166 lines
20 KiB
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166 lines
20 KiB
XML
<document id="8D68791F205E5915207DA0EAF0B0BEE9" ID-DOI="10.5281/zenodo.177488" ID-GBIF-Dataset="dae36369-9079-43cd-9a39-e837d69c81d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="177488" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459862853350" checkinUser="plazi" docAuthor="Shear, William A. & Leonard, William P." docDate="2007" docId="2C6887A3E03FFFA188E3FC5C8F55F80D" docLanguage="en" docName="zt01524p034.pdf" docOrigin="Zootaxa 1524" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Caseya borealis Gardner & Shelley 1989" docType="treatment" docVersion="10" lastPageNumber="31" masterDocId="D051FFDBE037FFA98874FFCB8E52FFE2" masterDocTitle="Additions to the milliped family Caseyidae. I. Caseya richarti, n. sp., and new records of previously described species in the genus Caseya Cook and Collins 1895 (Diplopoda, Chordeumatida, Caseyidae)" masterLastPageNumber="34" masterPageNumber="23" pageNumber="31" updateTime="1698226191230" updateUser="plazi">
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<mods:title id="909CA68E671B09D84A4E42D87E421ECE">Additions to the milliped family Caseyidae. I. Caseya richarti, n. sp., and new records of previously described species in the genus Caseya Cook and Collins 1895 (Diplopoda, Chordeumatida, Caseyidae)</mods:title>
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<mods:namePart id="CE708CBC714E00AD2F6D678FD22DA11D">Shear, William A.</mods:namePart>
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<mods:namePart id="06543E3D1C14D70AF0E8401B597FBFFF">Leonard, William P.</mods:namePart>
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<mods:title id="44D12616596942165D57BF64BABC8936">Zootaxa</mods:title>
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<mods:part id="85BE69FFC16E9315F27BCF5B85E3F476">
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<mods:date id="2ECDC1C65B644F00C514DC4636E3B030">2007</mods:date>
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<mods:detail id="DAE7A43303F6E0EAF3B01D9B19EBFA3F" type="volume">
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<mods:number id="5F639F8A808A8BCA105D9D1F82E6475A">1524</mods:number>
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<treatment id="2C6887A3E03FFFA188E3FC5C8F55F80D" ID-DOI="http://doi.org/10.5281/zenodo.5672582" ID-GBIF-Taxon="119368737" ID-Zenodo-Dep="5672582" LSID="urn:lsid:plazi:treatment:2C6887A3E03FFFA188E3FC5C8F55F80D" httpUri="http://treatment.plazi.org/id/2C6887A3E03FFFA188E3FC5C8F55F80D" lastPageNumber="31" pageId="8" pageNumber="31">
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<subSubSection id="ECDB653EE03FFFA188E3FC5C8CD9FC53" box="[151,651,919,945]" pageId="8" pageNumber="31" type="nomenclature">
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<paragraph id="A47E36B5E03FFFA188E3FC5C8CD9FC53" blockId="8.[151,651,919,945]" box="[151,651,919,945]" pageId="8" pageNumber="31">
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<heading id="FF3681D9E03FFFA188E3FC5C8CD9FC53" bold="true" box="[151,651,919,945]" fontSize="11" level="1" pageId="8" pageNumber="31" reason="1">
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<taxonomicName id="63C14D36E03FFFA188E3FC5C8CD9FC53" ID-CoL="RKDB" authority="Gardner & Shelley 1989" authorityName="Gardner & Shelley" authorityYear="1989" box="[151,651,919,945]" class="Diplopoda" family="Caseyidae" genus="Caseya" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="borealis">
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<emphasis id="96B5EAA7E03FFFA188E3FC5C8CD9FC53" bold="true" box="[151,651,919,945]" pageId="8" pageNumber="31">
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<emphasis id="96B5EAA7E03FFFA188E3FC5C8F07FC53" bold="true" box="[151,341,919,945]" italics="true" pageId="8" pageNumber="31">Caseya borealis</emphasis>
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<bibRefCitation id="C0504B44E03FFFA18928FC5C8CD9FC53" author="Gardner" box="[348,651,919,945]" pageId="8" pageNumber="34" refString="Gardner, M. R., & Shelley, R. M. (1989) New records, species, and genera of caseyid millipeds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan-Pacific Entomologist, 65 (2), 177 - 268." type="journal article" year="1989">Gardner & Shelley 1989</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="ECDB653EE03FFFA188E3FC138AFFF9ED" pageId="8" pageNumber="31" type="reference_group">
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<paragraph id="A47E36B5E03FFFA188E3FC138CD7FC0D" blockId="8.[151,645,984,1007]" box="[151,645,984,1007]" pageId="8" pageNumber="31">
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<treatmentCitationGroup id="84D1119BE03FFFA188E3FC138CD7FC0D" box="[151,645,984,1007]" pageId="8" pageNumber="31">
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<taxonomicName id="63C14D36E03FFFA188E3FC138CD3FC0D" ID-CoL="RKDB" authority="Gardner & Shelley 1989:231" authorityName="Gardner & Shelley" authorityPageNumber="231" authorityYear="1989" box="[151,641,984,1007]" class="Diplopoda" family="Caseyidae" genus="Caseya" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="borealis">
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<emphasis id="96B5EAA7E03FFFA188E3FC138F10FC0D" box="[151,322,984,1007]" italics="true" pageId="8" pageNumber="31">Caseya borealis</emphasis>
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<treatmentCitation id="256010A4E03FFFA1893DFC138CD3FC0D" author="Gardner" box="[329,641,984,1007]" page="231" pageId="8" pageNumber="31" year="1989">
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<bibRefCitation id="C0504B44E03FFFA1893DFC138C01FC0D" author="Gardner" box="[329,595,984,1007]" pageId="8" pageNumber="34" refString="Gardner, M. R., & Shelley, R. M. (1989) New records, species, and genera of caseyid millipeds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan-Pacific Entomologist, 65 (2), 177 - 268." type="journal article" year="1989">Gardner & Shelley 1989</bibRefCitation>
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:231
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="A47E36B5E03FFFA188E3FBDE8AFFF9ED" blockId="8.[151,1437,1045,2031]" pageId="8" pageNumber="31">
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The original localities given by
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<bibRefCitation id="C0504B44E03FFFA18A63FBDE8D02FBCD" author="Gardner" box="[535,848,1045,1071]" pageId="8" pageNumber="34" refString="Gardner, M. R., & Shelley, R. M. (1989) New records, species, and genera of caseyid millipeds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan-Pacific Entomologist, 65 (2), 177 - 268." type="journal article" year="1989">Gardner & Shelley (1989)</bibRefCitation>
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are included on the map (open symbols,
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<figureCitation id="3CFA2A30E03FFFA18D30FBDE8BD8FBCD" box="[1348,1418,1045,1071]" captionStart="FIGURE 1" captionStartId="1.[151,255,1361,1385]" captionTargetBox="[396,1190,795,1336]" captionTargetId="figure@1.[396,1190,794,1337]" captionTargetPageId="1" captionText="FIGURE 1. Map of Washington State, USA, showing distribution of Caseya richarti n. sp. (inverted triangles) and C. borealis Gardner & Shelley (filled dots are new records, open circles are literature records from Gardner and Shelley 1989)." httpUri="https://zenodo.org/record/177489/files/figure.png" pageId="8" pageNumber="31">Fig. 1</figureCitation>
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). The
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<typeStatus id="7B7A8817E03FFFA188BFFBF68EAFFBB5" box="[203,253,1085,1111]" pageId="8" pageNumber="31">type</typeStatus>
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locality is
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<quantity id="63399B50E03FFFA1890AFBF68F86FBB5" box="[382,468,1085,1111]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="8" pageNumber="31" unit="mi" value="4.0">4 miles</quantity>
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north of Shelton, Mason Co., Washington.
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<tableCitation id="E943030EE03FFFA18BBBFBF68A74FBB5" box="[975,1062,1085,1111]" captionStart="TABLE 1" captionStartId="6.[151,239,151,175]" captionText="TABLE 1. New collection records for Caseya borealis from Washington State." pageId="8" pageNumber="31">Table 1</tableCitation>
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provides the new records, all of which are based on the inclusion of at least one mature male specimen in the sample. It is a point of interest that the
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<collectingCountry id="DCD67625E03FFFA1888CFB468F3EFB45" box="[248,364,1165,1191]" name="United States of America" pageId="8" pageNumber="31">Columbia</collectingCountry>
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River appears to form a definite southern boundary to the range of this species. The most northerly Oregon record for any
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<taxonomicName id="63C14D36E03FFFA18A51FB7E8C29FB2C" ID-CoL="7NRZD" box="[549,635,1205,1230]" class="Diplopoda" family="Caseyidae" genus="Caseya" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="96B5EAA7E03FFFA18A51FB7E8C29FB2C" box="[549,635,1205,1230]" italics="true" pageId="8" pageNumber="31">Caseya</emphasis>
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</taxonomicName>
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is for
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<taxonomicName id="63C14D36E03FFFA18AA4FB7E8D20FB2C" ID-CoL="RKDS" box="[720,882,1205,1230]" class="Diplopoda" family="Caseyidae" genus="Caseya" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="megasoma">
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<emphasis id="96B5EAA7E03FFFA18AA4FB7E8D20FB2C" box="[720,882,1205,1230]" italics="true" pageId="8" pageNumber="31">C. megasoma</emphasis>
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</taxonomicName>
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in southern Tillamook Co., approximately
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<quantity id="63399B50E03FFFA18D0BFB7E8E85FB15" metricMagnitude="5" metricUnit="m" metricValue="1.1265408" pageId="8" pageNumber="31" unit="mi" value="70.0">70 miles</quantity>
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south of the river. Northeastern Oregon has been reasonably well-collected, and caseyids belonging to the genera
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<taxonomicName id="63C14D36E03FFFA1896FFACE8C24FAFD" ID-CoL="7Q67B" authority="Chamberlin 1941" authorityName="Chamberlin" authorityYear="1941" box="[283,630,1285,1311]" class="Diplopoda" family="Caseyidae" genus="Vasingtona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="96B5EAA7E03FFFA1896FFACE8FCCFAFC" box="[283,414,1285,1310]" italics="true" pageId="8" pageNumber="31">Vasingtona</emphasis>
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<bibRefCitation id="C0504B44E03FFFA189D3FACE8C24FAFD" author="Chamberlin" box="[423,630,1285,1311]" pageId="8" pageNumber="34" refString="Chamberlin, R. V. (1941) New western millipeds. Bulletin of the University of Utah Biological Series, 6, 1 - 23." type="journal article" year="1941">Chamberlin 1941</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="63C14D36E03FFFA18AF2FACE8DE4FAFD" ID-CoL="7PNGK" authority="Chamberlin 1951" authorityName="Chamberlin" authorityYear="1951" box="[646,950,1285,1311]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="96B5EAA7E03FFFA18AF2FACE8C8CFAFC" box="[646,734,1285,1310]" italics="true" pageId="8" pageNumber="31">Opiona</emphasis>
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Chamberlin 1951
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</taxonomicName>
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and
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<taxonomicName id="63C14D36E03FFFA18B86FACE8E80FAA5" ID-CoL="7PMVS" authority="Gardner and Shelley 1989" authorityName="Gardner and Shelley" authorityYear="1989" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="96B5EAA7E03FFFA18B86FACE8ACFFAFC" box="[1010,1181,1285,1310]" italics="true" pageId="8" pageNumber="31">Ochrogramma</emphasis>
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<bibRefCitation id="C0504B44E03FFFA18CD3FACE8E80FAA5" author="Gardner" pageId="8" pageNumber="34" refString="Gardner, M. R., & Shelley, R. M. (1989) New records, species, and genera of caseyid millipeds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan-Pacific Entomologist, 65 (2), 177 - 268." type="journal article" year="1989">Gardner and Shelley 1989</bibRefCitation>
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</taxonomicName>
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have all been found there. Some of the same species of these three genera are also both north and south of the
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<collectingCountry id="DCD67625E03FFFA18892FA9E8F0FFA8D" box="[230,349,1365,1391]" name="United States of America" pageId="8" pageNumber="31">Columbia</collectingCountry>
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. It appears therefore that the gap in the distribution of
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<taxonomicName id="63C14D36E03FFFA18B86FA9E8A1AFA8C" ID-CoL="7NRZD" box="[1010,1096,1365,1390]" class="Diplopoda" family="Caseyidae" genus="Caseya" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="96B5EAA7E03FFFA18B86FA9E8A1AFA8C" box="[1010,1096,1365,1390]" italics="true" pageId="8" pageNumber="31">Caseya</emphasis>
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</taxonomicName>
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species in northern Oregon, and the
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<collectingCountry id="DCD67625E03FFFA18880FAB68F3AFA75" box="[244,360,1405,1431]" name="United States of America" pageId="8" pageNumber="31">Columbia</collectingCountry>
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River as a barrier to the southerly dispersal of
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<taxonomicName id="63C14D36E03FFFA18BFBFAB68A42FA74" ID-CoL="RKDB" box="[911,1040,1405,1430]" class="Diplopoda" family="Caseyidae" genus="Caseya" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="borealis">
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<emphasis id="96B5EAA7E03FFFA18BFBFAB68A42FA74" box="[911,1040,1405,1430]" italics="true" pageId="8" pageNumber="31">C. borealis</emphasis>
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</taxonomicName>
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are real phenomena. It would not be a surprise to find
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<taxonomicName id="63C14D36E03FFFA189F2FA6E8C55FA5C" ID-CoL="RKDB" box="[390,519,1445,1470]" class="Diplopoda" family="Caseyidae" genus="Caseya" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="borealis">
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<emphasis id="96B5EAA7E03FFFA189F2FA6E8C55FA5C" box="[390,519,1445,1470]" italics="true" pageId="8" pageNumber="31">C. borealis</emphasis>
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</taxonomicName>
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on Vancouver Island;
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<taxonomicName id="63C14D36E03FFFA18B7AFA6E8A9EFA5D" ID-CoL="6SQSB" authority="Chamberlin 1951" authorityName="Chamberlin" authorityYear="1951" box="[782,1228,1445,1471]" class="Diplopoda" family="Caseyidae" genus="Opiona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="columbiana">
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<emphasis id="96B5EAA7E03FFFA18B7AFA6E8DA5FA5C" box="[782,1015,1445,1470]" italics="true" pageId="8" pageNumber="31">Opiona columbiana</emphasis>
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Chamberlin 1951
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</taxonomicName>
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has a distribution in Washington similar to that of
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<emphasis id="96B5EAA7E03FFFA18A60FA068CCFFA04" box="[532,669,1485,1510]" italics="true" pageId="8" pageNumber="31">
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<taxonomicName id="63C14D36E03FFFA18A60FA068CCBFA04" ID-CoL="RKDB" box="[532,665,1485,1510]" class="Diplopoda" family="Caseyidae" genus="Caseya" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="borealis">C. borealis</taxonomicName>
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,
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</emphasis>
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and the former species is found on Vancouver Island, the British
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<collectingCountry id="DCD67625E03FFFA188E3FA3E8F59F9ED" box="[151,267,1525,1551]" name="United States of America" pageId="8" pageNumber="31">Columbia</collectingCountry>
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mainland, the Queen Charlotte Islands and coastal Alaska (
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<bibRefCitation id="C0504B44E03FFFA18BB2FA3E8AF7F9ED" author="Shelley" box="[966,1189,1525,1551]" pageId="8" pageNumber="34" refString="Shelley, R. M., Shear, W. A., Leonard, W. P. & Ovaska, K. (2007) Diplopoda, Chordeumatida, Caseyidae, Opiona columbiana Chamberln 1951: distribution extensions into the Alexander Archipelago, Alaska, USA, Queen Charlotte Islands, British Columbia, Canada, and eastern and southern Washington State, USA; additional new records from British Columbia and Washington. Check List, 3, 14 - 17." type="journal article" year="2007">
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Shelley
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<emphasis id="96B5EAA7E03FFFA18C53FA3E8A09F9EC" box="[1063,1115,1525,1550]" italics="true" pageId="8" pageNumber="31">et al</emphasis>
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. 2007
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</bibRefCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="ECDB653EE03FFFA188B2F9D68F55F80D" pageId="8" pageNumber="31" type="description">
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<paragraph id="A47E36B5E03FFFA188B2F9D68DB7F87D" blockId="8.[151,1437,1045,2031]" pageId="8" pageNumber="31">
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Because these collections record only sexually mature individuals, they provide some hints of the life history of the species. Mature males first appear in the collections in September and October (single records in each month) become more abundant in November, December and January (7, 4 and 7 records respectively) and reach peak abundance in February (17 records) and March (10 records). Evidently some few males survive as late as June, when a single collection was made. These observations reinforce the point made earlier (
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<emphasis id="96B5EAA7E03FFFA188D5F92E8EEFF91C" box="[161,189,1765,1790]" italics="true" pageId="8" pageNumber="31">i.e</emphasis>
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.,
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<bibRefCitation id="C0504B44E03FFFA188A1F92E8FBBF91D" author="Shear" box="[213,489,1765,1791]" pageId="8" pageNumber="34" refString="Shear, W. A. & Leonard, W. P. (2003) Microlympiidae, a new milliped family from North America, and Microlympia echina, new genus and species (Diplopoda: Chordeumatida: Brannerioidea). Zootaxa, 243, 1 - 11." type="journal article" year="2003">Shear & Leonard 2003</bibRefCitation>
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) that chordeumatidan millipeds in the Pacific Northwest of North
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<collectingCountry id="DCD67625E03FFFA18D7EF92E8B3DF91D" box="[1290,1391,1765,1791]" name="United States of America" pageId="8" pageNumber="31">America</collectingCountry>
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are active and mature in the cool, wet winter, passing the warm, dry summer as juveniles, possibly aestivating deep in the soil. Specimens were collected at altitudes ranging from
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<quantity id="63399B50E03FFFA18BDBF8FE8DBBF8AD" box="[943,1001,1845,1871]" metricMagnitude="1" metricUnit="m" metricValue="1.2192" pageId="8" pageNumber="31" unit="ft" value="40.0">40 ft</quantity>
|
|
. (
|
|
<quantity id="63399B50E03FFFA18B8FF8FE8A1DF8AD" box="[1019,1103,1845,1871]" metricMagnitude="1" metricUnit="m" metricValue="1.22" pageId="8" pageNumber="31" unit="m" value="12.2">12.2 m</quantity>
|
|
) to
|
|
<quantity id="63399B50E03FFFA18C08F8FE8A80F8AD" box="[1148,1234,1845,1871]" metricMagnitude="2" metricUnit="m" metricValue="3.9441119999999996" pageId="8" pageNumber="31" unit="ft" value="1294.0">1294 ft</quantity>
|
|
. (
|
|
<quantity id="63399B50E03FFFA18C90F8FE8B15F8AD" box="[1252,1351,1845,1871]" metricMagnitude="2" metricUnit="m" metricValue="3.944" pageId="8" pageNumber="31" unit="m" value="394.4">394.4 m</quantity>
|
|
) above sea level (asl). Nearly all collections were associated with mixed forests including deciduous trees, and many were made close to streams, rivers, or other permanent sources of water.
|
|
</paragraph>
|
|
<paragraph id="A47E36B5E03FFFA188B2F8668F55F80D" blockId="8.[151,1437,1045,2031]" pageId="8" pageNumber="31">
|
|
All specimens listed in
|
|
<tableCitation id="E943030EE03FFFA1899BF8668C19F825" box="[495,587,1965,1991]" captionStart="TABLE 1" captionStartId="6.[151,239,151,175]" captionText="TABLE 1. New collection records for Caseya borealis from Washington State." pageId="8" pageNumber="31">Table 1</tableCitation>
|
|
will be deposited in the Field Museum of Natural History, Chicago (FMNH).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
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</document> |