259 lines
21 KiB
XML
259 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.876.36472" ID-GBIF-Dataset="b1a5930e-246e-4123-a748-f89ebb8a9cf3" ID-PMC="PMC6761209" ID-Pensoft-Pub="1313-2970-876-1" ID-Pensoft-UUID="81BC946DC3C752BDA2FF70619869204B" ID-PubMed="31582884" ID-ZooBank="036FEB7C534C47A988AD16C5B7DCE853" ModsDocID="1313-2970-876-1" checkinTime="1568862280139" checkinUser="pensoft" docAuthor="Bouaziz-Yahiatene, Houria, Inaebnit, Thomas, Medjdoub-Bensaad, Ferroudja, Colomba, Maria Stella, Sparacio, Ignazio, Gregorini, Armando, Liberto, Fabio & Neubert, Eike" docDate="2019" docId="2E3171AAAA64539F82DC2DFCA74FE2D8" docLanguage="en" docName="ZooKeys 876: 1-26" docOrigin="ZooKeys 876" docSource="http://dx.doi.org/10.3897/zookeys.876.36472" docTitle="Cantareus Risso 1826" docType="treatment" docVersion="4" id="81BC946DC3C752BDA2FF70619869204B" lastPageNumber="6" masterDocId="81BC946DC3C752BDA2FF70619869204B" masterDocTitle="Revisited - the species of Tweeting vineyard snails, genus Cantareus Risso, 1826 (Stylommatophora, Helicidae, Helicinae, Otalini)" masterLastPageNumber="26" masterPageNumber="1" pageNumber="4" updateTime="1668167815033" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revisited - the species of Tweeting vineyard snails, genus Cantareus Risso, 1826 (Stylommatophora, Helicidae, Helicinae, Otalini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bouaziz-Yahiatene, Houria</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Inaebnit, Thomas</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Medjdoub-Bensaad, Ferroudja</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Colomba, Maria Stella</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sparacio, Ignazio</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gregorini, Armando</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liberto, Fabio</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neubert, Eike</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>876</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>26</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.876.36472</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.876.36472</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-876-1</mods:identifier>
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<mods:identifier type="ZooBank">036FEB7C534C47A988AD16C5B7DCE853</mods:identifier>
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<mods:identifier type="Pensoft-UUID">81BC946DC3C752BDA2FF70619869204B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159412661" LSID="urn:lsid:plazi:treatment:2E3171AAAA64539F82DC2DFCA74FE2D8" httpUri="http://treatment.plazi.org/id/2E3171AAAA64539F82DC2DFCA74FE2D8" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
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<subSubSection pageId="3" pageNumber="4" type="nomenclature">
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<paragraph pageId="3" pageNumber="4">
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<taxonomicName LSID="2E3171AAAA64539F82DC2DFCA74FE2D8" authority="Risso, 1826" authorityName="Risso" authorityYear="1826" class="Gastropoda" family="Helicidae" genus="Cantareus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cantareus" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">Cantareus Risso, 1826</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="reference_group">
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<paragraph pageId="3" pageNumber="4">
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Cantareus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cantareus" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">Cantareus</taxonomicName>
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Risso, 1826, Histoire naturelle des principales productions de
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<normalizedToken originalValue="l’Europe">l'Europe</normalizedToken>
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<normalizedToken originalValue="Méridionale">Meridionale</normalizedToken>
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, IV: 64.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="notes">
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<paragraph pageId="3" pageNumber="4">Notes.</paragraph>
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<paragraph lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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In Table
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<tableCitation captionStartId="T3" httpUri="http://table.plazi.org/id/C9322C97B8908160F74CBCD55C5DACA4" pageId="3" pageNumber="4" tableUuid="C9322C97B8908160F74CBCD55C5DACA4">3</tableCitation>
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, the most important character states of the shells and the genital organs of the
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<taxonomicName lsidName="" pageId="3" pageNumber="4" rank="tribe" tribe="Otalini">Otalini</taxonomicName>
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subgroup according to
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<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2014.11.014" author="Razkin, O" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="14" pageNumber="15" pagination="99 - 117" refId="B31" refString="Razkin, O, Gomez-Moliner, BJ, Prieto, CE, Martinez-Orti, A, Arrebola, R, Munoz, B, Chueca, LJ, Madeira, MJ, 2015. Molecular phylogeny of the western Palaearctic Helicoidea (Gastropoda, Stylommatophora). Molecular Phylogenetics and Evolution 83: 99 - 117" title="Molecular phylogeny of the western Palaearctic Helicoidea (Gastropoda, Stylommatophora)." url="https://doi.org/10.1016/j.ympev.2014.11.014" volume="83" year="2015">Razkin et al. (2015</bibRefCitation>
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: 108,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cantareus apertus. Syntype of Helix aperta NHMW-MO 14005, shell diameter 28.75 mm. Shell in A frontal B dorsal C lateral and D apical views E labels of the syntype lot. Photographs NHMW, x 1.5." figureDoi="10.3897/zookeys.876.36472.figure2" httpUri="https://binary.pensoft.net/fig/340653" pageId="3" pageNumber="4" tableDoi="10.3897/zookeys.876.36472.figure2">Fig. 2</figureCitation>
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) including the genera
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Cantareus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cantareus" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="4">Cantareus</emphasis>
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</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Cornu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cornu" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="4">Cornu</emphasis>
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</taxonomicName>
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, and
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Rossmaessleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rossmaessleria" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="4">Rossmaessleria</emphasis>
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</taxonomicName>
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are shown. Within the
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<taxonomicName lsidName="" pageId="3" pageNumber="4" rank="tribe" tribe="Otalini">Otalini</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Cantareus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cantareus" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="4">Cantareus</emphasis>
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</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Cornu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cornu" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="4">Cornu</emphasis>
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</taxonomicName>
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, and
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Erctella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erctella" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="4">Erctella</emphasis>
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</taxonomicName>
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share the synapomorphy of a globular to slightly conical shell, other genera in the tribe tend to have more flattened shells (character 1). In all other shell traits, there is no apomorphy that discriminates between
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Cantareus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cantareus" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="4">Cantareus</emphasis>
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</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Cornu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cornu" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="4">Cornu</emphasis>
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</taxonomicName>
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, and
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Erctella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erctella" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="4">Erctella</emphasis>
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</taxonomicName>
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on generic level. On the level of the genital organs, the three genera share the synapomophy of the simple pore connecting epiphallus and penial chamber (char
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<pageBreakToken pageId="4" pageNumber="5" start="start">acter</pageBreakToken>
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8), while
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Rossmaessleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rossmaessleria" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="5">Rossmaessleria</emphasis>
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</taxonomicName>
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shows the plesiomophic state with two functional penial papillae. In
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Cornu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cornu" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="5">Cornu</emphasis>
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</taxonomicName>
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, the flagellum is much longer than in the other genera. The phylogenetic value of this character state is not clear within the
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<taxonomicName lsidName="" pageId="4" pageNumber="5" rank="tribe" tribe="Otalini">Otalini</taxonomicName>
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, within the
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<taxonomicName lsidName="" pageId="4" pageNumber="5" rank="tribe" tribe="Helicini">Helicini</taxonomicName>
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, it is considered a plesiomorphy (
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<bibRefCitation author="Neubert, E" journalOrPublisher="Contribution to Natural History" pageId="14" pageNumber="15" pagination="1 - 200" refId="B24" refString="Neubert, E, 2014. Revision of Helix (Linnaeus, 1758) in its eastern Mediterranean distribution area, and reassignment of Helixgodetiana (Kobelt, 1878) to Maltzanella (Hesse, 1917) (Gastropoda, Pulmonata, Helicidae). Contribution to Natural History 26: 1 - 200" title="Revision of Helix (Linnaeus, 1758) in its eastern Mediterranean distribution area, and reassignment of Helixgodetiana (Kobelt, 1878) to Maltzanella (Hesse, 1917) (Gastropoda, Pulmonata, Helicidae)." volume="26" year="2014">Neubert 2014</bibRefCitation>
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). Other character states like ratios in the bursa copulatrix complex (character 10). A massive atrial stimulator can be found in
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Cantareus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cantareus" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="5">Cantareus</emphasis>
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</taxonomicName>
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and
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Cornu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cornu" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="5">Cornu</emphasis>
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</taxonomicName>
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, while in
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Erctella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erctella" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="5">Erctella</emphasis>
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</taxonomicName>
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, it is relatively small (character 11). Large and massive stimulators are found in many taxa of
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<taxonomicName family="Helicidae" lsidName="" pageId="4" pageNumber="5" rank="family">Helicidae</taxonomicName>
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, so a reduction of this system is here interpreted as an apomorphic character state.
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</paragraph>
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<paragraph pageId="4" pageNumber="5">
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<emphasis bold="true" pageId="4" pageNumber="5">Remarks.</emphasis>
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The change of the status of
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Cantareus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cantareus" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="5">Cantareus</emphasis>
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</taxonomicName>
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from a monotypic to a polytypic genus causes some nomenclatorial problems. The type species of the genus is
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix naticoides" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="naticoides">
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<emphasis italics="true" pageId="4" pageNumber="5">Helix naticoides</emphasis>
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</taxonomicName>
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Draparnaud, 1801 from France, which so far has been considered a synonym of
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix aperta" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="aperta">
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<emphasis italics="true" pageId="4" pageNumber="5">Helix aperta</emphasis>
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</taxonomicName>
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Born, 1778, with the specimen preserved in the Born collection in the NHMW as the name bearing syntype of
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<taxonomicName genus="Columbrina" lsidName="Columbrina aperta" pageId="4" pageNumber="5" rank="species" species="aperta">
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<emphasis italics="true" pageId="4" pageNumber="5">aperta</emphasis>
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</taxonomicName>
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(
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cantareus apertus. Syntype of Helix aperta NHMW-MO 14005, shell diameter 28.75 mm. Shell in A frontal B dorsal C lateral and D apical views E labels of the syntype lot. Photographs NHMW, x 1.5." figureDoi="10.3897/zookeys.876.36472.figure2" httpUri="https://binary.pensoft.net/fig/340653" pageId="4" pageNumber="5" tableDoi="10.3897/zookeys.876.36472.figure2">Fig. 2</figureCitation>
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).The origin of
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<normalizedToken originalValue="Born’s">Born's</normalizedToken>
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specimen is unknown, and there are almost no shell morphological differences to the Algerian lineage, which proved to represent a separate species (
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Combined RaxML and Bayesian tree of Cantareus, Cornu, and Erctella, using COI, 16 S, H 3, 28 S, and ITS 2." figureDoi="10.3897/zookeys.876.36472.figure1" httpUri="https://binary.pensoft.net/fig/340677" pageId="4" pageNumber="5" tableDoi="10.3897/zookeys.876.36472.figure1">Fig. 1</figureCitation>
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). The correct origin of
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<normalizedToken originalValue="Born’s">Born's</normalizedToken>
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specimen could probably be clarified genetically by applying NGS methods using shell fragments of the syntype specimen, but this is beyond the scope of this paper.
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</paragraph>
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<caption doi="10.3897/zookeys.876.36472.figure2" httpUri="https://binary.pensoft.net/fig/340653" pageId="4" pageNumber="5" start="Figure 2" startId="F2">
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<paragraph pageId="4" pageNumber="5">
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<emphasis bold="true" pageId="4" pageNumber="5">Figure 2.</emphasis>
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Cantareus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cantareus apertus" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="apertus">
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<emphasis italics="true" pageId="4" pageNumber="5">Cantareus apertus</emphasis>
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</taxonomicName>
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. Syntype of
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix aperta" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="aperta">
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<emphasis italics="true" pageId="4" pageNumber="5">Helix aperta</emphasis>
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</taxonomicName>
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NHMW-MO 14005, shell diameter 28.75 mm. Shell in
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<emphasis bold="true" pageId="4" pageNumber="5">A</emphasis>
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frontal
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<emphasis bold="true" pageId="4" pageNumber="5">B</emphasis>
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dorsal
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<emphasis bold="true" pageId="4" pageNumber="5">C</emphasis>
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lateral and
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<emphasis bold="true" pageId="4" pageNumber="5">D</emphasis>
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apical views
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<emphasis bold="true" pageId="4" pageNumber="5">E</emphasis>
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labels of the syntype lot. Photographs NHMW,
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<normalizedToken originalValue="×">x</normalizedToken>
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1.5.
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</paragraph>
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</caption>
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<paragraph lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
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Anticipating a north African origin of the syntype NHMW-MO 14005 by fixing its type locality in Algeria ends up in a chaotic rearrangement of species names in the group. For Europe, the name
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<taxonomicName genus="Columbrina" lsidName="Columbrina naticoides" pageId="4" pageNumber="5" rank="species" species="naticoides">
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<emphasis italics="true" pageId="4" pageNumber="5">naticoides</emphasis>
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</taxonomicName>
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would be reactivated with its last use as an ac
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<pageBreakToken pageId="5" pageNumber="6" start="start">cepted</pageBreakToken>
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species in 1850 (!). The north African species would then be named
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<taxonomicName genus="Columbrina" lsidName="Columbrina apertus" pageId="5" pageNumber="6" rank="species" species="apertus">
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<emphasis italics="true" pageId="5" pageNumber="6">apertus</emphasis>
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</taxonomicName>
|
||
contradicting 170 years of permanent use. By fixing the use of the name
|
||
<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix koraegaelia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="koraegaelia">
|
||
<emphasis italics="true" pageId="5" pageNumber="6">Helix koraegaelia</emphasis>
|
||
</taxonomicName>
|
||
Bourguignat in Locard, 1882, to the north African lineage, this problem is resolved, and the stability or universality of names used in zoology is guaranteed.
|
||
</paragraph>
|
||
<paragraph pageId="5" pageNumber="6">
|
||
<taxonomicName class="Gastropoda" family="Helicidae" genus="Cantareus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cantareus apertus" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="apertus">
|
||
<emphasis italics="true" pageId="5" pageNumber="6">Cantareus apertus</emphasis>
|
||
</taxonomicName>
|
||
is well known for its protective behaviour, which gave the genus its name "
|
||
<taxonomicName class="Gastropoda" family="Helicidae" genus="Cantareus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cantareus" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="5" pageNumber="6">Cantareus</emphasis>
|
||
</taxonomicName>
|
||
: the singer". Once disturbed (
|
||
<figureCitation captionStart="Figures 21–25" captionStartId="F9" captionText="Figures 21 - 25. Pictures of living specimens of Cantareus species 21 Cantareus apertus: on the left a black specimen with brown shell from Roccapalumba, Sicily, Italy, 15. XI. 2009 (Coll. F. Liberto 5532); on the right a yellow specimen with green shell from Prizzi, Sicily, Italy, 15. XI. 2009 (Coll. F. Liberto 5545) 22 Cantareus apertus, Niscemi, Sicily Italy, 22. X. 2016 23 Cantareus apertus, Niscemi, Sicily, Italy, 22. X. 2016, specimens defending themselves by emitting bubbles of slime and a series of tweeting sounds 24 Cantareus apertus, Foggia, Ordona, 41.313889 N, 15.622222 E, 12.10.2018, leg. G. Martucci (photo / collection I. Sparacio 5031 / 9) 25 Cantareus subapertus, Algeria, Parc National du Djudjura, 1700 m, 11. X. 2008 (Photographs Vela Errol)." figureDoi="10.3897/zookeys.876.36472.figures21-25" httpUri="https://binary.pensoft.net/fig/340660" pageId="5" pageNumber="6" tableDoi="10.3897/zookeys.876.36472.figures21-25">Fig. 23</figureCitation>
|
||
), the species is able to press the air in its lung cavity through the pneumostome producing a series of tweeking sounds (https://youtu.be/CWOhZWLkd4o).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |