treatments-xml/data/03/BF/87/03BF87F4FFA92577FE9EFCD901CAFD46.xml
2024-06-21 12:22:17 +02:00

559 lines
68 KiB
XML
Raw Blame History

This file contains invisible Unicode characters

This file contains invisible Unicode characters that are indistinguishable to humans but may be processed differently by a computer. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="FE724865D9732F1225672DDE9752BA74" ID-DOI="10.5281/zenodo.171314" ID-GBIF-Dataset="9fcc3e54-c294-42b8-8f12-75919ae4f461" ID-ISSN="1175­5326" ID-Zenodo-Dep="171314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459720424464" checkinUser="plazi" docAuthor="Saether, Ole A." docDate="2005" docId="03BF87F4FFA92577FE9EFCD901CAFD46" docLanguage="en" docName="zt00974.pdf" docOrigin="Zootaxa 974" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Orthocladius (Mesorthocladius) lamellatus Saether, 2005, sp. n." docType="treatment" docVersion="8" lastPageNumber="36" masterDocId="FF86FF8CFFB52554FF96FF950554FF80" masterDocTitle="A new subgenus and new species of Orthocladius van der Wulp, with a phylogenetic evaluation of the validity of the subgenera of the genus (Diptera: Chironomidae)" masterLastPageNumber="56" masterPageNumber="1" pageNumber="29" updateTime="1698209064823" updateUser="plazi">
<mods:mods id="3A7262E922DA644142E2560F4C87AB31" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="95ACA89EB5F12944C0C4295AE1D480F7">
<mods:title id="ECF2DA2981F46B871A2F81C0DDB12428">A new subgenus and new species of Orthocladius van der Wulp, with a phylogenetic evaluation of the validity of the subgenera of the genus (Diptera: Chironomidae)</mods:title>
</mods:titleInfo>
<mods:name id="A3EF90F36A10594D06676975E8A44991" type="personal">
<mods:role id="4D25123290F7CFEE2D5E17F0CF56D8FF">
<mods:roleTerm id="EC873FEA1F47232F8342120806D8C446">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2EC861097D71D9FEECC4653F4C830F49">Saether, Ole A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="58B8C9E5A50367508B140AB819DF97B5">text</mods:typeOfResource>
<mods:relatedItem id="035186B0008857E633CEDA816C6F8189" type="host">
<mods:titleInfo id="F23D09B64C67E83826481C26D06A0B90">
<mods:title id="7401877A0E5D77CB050CADCAB49DF567">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C4201AF88490D9B07768EF1CCD128EC3">
<mods:date id="D8C604AB605C13F89C541A02B139FE00">2005</mods:date>
<mods:detail id="DEE957263C3DBA4AA52976C0F1A64A37" type="volume">
<mods:number id="68EA2C4C7F8DB1F983EAE5F898A63B60">974</mods:number>
</mods:detail>
<mods:extent id="FD160E7D4DF7D84596038ECE6838313D" unit="page">
<mods:start id="42A0994F33CCB59499D11BFF2F3EA81C">1</mods:start>
<mods:end id="84F1759A7BA522DF0A61D6D03ED7409D">56</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A661BCF57CFA88A6321C1B07604CC616">journal article</mods:classification>
<mods:identifier id="E4B980E8BE9A2906E2661363AAFFDD46" type="DOI">10.5281/zenodo.171314</mods:identifier>
<mods:identifier id="031BAA757B28B8EBC3606A434134BB5E" type="GBIF-Dataset">9fcc3e54-c294-42b8-8f12-75919ae4f461</mods:identifier>
<mods:identifier id="861AA6C1F11366679C469E7349BE83A5" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="B0365DE8EAA84CA0CA34F84973D2B039" type="Zenodo-Dep">171314</mods:identifier>
</mods:mods>
<treatment id="03BF87F4FFA92577FE9EFCD901CAFD46" ID-DOI="http://doi.org/10.5281/zenodo.6266202" ID-GBIF-Taxon="119335505" ID-Zenodo-Dep="6266202" LSID="urn:lsid:plazi:treatment:03BF87F4FFA92577FE9EFCD901CAFD46" httpUri="http://treatment.plazi.org/id/03BF87F4FFA92577FE9EFCD901CAFD46" lastPageId="35" lastPageNumber="36" pageId="28" pageNumber="29">
<subSubSection id="C30C6569FFA92548FE9EFCD907A0FC09" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BA936E2FFA92548FE9EFCD9060DFCE6" blockId="28.[264,857,844,905]" box="[264,857,844,870]" pageId="28" pageNumber="29">
<heading id="D0E1818EFFA92548FE9EFCD9060DFCE6" bold="true" box="[264,857,844,870]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<emphasis id="B962EAF0FFA92548FE9EFCD9060DFCE6" bold="true" box="[264,857,844,870]" pageId="28" pageNumber="29">
<taxonomicName id="4C164D61FFA92548FE9EFCD9065BFCE6" ID-CoL="8D49N" box="[264,783,844,870]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="lamellatus" status="sp. nov." subGenus="Mesorthocladius">
<emphasis id="B962EAF0FFA92548FE9EFCD904F2FCE6" bold="true" box="[264,422,844,870]" italics="true" pageId="28" pageNumber="29">Orthocladius</emphasis>
(
<emphasis id="B962EAF0FFA92548FE21FCD907D5FCE6" bold="true" box="[439,641,844,870]" italics="true" pageId="28" pageNumber="29">Mesorthocladius</emphasis>
)
<emphasis id="B962EAF0FFA92548FD04FCD9065BFCE6" bold="true" box="[658,783,844,870]" italics="true" pageId="28" pageNumber="29">lamellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251578BFFA92548FC81FCD9060DFCE6" box="[791,857,844,870]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA936E2FFA92548FE9EFCFA07A0FC09" blockId="28.[264,857,844,905]" box="[264,756,879,905]" pageId="28" pageNumber="29">
(
<figureCitation id="132D2A67FFA92548FE87FCFA04DAFC09" box="[273,398,879,905]" captionStart="FIGURES 8 12" captionStartId="29.[264,382,1778,1802]" captionTargetBox="[307,1302,317,1731]" captionTargetId="figure@29.[269,1318,307,1755]" captionTargetPageId="29" captionText="FIGURES 8 12. Orthocladius (Mesorthocladius) lamellatus sp. n., male imago, 8: head, 9: thorax, 10: wing, 11: hypopygium, 12: gonostylus, different view." httpUri="https://zenodo.org/record/171317/files/figure.png" pageId="28" pageNumber="29">Figs. 812</figureCitation>
,
<figureCitation id="132D2A67FFA92548FE0DFCFA04B0FC09" box="[411,484,879,905]" captionStart="FIGURES 13 22" captionStartId="31.[264,383,1768,1792]" captionTargetBox="[285,1303,643,1752]" captionTargetId="figure@31.[269,1318,618,1765]" captionTargetPageId="31" captionText="FIGURES 13 22. Orthocladius (Mesorthocladius) spp., female imagines, genitalia, 13 17: O. (M.) lamellatus sp. n., 13: ventral view, 14: ventrolateral lobe, 15: dorsomesal lobe, 16: apodeme lobe, 17: dorsal view, 18 22: O. (M.) nimidens sp. n., 18: ventral view, 19: ventrolateral lobe, 20: dorsomesal lobe, 21: apodeme lobe, 22 dorsal view." httpUri="https://zenodo.org/record/171318/files/figure.png" pageId="28" pageNumber="29">1317</figureCitation>
,
<figureCitation id="132D2A67FFA92548FE67FCFA0769FC09" box="[497,573,879,905]" captionStart="FIGURES 23 27" captionStartId="32.[264,382,1814,1838]" captionTargetBox="[362,1271,556,1802]" captionTargetId="figure@32.[312,1277,554,1804]" captionTargetPageId="32" captionText="FIGURES 23 27. Orthocladius (Mesorthocladius) spp., pupae, 23: O. (M.) lamellatus sp. n., tergites, 24: thoracic horn, 25 27: O. (M.) nimidens sp. n., 25: tergites, 26: thoracic horn, 27: frontal apotome." httpUri="https://zenodo.org/record/171319/files/figure.png" pageId="28" pageNumber="29">23, 24</figureCitation>
,
<figureCitation id="132D2A67FFA92548FDDFFCFA07C1FC09" box="[585,661,879,905]" captionStart="FIGURES 28 34" captionStartId="34.[264,383,1688,1712]" captionTargetBox="[308,1302,296,1678]" captionTargetId="figure@34.[269,1318,280,1682]" captionTargetPageId="34" captionText="FIGURES 28 34. Orthocladius spp., larvae, 28: Orthocladius (Mesorthocladius) frigidus (Zetterstedt) labrum, epipharyngeal area and premandible, 29, 30: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 29: antenna, 30: labrum, epipharyngeal area and premandible, 31, 32: Orthocladius (Mesorthocladius) lamellatus sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible, 33 34: Orthocladius (Mesorthocladius) nimidens sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible." httpUri="https://zenodo.org/record/171320/files/figure.png" pageId="28" pageNumber="29">31, 32</figureCitation>
,
<figureCitation id="132D2A67FFA92548FD37FCFA07BFFC09" box="[673,747,879,905]" captionStart="FIGURES 35 40" captionStartId="36.[264,383,1691,1715]" captionTargetBox="[281,1294,319,1672]" captionTargetId="figure@36.[269,1318,299,1687]" captionTargetPageId="36" captionText="FIGURES 35 40. Orthocladius spp., larvae, 35, 36: Orthocladius (Mesorthocladius) lamellatus sp. n., 35: mandible, 36: mentum, 37 38: Orthocladius (Mesorthocladius) nimidens sp. n., 37: mandible, 38: mentum, 39 40: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 39: mandible, 40: mentum" httpUri="https://zenodo.org/record/171321/files/figure.png" pageId="28" pageNumber="29">35, 36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C6569FFA92548FE9EFC2A0765F8EB" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BA936E2FFA92548FE9EFC2A04FDFC58" blockId="28.[264,1323,959,1265]" box="[264,425,959,984]" pageId="28" pageNumber="29">
<emphasis id="B962EAF0FFA92548FE9EFC2A04FDFC58" box="[264,425,959,984]" italics="true" pageId="28" pageNumber="29">
<typeStatus id="54AD8840FFA92548FE9EFC2A046BFC58" box="[264,319,959,984]" pageId="28" pageNumber="29">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="8BA936E2FFA92548FEAEFC720649FB71" blockId="28.[264,1323,959,1265]" pageId="28" pageNumber="29">
<typeStatus id="54AD8840FFA92548FEAEFC7204F1FB81" box="[312,421,999,1025]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
ɗ reared from larva,
<collectingCountry id="F3017672FFA92548FD32FC7207B7FB81" box="[676,739,999,1025]" name="United States of America" pageId="28" pageNumber="29">USA</collectingCountry>
: Ohio, Delaware Co., Olentangy R., N. of Winter Road,
<date id="FFA81022FFA92548FECEFB9A04D5FBA9" box="[344,385,1039,1065]" pageId="28" pageNumber="29">5 ii</date>
emerged
<date id="FFA81022FFA92548FE6DFB9A07D7FBA9" box="[507,643,1039,1065]" pageId="28" pageNumber="29" value="1987-03-14">14 iii 1987</date>
, M. J. Bolton (
<collectionCode id="ED07AE27FFA92548FCA9FB9A06C2FBA9" box="[831,918,1039,1065]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="28" pageNumber="29">ZMBN</collectionCode>
<typeStatus id="54AD8840FFA92548FC36FB9A068EFBA9" box="[928,986,1039,1065]" pageId="28" pageNumber="29">Type</typeStatus>
No. 406).
<typeStatus id="54AD8840FFA92548FBF5FB9A018EFBA9" box="[1123,1242,1039,1065]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F3017672FFA92548FB7EFB9A0073FBA9" box="[1256,1319,1039,1065]" name="United States of America" pageId="28" pageNumber="29">USA</collectingCountry>
: Ohio, Franklin Co., Sciota park, spring­stream, mature ɗ pupa reared from larva, mature Ψ pupa reared from larva,
<date id="FFA81022FFA92548FDB8FBCA07E0FBF9" box="[558,692,1119,1145]" pageId="28" pageNumber="29" value="1987-04-26">26 iv 1987</date>
, M. J. Bolton (
<collectionCode id="ED07AE27FFA92548FCF9FBCA069EFBF9" box="[879,970,1119,1145]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="28" pageNumber="29">ZMBN</collectionCode>
); Ohio, Franklin Co., springstream tributary to Sciota R.,
<specimenCount id="9D10FD6BFFA92548FDFDFB12079AFB21" box="[619,718,1159,1185]" pageId="28" pageNumber="29" type="larva">4 larvae</specimenCount>
,
<date id="FFA81022FFA92548FD4DFB12061CFB21" box="[731,840,1159,1185]" pageId="28" pageNumber="29" value="1986-05-01">1 v 1986</date>
, M. J. Bolton (
<collectionCode id="ED07AE27FFA92548FC68FB12010DFB21" box="[1022,1113,1159,1185]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="28" pageNumber="29">ZMBN</collectionCode>
); Ohio, Delaware Co., Highbanks Metro Park, small stream, 1ɗ, mature ɗ pupa reared from larva,
<specimenCount id="9D10FD6BFFA92548FB53FB3A0072FB49" box="[1221,1318,1199,1225]" pageId="28" pageNumber="29" type="larva">3 larvae</specimenCount>
,
<date id="FFA81022FFA92548FE9EFB42042DFB71" box="[264,377,1239,1265]" pageId="28" pageNumber="29" value="1987-04-04">4 iv 1987</date>
,
<date id="FFA81022FFA92548FE13FB42075CFB71" box="[389,520,1239,1265]" pageId="28" pageNumber="29" value="1989-03-26">26 iii 1989</date>
. M. J. Bolton (
<collectionCode id="ED07AE27FFA92548FD23FB420644FB71" box="[693,784,1239,1265]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="28" pageNumber="29">ZMBN</collectionCode>
).
</paragraph>
<paragraph id="8BA936E2FFA92548FE9EFAB2071CFA11" pageId="28" pageNumber="29">
<table id="F916C442FFA9DAABFE9EFAB2007EFA11" box="[264,1322,1319,1425]" colsContinueFrom="25.[264,1322,777,883]" gridcols="11" gridrows="3" pageId="28" pageNumber="29">
<tr id="352634A0FFA9DAABFE9EFAB2007EFAC0" box="[264,1322,1319,1344]" gridrow="0" pageId="28" pageNumber="29" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76F75DDCFFA9DAABFE9EFAB20726FAC0" box="[264,626,1319,1344]" gridcol="0" gridrow="0" pageId="28" pageNumber="29">
<emphasis id="B962EAF0FFA92548FE9EFAB204D1FAC0" box="[264,389,1319,1344]" italics="true" pageId="28" pageNumber="29">Etymology</emphasis>
</th>
</tr>
<tr id="352634A0FFA9DAABFE9EFADA007EFAE9" box="[264,1322,1359,1385]" gridrow="1" pageId="28" pageNumber="29">
<th id="76F75DDCFFA9DAABFE9EFADA0726FAE9" box="[264,626,1359,1385]" gridcol="0" gridrow="1" pageId="28" pageNumber="29">
From Latin,
<emphasis id="B962EAF0FFA92548FE5AFADA0770FAE8" box="[460,548,1359,1384]" italics="true" pageId="28" pageNumber="29">lamella</emphasis>
, small
</th>
<td id="76F75DDCFFA9DAABFDECFADA07E7FAE9" box="[634,691,1359,1385]" gridcol="1" gridrow="1" pageId="28" pageNumber="29">plate</td>
<td id="76F75DDCFFA9DAABFD2CFADA07B0FAE9" box="[698,740,1359,1385]" gridcol="2" gridrow="1" pageId="28" pageNumber="29">and</td>
<td id="76F75DDCFFA9DAABFD7AFADA067AFAE9" box="[748,814,1359,1385]" gridcol="3" gridrow="1" pageId="28" pageNumber="29">
<emphasis id="B962EAF0FFA92548FD7AFADA0673FAE8" box="[748,807,1359,1384]" italics="true" pageId="28" pageNumber="29">­atus</emphasis>
,
</td>
<td id="76F75DDCFFA9DAABFCA0FADA06F5FAE9" box="[822,929,1359,1385]" gridcol="4" gridrow="1" pageId="28" pageNumber="29">equipped</td>
<td id="76F75DDCFFA9DAABFC3FFADA06B0FAE9" box="[937,996,1359,1385]" gridcol="5" gridrow="1" pageId="28" pageNumber="29">with,</td>
<td id="76F75DDCFFA9DAABFC7AFADA0106FAE9" box="[1004,1106,1359,1385]" gridcol="6" gridrow="1" pageId="28" pageNumber="29">referring</td>
<td id="76F75DDCFFA9DAABFBCCFADA0125FAE9" box="[1114,1137,1359,1385]" gridcol="7" gridrow="1" pageId="28" pageNumber="29">to</td>
<td id="76F75DDCFFA9DAABFBEFFADA01C9FAE9" box="[1145,1181,1359,1385]" gridcol="8" gridrow="1" pageId="28" pageNumber="29">the</td>
<td id="76F75DDCFFA9DAABFB32FADA005FFAE9" box="[1188,1291,1359,1385]" gridcol="9" gridrow="1" pageId="28" pageNumber="29">presence</td>
<td id="76F75DDCFFA9DAABFA84FADA007EFAE9" box="[1298,1322,1359,1385]" gridcol="10" gridrow="1" pageId="28" pageNumber="29">of</td>
</tr>
<tr id="352634A0FFA9DAABFE9EFAE2007EFA11" box="[264,1322,1399,1425]" gridrow="2" pageId="28" pageNumber="29" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76F75DDCFFA9DAABFE9EFAE20726FA11" box="[264,626,1399,1425]" gridcol="0" gridrow="2" pageId="28" pageNumber="29">labral lamellae in the larva.</th>
</tr>
</table>
</paragraph>
<paragraph id="8BA936E2FFA92548FE9EFA540759FA5A" blockId="28.[264,1323,1473,1899]" box="[264,525,1473,1498]" pageId="28" pageNumber="29">
<emphasis id="B962EAF0FFA92548FE9EFA540759FA5A" box="[264,525,1473,1498]" italics="true" pageId="28" pageNumber="29">Diagnostic characters</emphasis>
</paragraph>
<paragraph id="8BA936E2FFA92548FEAEFA7C0765F8EB" blockId="28.[264,1323,1473,1899]" pageId="28" pageNumber="29">
The male imago is separable from the other species of the subgenus by having a well developed virga consisting of a cluster of spines combined with uni­biserial scutellars, robust anal point and sensilla chaetica on both mid and hind leg. The pear­shaped seminal capsule with apical rugulosity separates the female from other species. The pupa differs from other members of the subgenus by lacking pedes spurii B, from other members of the genus except
<taxonomicName id="4C164D61FFA92548FE30F9240700F94A" box="[422,596,1713,1738]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="chuzesextus">
<emphasis id="B962EAF0FFA92548FE30F9240700F94A" box="[422,596,1713,1738]" italics="true" pageId="28" pageNumber="29">O. chuzesextus</emphasis>
</taxonomicName>
by having median point band on tergites III and IV fused with posterior band and lacking apical spurs or lateral threads on the anal lobe. From
<taxonomicName id="4C164D61FFA92548FB2CF94C041BF89A" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="chuzesextus">
<emphasis id="B962EAF0FFA92548FB2CF94C041BF89A" italics="true" pageId="28" pageNumber="29">O. chuzesextus</emphasis>
</taxonomicName>
it differs only by its larger size (
<quantity id="4CEE9B07FFA92548FD40F8940630F89B" box="[726,868,1793,1819]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.4" metricValueMin="4.1" pageId="28" pageNumber="29" unit="mm" value="4.75" valueMax="5.4" valueMin="4.1">4.15.4 mm</quantity>
versus 3.0
<quantity id="4CEE9B07FFA92548FC67F8940105F89B" box="[1009,1105,1793,1819]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="28" pageNumber="29" unit="mm" value="3.6">3.6 mm</quantity>
). The larva can be separated from other members of the genus by the presence of labral lamellae combined with a bifid premandible.
</paragraph>
</subSubSection>
<caption id="DF69666AFFA82549FE9EF9670620F8B0" httpUri="https://zenodo.org/record/171317/files/figure.png" pageId="29" pageNumber="30" targetBox="[307,1302,317,1731]" targetPageId="29">
<paragraph id="8BA936E2FFA82549FE9EF9670620F8B0" blockId="29.[264,1323,1778,1840]" pageId="29" pageNumber="30">
<emphasis id="B962EAF0FFA82549FE9EF9670495F88A" bold="true" box="[264,449,1778,1802]" pageId="29" pageNumber="30">FIGURES 812.</emphasis>
<taxonomicName id="4C164D61FFA82549FE51F96706C0F889" box="[455,916,1778,1801]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="lamellatus" status="sp. nov." subGenus="Mesorthocladius">
<emphasis id="B962EAF0FFA82549FE51F9670707F889" box="[455,595,1778,1801]" italics="true" pageId="29" pageNumber="30">Orthocladius</emphasis>
(
<emphasis id="B962EAF0FFA82549FDF5F9670642F889" box="[611,790,1778,1801]" italics="true" pageId="29" pageNumber="30">Mesorthocladius</emphasis>
)
<emphasis id="B962EAF0FFA82549FCB3F96706C0F889" box="[805,916,1778,1801]" italics="true" pageId="29" pageNumber="30">lamellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251578BFFA82549FC0DF9670681F889" box="[923,981,1778,1801]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
, male imago, 8: head, 9: thorax, 10: wing, 11: hypopygium, 12: gonostylus, different view.
</paragraph>
</caption>
<subSubSection id="C30C6569FFAB2575FEAEFE89071DF8B6" lastPageId="33" lastPageNumber="34" pageId="30" pageNumber="31" type="description">
<paragraph id="8BA936E2FFAB254AFEAEFE8906CCFEB6" blockId="30.[264,1323,284,1123]" box="[312,920,284,310]" pageId="30" pageNumber="31">
<emphasis id="B962EAF0FFAB254AFEAEFE890490FEB5" box="[312,452,284,309]" italics="true" pageId="30" pageNumber="31">Male imago</emphasis>
(
<emphasis id="B962EAF0FFAB254AFE43FE8904B0FEB5" box="[469,484,284,309]" italics="true" pageId="30" pageNumber="31">n</emphasis>
= 23 except when otherwise stated)
</paragraph>
<paragraph id="8BA936E2FFAB254AFEAEFED10498FE2E" blockId="30.[264,1323,284,1123]" pageId="30" pageNumber="31">
Total length
<quantity id="4CEE9B07FFAB254AFE41FED107DDFEDE" box="[471,649,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="4.53" metricValueMax="4.57" metricValueMin="4.49" pageId="30" pageNumber="31" unit="mm" value="4.53" valueMax="4.57" valueMin="4.49">4.494.57 mm</quantity>
. Wing length
<quantity id="4CEE9B07FFAB254AFCABFED106BBFEDE" box="[829,1007,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="2.325" metricValueMax="2.41" metricValueMin="2.24" pageId="30" pageNumber="31" unit="mm" value="2.325" valueMax="2.41" valueMin="2.24">2.242.41 mm</quantity>
. Total length/wing length 1.902.00. Wing length/length of profemur 2.602.76. Coloration dark brown with blackish brown vittae.
</paragraph>
<paragraph id="8BA936E2FFAB254AFEAEFE290709FDF6" blockId="30.[264,1323,284,1123]" pageId="30" pageNumber="31">
Head (
<figureCitation id="132D2A67FFAB254AFE13FE290498FE56" box="[389,460,444,470]" captionStart="FIGURES 8 12" captionStartId="29.[264,382,1778,1802]" captionTargetBox="[307,1302,317,1731]" captionTargetId="figure@29.[269,1318,307,1755]" captionTargetPageId="29" captionText="FIGURES 8 12. Orthocladius (Mesorthocladius) lamellatus sp. n., male imago, 8: head, 9: thorax, 10: wing, 11: hypopygium, 12: gonostylus, different view." httpUri="https://zenodo.org/record/171317/files/figure.png" pageId="30" pageNumber="31">Fig. 8</figureCitation>
). AR 1.952.28. Ultimate flagellomere 775841 µm long. Temporal
<taxonomicName id="4C164D61FFAB254AFB66FE29007FFE56" box="[1264,1323,444,470]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
1215, including 36 inner verticals, 6 outer verticals, and 3 postorbitals. Clypeus with 10 12
<taxonomicName id="4C164D61FFAB254AFEBAFD99043EFDA6" box="[300,362,524,550]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Cibarial pump, tentorium and stipes as in
<figureCitation id="132D2A67FFAB254AFCCAFD9906F7FDA6" box="[860,931,524,550]" captionStart="FIGURES 8 12" captionStartId="29.[264,382,1778,1802]" captionTargetBox="[307,1302,317,1731]" captionTargetId="figure@29.[269,1318,307,1755]" captionTargetPageId="29" captionText="FIGURES 8 12. Orthocladius (Mesorthocladius) lamellatus sp. n., male imago, 8: head, 9: thorax, 10: wing, 11: hypopygium, 12: gonostylus, different view." httpUri="https://zenodo.org/record/171317/files/figure.png" pageId="30" pageNumber="31">Fig. 8</figureCitation>
. Tentorium 195199 µm long, 56 µm wide. Stipes 165184 µm long, 68 µm wide. Palpomere lengths (in µm): 4147, 53 64, 86109, 7590, 105135.
</paragraph>
<paragraph id="8BA936E2FFAB254AFEAEFD110049FD6E" blockId="30.[264,1323,284,1123]" pageId="30" pageNumber="31">
Thorax (
<figureCitation id="132D2A67FFAB254AFE0AFD1104B6FD1E" box="[412,482,644,670]" captionStart="FIGURES 8 12" captionStartId="29.[264,382,1778,1802]" captionTargetBox="[307,1302,317,1731]" captionTargetId="figure@29.[269,1318,307,1755]" captionTargetPageId="29" captionText="FIGURES 8 12. Orthocladius (Mesorthocladius) lamellatus sp. n., male imago, 8: head, 9: thorax, 10: wing, 11: hypopygium, 12: gonostylus, different view." httpUri="https://zenodo.org/record/171317/files/figure.png" pageId="30" pageNumber="31">Fig. 9</figureCitation>
,
<emphasis id="B962EAF0FFAB254AFE78FD1104A9FD1D" box="[494,509,644,669]" italics="true" pageId="30" pageNumber="31">n</emphasis>
= 4). Antepronotum with 68, 7
<taxonomicName id="4C164D61FFAB254AFCEAFD1106EEFD1E" box="[892,954,644,670]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Dorsocentrals 1014, 12; anterior dorsocentral 1579, 45 µm long, second 2698, 71 µm long, third 86106, 97 µm long; acrostichals 1318(3); prealars 79, 8. Scutellum with 1321, 17
<taxonomicName id="4C164D61FFAB254AFBD0FD4101D0FD6E" box="[1094,1156,724,750]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
, uni­biserial.
</paragraph>
<paragraph id="8BA936E2FFAB254AFEAEFD69041EFCE6" blockId="30.[264,1323,284,1123]" pageId="30" pageNumber="31">
Wing (
<figureCitation id="132D2A67FFAB254AFE1FFD6904B5FC96" box="[393,481,764,790]" captionStart="FIGURES 8 12" captionStartId="29.[264,382,1778,1802]" captionTargetBox="[307,1302,317,1731]" captionTargetId="figure@29.[269,1318,307,1755]" captionTargetPageId="29" captionText="FIGURES 8 12. Orthocladius (Mesorthocladius) lamellatus sp. n., male imago, 8: head, 9: thorax, 10: wing, 11: hypopygium, 12: gonostylus, different view." httpUri="https://zenodo.org/record/171317/files/figure.png" pageId="30" pageNumber="31">Fig. 10</figureCitation>
). Anal lobe moderately produced. VR 1.03 (1). Costal extension 1517 µm long. Brachiolum with 1 seta, R with 910
<taxonomicName id="4C164D61FFAB254AFCB9FCB10638FCBE" box="[815,876,804,830]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
, other veins bare. Squama with 2531
<taxonomicName id="4C164D61FFAB254AFE9EFCD90412FCE6" box="[264,326,844,870]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
.
</paragraph>
<paragraph id="8BA936E2FFAB254AFEAEFCE1070CFBE3" blockId="30.[264,1323,284,1123]" pageId="30" pageNumber="31">
Legs. Spur of front tibia 7179 µm long, spurs of middle tibia 30 and 2326 µm long, of hind tibia 75 and 2324 µm long. Width at apex of front and middle tibia each 56 µm, of hind tibia
<specimenCount id="9D10FD6BFFAB254AFE13FC510488FC5E" box="[389,476,964,990]" pageId="30" pageNumber="31" type="generic">
<quantity id="4CEE9B07FFAB254AFE13FC510483FC5E" box="[389,471,964,990]" metricMagnitude="1" metricUnit="m" metricValue="6.8" pageId="30" pageNumber="31" unit="m" value="68.0">68 m</quantity>
.
</specimenCount>
Pseudospurs present on ta1 of mid and hind leg, 2334 µm long; one specimen with additional pseudospur on ta2 of mid leg. Sensilla chaetica 3 at 0.100.13 to 0.220.24 of ta1 of mid leg, 68 at 0.100.16 to 0.250.33 on ta1 of hind leg. Lengths (in µm) and proportions of legs:
</paragraph>
<paragraph id="8BA936E2FFAB254AFEE3FB02019BFAA4" pageId="30" pageNumber="31">
<table id="F916C442FFABDAABFE83FB02019BFAAE" box="[277,1231,1175,1326]" gridcols="5" gridrows="4" pageId="30" pageNumber="31">
<tr id="352634A0FFABDAABFE83FB02019BFB34" box="[277,1231,1175,1204]" gridrow="0" pageId="30" pageNumber="31">
<th id="76F75DDCFFABDAABFE83FB020481FB34" box="[277,469,1175,1204]" gridcol="0" gridrow="0" pageId="30" pageNumber="31">fe</th>
<th id="76F75DDCFFABDAABFDA5FB0207C9FB34" box="[563,669,1175,1204]" gridcol="1" gridrow="0" pageId="30" pageNumber="31">ti</th>
<th id="76F75DDCFFABDAABFC97FB020601FB34" box="[769,853,1175,1204]" gridcol="2" gridrow="0" pageId="30" pageNumber="31">ta1</th>
<th id="76F75DDCFFABDAABFC28FB020146FB34" box="[958,1042,1175,1204]" gridcol="3" gridrow="0" pageId="30" pageNumber="31">ta2</th>
<th id="76F75DDCFFABDAABFBEDFB02019BFB34" box="[1147,1231,1175,1204]" gridcol="4" gridrow="0" pageId="30" pageNumber="31">ta3</th>
</tr>
<tr id="352634A0FFABDAABFE83FB2A019BFB5D" box="[277,1231,1215,1245]" gridrow="1" pageId="30" pageNumber="31">
<th id="76F75DDCFFABDAABFE83FB2A0481FB5D" box="[277,469,1215,1245]" gridcol="0" gridrow="1" pageId="30" pageNumber="31">p1 813926</th>
<td id="76F75DDCFFABDAABFDA5FB2A07C9FB5D" box="[563,669,1215,1245]" gridcol="1" gridrow="1" pageId="30" pageNumber="31">8881153</td>
<td id="76F75DDCFFABDAABFC97FB2A0601FB5D" box="[769,853,1215,1245]" gridcol="2" gridrow="1" pageId="30" pageNumber="31">614756</td>
<td id="76F75DDCFFABDAABFC28FB2A0146FB5D" box="[958,1042,1215,1245]" gridcol="3" gridrow="1" pageId="30" pageNumber="31">357482</td>
<td id="76F75DDCFFABDAABFBEDFB2A019BFB5D" box="[1147,1231,1215,1245]" gridcol="4" gridrow="1" pageId="30" pageNumber="31">293331</td>
</tr>
<tr id="352634A0FFABDAABFE83FB72019BFA85" box="[277,1231,1255,1285]" gridrow="2" pageId="30" pageNumber="31">
<th id="76F75DDCFFABDAABFE83FB720481FA85" box="[277,469,1255,1285]" gridcol="0" gridrow="2" pageId="30" pageNumber="31">p2 756954</th>
<td id="76F75DDCFFABDAABFDA5FB7207C9FA85" box="[563,669,1255,1285]" gridcol="1" gridrow="2" pageId="30" pageNumber="31">8881049</td>
<td id="76F75DDCFFABDAABFC97FB720601FA85" box="[769,853,1255,1285]" gridcol="2" gridrow="2" pageId="30" pageNumber="31">425482</td>
<td id="76F75DDCFFABDAABFC28FB720146FA85" box="[958,1042,1255,1285]" gridcol="3" gridrow="2" pageId="30" pageNumber="31">274302</td>
<td id="76F75DDCFFABDAABFBEDFB72019BFA85" box="[1147,1231,1255,1285]" gridcol="4" gridrow="2" pageId="30" pageNumber="31">217222</td>
</tr>
<tr id="352634A0FFABDAABFE83FA9A019BFAAE" box="[277,1231,1295,1326]" gridrow="3" pageId="30" pageNumber="31">
<th id="76F75DDCFFABDAABFE83FA9A0481FAAE" box="[277,469,1295,1326]" gridcol="0" gridrow="3" pageId="30" pageNumber="31">p3 8571068</th>
<td id="76F75DDCFFABDAABFDA5FA9A07C9FAAE" box="[563,669,1295,1326]" gridcol="1" gridrow="3" pageId="30" pageNumber="31">11341276</td>
<td id="76F75DDCFFABDAABFC97FA9A0601FAAE" box="[769,853,1295,1326]" gridcol="2" gridrow="3" pageId="30" pageNumber="31">605671</td>
<td id="76F75DDCFFABDAABFC28FA9A0146FAAE" box="[958,1042,1295,1326]" gridcol="3" gridrow="3" pageId="30" pageNumber="31">394416</td>
<td id="76F75DDCFFABDAABFBEDFA9A019BFAAE" box="[1147,1231,1295,1326]" gridcol="4" gridrow="3" pageId="30" pageNumber="31">284302</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA936E2FFAB254AFEE3FAF401B2FA6E" pageId="30" pageNumber="31">
<table id="F916C442FFABDAABFE83FAF401B2FA77" box="[277,1254,1377,1527]" gridcols="7" gridrows="4" pageId="30" pageNumber="31">
<tr id="352634A0FFABDAABFE83FAF401B2FAFE" box="[277,1254,1377,1406]" gridrow="0" pageId="30" pageNumber="31" rowspan-0="1">
<th id="76F75DDCFFABDAABFEE3FAF4049EFAFE" box="[373,458,1377,1406]" gridcol="1" gridrow="0" pageId="30" pageNumber="31">ta4</th>
<th id="76F75DDCFFABDAABFE69FAF40707FAFE" box="[511,595,1377,1406]" gridcol="2" gridrow="0" pageId="30" pageNumber="31">ta5</th>
<th id="76F75DDCFFABDAABFD0FFAF407AEFAFE" box="[665,762,1377,1406]" gridcol="3" gridrow="0" pageId="30" pageNumber="31">LR</th>
<th id="76F75DDCFFABDAABFCD3FAF406F2FAFE" box="[837,934,1377,1406]" gridcol="4" gridrow="0" pageId="30" pageNumber="31">BV</th>
<th id="76F75DDCFFABDAABFB94FAF40137FAFE" box="[1026,1123,1377,1406]" gridcol="5" gridrow="0" pageId="30" pageNumber="31">SV</th>
<th id="76F75DDCFFABDAABFB0BFAF401B2FAFE" box="[1181,1254,1377,1406]" gridcol="6" gridrow="0" pageId="30" pageNumber="31">BR</th>
</tr>
<tr id="352634A0FFABDAABFE83FA1D01B2FA26" box="[277,1254,1416,1446]" gridrow="1" pageId="30" pageNumber="31">
<th id="76F75DDCFFABDAABFE83FA1D047AFA26" box="[277,302,1416,1446]" gridcol="0" gridrow="1" pageId="30" pageNumber="31">p1</th>
<td id="76F75DDCFFABDAABFEE3FA1D049EFA26" box="[373,458,1416,1446]" gridcol="1" gridrow="1" pageId="30" pageNumber="31">148203</td>
<td id="76F75DDCFFABDAABFE69FA1D0707FA26" box="[511,595,1416,1446]" gridcol="2" gridrow="1" pageId="30" pageNumber="31">132142</td>
<td id="76F75DDCFFABDAABFD0FFA1D07AEFA26" box="[665,762,1416,1446]" gridcol="3" gridrow="1" pageId="30" pageNumber="31">0.660.69</td>
<td id="76F75DDCFFABDAABFCD3FA1D06F2FA26" box="[837,934,1416,1446]" gridcol="4" gridrow="1" pageId="30" pageNumber="31">2.272.45</td>
<td id="76F75DDCFFABDAABFB94FA1D0137FA26" box="[1026,1123,1416,1446]" gridcol="5" gridrow="1" pageId="30" pageNumber="31">2.752.77</td>
<td id="76F75DDCFFABDAABFB0BFA1D01B2FA26" box="[1181,1254,1416,1446]" gridcol="6" gridrow="1" pageId="30" pageNumber="31">1.62.</td>
</tr>
<tr id="352634A0FFABDAABFE83FA2401B2FA4F" box="[277,1254,1457,1487]" gridrow="2" pageId="30" pageNumber="31">
<th id="76F75DDCFFABDAABFE83FA24047AFA4F" box="[277,302,1457,1487]" gridcol="0" gridrow="2" pageId="30" pageNumber="31">p2</th>
<td id="76F75DDCFFABDAABFEE3FA24049EFA4F" box="[373,458,1457,1487]" gridcol="1" gridrow="2" pageId="30" pageNumber="31">151</td>
<td id="76F75DDCFFABDAABFE69FA240707FA4F" box="[511,595,1457,1487]" gridcol="2" gridrow="2" pageId="30" pageNumber="31">113132</td>
<td id="76F75DDCFFABDAABFD0FFA2407AEFA4F" box="[665,762,1457,1487]" gridcol="3" gridrow="2" pageId="30" pageNumber="31">0.460.49</td>
<td id="76F75DDCFFABDAABFCD3FA2406F2FA4F" box="[837,934,1457,1487]" gridcol="4" gridrow="2" pageId="30" pageNumber="31">3.30</td>
<td id="76F75DDCFFABDAABFB94FA240137FA4F" box="[1026,1123,1457,1487]" gridcol="5" gridrow="2" pageId="30" pageNumber="31">4.494.53</td>
<td id="76F75DDCFFABDAABFB0BFA2401B2FA4F" box="[1181,1254,1457,1487]" gridcol="6" gridrow="2" pageId="30" pageNumber="31">2.62.8</td>
</tr>
<tr id="352634A0FFABDAABFE83FA4C01B2FA77" box="[277,1254,1497,1527]" gridrow="3" pageId="30" pageNumber="31">
<th id="76F75DDCFFABDAABFE83FA4C047AFA77" box="[277,302,1497,1527]" gridcol="0" gridrow="3" pageId="30" pageNumber="31">p3</th>
<td id="76F75DDCFFABDAABFEE3FA4C049EFA77" box="[373,458,1497,1527]" gridcol="1" gridrow="3" pageId="30" pageNumber="31">175189</td>
<td id="76F75DDCFFABDAABFE69FA4C0707FA77" box="[511,595,1497,1527]" gridcol="2" gridrow="3" pageId="30" pageNumber="31">132142</td>
<td id="76F75DDCFFABDAABFD0FFA4C07AEFA77" box="[665,762,1497,1527]" gridcol="3" gridrow="3" pageId="30" pageNumber="31">0.53</td>
<td id="76F75DDCFFABDAABFCD3FA4C06F2FA77" box="[837,934,1497,1527]" gridcol="4" gridrow="3" pageId="30" pageNumber="31">2.963.10</td>
<td id="76F75DDCFFABDAABFB94FA4C0137FA77" box="[1026,1123,1497,1527]" gridcol="5" gridrow="3" pageId="30" pageNumber="31">3.143.49</td>
<td id="76F75DDCFFABDAABFB0BFA4C01B2FA77" box="[1181,1254,1497,1527]" gridcol="6" gridrow="3" pageId="30" pageNumber="31">2.93.0</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA936E2FFAB254AFEAEF9B90678F8DE" blockId="30.[264,1323,1580,1886]" pageId="30" pageNumber="31">
Hypopygium (
<figureCitation id="132D2A67FFAB254AFE71F9B9072BF9C6" box="[487,639,1580,1606]" captionStart="FIGURES 8 12" captionStartId="29.[264,382,1778,1802]" captionTargetBox="[307,1302,317,1731]" captionTargetId="figure@29.[269,1318,307,1755]" captionTargetPageId="29" captionText="FIGURES 8 12. Orthocladius (Mesorthocladius) lamellatus sp. n., male imago, 8: head, 9: thorax, 10: wing, 11: hypopygium, 12: gonostylus, different view." httpUri="https://zenodo.org/record/171317/files/figure.png" pageId="30" pageNumber="31">Figs. 11, 12</figureCitation>
). Anal point 86120, 109 µm long including triangular base; robust, tapering with rounded apex; with 1014, 13
<taxonomicName id="4C164D61FFAB254AFC26F9C106BFF9EE" box="[944,1003,1620,1646]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
including on base; laterosternite IX with 1011, 11
<taxonomicName id="4C164D61FFAB254AFDD3F9E907D7F916" box="[581,643,1660,1686]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Phallapodeme 5386, 74 µm long; transverse sternapodeme 8399, 95 µm long, oral projections moderately to well developed. Gonocoxite 308 364, 342 µm long; superior volsella collar­like; inferior volsella with ventral part not extended beyond dorsal part.. Gonostylus 135167, 152 µm long; crista dorsalis apical, rounded; megaseta 1019 µm (3) long. Virga 2636, 32 µm long; consisting of cluster of 7 spines. HR 2.182.37, 2.26; HV 2.743.12 (2).
</paragraph>
<paragraph id="8BA936E2FFAA254BFEAEFE890785FEB6" blockId="31.[264,1325,284,579]" box="[312,721,284,310]" pageId="31" pageNumber="32">
<emphasis id="B962EAF0FFAA254BFEAEFE8904B4FEB5" box="[312,480,284,309]" italics="true" pageId="31" pageNumber="32">Female imago</emphasis>
(
<emphasis id="B962EAF0FFAA254BFE67FE890754FEB5" box="[497,512,284,309]" italics="true" pageId="31" pageNumber="32">n</emphasis>
= 1 mature pupa)
</paragraph>
<paragraph id="8BA936E2FFAA254BFEAEFED101CDFEDE" blockId="31.[264,1325,284,579]" box="[312,1177,324,350]" pageId="31" pageNumber="32">
Total length about
<quantity id="4CEE9B07FFAA254BFD82FED1072BFEDE" box="[532,639,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="31" pageNumber="32" unit="mm" value="4.1">4.10 mm</quantity>
. Coloration apparently not as dark as in male.
</paragraph>
<paragraph id="8BA936E2FFAA254BFEAEFEF90695FE51" blockId="31.[264,1325,284,579]" pageId="31" pageNumber="32">
Head. AR 0.75. Flagellomere lengths (in µm): 56, 38, 38, 49, 154. Coronal suture 56 µm long. Temporal
<taxonomicName id="4C164D61FFAA254BFE65FE04077AFE2B" box="[499,558,401,427]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
not measurable. Clypeus with 14
<taxonomicName id="4C164D61FFAA254BFC54FE040154FE2B" box="[962,1024,401,427]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Tentorium 150 µm long, 41 µm wide. Palpomere lengths (in µm): 38, 45, 70, 86, 90.
</paragraph>
<paragraph id="8BA936E2FFAA254BFEAEFE49019BFE76" blockId="31.[264,1325,284,579]" box="[312,1231,476,502]" pageId="31" pageNumber="32">
Thorax. Dorsocentrals 12, acrostichals 11, prealars 6. Scutellum with 13
<taxonomicName id="4C164D61FFAA254BFB1BFE49019FFE76" box="[1165,1227,476,502]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
.
</paragraph>
<paragraph id="8BA936E2FFAA254BFEAEFD9407DAFD9B" blockId="31.[264,1325,284,579]" box="[312,654,513,539]" pageId="31" pageNumber="32">
Wing. Squama with 25
<taxonomicName id="4C164D61FFAA254BFDDAFD9407DEFD9B" box="[588,650,513,539]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
.
</paragraph>
<paragraph id="8BA936E2FFAA254BFEAEFDBC0766FDC3" blockId="31.[264,1325,284,579]" box="[312,562,553,579]" pageId="31" pageNumber="32">Legs not measurable.</paragraph>
<caption id="DF69666AFFAA254BFE9EF97D0665F8E0" httpUri="https://zenodo.org/record/171318/files/figure.png" pageId="31" pageNumber="32" targetBox="[285,1303,643,1752]" targetPageId="31">
<paragraph id="8BA936E2FFAA254BFE9EF97D0665F8E0" blockId="31.[264,1323,1768,1888]" pageId="31" pageNumber="32">
<emphasis id="B962EAF0FFAA254BFE9EF97D0480F880" bold="true" box="[264,468,1768,1792]" pageId="31" pageNumber="32">FIGURES 1322.</emphasis>
<taxonomicName id="4C164D61FFAA254BFE49F97C0669F880" authority="(Mesorthocladius)" baseAuthorityName="Mesorthocladius" box="[479,829,1769,1792]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B962EAF0FFAA254BFE49F97C073FF880" box="[479,619,1769,1792]" italics="true" pageId="31" pageNumber="32">Orthocladius</emphasis>
(
<emphasis id="B962EAF0FFAA254BFD16F97C0667F880" box="[640,819,1769,1792]" italics="true" pageId="31" pageNumber="32">Mesorthocladius</emphasis>
)
</taxonomicName>
spp., female imagines, genitalia, 1317:
<taxonomicName id="4C164D61FFAA254BFA87F97C04FAF8A0" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="lamellatus" status="sp. nov." subGenus="Mesorthocladius">
<emphasis id="B962EAF0FFAA254BFA87F97C007FF880" box="[1297,1323,1769,1792]" italics="true" pageId="31" pageNumber="32">O.</emphasis>
(
<emphasis id="B962EAF0FFAA254BFE87F89C047BF8A0" box="[273,303,1801,1824]" italics="true" pageId="31" pageNumber="32">M.</emphasis>
)
<emphasis id="B962EAF0FFAA254BFEA9F89C04FAF8A0" box="[319,430,1801,1824]" italics="true" pageId="31" pageNumber="32">lamellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251578BFFAA254BFE21F89C04A7F8A0" box="[439,499,1801,1824]" pageId="31" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
, 13: ventral view, 14: ventrolateral lobe, 15: dorsomesal lobe, 16: apodeme lobe, 17: dorsal view, 1822:
<taxonomicName id="4C164D61FFAA254BFDDDF8BC0650F8C0" box="[587,772,1833,1856]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nimidens" status="sp. nov." subGenus="Mesorthocladius">
<emphasis id="B962EAF0FFAA254BFDDDF8BC0731F8C0" box="[587,613,1833,1856]" italics="true" pageId="31" pageNumber="32">O.</emphasis>
(
<emphasis id="B962EAF0FFAA254BFDE0F8BC07C0F8C0" box="[630,660,1833,1856]" italics="true" pageId="31" pageNumber="32">M.</emphasis>
)
<emphasis id="B962EAF0FFAA254BFD32F8BC0650F8C0" box="[676,772,1833,1856]" italics="true" pageId="31" pageNumber="32">nimidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251578BFFAA254BFC9AF8BC061CF8C0" box="[780,840,1833,1856]" pageId="31" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
, 18: ventral view, 19: ventrolateral lobe, 20: dorsomesal lobe, 21: apodeme lobe, 22 dorsal view.
</paragraph>
</caption>
<paragraph id="8BA936E2FF952574FEAEFE890164FEB6" blockId="32.[264,1324,284,510]" box="[312,1072,284,310]" pageId="32" pageNumber="33">
Abdomen. Tergite VIII with 32
<taxonomicName id="4C164D61FF952574FD3BFE8907BEFEB6" box="[685,746,284,310]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
, sternite VIII with 23
<taxonomicName id="4C164D61FF952574FC78FE890178FEB6" box="[1006,1068,284,310]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
.
</paragraph>
<paragraph id="8BA936E2FF952574FEAEFED10047FE2E" blockId="32.[264,1324,284,510]" pageId="32" pageNumber="33">
Genitalia (
<figureCitation id="132D2A67FF952574FE23FED10717FEDE" box="[437,579,324,350]" captionStart="FIGURES 13 22" captionStartId="31.[264,383,1768,1792]" captionTargetBox="[285,1303,643,1752]" captionTargetId="figure@31.[269,1318,618,1765]" captionTargetPageId="31" captionText="FIGURES 13 22. Orthocladius (Mesorthocladius) spp., female imagines, genitalia, 13 17: O. (M.) lamellatus sp. n., 13: ventral view, 14: ventrolateral lobe, 15: dorsomesal lobe, 16: apodeme lobe, 17: dorsal view, 18 22: O. (M.) nimidens sp. n., 18: ventral view, 19: ventrolateral lobe, 20: dorsomesal lobe, 21: apodeme lobe, 22 dorsal view." httpUri="https://zenodo.org/record/171318/files/figure.png" pageId="32" pageNumber="33">Figs. 1317</figureCitation>
). Tergite IX divided, with 24
<taxonomicName id="4C164D61FF952574FC3FFED106B3FEDE" box="[937,999,324,350]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Gonocoxite with 19
<taxonomicName id="4C164D61FF952574FB7FFED10073FEDE" box="[1257,1319,324,350]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Cercus 131 µm long. Seminal capsule pear­shaped with apical rugulosity, 84 µm long, 53 µm wide. Spermathecal ducts with one bend and separate openings. Notum 98 µm long.
</paragraph>
<paragraph id="8BA936E2FF952574FEAEFE29061CFE56" blockId="32.[264,1324,284,510]" box="[312,840,444,470]" pageId="32" pageNumber="33">
<emphasis id="B962EAF0FF952574FEAEFE290422FE55" box="[312,374,444,469]" italics="true" pageId="32" pageNumber="33">Pupa</emphasis>
(
<emphasis id="B962EAF0FF952574FE10FE2904C1FE55" box="[390,405,444,469]" italics="true" pageId="32" pageNumber="33">n</emphasis>
= 34 except when otherwise stated)
</paragraph>
<paragraph id="8BA936E2FF952574FEAEFE710057FE7E" blockId="32.[264,1324,284,510]" box="[312,1283,484,510]" pageId="32" pageNumber="33">
Total length
<quantity id="4CEE9B07FF952574FE5DFE71072CFE7E" box="[459,632,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="4.775" metricValueMax="5.39" metricValueMin="4.16" pageId="32" pageNumber="33" unit="mm" value="4.775" valueMax="5.39" valueMin="4.16">4.165.39 mm</quantity>
. Exuviae pale greyish brown with brownish apophyses.
</paragraph>
<caption id="DF69666AFF952574FE9EF883043FF8ED" httpUri="https://zenodo.org/record/171319/files/figure.png" pageId="32" pageNumber="33" targetBox="[362,1271,556,1802]" targetPageId="32">
<paragraph id="8BA936E2FF952574FE9EF883043FF8ED" blockId="32.[264,1323,1814,1901]" pageId="32" pageNumber="33">
<emphasis id="B962EAF0FF952574FE9EF883049BF8AE" bold="true" box="[264,463,1814,1838]" pageId="32" pageNumber="33">FIGURES 2327.</emphasis>
<taxonomicName id="4C164D61FF952574FE40F883067AF8AD" authority="(Mesorthocladius)" baseAuthorityName="Mesorthocladius" box="[470,814,1814,1837]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B962EAF0FF952574FE40F8830737F8AD" box="[470,611,1814,1837]" italics="true" pageId="32" pageNumber="33">Orthocladius</emphasis>
(
<emphasis id="B962EAF0FF952574FDE4F8830671F8AD" box="[626,805,1814,1837]" italics="true" pageId="32" pageNumber="33">Mesorthocladius</emphasis>
)
</taxonomicName>
spp., pupae, 23:
<taxonomicName id="4C164D61FF952574FC73F88301FEF8AD" box="[997,1194,1814,1837]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lamellatus" status="sp. nov." subGenus="Mesorthocladius">
<emphasis id="B962EAF0FF952574FC73F88306ABF8AD" box="[997,1023,1814,1837]" italics="true" pageId="32" pageNumber="33">O.</emphasis>
(
<emphasis id="B962EAF0FF952574FB99F8830178F8AD" box="[1039,1068,1814,1837]" italics="true" pageId="32" pageNumber="33">M.</emphasis>
)
<emphasis id="B962EAF0FF952574FBADF88301FEF8AD" box="[1083,1194,1814,1837]" italics="true" pageId="32" pageNumber="33">lamellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251578BFF952574FB24F88301B8F8AD" box="[1202,1260,1814,1837]" pageId="32" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
, tergites, 24: thoracic horn, 2527:
<taxonomicName id="4C164D61FF952574FDCFF8A30647F8CD" box="[601,787,1846,1869]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nimidens" status="sp. nov." subGenus="Mesorthocladius">
<emphasis id="B962EAF0FF952574FDCFF8A30727F8CD" box="[601,627,1846,1869]" italics="true" pageId="32" pageNumber="33">O.</emphasis>
(
<emphasis id="B962EAF0FF952574FD12F8A307F6F8CD" box="[644,674,1846,1869]" italics="true" pageId="32" pageNumber="33">M.</emphasis>
)
<emphasis id="B962EAF0FF952574FD25F8A30647F8CD" box="[691,787,1846,1869]" italics="true" pageId="32" pageNumber="33">nimidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251578BFF952574FC8AF8A3060DF8CD" box="[796,857,1846,1869]" pageId="32" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
, 25: tergites, 26: thoracic horn, 27: frontal apotome.
</paragraph>
</caption>
<paragraph id="8BA936E2FF942575FEAEFE8906FBFD86" blockId="33.[264,1323,284,1846]" pageId="33" pageNumber="34">
Cephalothorax. Frontal
<taxonomicName id="4C164D61FF942575FDC7FE8907DAFEB6" box="[593,654,284,310]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
143150 µm long. Thoracic horn (
<figureCitation id="132D2A67FF942575FBB5FE89012EFEB6" box="[1059,1146,284,310]" captionStart="FIGURES 23 27" captionStartId="32.[264,382,1814,1838]" captionTargetBox="[362,1271,556,1802]" captionTargetId="figure@32.[312,1277,554,1804]" captionTargetPageId="32" captionText="FIGURES 23 27. Orthocladius (Mesorthocladius) spp., pupae, 23: O. (M.) lamellatus sp. n., tergites, 24: thoracic horn, 25 27: O. (M.) nimidens sp. n., 25: tergites, 26: thoracic horn, 27: frontal apotome." httpUri="https://zenodo.org/record/171319/files/figure.png" pageId="33" pageNumber="34">Fig. 24</figureCitation>
) 270428, 340 µm long; 4153, 49 µm wide; 4.148.77, 7.00 times as long as wide; 1.161, 54, 1.35 times as long as anal macrosetae. Precorneal
<taxonomicName id="4C164D61FF942575FC8EFEF90607FE06" box="[792,851,364,390]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
respectively 94128, 105 µm; 150171, 166 µm; and 68101, 91 µm long. Anterior dorsocentral (Dc1) 7990, 83 µm long; Dc2, 4979, 66 µm; Dc3 5675, 71 µm; and Dc4 6483, 71 µm long. Distances (in µm): Dc1 Dc
<date id="FFA81022FF942575FEBCFE6D04D0FD86" box="[298,388,492,521]" pageId="33" pageNumber="34" value="1949-02-26">2 2649</date>
, 40; Dc2Dc3 98131, 113; Dc3Dc
<date id="FFA81022FF942575FCB2FE6D062AFD86" box="[804,894,492,521]" pageId="33" pageNumber="34" value="1934-04-19">4 1934</date>
, 24.
</paragraph>
<paragraph id="8BA936E2FF942575FEAEFD8C0708FC1E" blockId="33.[264,1323,284,1846]" pageId="33" pageNumber="34">
Abdomen (
<figureCitation id="132D2A67FF942575FE2DFD8C0746FDB3" box="[443,530,537,563]" captionStart="FIGURES 23 27" captionStartId="32.[264,382,1814,1838]" captionTargetBox="[362,1271,556,1802]" captionTargetId="figure@32.[312,1277,554,1804]" captionTargetPageId="32" captionText="FIGURES 23 27. Orthocladius (Mesorthocladius) spp., pupae, 23: O. (M.) lamellatus sp. n., tergites, 24: thoracic horn, 25 27: O. (M.) nimidens sp. n., 25: tergites, 26: thoracic horn, 27: frontal apotome." httpUri="https://zenodo.org/record/171319/files/figure.png" pageId="33" pageNumber="34">Fig. 23</figureCitation>
). Tergite I bare, tergite II with weak mediolateral shagreen, stronger posterior shagreen on tergites IIIVII, shagreen relative extensive on tergites VIIIIX. Shagreen spinules about equal in size, anterior and posterior bands joined, anterior band not extending as far lateral as posterior bands. Sternites I and IX bare; IIIII with anterolateral plus posterolateral shagreen, IV with lateral, VVII with lateral plus anterolateral, VIII with anterolateral shagreen. Pedes spurii A on sternites IVVII. Pedes spurii B absent. Caudal hooklets on tergite
<date id="FFA81022FF942575FD30FC9C078BFCA3" box="[678,735,777,803]" pageId="33" pageNumber="34" value="1946-02">II 46</date>
69, 57; in 3 rows. Segments IIV each with 3
<taxonomicName id="4C164D61FF942575FA8FFC9C0411FCCB" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="setae">L setae</taxonomicName>
, VI and VII each with 34, and VIII with 4
<taxonomicName id="4C164D61FF942575FCC0FCA406FBFCCB" box="[854,943,817,843]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="setae">L setae</taxonomicName>
. Lengths of L1L4
<taxonomicName id="4C164D61FF942575FB06FCA4019FFCCB" box="[1168,1227,817,843]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
on segment VIII (in µm) as: 64113; 75124, 98; 38116, 65; 68101, 85. Occasionally a forked V seta present on sternite III.
</paragraph>
<paragraph id="8BA936E2FF942575FEAEFC390107FB96" blockId="33.[264,1323,284,1846]" pageId="33" pageNumber="34">Anal lobe 364443, 400 µm long. Anal macrosetae subequal in length; 225278, 249 µm long; 0.590.64, 0.62 as long as anal lobe. Male genital sac overreaching anal lobe by 30 µm (2), female genital sac ending 105 µm short of apex of anal lobe.</paragraph>
<paragraph id="8BA936E2FF942575FEAEFBB106AEFBBE" blockId="33.[264,1323,284,1846]" box="[312,1018,1060,1086]" pageId="33" pageNumber="34">
<emphasis id="B962EAF0FF942575FEAEFBB1074DFBBD" box="[312,537,1060,1085]" italics="true" pageId="33" pageNumber="34">Fourth instar larva</emphasis>
(
<emphasis id="B962EAF0FF942575FDBDFBB1076EFBBD" box="[555,570,1060,1085]" italics="true" pageId="33" pageNumber="34">n</emphasis>
= 911 except when otherwise stated)
</paragraph>
<paragraph id="8BA936E2FF942575FEAEFBD90607FB0E" blockId="33.[264,1323,284,1846]" pageId="33" pageNumber="34">
Total length 4.688.60,
<quantity id="4CEE9B07FF942575FDDAFBD907E3FBE6" box="[588,695,1100,1126]" metricMagnitude="-3" metricUnit="m" metricValue="6.06" pageId="33" pageNumber="34" unit="mm" value="6.06">6.06 mm</quantity>
. Head capsule 0.440.50,
<quantity id="4CEE9B07FF942575FC71FBD9011BFBE6" box="[999,1103,1100,1126]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="33" pageNumber="34" unit="mm" value="0.47">0.47 mm</quantity>
long. Postmentum 195212, 203 µm long. Head capsule dark brown.
</paragraph>
<paragraph id="8BA936E2FF942575FEAEFB09010DF9C6" blockId="33.[264,1323,284,1846]" pageId="33" pageNumber="34">
Head. Antenna as in
<figureCitation id="132D2A67FF942575FDB8FB0907D1FB36" box="[558,645,1180,1206]" captionStart="FIGURES 28 34" captionStartId="34.[264,383,1688,1712]" captionTargetBox="[308,1302,296,1678]" captionTargetId="figure@34.[269,1318,280,1682]" captionTargetPageId="34" captionText="FIGURES 28 34. Orthocladius spp., larvae, 28: Orthocladius (Mesorthocladius) frigidus (Zetterstedt) labrum, epipharyngeal area and premandible, 29, 30: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 29: antenna, 30: labrum, epipharyngeal area and premandible, 31, 32: Orthocladius (Mesorthocladius) lamellatus sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible, 33 34: Orthocladius (Mesorthocladius) nimidens sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible." httpUri="https://zenodo.org/record/171320/files/figure.png" pageId="33" pageNumber="34">Fig. 31</figureCitation>
. Length of antennal segments (in µm): 4964, 55; 1215, 13; 47, 5; 34, 4; 34, 4. AR 1.752.32, 1.96. Ring organ 1019, 15 µm from base; basal setal mark 1422, 19 µm from base; apical setal mark 2037, 26 µm from base. Basal antennal segment 1923, 20 µm wide; blade 2026, 23 µm (7) long; accessory blade12 17, 15 µm (7) long. Lauterborn organs 59, 7 µm long; style 59, 7 µm long. Two labral lamellae (
<figureCitation id="132D2A67FF942575FEEAFAF10487FAFE" box="[380,467,1380,1406]" captionStart="FIGURES 28 34" captionStartId="34.[264,383,1688,1712]" captionTargetBox="[308,1302,296,1678]" captionTargetId="figure@34.[269,1318,280,1682]" captionTargetPageId="34" captionText="FIGURES 28 34. Orthocladius spp., larvae, 28: Orthocladius (Mesorthocladius) frigidus (Zetterstedt) labrum, epipharyngeal area and premandible, 29, 30: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 29: antenna, 30: labrum, epipharyngeal area and premandible, 31, 32: Orthocladius (Mesorthocladius) lamellatus sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible, 33 34: Orthocladius (Mesorthocladius) nimidens sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible." httpUri="https://zenodo.org/record/171320/files/figure.png" pageId="33" pageNumber="34">Fig. 32</figureCitation>
) each with 37, 5 spines. With 12 chaetae and 23 spinulae. One pair of chaetulae laterales slightly serrated, chaetulae basales tapering, bifid, with 1 long and 1 short apical tooth. Premandible (
<figureCitation id="132D2A67FF942575FD18FA2107B2FA4E" box="[654,742,1460,1486]" captionStart="FIGURES 28 34" captionStartId="34.[264,383,1688,1712]" captionTargetBox="[308,1302,296,1678]" captionTargetId="figure@34.[269,1318,280,1682]" captionTargetPageId="34" captionText="FIGURES 28 34. Orthocladius spp., larvae, 28: Orthocladius (Mesorthocladius) frigidus (Zetterstedt) labrum, epipharyngeal area and premandible, 29, 30: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 29: antenna, 30: labrum, epipharyngeal area and premandible, 31, 32: Orthocladius (Mesorthocladius) lamellatus sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible, 33 34: Orthocladius (Mesorthocladius) nimidens sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible." httpUri="https://zenodo.org/record/171320/files/figure.png" pageId="33" pageNumber="34">Fig. 32</figureCitation>
) bifid, 79109, 89 µm long; brush consisting of 1116, 14 (7) spines present. Mandible (
<figureCitation id="132D2A67FF942575FD6BFA490603FA76" box="[765,855,1500,1526]" captionStart="FIGURES 35 40" captionStartId="36.[264,383,1691,1715]" captionTargetBox="[281,1294,319,1672]" captionTargetId="figure@36.[269,1318,299,1687]" captionTargetPageId="36" captionText="FIGURES 35 40. Orthocladius spp., larvae, 35, 36: Orthocladius (Mesorthocladius) lamellatus sp. n., 35: mandible, 36: mentum, 37 38: Orthocladius (Mesorthocladius) nimidens sp. n., 37: mandible, 38: mentum, 39 40: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 39: mandible, 40: mentum" httpUri="https://zenodo.org/record/171321/files/figure.png" pageId="33" pageNumber="34">Fig. 35</figureCitation>
) 146165, 157 µm long, without seta interna. Mentum (
<figureCitation id="132D2A67FF942575FE4AF9910766F99E" box="[476,562,1540,1566]" captionStart="FIGURES 35 40" captionStartId="36.[264,383,1691,1715]" captionTargetBox="[281,1294,319,1672]" captionTargetId="figure@36.[269,1318,299,1687]" captionTargetPageId="36" captionText="FIGURES 35 40. Orthocladius spp., larvae, 35, 36: Orthocladius (Mesorthocladius) lamellatus sp. n., 35: mandible, 36: mentum, 37 38: Orthocladius (Mesorthocladius) nimidens sp. n., 37: mandible, 38: mentum, 39 40: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 39: mandible, 40: mentum" httpUri="https://zenodo.org/record/171321/files/figure.png" pageId="33" pageNumber="34">Fig. 36</figureCitation>
) 120144, 133 µm wide; median tooth 1923, 20 µm wide; ventromental plate 412, 8 µm wide; with 57, 7
<taxonomicName id="4C164D61FF942575FCB6F9B9060FF9C6" box="[800,859,1580,1606]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
underneath on cardo.
</paragraph>
<paragraph id="8BA936E2FF942575FEAEF9C1071DF8B6" blockId="33.[264,1323,284,1846]" pageId="33" pageNumber="34">
Abdomen. Claws of anterior parapods strongly serrated. Longest body
<taxonomicName id="4C164D61FF942575FBEAF9C101E3F9EE" box="[1148,1207,1620,1646]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
131278, 186 µm long. Posterior parapods 293450, 352 µm (7) long. Procercus 3853, 46 µm high; 3445, 41 µm wide. Anal
<taxonomicName id="4C164D61FF942575FDA9F931072EF93E" box="[575,634,1700,1726]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
506675, 595 µm long; supraanal seta 64105, 84 µm (7) long, 0.110.20, 0.15 (8) as long as anal
<taxonomicName id="4C164D61FF942575FD79F9590679F966" box="[751,813,1740,1766]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Longer anal tubules 150278, 183 µm (6) long, 5683, 72 µm (4) wide at base; shorter anal tubules 113184, 153 µm (6) long, 41 60, 54 µm (7) wide at base.
</paragraph>
</subSubSection>
<caption id="DF69666AFF972576FE9EF90D07C7F8DD" httpUri="https://zenodo.org/record/171320/files/figure.png" pageId="34" pageNumber="35" targetBox="[308,1302,296,1678]" targetPageId="34">
<paragraph id="8BA936E2FF972576FE9EF90D07C7F8DD" blockId="34.[264,1323,1688,1885]" pageId="34" pageNumber="35">
<emphasis id="B962EAF0FF972576FE9EF90D0484F930" bold="true" box="[264,464,1688,1712]" pageId="34" pageNumber="35">FIGURES 2834.</emphasis>
<taxonomicName id="4C164D61FF972576FE41F90C0730F930" box="[471,612,1689,1712]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B962EAF0FF972576FE41F90C0730F930" box="[471,612,1689,1712]" italics="true" pageId="34" pageNumber="35">Orthocladius</emphasis>
</taxonomicName>
spp., larvae, 28:
<taxonomicName id="4C164D61FF972576FC88F90C0417F952" authority="Zetterstedt" authorityName="Zetterstedt" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="frigidus" subGenus="Mesorthocladius">
<emphasis id="B962EAF0FF972576FC88F90C06FEF930" box="[798,938,1689,1712]" italics="true" pageId="34" pageNumber="35">Orthocladius</emphasis>
(
<emphasis id="B962EAF0FF972576FC2DF90C013AF930" box="[955,1134,1689,1712]" italics="true" pageId="34" pageNumber="35">Mesorthocladius</emphasis>
)
<emphasis id="B962EAF0FF972576FBE9F90C0186F930" box="[1151,1234,1689,1712]" italics="true" pageId="34" pageNumber="35">frigidus</emphasis>
(Zetterstedt)
</taxonomicName>
labrum, epipharyngeal area and premandible, 29, 30:
<taxonomicName id="4C164D61FF972576FC11F92E048AF975" authority="Lundstrom" authorityName="Lundstrom" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="nitidoscutellatus" subGenus="Orthocladius">
<emphasis id="B962EAF0FF972576FC11F92E0140F952" box="[903,1044,1723,1746]" italics="true" pageId="34" pageNumber="35">Orthocladius</emphasis>
(
<emphasis id="B962EAF0FF972576FBB5F92E01E4F952" box="[1059,1200,1723,1746]" italics="true" pageId="34" pageNumber="35">Orthocladius</emphasis>
)
<emphasis id="B962EAF0FF972576FB56F92E0402F975" italics="true" pageId="34" pageNumber="35">nitidoscutellatus</emphasis>
Lundström
</taxonomicName>
, 29: antenna, 30: labrum, epipharyngeal area and premandible, 31, 32:
<taxonomicName id="4C164D61FF972576FE9EF8940783F898" box="[264,727,1793,1816]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lamellatus" status="sp. nov." subGenus="Mesorthocladius">
<emphasis id="B962EAF0FF972576FE9EF89404C0F898" box="[264,404,1793,1816]" italics="true" pageId="34" pageNumber="35">Orthocladius</emphasis>
(
<emphasis id="B962EAF0FF972576FE33F894070CF898" box="[421,600,1793,1816]" italics="true" pageId="34" pageNumber="35">Mesorthocladius</emphasis>
)
<emphasis id="B962EAF0FF972576FDFEF8940783F898" box="[616,727,1793,1816]" italics="true" pageId="34" pageNumber="35">lamellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251578BFF972576FD48F894064DF898" box="[734,793,1793,1816]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
, 31: antenna, 32: labrum, epipharyngeal area and premandible, 3334:
<taxonomicName id="4C164D61FF972576FE62F8B606E9F8BA" box="[500,957,1827,1850]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="nimidens" status="sp. nov." subGenus="Mesorthocladius">
<emphasis id="B962EAF0FF972576FE62F8B607D5F8BA" box="[500,641,1827,1850]" italics="true" pageId="34" pageNumber="35">Orthocladius</emphasis>
(
<emphasis id="B962EAF0FF972576FD00F8B6061DF8BA" box="[662,841,1827,1850]" italics="true" pageId="34" pageNumber="35">Mesorthocladius</emphasis>
)
<emphasis id="B962EAF0FF972576FCCBF8B606E9F8BA" box="[861,957,1827,1850]" italics="true" pageId="34" pageNumber="35">nimidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251578BFF972576FC5FF8B6015CF8BA" box="[969,1032,1827,1850]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
, 31: antenna, 32: labrum, epipharyngeal area and premandible.
</paragraph>
</caption>
<subSubSection id="C30C6569FF962577FE9EFE8901CAFD46" pageId="35" pageNumber="36" type="discussion">
<paragraph id="8BA936E2FF962577FE9EFE8904D7FEB5" blockId="35.[264,1323,284,590]" box="[264,387,284,309]" pageId="35" pageNumber="36">
<emphasis id="B962EAF0FF962577FE9EFE8904D7FEB5" box="[264,387,284,309]" italics="true" pageId="35" pageNumber="36">Comments</emphasis>
</paragraph>
<paragraph id="8BA936E2FF962577FEAEFED107FFFDCE" blockId="35.[264,1323,284,590]" pageId="35" pageNumber="36">
The pupa of this species shows no synapomorphies with the other members of the subgenus and is practically inseparable from
<taxonomicName id="4C164D61FF962577FC9CFEF90122FE05" box="[778,1142,364,390]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="chuzesextus" subGenus="Orthocladius">
<emphasis id="B962EAF0FF962577FC9CFEF90673FE05" box="[778,807,364,389]" italics="true" pageId="35" pageNumber="36">O.</emphasis>
(
<emphasis id="B962EAF0FF962577FCAAFEF90683FE05" box="[828,983,364,389]" italics="true" pageId="35" pageNumber="36">Orthocladius</emphasis>
)
<emphasis id="B962EAF0FF962577FC7AFEF90122FE05" box="[1004,1142,364,389]" italics="true" pageId="35" pageNumber="36">chuzesextus</emphasis>
</taxonomicName>
. However, the male imago has the characteristic anal point of the other species except
<taxonomicName id="4C164D61FF962577FBF6FE01007FFE2D" box="[1120,1323,404,430]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="nimidens" subGenus="Mesorthocladius">
<emphasis id="B962EAF0FF962577FBF6FE010128FE2D" box="[1120,1148,404,429]" italics="true" pageId="35" pageNumber="36">O.</emphasis>
(
<emphasis id="B962EAF0FF962577FB18FE0101FBFE2D" box="[1166,1199,404,429]" italics="true" pageId="35" pageNumber="36">M.</emphasis>
)
<emphasis id="B962EAF0FF962577FB57FE01007FFE2D" box="[1217,1323,404,429]" italics="true" pageId="35" pageNumber="36">nimidens</emphasis>
</taxonomicName>
and the larva shares several apparent synapomorphies such as presence of labral lamellae, absence of seta interna, and bifid premandible with one or more of the other species of the genus. The phylogenetic placement of this and the next species, however, cannot be regarded as completely ascertained.
</paragraph>
<paragraph id="8BA936E2FF962577FE9EFD110778FD1D" blockId="35.[264,1182,644,710]" box="[264,556,644,669]" pageId="35" pageNumber="36">
<emphasis id="B962EAF0FF962577FE9EFD110778FD1D" box="[264,556,644,669]" italics="true" pageId="35" pageNumber="36">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="8BA936E2FF962577FEAEFD3901CAFD46" blockId="35.[264,1182,644,710]" box="[312,1182,684,710]" pageId="35" pageNumber="36">The species is known only from Ohio, inhabiting small streams and seeps.</paragraph>
</subSubSection>
</treatment>
</document>