-94-13" checkinTime="1671601504499" checkinUser="pensoft" docAuthor="Burks, Roger, Mitroiu, Mircea-Dan, Fusu, Lucian, Heraty, John M., Jansta, Petr, Heydon, Steve, Papilloud, Natalie Dale-Skey, Peters, Ralph S., Tselikh, Ekaterina V., Woolley, James B., van Noort, Simon, Baur, Hannes, Cruaud, Astrid, Darling, Christopher, Haas, Michael, Hanson, Paul, Krogmann, Lars &amp; Rasplus, Jean-Yves" docDate="2022" docId="7FC5FA47F47D587C85B2F418DD3B562B" docLanguage="en" docName="JourHymenoptRes 94: 13-88" docOrigin="Journal of Hymenoptera Research 94" docPubDate="2022-12-20" docSource="http://dx.doi.org/10.3897/jhr.94.94263" docTitle="Diparisca Hedqvist 1964" docType="treatment" docVersion="1" id="ADCFB8021287566FB2D7E8A8711D5CAE" lastPageNumber="13" masterDocId="ADCFB8021287566FB2D7E8A8711D5CAE" masterDocTitle="From hell's heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera)" masterLastPageNumber="88" masterPageNumber="13" pageNumber="13" updateTime="1671601504499" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>From hell's heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera)</mods:title>
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<mods:namePart>Burks, Roger</mods:namePart>
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<mods:affiliation>Department of Entomology, University of California Riverside, Riverside, CA, USA</mods:affiliation>
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<mods:namePart>Mitroiu, Mircea-Dan</mods:namePart>
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<mods:affiliation>Faculty of Biology, Alexandru Ioan Cuza University, Iasi, Romania</mods:affiliation>
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<mods:namePart>Fusu, Lucian</mods:namePart>
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<mods:affiliation>Faculty of Biology, Alexandru Ioan Cuza University, Iasi, Romania</mods:affiliation>
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<mods:namePart>Heraty, John M.</mods:namePart>
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<mods:affiliation>Department of Entomology, University of California Riverside, Riverside, CA, USA</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jansta, Petr</mods:namePart>
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<mods:affiliation>Department of Zoology, Faculty of Science, Charles University, Prague, Czech Republic &amp; Department of Entomology, State Museum of Natural History, Stuttgart, Germany</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Heydon, Steve</mods:namePart>
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<mods:namePart>Papilloud, Natalie Dale-Skey</mods:namePart>
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<mods:affiliation>Insects Division, Natural History Museum, London, UK</mods:affiliation>
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<mods:namePart>Peters, Ralph S.</mods:namePart>
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<mods:namePart>Tselikh, Ekaterina V.</mods:namePart>
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<mods:affiliation>Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia</mods:affiliation>
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<mods:namePart>Woolley, James B.</mods:namePart>
<mods:affiliation>Department of Entomology, Texas A &amp; M University, College Station, TX, USA</mods:affiliation>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>van Noort, Simon</mods:namePart>
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<mods:affiliation>Research and Exhibitions Department, South African Museum, Iziko Museums of South Africa, PO Box 61, Cape Town 8000 South Africa &amp; Department of Biological Sciences, University of Cape Town, Private Bag, Rondebosch, 7701, South Africa</mods:affiliation>
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<mods:namePart>Baur, Hannes</mods:namePart>
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<mods:affiliation>Department of Invertebrates, Natural History Museum Bern, Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, Bern, Switzerland</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cruaud, Astrid</mods:namePart>
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<mods:affiliation>CBGP, INRAE, CIRAD, IRD, Montpellier SupAgro, University of Montpellier, Montpellier, France</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Darling, Christopher</mods:namePart>
<mods:affiliation>Department of Natural History, Royal Ontario Museum, Toronto, ON, M 5 S 2 C 6, Canada &amp; Department of Ecology and Evolutionary Biology, University of Toronto, Toronto, ON, M 5 S 1 A 1, Canada</mods:affiliation>
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<mods:namePart>Haas, Michael</mods:namePart>
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<mods:affiliation>Department of Entomology, State Museum of Natural History, Stuttgart, Germany</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hanson, Paul</mods:namePart>
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<mods:namePart>Krogmann, Lars</mods:namePart>
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<mods:affiliation>Department of Entomology, State Museum of Natural History, Stuttgart, Germany &amp; Institute of Biology, Biological Systematics (190 n) University of Hohenheim, Stuttgart, Germany</mods:affiliation>
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<mods:namePart>Rasplus, Jean-Yves</mods:namePart>
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<mods:affiliation>CBGP, INRAE, CIRAD, IRD, Montpellier SupAgro, University of Montpellier, Montpellier, France</mods:affiliation>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-12-20</mods:number>
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<mods:detail type="volume">
<mods:number>94</mods:number>
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<mods:extent unit="page">
<mods:start>13</mods:start>
<mods:end>88</mods:end>
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<mods:url>http://dx.doi.org/10.3897/jhr.94.94263</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.94.94263</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-94-13</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:7FC5FA47F47D587C85B2F418DD3B562B" httpUri="http://treatment.plazi.org/id/7FC5FA47F47D587C85B2F418DD3B562B" lastPageNumber="13" pageId="0" pageNumber="13">
<subSubSection pageId="0" pageNumber="13" type="nomenclature">
<paragraph pageId="0" pageNumber="13">
<taxonomicName LSID="7FC5FA47-F47D-587C-85B2-F418DD3B562B" authority="new status" authorityName="Hedqvist" authorityYear="1964" family="Epichrysomallidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="family">Epichrysomallidae new status</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="reference_group">
<paragraph pageId="0" pageNumber="13">
<taxonomicName authorityName="Hill &amp; Riek" authorityYear="1967" family="Epichrysomallidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="subFamily" subFamily="Epichrysomallinae">Epichrysomallinae</taxonomicName>
Hill &amp; Riek, 1967. Type genus:
<taxonomicName authorityName="Girault" authorityYear="1915" class="Insecta" family="Epichrysomallidae" genus="Epichrysomalla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epichrysomalla" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">Epichrysomalla</taxonomicName>
Girault, 1915.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="diagnosis">
<paragraph pageId="0" pageNumber="13">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="13">
Antenna with 10-12 flagellomeres, including a small 4th clavomere. Eyes not ventrally divergent. Labrum hidden behind clypeus, flexible. Mandibles with 3 teeth. Subforaminal bridge with postgenal bridge separating secondary posterior tentorial pits from hypostoma. Notauli complete. Mesoscutellum with frenum indicated laterally, without axillular sulcus (Fig.
<figureCitation captionStart="Figures 1318" captionStartId="F3" captionText="Figures 13 - 18. 13 Coelocyboides sp. (Coelocybidae): mesosoma in dorsal view, arrow indicating setae on or adjacent to frenal groove 14 Ormyromorpha trifasciata Girault (Coelocybidae): mesosoma in lateral view 15, 16 Lelaps sp. (Diparidae) 15 head posterior view 16 cercal brush 17 Eufroggattisca polita (Ashmead) (Epichrysomallidae): mesosoma lateral view 18 Odontofroggattia sp. (Epichrysomallidae): propodeum female dorsal view." figureDoi="10.3897/jhr.94.94263.figures13-18" httpUri="https://binary.pensoft.net/fig/781066" pageId="0" pageNumber="13">17</figureCitation>
). Mesopleural area without an expanded acropleuron; mesepimeron extending over anterior margin of metapleuron (Fig.
<figureCitation captionStart="Figures 1318" captionStartId="F3" captionText="Figures 13 - 18. 13 Coelocyboides sp. (Coelocybidae): mesosoma in dorsal view, arrow indicating setae on or adjacent to frenal groove 14 Ormyromorpha trifasciata Girault (Coelocybidae): mesosoma in lateral view 15, 16 Lelaps sp. (Diparidae) 15 head posterior view 16 cercal brush 17 Eufroggattisca polita (Ashmead) (Epichrysomallidae): mesosoma lateral view 18 Odontofroggattia sp. (Epichrysomallidae): propodeum female dorsal view." figureDoi="10.3897/jhr.94.94263.figures13-18" httpUri="https://binary.pensoft.net/fig/781066" pageId="0" pageNumber="13">17</figureCitation>
). All legs with 5 tarsomeres in most, except tarsi 4-segmented in
<taxonomicName authorityName="Ishii" authorityYear="1934" class="Insecta" family="Agaonidae" genus="Odontofroggatia" kingdom="Animalia" lsidName="Odontofroggatia" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Odontofroggatia</emphasis>
</taxonomicName>
Ishii and
<taxonomicName authorityName="Narendran" authorityYear="1994" class="Insecta" family="Agaonidae" genus="Josephiella" kingdom="Animalia" lsidName="Josephiella" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Josephiella</emphasis>
</taxonomicName>
Narendran; protibial spur stout and curved; basitarsal comb longitudinal. Metasoma with syntergum, therefore without epipygium.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="discussion">
<paragraph pageId="0" pageNumber="13">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="13">
<taxonomicName family="Epichrysomallidae" lsidName="" pageId="0" pageNumber="13" rank="family">Epichrysomallidae</taxonomicName>
mostly resemble
<taxonomicName family="Melanosomellidae" lsidName="" pageId="0" pageNumber="13" rank="family">Melanosomellidae</taxonomicName>
in habitus, but do not have a linear mesopleural sulcus. They also have different fore wing venation with a stigmal vein arising at a right angle (excepted in
<taxonomicName authorityName="Ishii" authorityYear="1934" class="Insecta" family="Eunotidae" genus="Acophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acophila" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Acophila</emphasis>
</taxonomicName>
Ishii) and a postmarginal vein that is shorter than the stigmal vein.
<taxonomicName family="Epichrysomallidae" lsidName="" pageId="0" pageNumber="13" rank="family">Epichrysomallidae</taxonomicName>
have a characteristic flap-like expansion of cuticle from the lateral edge of the propodeal spiracle, partially covering the spiracle in dorsal view (Fig.
<figureCitation captionStart="Figures 1318" captionStartId="F3" captionText="Figures 13 - 18. 13 Coelocyboides sp. (Coelocybidae): mesosoma in dorsal view, arrow indicating setae on or adjacent to frenal groove 14 Ormyromorpha trifasciata Girault (Coelocybidae): mesosoma in lateral view 15, 16 Lelaps sp. (Diparidae) 15 head posterior view 16 cercal brush 17 Eufroggattisca polita (Ashmead) (Epichrysomallidae): mesosoma lateral view 18 Odontofroggattia sp. (Epichrysomallidae): propodeum female dorsal view." figureDoi="10.3897/jhr.94.94263.figures13-18" httpUri="https://binary.pensoft.net/fig/781066" pageId="0" pageNumber="13">18</figureCitation>
) that neither
<taxonomicName family="Melanosomellidae" lsidName="" pageId="0" pageNumber="13" rank="family">Melanosomellidae</taxonomicName>
nor
<taxonomicName authorityName="Forster" authorityYear="1856" family="Ormyridae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="family">Ormyridae</taxonomicName>
have.
<taxonomicName authorityName="Forster" authorityYear="1856" family="Ormyridae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="family">Ormyridae</taxonomicName>
differ further from
<taxonomicName family="Epichrysomallidae" lsidName="" pageId="0" pageNumber="13" rank="family">Epichrysomallidae</taxonomicName>
by having a more conventional fore wing venation, with longer marginal and postmarginal veins, and iridescent coloration in most species.
</paragraph>
</subSubSection>
</treatment>
</document>