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<document id="D9278AF671B3FF4E5BDC50B1A53462D8" ID-DOI="10.5281/zenodo.176627" ID-GBIF-Dataset="7a1d86bd-1bb6-4050-adec-69d9a7ef0b71" ID-ISSN="1175-5326" ID-Zenodo-Dep="176627" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459836353796" checkinUser="plazi" docAuthor="Packer, Laurence &amp; Genaro, Julio A." docDate="2007" docId="59368781A446FFE6FF7DFAFBE338FBAF" docLanguage="en" docName="zt01468p056.pdf" docOrigin="Zootaxa 1468" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Chilicola biguttata Packer, n. sp." docType="treatment" docVersion="9" lastPageNumber="29" masterDocId="A50FFFF9A45EFFFAFFEAFF93E628FFDD" masterDocTitle="Fifteen new species of Chilicola (Hymenoptera: Apoidea; Colletidae)" masterLastPageNumber="55" masterPageNumber="1" pageNumber="25" updateTime="1698223746702" updateUser="plazi">
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<mods:titleInfo id="B600CE6682A20F4F78799EE00B249FCC">
<mods:title id="5F45D563BED395B125AF131A35A2E687">Fifteen new species of Chilicola (Hymenoptera: Apoidea; Colletidae)</mods:title>
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<mods:namePart id="49CE445BEB533D1AE9AB4775B54B114B">Packer, Laurence</mods:namePart>
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<mods:namePart id="A5F075DDBF92174A311088E115837DAB">Genaro, Julio A.</mods:namePart>
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<mods:title id="61D6706DB8FBD19E05C5DA6601F03997">Zootaxa</mods:title>
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<mods:date id="D7474615096C12718C37FCA3708FBE60">2007</mods:date>
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<mods:number id="E03E8ABC59AB240DD4D308DC8758AC72">1468</mods:number>
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<mods:classification id="2F11073691995405349A2D9AA56D2618">journal article</mods:classification>
<mods:identifier id="AC53EDBFB5BF1CC372B6705542BB46DB" type="DOI">10.5281/zenodo.176627</mods:identifier>
<mods:identifier id="9FD7EA02D8FD598007807CAF48A5307F" type="GBIF-Dataset">7a1d86bd-1bb6-4050-adec-69d9a7ef0b71</mods:identifier>
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<treatment id="59368781A446FFE6FF7DFAFBE338FBAF" ID-DOI="http://doi.org/10.5281/zenodo.6249377" ID-GBIF-Taxon="119359066" ID-Zenodo-Dep="6249377" LSID="urn:lsid:plazi:treatment:59368781A446FFE6FF7DFAFBE338FBAF" httpUri="http://treatment.plazi.org/id/59368781A446FFE6FF7DFAFBE338FBAF" lastPageId="28" lastPageNumber="29" pageId="24" pageNumber="25">
<subSubSection id="9985651CA446FFE2FF7DFAFBE71BFA74" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="D1203697A446FFE2FF7DFAFBE40DFA5F" blockId="24.[151,549,1384,1449]" box="[151,549,1384,1410]" pageId="24" pageNumber="25">
<heading id="8A6881FBA446FFE2FF7DFAFBE40DFA5F" bold="true" box="[151,549,1384,1410]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="E3EBEA85A446FFE2FF7DFAFBE40DFA5F" bold="true" box="[151,549,1384,1410]" pageId="24" pageNumber="25">
<taxonomicName id="169F4D14A446FFE2FF7DFAFBE7FEFA5F" ID-CoL="5Y35C" authority="Packer" authorityName="Packer" box="[151,470,1384,1410]" class="Insecta" family="Colletidae" genus="Chilicola" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="biguttata" status="sp. nov.">
<emphasis id="E3EBEA85A446FFE2FF7DFAFBE750FA5F" bold="true" box="[151,376,1384,1410]" italics="true" pageId="24" pageNumber="25">Chilicola biguttata</emphasis>
Packer
</taxonomicName>
,
<taxonomicNameLabel id="F8D857FEA446FFE2FE08FAFBE40DFA5F" box="[482,549,1384,1410]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D1203697A446FFE2FF7DFA1CE71BFA74" blockId="24.[151,549,1384,1449]" box="[151,307,1423,1449]" pageId="24" pageNumber="25">
(
<figureCitation id="49A42A12A446FFE2FF75FA1CE703FA74" box="[159,299,1423,1449]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="24" pageNumber="25">Figs. 9A–O</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9985651CA446FFE2FF7DFA4CE723F881" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="D1203697A446FFE2FF7DFA4CE723F881" blockId="24.[151,1436,1503,1924]" pageId="24" pageNumber="25">
<emphasis id="E3EBEA85A446FFE2FF7DFA4CE738FA24" bold="true" box="[151,272,1503,1529]" pageId="24" pageNumber="25">Diagnosis</emphasis>
: This is a highly distinctive species that appears to warrant subgeneric status (Packer in press). The large areas of yellow on the face (clypeus, lower portion of supraclypeus, lower paraocular area to antennal socket, mandible) combined with the black labrum (with small yellow spot in one specimen) is alone sufficient to separate males from all other species of
<taxonomicName id="169F4D14A446FFE2FD37F9C5E562F9B2" box="[733,842,1622,1647]" class="Insecta" family="Colletidae" genus="Chilicola" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="E3EBEA85A446FFE2FD37F9C5E562F9B2" box="[733,842,1622,1647]" italics="true" pageId="24" pageNumber="25">Chilicola</emphasis>
</taxonomicName>
: all others with considerable yellow on the lower face also have a yellow labrum. The broad unflattened gonostylus of the male is also unique (
<figureCitation id="49A42A12A446FFE2FB0AF9EEE3A6F94A" box="[1248,1422,1661,1687]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="24" pageNumber="25">Figs. 9J and K</figureCitation>
).
<taxonomicName id="169F4D14A446FFE2FF7DF937E4E1F963" authority="Michener" authorityName="Michener" box="[151,713,1700,1726]" class="Insecta" family="Colletidae" genus="Chilicola" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pedunculata" subGenus="Anoediscelis">
<emphasis id="E3EBEA85A446FFE2FF7DF937E72CF960" box="[151,260,1700,1725]" italics="true" pageId="24" pageNumber="25">Chilicola</emphasis>
(
<emphasis id="E3EBEA85A446FFE2FEFDF937E783F960" box="[279,427,1700,1725]" italics="true" pageId="24" pageNumber="25">Anoediscelis</emphasis>
)
<emphasis id="E3EBEA85A446FFE2FE55F937E478F960" box="[447,592,1700,1725]" italics="true" pageId="24" pageNumber="25">pedunculata</emphasis>
Michener
</taxonomicName>
also has the gonostyli broader than the adjacent gonocoxite but has them strongly flattened and is very different from
<taxonomicName id="169F4D14A446FFE2FCA0F95FE5F0F938" box="[842,984,1740,1765]" class="Insecta" family="Colletidae" genus="Chilicola" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="biguttata">
<emphasis id="E3EBEA85A446FFE2FCA0F95FE5F0F938" box="[842,984,1740,1765]" italics="true" pageId="24" pageNumber="25">C. biguttata</emphasis>
</taxonomicName>
in many ways. The form of the facial fovea (
<figureCitation id="49A42A12A446FFE2FF02F960E76CF8D0" box="[232,324,1779,1805]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="24" pageNumber="25">Fig. 9C</figureCitation>
), small yellow spot on the lower paraocular area and the capitate to blunt, unbranched metasomal scopal hairs of the female are unusual and readily permit separation of this new species from any other
<taxonomicName id="169F4D14A446FFE2FF7DF8D1E72CF886" box="[151,260,1858,1883]" class="Insecta" family="Colletidae" genus="Chilicola" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="E3EBEA85A446FFE2FF7DF8D1E72CF886" box="[151,260,1858,1883]" italics="true" pageId="24" pageNumber="25">Chilicola</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9985651CA446FFE6FF2DF8FAE5F1FF07" lastPageId="28" lastPageNumber="29" pageId="24" pageNumber="25" type="description">
<paragraph id="D1203697A446FFE2FF2DF8FAE26EF859" blockId="24.[151,1436,1503,1924]" box="[199,1094,1897,1924]" pageId="24" pageNumber="25">
<emphasis id="E3EBEA85A446FFE2FF2DF8FAE78DF85E" bold="true" box="[199,421,1897,1923]" pageId="24" pageNumber="25">Description. Male</emphasis>
: Length
<quantity id="16679B72A446FFE2FDFAF8F9E44CF859" box="[528,612,1898,1924]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="24" pageNumber="25" unit="mm" value="5.1">5.1mm</quantity>
, wing length
<quantity id="16679B72A446FFE2FCE9F8F9E570F859" box="[771,856,1898,1924]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="24" pageNumber="25" unit="mm" value="3.7">3.7mm</quantity>
, head width
<quantity id="16679B72A446FFE2FC06F8F9E26BF859" box="[1004,1091,1898,1924]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="24" pageNumber="25" unit="mm" value="1.3">1.3mm</quantity>
.
</paragraph>
<caption id="85E0661FA447FFE3FF7DF88BE755F80D" httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="25" pageNumber="26" targetBox="[375,1187,186,1815]" targetPageId="25">
<paragraph id="D1203697A447FFE3FF7DF88BE755F80D" blockId="25.[151,1436,1816,2000]" pageId="25" pageNumber="26">
<emphasis id="E3EBEA85A447FFE3FF7DF88BE770F8ED" bold="true" box="[151,344,1816,1840]" pageId="25" pageNumber="26">FIGURES 9A–O</emphasis>
.
<taxonomicName id="169F4D14A447FFE3FE8DF88AE4ADF8ED" authority="Packer" authorityName="Packer" box="[359,645,1817,1840]" class="Insecta" family="Colletidae" genus="Chilicola" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="biguttata" status="sp. nov.">
<emphasis id="E3EBEA85A447FFE3FE8DF88AE41BF8ED" box="[359,563,1817,1840]" italics="true" pageId="25" pageNumber="26">Chilicola biguttata</emphasis>
Packer
</taxonomicName>
,
<taxonomicNameLabel id="F8D857FEA447FFE3FD78F88BE4E7F8ED" box="[658,719,1816,1840]" pageId="25" pageNumber="26" rank="species">
<emphasis id="E3EBEA85A447FFE3FD78F88BE4E7F8ED" bold="true" box="[658,719,1816,1840]" pageId="25" pageNumber="26">n. sp.</emphasis>
</taxonomicNameLabel>
Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H–K. Male
<taxonomicName id="169F4D14A447FFE3FB35F8AAE367F88D" box="[1247,1359,1849,1872]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="25" pageNumber="52" phylum="Tracheophyta" rank="genus">terminalia</taxonomicName>
. H. S7. I. S8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L–P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O–P. Furcula in dorsal and lateral views respectively. Stippling in A–C and F indicates pale colouration, elsewhere it indicates membranous regions.
</paragraph>
</caption>
<paragraph id="D1203697A444FFE0FF2DFF0BE37DFE42" blockId="26.[151,1437,152,2016]" pageId="26" pageNumber="27">
<emphasis id="E3EBEA85A444FFE0FF2DFF0BE77EFF6C" box="[199,342,152,177]" italics="true" pageId="26" pageNumber="27">Colouration</emphasis>
: Black, with following parts yellow: mandible (except apex, red-brown), clypeus,
<emphasis id="E3EBEA85A444FFE0FAC7FF0BE502FF05" italics="true" pageId="26" pageNumber="27">supraclypeal area up to just below mid level of antennal socket</emphasis>
, paraocular area to just above mid level of antennal socket, hind tibia except for brown spot on inner and outer surfaces, hind basitarsus. Following parts pale orange: apical ring on forefemur, foretibia (except for brown spot on posterior surface), apical ring on midfemur, basal and apical rings on mid tibia, narrow apical ring on hind femur, all tarsi. Following parts dark brown: antennae, rest of legs and apical impressed regions of metasomal terga. Tegula translucent with anterior and posterior yellow spots. Wing veins brown, basal portions of costa, M+Cu and V orange-brown.
</paragraph>
<paragraph id="D1203697A444FFE0FF2DFE3DE423FC8A" blockId="26.[151,1437,152,2016]" pageId="26" pageNumber="27">
<emphasis id="E3EBEA85A444FFE0FF2DFE3DE7B2FE1A" box="[199,410,430,455]" italics="true" pageId="26" pageNumber="27">Surface Sculpture</emphasis>
: Labrum shining, punctures deep, distinct and moderately dense (i~d). Clypeus dull with punctures irregular (i=1–3d). Supraclypeus dull, punctures somewhat denser, i=1–2d. Lower paraocular area shinier, punctures more distinct (i=1–3d). Frons with dense punctures (i~0.5d) appearing almost rugulose. Vertex more finely punctate (i&lt;d). Genal area with punctures and microsculpture increasingly weak anterolaterally, shiny with longitudinal microstriation, punctures effaced. Pronotal collar with shallow but distinct punctures (i~d). Mesoscutum somewhat shiny with small but deep and dense punctures (i&lt;d). Scutellum, metanotum and mesopleuron with punctures slightly larger and sparser than on mesoscutum and microsculpture weaker, surface shinier. Dorsal surface of propodeum with few, strong, radiating striae; lateral surface shiny, weakly microsculptured, impunctate; dorsolateral area coarsely rugose. Metasomal terga with few small, shallow and obscure punctures; surface moderately shiny despite strong microsculpture; apical impressed areas with very weak microsculpture.
</paragraph>
<paragraph id="D1203697A444FFE0FF2DFCF5E53AFB9A" blockId="26.[151,1437,152,2016]" pageId="26" pageNumber="27">
<emphasis id="E3EBEA85A444FFE0FF2DFCF5E779FCA2" box="[199,337,870,895]" italics="true" pageId="26" pageNumber="27">Pubescence</emphasis>
: Whitish, short, sparse and weakly plumose, hairs especially short on mesoscutum (&lt;0.5 MOD), longest on head just above antennae (2MOD). Genal beard moderately strongly developed (2MOD posteriorly, 0.5MOD anteriorly). Lateral surface of propodeum with short (0.5MOD), sparse hairs; those of dorsolateral area slightly longer (~0.75MOD) and denser. Metasomal terga with extremely short hairs on more anterior segments, becoming somewhat longer posteriorly, lacking posterolateral patches. Metasomal sterna lacking patches of erect hair, almost entirely glabrous.
</paragraph>
<paragraph id="D1203697A444FFE0FF2DFBC5E5A8F8C5" blockId="26.[151,1437,152,2016]" pageId="26" pageNumber="27">
<emphasis id="E3EBEA85A444FFE0FF2DFBC5E7A9FBB2" box="[199,385,1110,1135]" italics="true" pageId="26" pageNumber="27">Structure: Head</emphasis>
: Slightly shorter than broad (86:88) (
<figureCitation id="49A42A12A444FFE0FCDEFBC5E5BBFBAD" box="[820,915,1110,1136]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="26" pageNumber="27">Fig. 9A</figureCitation>
). Labrum almost
<date id="A5211057A444FFE0FBB5FBC5E2ABFBAD" box="[1119,1155,1110,1136]" pageId="26" pageNumber="27">2X</date>
as broad as long (9:17); apical margin very weakly angulate medially. Mandible short and deep, length to basal depth 21:14. Clypeus slightly broader than long (29:26) (
<figureCitation id="49A42A12A444FFE0FDA5FB35E49AFB1D" box="[591,690,1190,1216]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="26" pageNumber="27">Fig. 9A</figureCitation>
), lower one quarter extending beyond lower ocular tangent, slightly protuberant in profile, with weak median longitudinal groove in upper half; lateral margin gently sinuate (
<figureCitation id="49A42A12A444FFE0FF30FB65E711FACD" box="[218,313,1270,1296]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="26" pageNumber="27">Fig. 9B</figureCitation>
). Epistomal suture beneath anterior tentorial pit expanded into a groove such that pit appears comma-shaped. Supraclypeal area strongly protruberant dorsally, length to breadth 22:16. Subantennal suture very slightly concave outwardly. Frons without marked swellings or depressions. Frontal line strongly raised for approximately lower half of distance between dorsal margin of supraclypeal area and median ocellus. Facial fovea absent. Eyes emarginate, converging below (
<figureCitation id="49A42A12A444FFE0FCD0FA05E5BEFA6D" box="[826,918,1430,1456]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="26" pageNumber="27">Fig. 9A</figureCitation>
) (UOD:LOD 51:33). Lateral ocelli closer to eye than each other (
<figureCitation id="49A42A12A444FFE0FE78FA2DE7DCFA05" box="[402,500,1470,1496]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="26" pageNumber="27">Fig. 9A</figureCitation>
); OOC:IOC 10:19. OOC 1.3X LOL. Vertex behind lateral ocellus subequal to LOL, evenly convex in anterior view. Head not strongly developed above summits of eyes (
<figureCitation id="49A42A12A444FFE0FB09FA75E36AF9DD" box="[1251,1346,1510,1536]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="26" pageNumber="27">Fig. 9A</figureCitation>
), upper ocular tangent passing through middle of median ocellus. Occipital margin sharp but not strongly carinate. Scape slightly longer than pedicel + F1 + F2 combined; F1 shorter than F2 and F3 combined on longer surface (11:13); all flagellomeres other than F1 and F11 shorter than broad (for F8 8:11) and expanded apically on anterior surface giving undulating appearance in profile; F11 similar in length to F10; flagellum becoming markedly deeper but less distinctly broader from base to apex (
<figureCitation id="49A42A12A444FFE0FC95F93DE5F6F915" box="[895,990,1710,1736]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="26" pageNumber="27">Fig. 9D</figureCitation>
). Malar space linear making presence or absence of malar suture undetectable. Gena slightly greater than 0.5X width of eye in lateral view (16:30) (
<figureCitation id="49A42A12A444FFE0FF75F96DE6D4F8C5" box="[159,252,1790,1816]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="26" pageNumber="27">Fig. 9B</figureCitation>
). Apical portion of genal area upwardly curved mesad.
</paragraph>
<paragraph id="D1203697A444FFE1FF2DF8B5E7A8FE7F" blockId="26.[151,1437,152,2016]" lastBlockId="27.[151,1437,152,2020]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<taxonomicName id="169F4D14A444FFE0FF2DF8B5E7A1F89D" authority="Less" authorityName="Less" box="[199,393,1830,1856]" class="Crustacea" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="E3EBEA85A444FFE0FF2DF8B5E76CF8E2" box="[199,324,1830,1855]" italics="true" pageId="26" pageNumber="27">Mesosoma</emphasis>
: Less
</taxonomicName>
than
<date id="A5211057A444FFE0FE26F8B5E7D8F89D" box="[460,496,1830,1856]" pageId="26" pageNumber="27">2X</date>
as long as greatest depth, 125:70. Pronotal collar short, medial length subequal to LOL, anterolateral margins rounded. Episternal groove distinct and complete; scrobal groove evident only posterior to scrobe. Propodeum less than 1.5X as long as metanotum (16:11), shorter than scutellum (22) and its own posterior depth (20); propodeal sulcus shallow, shiny, traversed by numerous striae.
<collectingCountry id="A9887607A444FFE0FB26F80DE32FF865" box="[1228,1287,1950,1976]" name="India" pageId="26" pageNumber="27">Hind</collectingCountry>
leg unmodified, lacking swellings, angles or carinae; femur almost
<date id="A5211057A444FFE0FCD0F855E576F83D" box="[826,862,1990,2016]" pageId="26" pageNumber="27">3X</date>
as long as greatest depth 90:35, ventral surface convex; tibia gradually expanding towards apex length,
<date id="A5211057A445FFE1FCC2FF0BE564FF6F" box="[808,844,152,178]" pageId="27" pageNumber="28">4X</date>
apical depth, apical one half weakly concave ventrally, attaining mid length of trochanter when folded (
<figureCitation id="49A42A12A445FFE1FCFEFF53E545FF07" box="[788,877,192,218]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="27" pageNumber="28">Fig. 9F</figureCitation>
); tibial spurs not strongly sclerotised or curved; basitarsus slightly more than
<date id="A5211057A445FFE1FE1FFF7BE431FEDF" box="[501,537,232,258]" pageId="27" pageNumber="28">4X</date>
as long as greatest depth, which is near base, gradually narrowing to apex; tarsal claws bifurcate. Basal vein evenly curved; distal stigmal perpendicular traversing second submarginal cell near middle or closer to base; stigma shorter than length of marginal cell on wing margin (35:45); stigma in marginal cell strongly convex towards apex; first recurrent vein slightly apical to first submarginal crossvein on vein Rs+M.
</paragraph>
<paragraph id="D1203697A445FFE1FF2DFE23E315FDC7" blockId="27.[151,1437,152,2020]" pageId="27" pageNumber="28">
<emphasis id="E3EBEA85A445FFE1FF2DFE23E769FE14" box="[199,321,432,457]" italics="true" pageId="27" pageNumber="28">Metasoma</emphasis>
: T1 shorter than broad (55:60). Apical impressed areas approximately one quarter as long as entire tergum. S1–S6 without modifications except S1 weakly concave with subapical obtuse angle; sternal gradulus of S2 with long posteriorly directed lateral portion, S3–S5 with tiny circular gradular marks;
</paragraph>
<paragraph id="D1203697A445FFE1FF2DFDBBE33CFC87" blockId="27.[151,1437,152,2020]" pageId="27" pageNumber="28">
<taxonomicName id="169F4D14A445FFE1FF2DFDBBE76CFD9C" box="[199,324,552,577]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="E3EBEA85A445FFE1FF2DFDBBE76CFD9C" box="[199,324,552,577]" italics="true" pageId="27" pageNumber="28">Terminalia</emphasis>
</taxonomicName>
: S7 with two pairs of lateral lobes; ventral lobe broad and membranous with outer margin concave and posterior margin curved dorsally; dorsal lobe elongate, laterally oriented, narrowing abruptly at midlength from broad base (
<figureCitation id="49A42A12A445FFE1FE0FFDEBE416FD4F" box="[485,574,632,658]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="27" pageNumber="28">Fig 9H</figureCitation>
). S8 with apical process very large, emarginate apically, broadest subapically (
<figureCitation id="49A42A12A445FFE1FF0BFD33E71DFD67" box="[225,309,672,698]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="27" pageNumber="28">
Fig.
<date id="A5211057A445FFE1FEF0FD33E71DFD67" box="[282,309,672,698]" pageId="27" pageNumber="28">9I</date>
</figureCitation>
). Gonobase with foramen very large, apicoventral process broadly concave. Volsella U-shaped with outer margin concave before apex. Inner margin of gonocoxite undulate, lacking marked angulation; ventrobasal lobe slightly less than right angular, directed mesad. G
<emphasis id="E3EBEA85A445FFE1FC5BFD63E58AFCEC" italics="true" pageId="27" pageNumber="28">onostylus very broad, clearly demarcated from gonocoxite and not flattened or markedly concave beneath</emphasis>
. Penis valve with two large membranous lobes, inner one dorsolaterally oriented, outer one partially folded over inner one (
<figureCitation id="49A42A12A445FFE1FBB3FCD3E32FFC87" box="[1113,1287,832,858]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="27" pageNumber="28">Figs. 9J and K</figureCitation>
).
</paragraph>
<paragraph id="D1203697A445FFE1FF2DFCF4E218FC5F" blockId="27.[151,1437,152,2020]" box="[199,1072,871,898]" pageId="27" pageNumber="28">
<emphasis id="E3EBEA85A445FFE1FF2DFCF4E703FC5C" bold="true" box="[199,299,871,897]" pageId="27" pageNumber="28">Female:</emphasis>
Body length
<quantity id="16679B72A445FFE1FE20FCFBE408FC5F" box="[458,544,872,898]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="27" pageNumber="28" unit="mm" value="4.6">4.6mm</quantity>
, forewing length
<quantity id="16679B72A445FFE1FD07FCFBE56AFC5F" box="[749,834,872,898]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="27" pageNumber="28" unit="mm" value="3.1">3.1mm</quantity>
, head width
<quantity id="16679B72A445FFE1FC3CFCFBE205FC5F" box="[982,1069,872,898]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="27" pageNumber="28" unit="mm" value="1.3">1.3mm</quantity>
.
</paragraph>
<paragraph id="D1203697A445FFE1FF2DFC03E2EAFC0F" blockId="27.[151,1437,152,2020]" pageId="27" pageNumber="28">
<emphasis id="E3EBEA85A445FFE1FF2DFC03E77EFC74" box="[199,342,912,937]" italics="true" pageId="27" pageNumber="28">Colouration</emphasis>
: Black,
<emphasis id="E3EBEA85A445FFE1FE53FC03E300FC74" box="[441,1320,912,937]" italics="true" pageId="27" pageNumber="28">with yellow spot on lower paraocular area just below anterior tentorial pit</emphasis>
(
<figureCitation id="49A42A12A445FFE1FAD3FC03E3BBFC77" box="[1337,1427,912,938]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="27" pageNumber="28">Fig. 9C</figureCitation>
) and basal yellow spot on all tibiae; tarsi brown; tegula brown with outer margin colourless.
</paragraph>
<paragraph id="D1203697A445FFE1FF2DFC73E28EFB97" blockId="27.[151,1437,152,2020]" pageId="27" pageNumber="28">
<emphasis id="E3EBEA85A445FFE1FF2DFC73E7B2FC24" box="[199,410,992,1017]" italics="true" pageId="27" pageNumber="28">Surface Sculpture</emphasis>
: As in male but with more strongly imbricate microsculpture on face; punctures on lower face shallower, those of frons sparser and more distinct (i~d), punctures of vertex less distinct than those of frons; pronotum with punctures obscure; mesoscutum with punctures shallower.
</paragraph>
<paragraph id="D1203697A445FFE1FF2DFBCBE48FFACF" blockId="27.[151,1437,152,2020]" pageId="27" pageNumber="28">
<emphasis id="E3EBEA85A445FFE1FF2DFBCBE779FBAC" box="[199,337,1112,1137]" italics="true" pageId="27" pageNumber="28">Pubescence</emphasis>
: As in male but hairs above antennae shorter (1MOD). Most hairs on foretarsi thick and strongly capitate, lacking branches, 1.25 MOD, (
<figureCitation id="49A42A12A445FFE1FD12FB13E574FB47" box="[760,860,1152,1178]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="27" pageNumber="28">Fig. 9G</figureCitation>
).
<collectingCountry id="A9887607A445FFE1FC9DFB13E59AFB47" box="[887,946,1152,1178]" name="India" pageId="27" pageNumber="28">Hind</collectingCountry>
femur with scopal hairs with branches towards apex on anterior surface only, &lt;1.5 MOD.
<collectingCountry id="A9887607A445FFE1FD1BFB3BE504FB1F" box="[753,812,1192,1218]" name="India" pageId="27" pageNumber="28">Hind</collectingCountry>
tibia with scopal hairs unbranched but not capitate, ~ 2MOD. Terga lacking posterolateral hair patches.
<emphasis id="E3EBEA85A445FFE1FD1AFB43E6FBFACC" italics="true" pageId="27" pageNumber="28">Scopa on S2 with comparatively sparse capitate or blunt hairs</emphasis>
&lt;3.5MOD, scopa in form of corbicula.
</paragraph>
<paragraph id="D1203697A445FFE1FF2DFAB0E317F989" blockId="27.[151,1437,152,2020]" pageId="27" pageNumber="28">
<emphasis id="E3EBEA85A445FFE1FF2DFAB0E719FAE1" box="[199,305,1315,1340]" italics="true" pageId="27" pageNumber="28">Structure</emphasis>
: Maxillary palpus unmodified, 0.5X as long as prementum. Prementum 1/5th as long as greatest breadth; fovea very large, almost entirely covering ventral surface, margin carinate. Lacinia broadly triangular, 0.33X as long as greatest breadth. Lorum elongate, 0.4X as long as cardo, parallel-sided. Rest of body as in male except for usual sexual differences and as follows: Head slightly longer than broad (85:82) (
<figureCitation id="49A42A12A445FFE1FADBFA09E3A6FA69" box="[1329,1422,1434,1460]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="27" pageNumber="28">Fig. 9C</figureCitation>
). Length and breadth of clypeus subequal (28:27); UOD:LOD 50:36. OOC:IOC 9:20. Occipital margin rounded. Facial fovea large, broad and shallow, extending ventrally almost to upper tangent of antennal socket, dull, covered in somewhat plumose, woolly, laterally directed hairs 0.75MOD. Ratio of width of gena to eye 15:27. Propodeal sulcus weakly impressed. Apical lunule of S5 more than
<date id="A5211057A445FFE1FBA5F9A9E25BF989" box="[1103,1139,1594,1620]" pageId="27" pageNumber="28">2X</date>
as broad as long.
</paragraph>
<paragraph id="D1203697A445FFE6FF2DF9F0E5F1FF07" blockId="27.[151,1437,152,2020]" lastBlockId="28.[151,1437,152,1138]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="E3EBEA85A445FFE1FF2DF9F0E7A9F9A1" box="[199,385,1635,1660]" italics="true" pageId="27" pageNumber="28">Sting apparatus</emphasis>
: As in
<figureCitation id="49A42A12A445FFE1FE33F9F1E44BF9A1" box="[473,611,1634,1660]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="27" pageNumber="28">Figs. 9L–O</figureCitation>
. Hemitergite 7 with lateral portion of marginal ridge somewhat thickened between apodemal region and close to apex, with slight obtuse angle close to base of lateral process; lateral process short and acute; lateral lamella originating slightly anterior to lateral process; medial portion of marginal ridge gently concave; spiracle separated from evenly concave posterior margin of lamina spiracularis, closer to lateral than medial portion of marginal ridge, set in depression in lamina spiracularis (
<figureCitation id="49A42A12A445FFE1FAD9F891E3A6F8C1" box="[1331,1422,1794,1820]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="27" pageNumber="28">Fig. 9L</figureCitation>
). Hemitergite 8 with anterior ridge straight and strong to close to apex, apodeme larger than plate, carina separating apodeme and plate straight (
<figureCitation id="49A42A12A445FFE1FDDAF8C1E4BCF8B1" box="[560,660,1874,1900]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="27" pageNumber="28">Fig. 9M</figureCitation>
). First valvifer with dorsal process narrow but similar in length to ventral process (
<figureCitation id="49A42A12A445FFE1FEB4F8E9E7EEF849" box="[350,454,1914,1940]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="27" pageNumber="28">Fig. 9M</figureCitation>
). Second valvifer with apodemal ridge straight, apical process poorly developed, pars articularis rounded, incisura postarticularis strongly narrowed towards pars articularis, region basal to gonostylus membranous, gonostylus converging towards apex from broad base. Sting shaft almost straight ventrally, processus muscularis not developed, processus medianus very long (
<figureCitation id="49A42A12A442FFE6FBD2FF0BE2B0FF6F" box="[1080,1176,152,178]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="28" pageNumber="29">Fig. 9N</figureCitation>
). Furcula with ventral arms broadest close to apex, crescentic in lateral view (
<figureCitation id="49A42A12A442FFE6FCF7FF53E550FF07" box="[797,888,192,218]" captionStart="FIGURES 9 A – O" captionStartId="25.[151,270,1816,1840]" captionTargetBox="[375,1187,186,1815]" captionTargetId="figure@25.[343,1238,148,1304]" captionTargetPageId="25" captionText="FIGURES 9 A – O. Chilicola biguttata Packer, n. sp. Head of male: A. frontal view, B. lateral view. C. frontal view of female head. Male: D. male antenna. E. pronotum. F. Male hind leg. G. Female foretarsus. H – K. Male terminalia. H. S 7. I. S 8. J. genital capsule lateral view, K. same. Dorsal views are to the left. L – P. Sting apparatus of female: L. hemitergite 7, M. hemitergite 8, N. First and second valvifer with gonostylus and sting shaft (latter broken in specimen), O – P. Furcula in dorsal and lateral views respectively. Stippling in A – C and F indicates pale colouration, elsewhere it indicates membranous regions." httpUri="https://zenodo.org/record/176635/files/figure.png" pageId="28" pageNumber="29">Fig. 9O</figureCitation>
and P).
</paragraph>
</subSubSection>
<subSubSection id="9985651CA442FFE6FF2DFF74E30BFCD7" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="D1203697A442FFE6FF2DFF74E30BFCD7" blockId="28.[151,1437,152,1138]" pageId="28" pageNumber="29">
<emphasis id="E3EBEA85A442FFE6FF2DFF74E788FEDC" bold="true" box="[199,416,231,257]" pageId="28" pageNumber="29">Material studied.</emphasis>
<typeStatus id="0E248835A442FFE6FE5EFF7BE409FEDF" box="[436,545,232,258]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="A9887607A442FFE6FD9CFF7BE50CFEDF" box="[630,804,232,258]" name="Argentina" pageId="28" pageNumber="29">ARGENTINA</collectingCountry>
,
<emphasis id="E3EBEA85A442FFE6FCD9FF7BE59EFEDC" box="[819,950,232,257]" italics="true" pageId="28" pageNumber="29">Catamarca</emphasis>
; Andalgala, km marker 1510 on Andalgala-Belen road;
<date id="A5211057A442FFE6FE8DFE83E7F6FEF7" box="[359,478,272,298]" pageId="28" pageNumber="29" value="1973-10-15">15.x.1973</date>
, J.L. Neff, ex
<taxonomicName id="169F4D14A442FFE6FD79FE83E5D2FEF7" box="[659,1018,272,298]" class="Magnoliopsida" family="Fabaceae" genus="Prosopis" kingdom="Animalia" order="Fabales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="chilensis">
<emphasis id="E3EBEA85A442FFE6FD79FE83E544FEF4" box="[659,876,272,297]" italics="true" pageId="28" pageNumber="29">Prosopis chilensis</emphasis>
(Fabaceae)
</taxonomicName>
; allotype female same data except
<date id="A5211057A442FFE6FF7DFEABE727FE8F" box="[151,271,312,338]" pageId="28" pageNumber="29" value="1973-10-18">18.x.1973</date>
;
<typeStatus id="0E248835A442FFE6FEC9FEABE7A0FE8F" box="[291,392,312,338]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
male same data except
<date id="A5211057A442FFE6FD2FFEABE512FE8F" box="[709,826,312,338]" pageId="28" pageNumber="29" value="1973-10-29">29.x.1973</date>
and ex
<taxonomicName id="169F4D14A442FFE6FC46FEABE364FE8F" box="[940,1356,312,338]" class="Magnoliopsida" family="Solanaceae" genus="Nicotiana" kingdom="Plantae" order="Solanales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="noctiflora">
<emphasis id="E3EBEA85A442FFE6FC46FEABE28AFE8C" box="[940,1186,312,337]" italics="true" pageId="28" pageNumber="29">Nicotiana noctiflora</emphasis>
(Solanaceae)
</taxonomicName>
; other
<typeStatus id="0E248835A442FFE6FF7DFEF3E72FFEA7" box="[151,263,352,378]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
as follows:
<emphasis id="E3EBEA85A442FFE6FE7EFEF3E436FEA4" box="[404,542,352,377]" italics="true" pageId="28" pageNumber="29">Catamarca,</emphasis>
Andalgala desert site, no date data, J.L. Neff, one male, one female;
<emphasis id="E3EBEA85A442FFE6FAAEFEF3E6E3FE7C" italics="true" pageId="28" pageNumber="29">Catamarca</emphasis>
; Cuesta de la Chilca (base),
<date id="A5211057A442FFE6FDC2FE1BE4B1FE7F" box="[552,665,392,418]" pageId="28" pageNumber="29" value="1972-11-07">7.xi.1972</date>
, J.L. Neff, ex
<taxonomicName id="169F4D14A442FFE6FCA2FE1BE208FE7C" box="[840,1056,392,417]" class="Magnoliopsida" family="Fabaceae" genus="Prosopis" kingdom="Animalia" order="Fabales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="chilensis">
<emphasis id="E3EBEA85A442FFE6FCA2FE1BE208FE7C" box="[840,1056,392,417]" italics="true" pageId="28" pageNumber="29">Prosopis chilensis</emphasis>
</taxonomicName>
, one female;
<emphasis id="E3EBEA85A442FFE6FB2FFE1BE360FE7C" box="[1221,1352,392,417]" italics="true" pageId="28" pageNumber="29">Catamarca</emphasis>
, Punta Balasto,
<quantity id="16679B72A442FFE6FF17FE23E768FE17" box="[253,320,432,458]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="29" unit="km" value="30.0">30km</quantity>
from Santa Maria,
<date id="A5211057A442FFE6FDC9FE23E48BFE17" box="[547,675,432,458]" pageId="28" pageNumber="29" value="1966-11-24">24.xi.1966</date>
, Willink coll. one female;
<emphasis id="E3EBEA85A442FFE6FC32FE23E249FE14" box="[984,1121,432,457]" italics="true" pageId="28" pageNumber="29">Catamarca,</emphasis>
San Fernando,
<date id="A5211057A442FFE6FAF2FE23E3BFFE17" box="[1304,1431,432,458]" pageId="28" pageNumber="29" value="1991-11-11">11.xi.1991</date>
, J.G. Rozen, L. Pena &amp; A. Ugarte one male;
<emphasis id="E3EBEA85A442FFE6FD5BFE4BE531FE2C" box="[689,793,472,497]" italics="true" pageId="28" pageNumber="29">La Rioja</emphasis>
; Dept. Rosario V., Penaloza, Sierra de Arganaraz, El Rocillo,
<date id="A5211057A442FFE6FF15FD93E75EFDC7" box="[255,374,512,538]" pageId="28" pageNumber="29" value="1997-10-22">22.x.1997</date>
, M.E. Irwin, F.D. Parker, S. Roig,
<geoCoordinate id="B4AB5050A442FFE6FCC8FD93E5BDFDC7" box="[802,917,512,538]" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="5" value="-31.2066">S31.2066</geoCoordinate>
,
<geoCoordinate id="B4AB5050A442FFE6FC49FD93E20AFDC7" box="[931,1058,512,538]" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="5" value="-66.7126">W66.7126</geoCoordinate>
,
<quantity id="16679B72A442FFE6FBDAFD93E2A8FDC7" box="[1072,1152,512,538]" metricMagnitude="2" metricUnit="m" metricValue="7.22376" pageId="28" pageNumber="29" unit="ft" value="2370.0">2370ft</quantity>
, one female;
<emphasis id="E3EBEA85A442FFE6FACFFD93E3BDFDC4" box="[1317,1429,512,537]" italics="true" pageId="28" pageNumber="29">San Juan</emphasis>
, Dept. Caucete,
<quantity id="16679B72A442FFE6FEB8FDBBE7BCFD9F" box="[338,404,552,578]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="28" pageNumber="29" unit="km" value="23.0">23km</quantity>
SW Bermejo,
<date id="A5211057A442FFE6FDA6FDBBE4EBFD9F" box="[588,707,552,578]" pageId="28" pageNumber="29" value="1997-10-23">23.x.1997</date>
, S31 7472W67 8752, M.E. Irwin, F.D. Parker, S. Roig, one male;
<emphasis id="E3EBEA85A442FFE6FF0AFDC3E7EFFDB4" box="[224,455,592,617]" italics="true" pageId="28" pageNumber="29">Santiago del Estero</emphasis>
; Departamento Matará, Desvio 511,
<date id="A5211057A442FFE6FC91FDC3E5E6FDB7" box="[891,974,592,618]" pageId="28" pageNumber="29" value="1928-10">x.1928</date>
, M. Gómez. The
<typeStatus id="0E248835A442FFE6FB74FDC3E32CFDB7" box="[1182,1284,592,618]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
and allotype are housed at
<collectionCode id="B78EAE52A442FFE6FED1FDEBE7AFFD4F" box="[315,391,632,658]" pageId="28" pageNumber="29">CTMI</collectionCode>
, the Cuesta de la Chilca
<typeStatus id="0E248835A442FFE6FD45FDEBE537FD4F" box="[687,799,632,658]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
are at PYU, the Punta Balasto one is at
<collectionCode id="B78EAE52A442FFE6FB15FDEBE310FD4F" box="[1279,1336,632,658]" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="28" pageNumber="29">IML</collectionCode>
, the two collected by Irwin
<emphasis id="E3EBEA85A442FFE6FE99FD33E78FFD64" box="[371,423,672,697]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
., are at Logan and the oldest collected specimen is at
<collectionCode id="B78EAE52A442FFE6FBF5FD33E253FD67" box="[1055,1147,672,698]" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="28" pageNumber="29">MACN</collectionCode>
. The locality for this latter specimen almost certainly refers to a temporary detour on a highway, rather than actually listing a site that can be found on contemporary maps: the Spanish word for “detour” used on road signs is “desvio”.
</paragraph>
</subSubSection>
<subSubSection id="9985651CA442FFE6FF2DFC84E29DFCEF" box="[199,1205,791,818]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="D1203697A442FFE6FF2DFC84E29DFCEF" blockId="28.[151,1437,152,1138]" box="[199,1205,791,818]" pageId="28" pageNumber="29">
<emphasis id="E3EBEA85A442FFE6FF2DFC84E77BFCEC" bold="true" box="[199,339,791,817]" pageId="28" pageNumber="29">Etymology.</emphasis>
The specific epithet refers to the two yellow spots on the face of females.
</paragraph>
</subSubSection>
<subSubSection id="9985651CA442FFE6FF2DFCACE338FBAF" pageId="28" pageNumber="29" type="discussion">
<paragraph id="D1203697A442FFE6FF2DFCACE714FBFF" blockId="28.[151,1437,152,1138]" pageId="28" pageNumber="29">
<emphasis id="E3EBEA85A442FFE6FF2DFCACE77DFC84" bold="true" box="[199,341,831,857]" pageId="28" pageNumber="29">Comments.</emphasis>
The yellow spots on the face of the female are found very rarely in other species of
<taxonomicName id="169F4D14A442FFE6FACDFCD3E3BCFC84" box="[1319,1428,832,857]" class="Insecta" family="Colletidae" genus="Chilicola" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="E3EBEA85A442FFE6FACDFCD3E3BCFC84" box="[1319,1428,832,857]" italics="true" pageId="28" pageNumber="29">Chilicola</emphasis>
</taxonomicName>
; they are comparatively small and obscure in some females of
<taxonomicName id="169F4D14A442FFE6FC9CFCFBE30CFC5F" authority="Michener" authorityName="Michener" box="[886,1316,872,898]" class="Insecta" family="Colletidae" genus="Chilicola" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="brzoskai" subGenus="Oroediscelis">
<emphasis id="E3EBEA85A442FFE6FC9CFCFBE5B9FC5C" box="[886,913,872,897]" italics="true" pageId="28" pageNumber="29">C.</emphasis>
(
<emphasis id="E3EBEA85A442FFE6FC49FCFBE21EFC5C" box="[931,1078,872,897]" italics="true" pageId="28" pageNumber="29">Oroediscelis</emphasis>
)
<emphasis id="E3EBEA85A442FFE6FBA2FCFBE283FC5C" box="[1096,1195,872,897]" italics="true" pageId="28" pageNumber="29">brzoskai</emphasis>
Michener
</taxonomicName>
and an as yet unidentified species of
<emphasis id="E3EBEA85A442FFE6FE39FC03E465FC74" box="[467,589,912,937]" italics="true" pageId="28" pageNumber="29">Oediscelis</emphasis>
from southern
<collectingCountry id="A9887607A442FFE6FCE8FC03E56EFC77" box="[770,838,912,938]" name="Chile" pageId="28" pageNumber="29">Chile</collectingCountry>
. Orange markings on the lower paraocular area are more common, but are associated with a generally testaceous-orange colouration on the lower face as found, for example, in
<taxonomicName id="169F4D14A442FFE6FEA4FC73E7D1FC24" box="[334,505,992,1017]" class="Insecta" family="Colletidae" genus="Chilicola" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="unicarinata">
<emphasis id="E3EBEA85A442FFE6FEA4FC73E7D1FC24" box="[334,505,992,1017]" italics="true" pageId="28" pageNumber="29">C. unicarinata</emphasis>
</taxonomicName>
described above. One
<typeStatus id="0E248835A442FFE6FCE8FC73E54FFC27" box="[770,871,992,1018]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
male has a small yellow medial spot on the base of the labrum.
</paragraph>
<paragraph id="D1203697A442FFE6FF2DFBA3E518FB97" blockId="28.[151,1437,152,1138]" box="[199,816,1072,1098]" pageId="28" pageNumber="29">This is an apparently widespread, but scarce species.</paragraph>
<paragraph id="D1203697A442FFE6FF2DFBCBE338FBAF" blockId="28.[151,1437,152,1138]" box="[199,1296,1112,1138]" pageId="28" pageNumber="29">
Some specimens bear a label stating
<taxonomicName id="169F4D14A442FFE6FD9CFBCBE553FBAC" box="[630,891,1112,1137]" class="Magnoliopsida" family="Solanaceae" genus="Oediscelis" kingdom="Plantae" order="Solanales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="bimaculata">
<emphasis id="E3EBEA85A442FFE6FD9CFBCBE553FBAC" box="[630,891,1112,1137]" italics="true" pageId="28" pageNumber="29">Oediscelis bimaculata</emphasis>
</taxonomicName>
, another Moure manuscript name.
</paragraph>
</subSubSection>
</treatment>
</document>
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