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<document id="3BCF24A17990B9F5DED2B583175B19C3" ID-CLB-Dataset="187956" ID-DOI="10.11646/zootaxa.5205.6.1" ID-GBIF-Dataset="4b6eb79c-018f-470b-ac46-6646270ec7ee" ID-ISSN="1175-5326" ID-Zenodo-Dep="7318605" ID-ZooBank="9D1DEC25-5FA2-4D64-807E-F103C3FCB5CA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668422045351" checkinUser="plazi" docAuthor="Shear, William A. &amp; Marek, Paul E." docDate="2022" docId="03F89303FFB4FFB99AF92CD9E1B4F93D" docLanguage="en" docName="zootaxa.5205.6.1.pdf" docOrigin="Zootaxa 5205 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Maraplia schusteri Shear &amp; Marek 2022, n. sp." docType="treatment" docVersion="6" lastPageNumber="509" masterDocId="FFC1EB7BFFBCFFB19A6E2F53E208FFDC" masterDocTitle="The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea)" masterLastPageNumber="531" masterPageNumber="501" pageNumber="509" updateTime="1699411384392" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F9C6A56B739EFD718ED7F198C944E9FE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E0A51B4C2A494AA1DEBBDE581FDB70B4">
<mods:title id="7751DAB7554681BEDAA0D28BE461D1BF">The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea)</mods:title>
</mods:titleInfo>
<mods:name id="F85B7D362635E2D34CBCCD7AC905FF71" type="personal">
<mods:role id="F51765E2C14FB0D1E7FEDEA293C05821">
<mods:roleTerm id="091FCC7DF33E0ADABF0BD58204010F10">Author</mods:roleTerm>
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<mods:namePart id="2FCAD0BC0B7C387F7A2E059D79002860">Shear, William A.</mods:namePart>
<mods:nameIdentifier id="2C15492014A07CEB02D76DB29540309B" type="ORCID">0000-0002-5887-7003</mods:nameIdentifier>
<mods:affiliation id="41DB51FFBC215819CD98B66377E99F9C">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943 USA, current address: 1950 Price Drive, Farmville VA 23901 USA. wshear @ hsc. edu; https: // orcid. org / 0000 - 0002 - 5887 - 7003</mods:affiliation>
<mods:nameIdentifier id="D538DA851B5C189165B209C5C8E345DE" type="email">wshear@hsc.edu</mods:nameIdentifier>
</mods:name>
<mods:name id="637A8EDE3BFEC3425F9FBA8EC0FB3A82" type="personal">
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</mods:role>
<mods:namePart id="0270F3882B942961F7122D394D656405">Marek, Paul E.</mods:namePart>
<mods:affiliation id="9D2B88A7E23FA3832710F5BF511BAE0E">Department of Entomology, Virginia Tech, Blacksburg VA 24061 USA.</mods:affiliation>
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<mods:typeOfResource id="FB9957A51D535637095F6AEE7B3473F4">text</mods:typeOfResource>
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<mods:title id="AF06791879BEC1C0C0128C47E82E91D5">Zootaxa</mods:title>
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<mods:part id="727F781FE4B8F528A151D15B191E0F1E">
<mods:date id="67E5892C6CE047110B27D15294440A63">2022</mods:date>
<mods:detail id="51FD01F278327D6BDAAC92E80E964B83" type="pubDate">
<mods:number id="6A1451015650B461DB475DF6BA77E91C">2022-11-11</mods:number>
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<mods:detail id="C0ABE962B6A8B0509D75E6C186227EAD" type="volume">
<mods:number id="ED7CE206493ECA6EAC2ADE50FD1664C1">5205</mods:number>
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<mods:classification id="4666109EF0B72A928105E667D8B365B2">journal article</mods:classification>
<mods:identifier id="34E612E58EA74C65B22D57FC64EA5B1E" type="CLB-Dataset">187956</mods:identifier>
<mods:identifier id="D8C3B15CBDBEEF81A308A00536107018" type="DOI">10.11646/zootaxa.5205.6.1</mods:identifier>
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<treatment id="03F89303FFB4FFB99AF92CD9E1B4F93D" ID-DOI="http://doi.org/10.5281/zenodo.7318635" ID-GBIF-Taxon="205076367" ID-Zenodo-Dep="7318635" LSID="urn:lsid:plazi:treatment:03F89303FFB4FFB99AF92CD9E1B4F93D" httpUri="http://treatment.plazi.org/id/03F89303FFB4FFB99AF92CD9E1B4F93D" lastPageNumber="509" pageId="8" pageNumber="509">
<subSubSection id="C34B719EFFB4FFB99AF92CD9E091FC79" box="[151,665,906,933]" pageId="8" pageNumber="509" type="nomenclature">
<paragraph id="8BEE2215FFB4FFB99AF92CD9E091FC79" blockId="8.[151,665,906,969]" box="[151,665,906,933]" pageId="8" pageNumber="509">
<heading id="D0A69579FFB4FFB99AF92CD9E091FC79" bold="true" box="[151,665,906,933]" fontSize="11" level="1" pageId="8" pageNumber="509" reason="1">
<emphasis id="B925FE07FFB4FFB99AF92CD9E091FC79" bold="true" box="[151,665,906,933]" pageId="8" pageNumber="509">
<taxonomicName id="4C515996FFB4FFB99AF92CD9E041FC79" authority="Shear &amp; Marek" authorityName="Shear &amp; Marek" authorityYear="2022" box="[151,585,906,933]" class="Diplopoda" family="Striariidae" genus="Maraplia" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="schusteri" status="sp. nov.">
<emphasis id="B925FE07FFB4FFB99AF92CD9E374FC79" bold="true" box="[151,380,906,933]" italics="true" pageId="8" pageNumber="509">Maraplia schusteri</emphasis>
Shear &amp; Marek
</taxonomicName>
,
<taxonomicNameLabel id="A216437CFFB4FFB998382CD8E091FC79" box="[598,665,907,933]" pageId="8" pageNumber="509" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34B719EFFB4FFB99AF92CFCE0EAFB85" pageId="8" pageNumber="509" type="description">
<paragraph id="8BEE2215FFB4FFB99AF92CFCE360FC15" blockId="8.[151,665,906,969]" box="[151,360,942,969]" pageId="8" pageNumber="509">
<figureCitation id="136A3E90FFB4FFB99AF92CFCE31DFC15" box="[151,277,943,969]" captionStart="FIGURES 10–15" captionStartId="9.[152,270,1795,1819]" captionTargetBox="[211,1378,183,1770]" captionTargetId="figure-17@9.[210,1378,181,1771]" captionTargetPageId="9" captionText="FIGURES 10–15. Striariid millipedes. Figs 10, 11. Maraplia schusteri, n. gen., n. sp. 10. Gonopods, legpair 9 and paratergite 7, oblique lateral view. 11. Coxae of legpair 2, ventral view. Figs 12–15. Maraplia chico, n. gen., n. sp. 12. Head and first two legpairs, ventral view. 13. Coxal flasks of legpair 3, ventral view. 14. Gonopods, ventral view. 15. Posterior end of male, dorsal view. Abbreviations: aac, anterior angiocoxite; c4, coxa of leg 4; c5, coxa of leg 5; cf, coxal flask of leg 3; cx2, coxa of leg 2; f1, femur of leg 1; lab, labrum; m, mandible; numerals, ring numbers; pac, posterior angiocoxite; pf, prefemur of leg 3; pt7, paratergite of ring 7; s, gonopod sternum; t9, telopodite of ninth leg; tp9, telopodite process; vcp, common opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7318615" httpUri="https://zenodo.org/record/7318615/files/figure.png" pageId="8" pageNumber="509">Figs 10, 11</figureCitation>
,
<figureCitation id="136A3E90FFB4FFB99B4F2CFDE360FC15" box="[289,360,942,969]" captionStart-0="FIGURES 49–55" captionStart-1="FIGURES 56–62" captionStartId-0="24.[152,270,1824,1848]" captionStartId-1="25.[152,270,1668,1692]" captionTargetBox-0="[158,1433,279,1801]" captionTargetBox-1="[163,1423,198,1639]" captionTargetId-0="figure-56@24.[151,1436,273,1801]" captionTargetId-1="figure-17@25.[151,1436,181,1644]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURES 49–55. Striariid millipedes. Figs 49–51. Nototrisaria ornata, n. gen., n. sp., right gonopod, lateral view. 50. Same, mesal view. 51. Legpair 9 of male, anterior view. Figs 52–54. Maraplia napa, n. gen., n. sp. 52. Right gonopod, mesal view. 53. Right gonopod, lateral view. 54. Right leg 9 of male, posterior view. 55. Right gonopod of Maraplia schusteri, n. gen., n. sp., lateral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." captionText-1="FIGURES 56–62. Striariid millipedes. Figs 56, 57. Maraplia schusteri, n. gen., n. sp. 56. Right gonopod, mesal view. 57. Right leg 9 of male, anterior view. Figs 58–60. Maraplia chico, n. gen., n. sp. 58. Right gonopod, lateral view. 59. Right gonopod, mesal view. 60. Right leg 9 of male, posterior view. Figs 61, 62. Lamparia curryensis, n. gen., n. sp. 61. Right gonopod, lateral view. 62. Right gonopod, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; cxs, coxosternite of legs 9; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi-0="http://doi.org/10.5281/zenodo.7318639" figureDoi-1="http://doi.org/10.5281/zenodo.7318645" httpUri-0="https://zenodo.org/record/7318639/files/figure.png" httpUri-1="https://zenodo.org/record/7318645/files/figure.png" pageId="8" pageNumber="509">55–57</figureCitation>
</paragraph>
<paragraph id="8BEE2215FFB4FFB99AF92CA4E0EAFB85" blockId="8.[151,1437,1015,1761]" pageId="8" pageNumber="509">
<emphasis id="B925FE07FFB4FFB99AF92CA4E2EEFBCD" bold="true" box="[151,230,1015,1041]" pageId="8" pageNumber="509">Types:</emphasis>
<materialsCitation id="3B392848FFB4FFB99A812CA4E0D6FB85" ID-GBIF-Occurrence="3965952325" collectingDate="1956-02-09" collectionCode="FSCA" collectorName="R. O. Schuster. Types &amp; Parts" county="Butte Co." latitude="39.5219" location="Stringtown Hill" longLatPrecision="6" longitude="-121.3909" municipality="Male" pageId="8" pageNumber="509" specimenCount="2" specimenCount-male="1" stateProvince="California" typeStatus="holotype">
<collectingMunicipality id="6B8AB86FFFB4FFB99A812CA4E321FBCD" box="[239,297,1015,1041]" pageId="8" pageNumber="509">Male</collectingMunicipality>
<typeStatus id="54EA9CB7FFB4FFB99B5F2CA4E39BFBCD" box="[305,403,1015,1041]" pageId="8" pageNumber="509" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D57E99CFFB4FFB99BA32CA4E03EFBCD" box="[461,566,1015,1041]" pageId="8" pageNumber="509" type="male">two male</specimenCount>
<typeStatus id="54EA9CB7FFB4FFB998512CA4E0A2FBCD" box="[575,682,1015,1041]" pageId="8" pageNumber="509" type="paratype">paratypes</typeStatus>
from near
<location id="8E8E74CEFFB4FFB999452CA4E1D6FBCD" LSID="urn:lsid:plazi:treatment:03F89303FFB4FFB99AF92CD9E1B4F93D:8E8E74CEFFB4FFB999452CA4E1D6FBCD" box="[811,990,1015,1041]" county="Butte Co." latitude="39.5219" longLatPrecision="6" longitude="-121.3909" municipality="Male" name="Stringtown Hill" pageId="8" pageNumber="509" stateProvince="California">Stringtown Hill</location>
, NE of
<location id="8E8E74CEFFB4FFB99E542CA4E692FBCD" LSID="urn:lsid:plazi:treatment:03F89303FFB4FFB99AF92CD9E1B4F93D:8E8E74CEFFB4FFB99E542CA4E692FBCD" box="[1082,1178,1015,1041]" county="Butte Co." latitude="39.5219" longLatPrecision="6" longitude="-121.3909" municipality="Male" name="Oroville" pageId="8" pageNumber="509" stateProvince="California">Oroville</location>
,
<collectingCounty id="628F5A99FFB4FFB99EC92CA4E71EFBCD" box="[1191,1302,1015,1041]" pageId="8" pageNumber="509">Butte Co.</collectingCounty>
,
<collectingRegion id="4995ECF7FFB4FFB99F4A2CA4E790FBCD" box="[1316,1432,1015,1041]" country="United States of America" name="California" pageId="8" pageNumber="509">California</collectingRegion>
,
<geoCoordinate id="EE6544D2FFB4FFB99AF92B49E31CFBE9" box="[151,276,1050,1077]" degrees="39.5219" direction="north" orientation="latitude" pageId="8" pageNumber="509" precision="5" value="39.5219">39.5219°N</geoCoordinate>
, -
<geoCoordinate id="EE6544D2FFB4FFB99B492B48E3B0FBE9" box="[295,440,1051,1077]" degrees="121.3909" direction="west" orientation="longitude" pageId="8" pageNumber="509" precision="5" value="-121.3909">121.3909°W</geoCoordinate>
, 1440’ asl, collected
<date id="FFEF04D5FFB4FFB998CD2B48E155FBE9" box="[675,861,1050,1077]" pageId="8" pageNumber="509" value="1956-02-09">
<collectingDate id="EFABFD3DFFB4FFB998CD2B48E155FBE9" box="[675,861,1050,1077]" pageId="8" pageNumber="509" value="1956-02-09">9 February 1956</collectingDate>
</date>
by
<collectorName id="26A447C3FFB4FFB999E92B48E671FBE9" box="[903,1145,1051,1077]" pageId="8" pageNumber="509">R. O. Schuster. Types</collectorName>
deposited in
<collectionCode id="ED40BAD0FFB4FFB99F7E2B48E752FBE9" box="[1296,1370,1051,1077]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="8" pageNumber="509" type="Museum">FSCA</collectionCode>
.
<collectorName id="26A447C3FFB4FFB99F0A2B48E794FBE9" box="[1380,1436,1051,1077]" pageId="8" pageNumber="509">Parts</collectorName>
of
<typeStatus id="54EA9CB7FFB4FFB99AD82B6CE310FB85" box="[182,280,1087,1113]" pageId="8" pageNumber="509" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D57E99CFFB4FFB99B202B6CE3D5FB85" box="[334,477,1087,1113]" pageId="8" pageNumber="509" type="generic" typeStatus="paratype">one paratype</specimenCount>
on SEM stub WS35-4
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B719EFFB4FFB99AA92B30E68AFBA1" box="[199,1154,1123,1149]" pageId="8" pageNumber="509" type="etymology">
<paragraph id="8BEE2215FFB4FFB99AA92B30E68AFBA1" blockId="8.[151,1437,1015,1761]" box="[199,1154,1123,1149]" pageId="8" pageNumber="509">
<emphasis id="B925FE07FFB4FFB99AA92B30E340FBA1" bold="true" box="[199,328,1123,1149]" pageId="8" pageNumber="509">Etymology</emphasis>
: The species is named for entomologist Robert O. Schuster (1927–1989).
</paragraph>
</subSubSection>
<subSubSection id="C34B719EFFB4FFB99AA92BD5E094FAD1" pageId="8" pageNumber="509" type="diagnosis">
<paragraph id="8BEE2215FFB4FFB99AA92BD5E094FAD1" blockId="8.[151,1437,1015,1761]" pageId="8" pageNumber="509">
<emphasis id="B925FE07FFB4FFB99AA92BD5E332FB7C" bold="true" box="[199,314,1158,1184]" pageId="8" pageNumber="509">Diagnosis</emphasis>
: Distinct from the two other species of
<taxonomicName id="4C515996FFB4FFB999632BD4E17FFB7D" authority="Shear &amp; Marek, 2022" authorityName="Shear &amp; Marek" authorityYear="2022" box="[781,887,1159,1185]" class="Diplopoda" family="Striariidae" genus="Maraplia" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="509" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B925FE07FFB4FFB999632BD4E17FFB7D" box="[781,887,1159,1185]" italics="true" pageId="8" pageNumber="509">Maraplia</emphasis>
</taxonomicName>
<emphasis id="B925FE07FFB4FFB999EF2BD4E1DAFB7D" bold="true" box="[897,978,1159,1185]" pageId="8" pageNumber="509">
<taxonomicNameLabel id="A216437CFFB4FFB999EF2BD4E1DAFB7D" box="[897,978,1159,1185]" pageId="8" pageNumber="509" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
in the form of the gonopods.
<taxonomicName id="4C515996FFB4FFB99F5C2BD4E2F4FB19" authority="Shear &amp; Marek, 2022" authorityName="Shear &amp; Marek" authorityYear="2022" class="Diplopoda" family="Striariidae" genus="Maraplia" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="schusteri" status="sp. nov.">
<emphasis id="B925FE07FFB4FFB99F5C2BD4E2F4FB19" italics="true" pageId="8" pageNumber="509">Maraplia schusteri</emphasis>
</taxonomicName>
<emphasis id="B925FE07FFB4FFB99B6B2BF8E34EFB19" bold="true" box="[261,326,1195,1221]" pageId="8" pageNumber="509">
<taxonomicNameLabel id="A216437CFFB4FFB99B6B2BF8E34EFB19" box="[261,326,1195,1221]" pageId="8" pageNumber="509" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, has long, thin, and evenly curved angiocoxites, rather than the sharply deflexed, relatively short anterior angiocoxites seen in the other species (
<figureCitation id="136A3E90FFB4FFB998CC2B9CE0F0FB35" box="[674,760,1231,1257]" captionStart="FIGURES 10–15" captionStartId="9.[152,270,1795,1819]" captionTargetBox="[211,1378,183,1770]" captionTargetId="figure-17@9.[210,1378,181,1771]" captionTargetPageId="9" captionText="FIGURES 10–15. Striariid millipedes. Figs 10, 11. Maraplia schusteri, n. gen., n. sp. 10. Gonopods, legpair 9 and paratergite 7, oblique lateral view. 11. Coxae of legpair 2, ventral view. Figs 12–15. Maraplia chico, n. gen., n. sp. 12. Head and first two legpairs, ventral view. 13. Coxal flasks of legpair 3, ventral view. 14. Gonopods, ventral view. 15. Posterior end of male, dorsal view. Abbreviations: aac, anterior angiocoxite; c4, coxa of leg 4; c5, coxa of leg 5; cf, coxal flask of leg 3; cx2, coxa of leg 2; f1, femur of leg 1; lab, labrum; m, mandible; numerals, ring numbers; pac, posterior angiocoxite; pf, prefemur of leg 3; pt7, paratergite of ring 7; s, gonopod sternum; t9, telopodite of ninth leg; tp9, telopodite process; vcp, common opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7318615" httpUri="https://zenodo.org/record/7318615/files/figure.png" pageId="8" pageNumber="509">Figs 10</figureCitation>
,
<figureCitation id="136A3E90FFB4FFB9996D2B9DE129FB35" box="[771,801,1230,1257]" captionStart="FIGURES 49–55" captionStartId="24.[152,270,1824,1848]" captionTargetBox="[158,1433,279,1801]" captionTargetId="figure-56@24.[151,1436,273,1801]" captionTargetPageId="24" captionText="FIGURES 49–55. Striariid millipedes. Figs 49–51. Nototrisaria ornata, n. gen., n. sp., right gonopod, lateral view. 50. Same, mesal view. 51. Legpair 9 of male, anterior view. Figs 52–54. Maraplia napa, n. gen., n. sp. 52. Right gonopod, mesal view. 53. Right gonopod, lateral view. 54. Right leg 9 of male, posterior view. 55. Right gonopod of Maraplia schusteri, n. gen., n. sp., lateral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318639" httpUri="https://zenodo.org/record/7318639/files/figure.png" pageId="8" pageNumber="509">55</figureCitation>
,
<figureCitation id="136A3E90FFB4FFB999432B9DE143FB35" box="[813,843,1230,1257]" captionStart="FIGURES 56–62" captionStartId="25.[152,270,1668,1692]" captionTargetBox="[163,1423,198,1639]" captionTargetId="figure-17@25.[151,1436,181,1644]" captionTargetPageId="25" captionText="FIGURES 56–62. Striariid millipedes. Figs 56, 57. Maraplia schusteri, n. gen., n. sp. 56. Right gonopod, mesal view. 57. Right leg 9 of male, anterior view. Figs 58–60. Maraplia chico, n. gen., n. sp. 58. Right gonopod, lateral view. 59. Right gonopod, mesal view. 60. Right leg 9 of male, posterior view. Figs 61, 62. Lamparia curryensis, n. gen., n. sp. 61. Right gonopod, lateral view. 62. Right gonopod, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; cxs, coxosternite of legs 9; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318645" httpUri="https://zenodo.org/record/7318645/files/figure.png" pageId="8" pageNumber="509">56</figureCitation>
,
<emphasis id="B925FE07FFB4FFB999382B9CE176FB35" bold="true" box="[854,894,1231,1257]" pageId="8" pageNumber="509">aac</emphasis>
). The coxal process is not flattened and platelike but narrower and distinctly bifid (
<figureCitation id="136A3E90FFB4FFB9987F2BA0E06BFAD1" box="[529,611,1266,1293]" captionStart="FIGURES 49–55" captionStartId="24.[152,270,1824,1848]" captionTargetBox="[158,1433,279,1801]" captionTargetId="figure-56@24.[151,1436,273,1801]" captionTargetPageId="24" captionText="FIGURES 49–55. Striariid millipedes. Figs 49–51. Nototrisaria ornata, n. gen., n. sp., right gonopod, lateral view. 50. Same, mesal view. 51. Legpair 9 of male, anterior view. Figs 52–54. Maraplia napa, n. gen., n. sp. 52. Right gonopod, mesal view. 53. Right gonopod, lateral view. 54. Right leg 9 of male, posterior view. 55. Right gonopod of Maraplia schusteri, n. gen., n. sp., lateral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318639" httpUri="https://zenodo.org/record/7318639/files/figure.png" pageId="8" pageNumber="509">Fig. 55</figureCitation>
,
<emphasis id="B925FE07FFB4FFB998012BA0E083FAD1" bold="true" box="[623,651,1267,1293]" pageId="8" pageNumber="509">cp</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B719EFFB4FFB99AA92A45E3CBF961" pageId="8" pageNumber="509" type="description">
<paragraph id="8BEE2215FFB4FFB99AA92A45E2E8FAA5" blockId="8.[151,1437,1015,1761]" pageId="8" pageNumber="509">
<emphasis id="B925FE07FFB4FFB99AA92A45E359FAEC" bold="true" box="[199,337,1302,1328]" pageId="8" pageNumber="509">Description</emphasis>
:
<emphasis id="B925FE07FFB4FFB99B312A44E00FFAED" box="[351,519,1303,1329]" italics="true" pageId="8" pageNumber="509">
Male
<typeStatus id="54EA9CB7FFB4FFB99BF12A44E00BFAED" box="[415,515,1303,1329]" pageId="8" pageNumber="509" type="holotype">holotype</typeStatus>
.
</emphasis>
Length
<quantity id="4CA98FF0FFB4FFB9980C2A45E0B1FAED" box="[610,697,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="509" unit="mm" value="3.5">3.5 mm</quantity>
, width about
<quantity id="4CA98FF0FFB4FFB999232A44E1BCFAED" box="[845,948,1303,1329]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="509" unit="mm" value="0.32">0.32 mm</quantity>
. Two black ommatidia on each side of head. Color white. Sixth crests about twice height of C5, as short paranota. Telson lobes obscure. Other characters as for genus.
</paragraph>
<paragraph id="8BEE2215FFB4FFB99AA92AD0E72EF945" blockId="8.[151,1437,1015,1761]" pageId="8" pageNumber="509">
Gonopods (
<figureCitation id="136A3E90FFB4FFB99B282AD0E392FA41" box="[326,410,1411,1437]" captionStart="FIGURES 10–15" captionStartId="9.[152,270,1795,1819]" captionTargetBox="[211,1378,183,1770]" captionTargetId="figure-17@9.[210,1378,181,1771]" captionTargetPageId="9" captionText="FIGURES 10–15. Striariid millipedes. Figs 10, 11. Maraplia schusteri, n. gen., n. sp. 10. Gonopods, legpair 9 and paratergite 7, oblique lateral view. 11. Coxae of legpair 2, ventral view. Figs 12–15. Maraplia chico, n. gen., n. sp. 12. Head and first two legpairs, ventral view. 13. Coxal flasks of legpair 3, ventral view. 14. Gonopods, ventral view. 15. Posterior end of male, dorsal view. Abbreviations: aac, anterior angiocoxite; c4, coxa of leg 4; c5, coxa of leg 5; cf, coxal flask of leg 3; cx2, coxa of leg 2; f1, femur of leg 1; lab, labrum; m, mandible; numerals, ring numbers; pac, posterior angiocoxite; pf, prefemur of leg 3; pt7, paratergite of ring 7; s, gonopod sternum; t9, telopodite of ninth leg; tp9, telopodite process; vcp, common opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7318615" httpUri="https://zenodo.org/record/7318615/files/figure.png" pageId="8" pageNumber="509">Figs 10</figureCitation>
,
<figureCitation id="136A3E90FFB4FFB99BCD2AD1E3C9FA41" box="[419,449,1410,1437]" captionStart="FIGURES 49–55" captionStartId="24.[152,270,1824,1848]" captionTargetBox="[158,1433,279,1801]" captionTargetId="figure-56@24.[151,1436,273,1801]" captionTargetPageId="24" captionText="FIGURES 49–55. Striariid millipedes. Figs 49–51. Nototrisaria ornata, n. gen., n. sp., right gonopod, lateral view. 50. Same, mesal view. 51. Legpair 9 of male, anterior view. Figs 52–54. Maraplia napa, n. gen., n. sp. 52. Right gonopod, mesal view. 53. Right gonopod, lateral view. 54. Right leg 9 of male, posterior view. 55. Right gonopod of Maraplia schusteri, n. gen., n. sp., lateral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318639" httpUri="https://zenodo.org/record/7318639/files/figure.png" pageId="8" pageNumber="509">55</figureCitation>
,
<figureCitation id="136A3E90FFB4FFB99BA42AD1E3EFFA41" box="[458,487,1410,1437]" captionStart="FIGURES 56–62" captionStartId="25.[152,270,1668,1692]" captionTargetBox="[163,1423,198,1639]" captionTargetId="figure-17@25.[151,1436,181,1644]" captionTargetPageId="25" captionText="FIGURES 56–62. Striariid millipedes. Figs 56, 57. Maraplia schusteri, n. gen., n. sp. 56. Right gonopod, mesal view. 57. Right leg 9 of male, anterior view. Figs 58–60. Maraplia chico, n. gen., n. sp. 58. Right gonopod, lateral view. 59. Right gonopod, mesal view. 60. Right leg 9 of male, posterior view. Figs 61, 62. Lamparia curryensis, n. gen., n. sp. 61. Right gonopod, lateral view. 62. Right gonopod, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; cxs, coxosternite of legs 9; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318645" httpUri="https://zenodo.org/record/7318645/files/figure.png" pageId="8" pageNumber="509">56</figureCitation>
) small, compact. In lateral view, coxa (
<figureCitation id="136A3E90FFB4FFB999E22AD0E1D4FA41" box="[908,988,1410,1437]" captionStart="FIGURES 49–55" captionStartId="24.[152,270,1824,1848]" captionTargetBox="[158,1433,279,1801]" captionTargetId="figure-56@24.[151,1436,273,1801]" captionTargetPageId="24" captionText="FIGURES 49–55. Striariid millipedes. Figs 49–51. Nototrisaria ornata, n. gen., n. sp., right gonopod, lateral view. 50. Same, mesal view. 51. Legpair 9 of male, anterior view. Figs 52–54. Maraplia napa, n. gen., n. sp. 52. Right gonopod, mesal view. 53. Right gonopod, lateral view. 54. Right leg 9 of male, posterior view. 55. Right gonopod of Maraplia schusteri, n. gen., n. sp., lateral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318639" httpUri="https://zenodo.org/record/7318639/files/figure.png" pageId="8" pageNumber="509">Fig. 55</figureCitation>
, cx) with about 6 or 7 setae, coxal process narrow, distally bifid, anterior branch longer, posterior branch slightly recurved (
<figureCitation id="136A3E90FFB4FFB99E4D2AF4E67EFA1D" box="[1059,1142,1446,1473]" captionStart="FIGURES 49–55" captionStartId="24.[152,270,1824,1848]" captionTargetBox="[158,1433,279,1801]" captionTargetId="figure-56@24.[151,1436,273,1801]" captionTargetPageId="24" captionText="FIGURES 49–55. Striariid millipedes. Figs 49–51. Nototrisaria ornata, n. gen., n. sp., right gonopod, lateral view. 50. Same, mesal view. 51. Legpair 9 of male, anterior view. Figs 52–54. Maraplia napa, n. gen., n. sp. 52. Right gonopod, mesal view. 53. Right gonopod, lateral view. 54. Right leg 9 of male, posterior view. 55. Right gonopod of Maraplia schusteri, n. gen., n. sp., lateral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318639" httpUri="https://zenodo.org/record/7318639/files/figure.png" pageId="8" pageNumber="509">Fig. 55</figureCitation>
, cp). Anterior angiocoxite (
<figureCitation id="136A3E90FFB4FFB99AF12A98E2F1FA39" box="[159,249,1483,1509]" captionStart="FIGURES 10–15" captionStartId="9.[152,270,1795,1819]" captionTargetBox="[211,1378,183,1770]" captionTargetId="figure-17@9.[210,1378,181,1771]" captionTargetPageId="9" captionText="FIGURES 10–15. Striariid millipedes. Figs 10, 11. Maraplia schusteri, n. gen., n. sp. 10. Gonopods, legpair 9 and paratergite 7, oblique lateral view. 11. Coxae of legpair 2, ventral view. Figs 12–15. Maraplia chico, n. gen., n. sp. 12. Head and first two legpairs, ventral view. 13. Coxal flasks of legpair 3, ventral view. 14. Gonopods, ventral view. 15. Posterior end of male, dorsal view. Abbreviations: aac, anterior angiocoxite; c4, coxa of leg 4; c5, coxa of leg 5; cf, coxal flask of leg 3; cx2, coxa of leg 2; f1, femur of leg 1; lab, labrum; m, mandible; numerals, ring numbers; pac, posterior angiocoxite; pf, prefemur of leg 3; pt7, paratergite of ring 7; s, gonopod sternum; t9, telopodite of ninth leg; tp9, telopodite process; vcp, common opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7318615" httpUri="https://zenodo.org/record/7318615/files/figure.png" pageId="8" pageNumber="509">Figs 10</figureCitation>
,
<figureCitation id="136A3E90FFB4FFB99B692A99E32DFA39" box="[263,293,1482,1509]" captionStart="FIGURES 49–55" captionStartId="24.[152,270,1824,1848]" captionTargetBox="[158,1433,279,1801]" captionTargetId="figure-56@24.[151,1436,273,1801]" captionTargetPageId="24" captionText="FIGURES 49–55. Striariid millipedes. Figs 49–51. Nototrisaria ornata, n. gen., n. sp., right gonopod, lateral view. 50. Same, mesal view. 51. Legpair 9 of male, anterior view. Figs 52–54. Maraplia napa, n. gen., n. sp. 52. Right gonopod, mesal view. 53. Right gonopod, lateral view. 54. Right leg 9 of male, posterior view. 55. Right gonopod of Maraplia schusteri, n. gen., n. sp., lateral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318639" httpUri="https://zenodo.org/record/7318639/files/figure.png" pageId="8" pageNumber="509">55</figureCitation>
,
<figureCitation id="136A3E90FFB4FFB99B5A2A99E35AFA39" box="[308,338,1482,1509]" captionStart="FIGURES 56–62" captionStartId="25.[152,270,1668,1692]" captionTargetBox="[163,1423,198,1639]" captionTargetId="figure-17@25.[151,1436,181,1644]" captionTargetPageId="25" captionText="FIGURES 56–62. Striariid millipedes. Figs 56, 57. Maraplia schusteri, n. gen., n. sp. 56. Right gonopod, mesal view. 57. Right leg 9 of male, anterior view. Figs 58–60. Maraplia chico, n. gen., n. sp. 58. Right gonopod, lateral view. 59. Right gonopod, mesal view. 60. Right leg 9 of male, posterior view. Figs 61, 62. Lamparia curryensis, n. gen., n. sp. 61. Right gonopod, lateral view. 62. Right gonopod, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; cxs, coxosternite of legs 9; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318645" httpUri="https://zenodo.org/record/7318645/files/figure.png" pageId="8" pageNumber="509">56</figureCitation>
, aac) narrow, long, evenly curved, apically acute, with smaller basal triangular tooth. Posterior angiocoxite strongly reduced to single blunt rod, apparently not sheathing the very short, straight flagellocoxite. Colpocoxite (
<figureCitation id="136A3E90FFB4FFB99B412940E38DF9F1" box="[303,389,1554,1581]" captionStart="FIGURES 49–55" captionStartId="24.[152,270,1824,1848]" captionTargetBox="[158,1433,279,1801]" captionTargetId="figure-56@24.[151,1436,273,1801]" captionTargetPageId="24" captionText="FIGURES 49–55. Striariid millipedes. Figs 49–51. Nototrisaria ornata, n. gen., n. sp., right gonopod, lateral view. 50. Same, mesal view. 51. Legpair 9 of male, anterior view. Figs 52–54. Maraplia napa, n. gen., n. sp. 52. Right gonopod, mesal view. 53. Right gonopod, lateral view. 54. Right leg 9 of male, posterior view. 55. Right gonopod of Maraplia schusteri, n. gen., n. sp., lateral view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318639" httpUri="https://zenodo.org/record/7318639/files/figure.png" pageId="8" pageNumber="509">Figs 55</figureCitation>
,
<figureCitation id="136A3E90FFB4FFB99BFF2941E3A7F9F1" box="[401,431,1554,1581]" captionStart="FIGURES 56–62" captionStartId="25.[152,270,1668,1692]" captionTargetBox="[163,1423,198,1639]" captionTargetId="figure-17@25.[151,1436,181,1644]" captionTargetPageId="25" captionText="FIGURES 56–62. Striariid millipedes. Figs 56, 57. Maraplia schusteri, n. gen., n. sp. 56. Right gonopod, mesal view. 57. Right leg 9 of male, anterior view. Figs 58–60. Maraplia chico, n. gen., n. sp. 58. Right gonopod, lateral view. 59. Right gonopod, mesal view. 60. Right leg 9 of male, posterior view. Figs 61, 62. Lamparia curryensis, n. gen., n. sp. 61. Right gonopod, lateral view. 62. Right gonopod, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; cxs, coxosternite of legs 9; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318645" httpUri="https://zenodo.org/record/7318645/files/figure.png" pageId="8" pageNumber="509">56</figureCitation>
, cc) shifted anteriomesally, with anterior branch densely fimbriate, posterior part swollen, with a few small, fine triangular fimbriae. Ninth legs (
<figureCitation id="136A3E90FFB4FFB998992964E140F98D" box="[759,840,1590,1617]" captionStart="FIGURES 56–62" captionStartId="25.[152,270,1668,1692]" captionTargetBox="[163,1423,198,1639]" captionTargetId="figure-17@25.[151,1436,181,1644]" captionTargetPageId="25" captionText="FIGURES 56–62. Striariid millipedes. Figs 56, 57. Maraplia schusteri, n. gen., n. sp. 56. Right gonopod, mesal view. 57. Right leg 9 of male, anterior view. Figs 58–60. Maraplia chico, n. gen., n. sp. 58. Right gonopod, lateral view. 59. Right gonopod, mesal view. 60. Right leg 9 of male, posterior view. Figs 61, 62. Lamparia curryensis, n. gen., n. sp. 61. Right gonopod, lateral view. 62. Right gonopod, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; cxs, coxosternite of legs 9; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318645" httpUri="https://zenodo.org/record/7318645/files/figure.png" pageId="8" pageNumber="509">Fig. 57</figureCitation>
) with mesal coxosternal process low, triangular (
<figureCitation id="136A3E90FFB4FFB99F012964E2BDF9A9" captionStart="FIGURES 56–62" captionStartId="25.[152,270,1668,1692]" captionTargetBox="[163,1423,198,1639]" captionTargetId="figure-17@25.[151,1436,181,1644]" captionTargetPageId="25" captionText="FIGURES 56–62. Striariid millipedes. Figs 56, 57. Maraplia schusteri, n. gen., n. sp. 56. Right gonopod, mesal view. 57. Right leg 9 of male, anterior view. Figs 58–60. Maraplia chico, n. gen., n. sp. 58. Right gonopod, lateral view. 59. Right gonopod, mesal view. 60. Right leg 9 of male, posterior view. Figs 61, 62. Lamparia curryensis, n. gen., n. sp. 61. Right gonopod, lateral view. 62. Right gonopod, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; cxs, coxosternite of legs 9; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318645" httpUri="https://zenodo.org/record/7318645/files/figure.png" pageId="8" pageNumber="509">Fig. 57</figureCitation>
, cp), telopodite fungiform (
<figureCitation id="136A3E90FFB4FFB99B9A2908E045F9A9" box="[500,589,1627,1653]" captionStart="FIGURES 10–15" captionStartId="9.[152,270,1795,1819]" captionTargetBox="[211,1378,183,1770]" captionTargetId="figure-17@9.[210,1378,181,1771]" captionTargetPageId="9" captionText="FIGURES 10–15. Striariid millipedes. Figs 10, 11. Maraplia schusteri, n. gen., n. sp. 10. Gonopods, legpair 9 and paratergite 7, oblique lateral view. 11. Coxae of legpair 2, ventral view. Figs 12–15. Maraplia chico, n. gen., n. sp. 12. Head and first two legpairs, ventral view. 13. Coxal flasks of legpair 3, ventral view. 14. Gonopods, ventral view. 15. Posterior end of male, dorsal view. Abbreviations: aac, anterior angiocoxite; c4, coxa of leg 4; c5, coxa of leg 5; cf, coxal flask of leg 3; cx2, coxa of leg 2; f1, femur of leg 1; lab, labrum; m, mandible; numerals, ring numbers; pac, posterior angiocoxite; pf, prefemur of leg 3; pt7, paratergite of ring 7; s, gonopod sternum; t9, telopodite of ninth leg; tp9, telopodite process; vcp, common opening of vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7318615" httpUri="https://zenodo.org/record/7318615/files/figure.png" pageId="8" pageNumber="509">Figs 10</figureCitation>
,
<figureCitation id="136A3E90FFB4FFB998352909E071F9A9" box="[603,633,1626,1653]" captionStart="FIGURES 56–62" captionStartId="25.[152,270,1668,1692]" captionTargetBox="[163,1423,198,1639]" captionTargetId="figure-17@25.[151,1436,181,1644]" captionTargetPageId="25" captionText="FIGURES 56–62. Striariid millipedes. Figs 56, 57. Maraplia schusteri, n. gen., n. sp. 56. Right gonopod, mesal view. 57. Right leg 9 of male, anterior view. Figs 58–60. Maraplia chico, n. gen., n. sp. 58. Right gonopod, lateral view. 59. Right gonopod, mesal view. 60. Right leg 9 of male, posterior view. Figs 61, 62. Lamparia curryensis, n. gen., n. sp. 61. Right gonopod, lateral view. 62. Right gonopod, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cp, coxal process; cx, gonopod coxa; cxs, coxosternite of legs 9; fc, flagellocoxite; pac, posterior angiocoxite; s9, sternum of ninth legs; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7318645" httpUri="https://zenodo.org/record/7318645/files/figure.png" pageId="8" pageNumber="509">57</figureCitation>
, t
<figureCitation id="136A3E90FFB4FFB998FF2908E0A8F9A9" box="[657,672,1627,1653]" captionStart="FIGURES 7–9" captionStartId="7.[152,270,1919,1943]" captionTargetBox="[159,1431,273,1895]" captionTargetId="figure-60@7.[157,1431,273,1896]" captionTargetPageId="7" captionText="FIGURES 7–9. Maraplia napa, n. gen., n. sp. 7. Female, lateral view. 8. Head and first legpair of male, ventral view. 9. Coxal flask of leg 3 of male, lateral view. Abbreviation: cf, coxal flask of leg 3." figureDoi="http://doi.org/10.5281/zenodo.7318611" httpUri="https://zenodo.org/record/7318611/files/figure.png" pageId="8" pageNumber="509">9</figureCitation>
), bearing mesal process set with 6 or 7 ensiform setae and second curved, roughly triangular posterior process. Tenth leg coxae slightly enlarged, with glandular openings.
</paragraph>
<paragraph id="8BEE2215FFB4FFB99AA929F0E3CBF961" blockId="8.[151,1437,1015,1761]" box="[199,451,1699,1725]" pageId="8" pageNumber="509">Females not collected.</paragraph>
</subSubSection>
<subSubSection id="C34B719EFFB4FFB99AA92995E1B4F93D" box="[199,956,1734,1761]" pageId="8" pageNumber="509" type="distribution">
<paragraph id="8BEE2215FFB4FFB99AA92995E1B4F93D" blockId="8.[151,1437,1015,1761]" box="[199,956,1734,1761]" pageId="8" pageNumber="509">
<emphasis id="B925FE07FFB4FFB99AA92995E351F93C" bold="true" box="[199,345,1734,1760]" pageId="8" pageNumber="509">Distribution</emphasis>
: Presently known only from Butte County,
<collectingRegion id="4995ECF7FFB4FFB9992D2994E1B0F93D" box="[835,952,1735,1761]" country="United States of America" name="California" pageId="8" pageNumber="509">California</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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