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<document id="C57018F13FECF9F497C2084331F9BDD7" ID-DOI="10.24199/j.mmv.2008.65.3" ID-ISSN="1447-2554" ID-Zenodo-Dep="10665901" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708004926147" checkinUser="felipe" docAuthor="Dean, J. C., Forteath, G. N. R. &amp; Osborn, A. W." docDate="2008" docId="039D8005AC229D49FCA1FC45FD55F2A7" docLanguage="en" docName="MemMusVictoria.65.43-50.pdf" docOrigin="Memoirs of Museum Victoria 65" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-65-2008/pages-43-50/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Marmenuera tillyardi Dean &amp; Forteath &amp; Osborn 2008, sp.nov." docType="treatment" docVersion="2" lastPageNumber="47" masterDocId="FFA4F87DAC239D4DFF90FFE5FFF4F67C" masterDocTitle="Marmenuera, a new genus of leptophlebiid mayfly (Insecta: Ephemeroptera) endemic to Tasmania." masterLastPageNumber="50" masterPageNumber="43" pageNumber="44" updateTime="1708005457284" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="A2CE3346C9434E1F77DC511D8AFFD6E4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9534DFB9ACF4A5DBB4E7487DE42D8C6B">Marmenuera, a new genus of leptophlebiid mayfly (Insecta: Ephemeroptera) endemic to Tasmania.</mods:title>
</mods:titleInfo>
<mods:name id="33D39023E1641594FAD83F60E12F6ABD" type="personal">
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<mods:namePart id="C440B1F012953A83B343C02C25B64943">Dean, J. C.</mods:namePart>
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</mods:role>
<mods:namePart id="01A6C77577F56C1CF1FC6F8857B8095D">Forteath, G. N. R.</mods:namePart>
</mods:name>
<mods:name id="0B1A18D1AF4EC989C8674508EBA1E042" type="personal">
<mods:role id="1A0EE19A28E5C5372A28E18A4EFAF5EC">
<mods:roleTerm id="3DE86FBA4D68234F186544D1D9A1071A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="201628C5698E964B95966D2F8CDCFED6">Osborn, A. W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="41A2CE7D7408BC863DED2DEF79F70CA1">text</mods:typeOfResource>
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<mods:title id="773EA6308637AB24789847DA2F685B3A">Memoirs of Museum Victoria</mods:title>
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<mods:part id="7C6498761988C26C3497F4431DB813CF">
<mods:date id="AFB1390D4C624EEF95A233B1403E6A99">2008</mods:date>
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<mods:number id="A26027FF5D08B8610142EF2C9B3EAE5C">2008-12-31</mods:number>
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<mods:number id="26326A56F4B8AC91336E23182744431B">65</mods:number>
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<mods:start id="EFE55B9EE43D919DB83AC97EE0E4BA33">43</mods:start>
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<mods:url id="0515AD11B2BA1A4DE47A55B2EFCC4216">https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-65-2008/pages-43-50/</mods:url>
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<mods:classification id="01E83DC809141F90204376A431803BB3">journal article</mods:classification>
<mods:identifier id="4EE3CFCFB104786F0C42A94CDF2C21DA" type="DOI">10.24199/j.mmv.2008.65.3</mods:identifier>
<mods:identifier id="B848E1A7ECE7E7D5B7BCAA7875776201" type="ISSN">1447-2554</mods:identifier>
<mods:identifier id="50E2F89AC85E03FFF6FF601D7AC1DEDF" type="Zenodo-Dep">10665901</mods:identifier>
</mods:mods>
<treatment id="039D8005AC229D49FCA1FC45FD55F2A7" LSID="urn:lsid:plazi:treatment:039D8005AC229D49FCA1FC45FD55F2A7" httpUri="http://treatment.plazi.org/id/039D8005AC229D49FCA1FC45FD55F2A7" lastPageId="4" lastPageNumber="47" pageId="1" pageNumber="44">
<subSubSection id="C32E6298AC229D4CFCA1FC45FB96F5C9" box="[817,1122,928,950]" pageId="1" pageNumber="44" type="nomenclature">
<paragraph id="8B8B3113AC229D4CFCA1FC45FB96F5C9" blockId="1.[817,1122,928,950]" box="[817,1122,928,950]" pageId="1" pageNumber="44">
<heading id="D0C3867FAC229D4CFCA1FC45FB96F5C9" box="[817,1122,928,950]" fontSize="9" level="2" pageId="1" pageNumber="44" reason="5">
<taxonomicName id="4C344A90AC229D4CFCA1FC45FBE2F5CA" ID-CoL="3Y7Y7" authority="Dean &amp; Forteath &amp; Osborn, 2008" authorityName="Dean &amp; Forteath &amp; Osborn" authorityYear="2008" box="[817,1046,928,950]" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="tillyardi" status="sp. nov.">
<emphasis id="B940ED01AC229D4CFCA1FC45FBE2F5CA" bold="true" box="[817,1046,928,950]" italics="true" pageId="1" pageNumber="44">Marmenuera tillyardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273507AAC229D4CFB8DFC45FB96F5C9" box="[1053,1122,928,949]" pageId="1" pageNumber="44" rank="species">sp.nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E6298AC229D49FCA1FC29FD55F2A7" lastPageId="4" lastPageNumber="47" pageId="1" pageNumber="44" type="description">
<paragraph id="8B8B3113AC229D4CFCA1FC29FC47F59E" blockId="1.[817,947,972,994]" box="[817,947,972,994]" pageId="1" pageNumber="44">
<figureCitation id="130F2D96AC229D4CFCA1FC29FC47F59E" box="[817,947,972,994]" captionStart-0="Figures 14" captionStart-1="Figures 510" captionStart-2="Figures 1116" captionStartId-0="2.[140,208,950,969]" captionStartId-1="2.[140,208,1858,1877]" captionStartId-2="3.[148,216,1155,1174]" captionTargetBox-0="[149,1426,223,903]" captionTargetBox-1="[140,1415,1032,1832]" captionTargetBox-2="[147,1424,227,1127]" captionTargetId-0="figure-15@2.[141,1441,209,939]" captionTargetId-1="figure-36@2.[140,1438,1023,1841]" captionTargetId-2="figure-384@3.[147,1446,227,1138]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Figures 14. Marmenuera tillyardi. Male imago: 1, forewing; 2, hind wing. Female imago: 3, forewing; 4, hind wing." captionText-1="Figures 510. Marmenuera tillyardi. Male imago: 5, abdomen, dorsal; 6, abdomen, ventral; 7, penes, dorsal; 8, penes, ventral, slide preparation; 9, foretarsal claw. Female imago: 10, sternum nine." captionText-2="Figures 1116. Marmenuera tillyardi. Nymph: 11, abdomen, dorsal; 12, labrum; 13, forefemur; 14, foretarsus; 15, posterior margin of abdominal segment four; 16, gill, abdominal segment four." figureDoi-0="http://doi.org/10.5281/zenodo.10665903" figureDoi-1="http://doi.org/10.5281/zenodo.10665907" figureDoi-2="http://doi.org/10.5281/zenodo.10665909" httpUri-0="https://zenodo.org/record/10665903/files/figure.png" httpUri-1="https://zenodo.org/record/10665907/files/figure.png" httpUri-2="https://zenodo.org/record/10665909/files/figure.png" pageId="1" pageNumber="44">Figures 116</figureCitation>
</paragraph>
<paragraph id="8B8B3113AC229D4CFCC7FC1DFAF8F277" blockId="1.[855,1292,1015,1035]" box="[855,1292,1015,1035]" pageId="1" pageNumber="44">Genus W sp.AV1 Dean 1999, p. 86, figs 23941.</paragraph>
<paragraph id="8B8B3113AC229D4CFCA1FBC7FAE1F340" blockId="1.[817,1449,1057,1683]" pageId="1" pageNumber="44">
<emphasis id="B940ED01AC229D4CFCA1FBC7FC90F248" box="[817,868,1058,1076]" italics="true" pageId="1" pageNumber="44">Types</emphasis>
.
<materialsCitation id="3B5C3B4EAC229D4CFCE3FBC4FC73F2E2" collectingDate="2006-01-28" collectingDateMax="2006-01-30" collectingDateMin="2006-01-28" collectionCode="MV" collectorName="A. Osborn &amp; N. Forteath" location="Lake Pedder" municipality="Male" pageId="1" pageNumber="44" specimenCount="1" stateProvince="Tasmania" typeStatus="holotype">
<typeStatus id="548F8FB1AC229D4CFCE3FBC4FC3EF248" box="[883,970,1057,1076]" pageId="1" pageNumber="44" type="holotype">Holotype</typeStatus>
:
<collectingMunicipality id="6BEFAB69AC229D4CFC47FBC4FBF2F248" box="[983,1030,1057,1076]" pageId="1" pageNumber="44">Male</collectingMunicipality>
imago (reared from nymph),
<collectingRegion id="49F0FFF1AC229D4CFAB3FBC4FA89F248" box="[1315,1405,1057,1076]" country="Australia" name="Tasmania" pageId="1" pageNumber="44">Tasmania</collectingRegion>
, un-named creek flowing into
<location id="8EEB67C8AC229D4CFBA3FBDEFB5EF232" LSID="urn:lsid:plazi:treatment:039D8005AC229D49FCA1FC45FD55F2A7:8EEB67C8AC229D4CFBA3FBDEFB5EF232" box="[1075,1194,1083,1102]" municipality="Male" name="Lake Pedder" pageId="1" pageNumber="44" stateProvince="Tasmania">Lake Pedder</location>
(Tasmap 1:25 000 series, Serpentine 4026, Grid reference 55G 4184 52606), collected
<date id="FF8A17D3AC229D4CFAFBFBB3FC95F2FF" pageId="1" pageNumber="44" value="2006-01-28">
<collectingDate id="EFCEEE3BAC229D4CFAFBFBB3FC95F2FF" pageId="1" pageNumber="44" value="2006-01-28">28 Jan 2006</collectingDate>
</date>
, emerged
<date id="FF8A17D3AC229D4CFC5CFB95FBBEF2FF" box="[972,1098,1136,1156]" pageId="1" pageNumber="44" value="2006-01-30">
<collectingDate id="EFCEEE3BAC229D4CFC5CFB95FBBEF2FF" box="[972,1098,1136,1156]" pageId="1" pageNumber="44" value="2006-01-30">30 Jan 2006</collectingDate>
</date>
,
<collectorName id="26C154C5AC229D4CFBCBFB95FB35F2FF" box="[1115,1217,1136,1155]" pageId="1" pageNumber="44">A. Osborn</collectorName>
and
<collectorName id="26C154C5AC229D4CFB6CFB94FA9FF2FF" box="[1276,1387,1136,1156]" pageId="1" pageNumber="44">N. Forteath</collectorName>
(
<collectionCode id="ED25A9D6AC229D4CFAEEFB95FA50F2FF" box="[1406,1444,1136,1155]" country="0" name="University of Montana Museum" pageId="1" pageNumber="44">MV</collectionCode>
, T-20000)
</materialsCitation>
.
<materialsCitation id="3B5C3B4EAC229D4CFC1FFB6EFAE4F340" collectingDate="2006-01-28" collectingDateMax="2006-01-30" collectingDateMin="2006-01-28" collectionCode="MV, ANIC" collectorName="Male &amp; A. Osborn &amp; N. Forteath" location="Lake Pedder" municipality="Male" pageId="1" pageNumber="44" specimenCount="2" stateProvince="Tasmania" typeStatus="paratype">
<typeStatus id="548F8FB1AC229D4CFC1FFB6EFC1FF2E2" box="[911,1003,1163,1182]" pageId="1" pageNumber="44" type="paratype">Paratypes</typeStatus>
:
<collectorName id="26C154C5AC229D4CFC63FB6EFBD5F2E2" box="[1011,1057,1163,1182]" pageId="1" pageNumber="44">Male</collectorName>
imago and female imago (both reared from nymph), same location as holotype, collected
<date id="FF8A17D3AC229D4CFB59FB40FAC3F2C4" box="[1225,1335,1189,1208]" pageId="1" pageNumber="44" value="2006-01-28">
<collectingDate id="EFCEEE3BAC229D4CFB59FB40FAC3F2C4" box="[1225,1335,1189,1208]" pageId="1" pageNumber="44" value="2006-01-28">28 Jan 2006</collectingDate>
</date>
, emerged
<collectingDate id="EFCEEE3BAC229D4CFA02FB40FBD7F2AE" isEnumeration="true" pageId="1" pageNumber="44">
<date id="FF8A17D3AC229D4CFA02FB40FC7EF2AE" pageId="1" pageNumber="44" value="2006-03-19">19 Mar 2006</date>
and
<date id="FF8A17D3AC229D4CFC24FB25FBD7F2AE" box="[948,1059,1215,1235]" pageId="1" pageNumber="44" value="2006-02-23">23 Feb 2006</date>
</collectingDate>
respectively,
<collectorName id="26C154C5AC229D4CFB0DFB5AFB02F2AE" box="[1181,1270,1215,1234]" pageId="1" pageNumber="44">A.Osborn</collectorName>
&amp;
<collectorName id="26C154C5AC229D4CFA81FB5AFA87F2AE" box="[1297,1395,1215,1234]" pageId="1" pageNumber="44">N.Forteath</collectorName>
(
<collectionCode id="ED25A9D6AC229D4CFAEEFB25FA50F2AF" box="[1406,1444,1216,1235]" country="0" name="University of Montana Museum" pageId="1" pageNumber="44">MV</collectionCode>
, T-20001 and T-20002 respectively); male imago and female imago (both reared from nymph), same location as
<typeStatus id="548F8FB1AC229D4CFB55FB11FAE3F37B" box="[1221,1303,1268,1287]" pageId="1" pageNumber="44" type="holotype">holotype</typeStatus>
, collected
<collectingDate id="EFCEEE3BAC229D4CFAE8FB11FC08F35E" isEnumeration="true" pageId="1" pageNumber="44">
<date id="FF8A17D3AC229D4CFAE8FB11FCABF35E" pageId="1" pageNumber="44" value="2006-01-05">5 Jan 2006</date>
and
<date id="FF8A17D3AC229D4CFC1DFAEAFC08F35E" box="[909,1020,1295,1314]" pageId="1" pageNumber="44" value="2006-01-28">28 Jan 2006</date>
</collectingDate>
respectively, emerged
<collectingDate id="EFCEEE3BAC229D4CFB5EFAEAFCABF340" isEnumeration="true" pageId="1" pageNumber="44">
<date id="FF8A17D3AC229D4CFB5EFAEAFAC8F35E" box="[1230,1340,1295,1314]" pageId="1" pageNumber="44" value="2006-01-14">14 Jan 2006</date>
and
<date id="FF8A17D3AC229D4CFAFAFAEAFCABF340" pageId="1" pageNumber="44" value="2006-02-13">13 Feb 2006</date>
</collectingDate>
respectively,
<collectorName id="26C154C5AC229D4CFC4BFACCFBC0F340" box="[987,1076,1321,1340]" pageId="1" pageNumber="44">A.Osborn</collectorName>
and
<collectorName id="26C154C5AC229D4CFBF0FACCFB36F340" box="[1120,1218,1321,1340]" pageId="1" pageNumber="44">N.Forteath</collectorName>
(
<collectionCode id="ED25A9D6AC229D4CFB5EFACCFAFEF340" box="[1230,1290,1321,1340]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="1" pageNumber="44">ANIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8B3113AC229D4CFCC7FAA6FC27F0EF" blockId="1.[817,1449,1057,1683]" pageId="1" pageNumber="44">
<emphasis id="B940ED01AC229D4CFCC7FAA6FBE7F32A" box="[855,1043,1347,1366]" italics="true" pageId="1" pageNumber="44">Material examined.</emphasis>
<materialsCitation id="3B5C3B4EAC229D4CFB8DFAA6FC3AF0EF" collectingDate="1994-10-27" collectingDateMax="2006-01-28" collectingDateMin="1994-10-27" collectionCode="MRHI" collectorName="A. Osborn &amp; N. Forteath &amp; Heamskirk Rd &amp; Serpentine Arm" county="Lake Pedder" location="Lake Pedder" municipality="Piney Creek" pageId="1" pageNumber="44" specimenCount="1" stateProvince="Tasmania">
<collectingRegion id="49F0FFF1AC229D4CFB8DFAA6FB8CF32A" box="[1053,1144,1347,1366]" country="Australia" name="Tasmania" pageId="1" pageNumber="44">Tasmania</collectingRegion>
. 6 MI, 12 FI (all reared from nymphs), un-named creek flowing into
<collectingCounty id="62EA499FAC229D4CFB01FABBFAF7F30D" box="[1169,1283,1374,1393]" pageId="1" pageNumber="44">Lake Pedder</collectingCounty>
(
<typeStatus id="548F8FB1AC229D4CFA9FFABBFAC3F30D" box="[1295,1335,1374,1393]" pageId="1" pageNumber="44">type</typeStatus>
locality),
<date id="FF8A17D3AC229D4CFA01FABBFC71F3F7" pageId="1" pageNumber="44" value="2006-01-28">
<collectingDate id="EFCEEE3BAC229D4CFA01FABBFC71F3F7" pageId="1" pageNumber="44" value="2006-01-28">28 Jan 2006</collectingDate>
</date>
, emerged various dates,
<collectorName id="26C154C5AC229D4CFBF5FA9DFB30F3F7" box="[1125,1220,1400,1419]" pageId="1" pageNumber="44">A. Osborn</collectorName>
and
<collectorName id="26C154C5AC229D4CFB60FA9CFAAFF3F7" box="[1264,1371,1400,1420]" pageId="1" pageNumber="44">N. Forteath</collectorName>
; 3 MI, 1 FI (all reared from nymphs), same location,
<date id="FF8A17D3AC229D4CFB52FA77FAC4F3DA" box="[1218,1328,1426,1446]" pageId="1" pageNumber="44" value="1998-01-25">
<collectingDate id="EFCEEE3BAC229D4CFB52FA77FAC4F3DA" box="[1218,1328,1426,1446]" pageId="1" pageNumber="44" value="1998-01-25">25 Jan 1998</collectingDate>
</date>
,
<collectorName id="26C154C5AC229D4CFAA9FA76FA50F3DA" box="[1337,1444,1427,1446]" pageId="1" pageNumber="44">N. Forteath</collectorName>
; 1 FI (reared from nymph), same location,
<date id="FF8A17D3AC229D4CFB3DFA48FAEBF3BC" box="[1197,1311,1453,1472]" pageId="1" pageNumber="44" value="1998-02-23">
<collectingDate id="EFCEEE3BAC229D4CFB3DFA48FAEBF3BC" box="[1197,1311,1453,1472]" pageId="1" pageNumber="44" value="1998-02-23">23 Feb 1998</collectingDate>
</date>
,
<collectorName id="26C154C5AC229D4CFAB9FA48FA67F3BC" box="[1321,1427,1453,1472]" pageId="1" pageNumber="44">N. Forteath</collectorName>
; 1 MI, 1 FI (both reared from nymphs), same location,
<date id="FF8A17D3AC229D4CFA89FA2DFA73F3A7" box="[1305,1415,1480,1499]" pageId="1" pageNumber="44" value="1998-03-04">
<collectingDate id="EFCEEE3BAC229D4CFA89FA2DFA73F3A7" box="[1305,1415,1480,1499]" pageId="1" pageNumber="44" value="1998-03-04">4 Mar 1998</collectingDate>
</date>
,
<collectorName id="26C154C5AC229D4CFA02FA2DFC74F389" pageId="1" pageNumber="44">N. Forteath</collectorName>
; 2 MI, 2 FI (all reared from nymphs), same location,
<date id="FF8A17D3AC229D4CFAE7FA07FC95F073" pageId="1" pageNumber="44" value="2006-01-05">
<collectingDate id="EFCEEE3BAC229D4CFAE7FA07FC95F073" pageId="1" pageNumber="44" value="2006-01-05">5 Jan 2006</collectingDate>
</date>
,
<collectorName id="26C154C5AC229D4CFCFFFA19FC25F073" box="[879,977,1532,1551]" pageId="1" pageNumber="44">A. Osborn</collectorName>
and
<collectorName id="26C154C5AC229D4CFB96FA18FB80F073" box="[1030,1140,1532,1552]" pageId="1" pageNumber="44">N. Forteath</collectorName>
; 1 MS, 1 FS (both reared from nymphs), 4 N, same location,
<date id="FF8A17D3AC229D4CFBDCF9F2FB31F056" box="[1100,1221,1559,1578]" pageId="1" pageNumber="44" value="1997-12-27">
<collectingDate id="EFCEEE3BAC229D4CFBDCF9F2FB31F056" box="[1100,1221,1559,1578]" pageId="1" pageNumber="44" value="1997-12-27">27 Dec 1997</collectingDate>
</date>
,
<collectorName id="26C154C5AC229D4CFB41F9F2FAC2F056" box="[1233,1334,1559,1578]" pageId="1" pageNumber="44">N.Forteath</collectorName>
; 2 N,
<collectingMunicipality id="6BEFAB69AC229D4CFAE4F9F2FC9EF038" pageId="1" pageNumber="44">Piney Creek</collectingMunicipality>
,
<collectorName id="26C154C5AC229D4CFCE4F9D4FC08F038" box="[884,1020,1585,1604]" pageId="1" pageNumber="44">Heamskirk Rd</collectorName>
,
<date id="FF8A17D3AC229D4CFB97F9D4FB88F038" box="[1031,1148,1585,1604]" pageId="1" pageNumber="44" value="1994-10-27">
<collectingDate id="EFCEEE3BAC229D4CFB97F9D4FB88F038" box="[1031,1148,1585,1604]" pageId="1" pageNumber="44" value="1994-10-27">27 Oct 1994</collectingDate>
</date>
,
<collectionCode id="ED25A9D6AC229D4CFB16F9D4FB31F038" box="[1158,1221,1585,1604]" pageId="1" pageNumber="44">MRHI</collectionCode>
; 1 FI, 1 FS (both reared from nymphs),
<location id="8EEB67C8AC229D4CFC2BF9AEFBC4F022" LSID="urn:lsid:plazi:treatment:039D8005AC229D49FCA1FC45FD55F2A7:8EEB67C8AC229D4CFC2BF9AEFBC4F022" box="[955,1072,1611,1630]" county="Lake Pedder" municipality="Piney Creek" name="Lake Pedder" pageId="1" pageNumber="44" stateProvince="Tasmania">Lake Pedder</location>
,
<location id="8EEB67C8AC229D4CFBA9F9A9FB5FF023" LSID="urn:lsid:plazi:treatment:039D8005AC229D49FCA1FC45FD55F2A7:8EEB67C8AC229D4CFBA9F9A9FB5FF023" box="[1081,1195,1612,1631]" county="Lake Pedder" municipality="Piney Creek" name="Trappes Bay" pageId="1" pageNumber="44" stateProvince="Tasmania">Trappes Bay</location>
,
<date id="FF8A17D3AC229D4CFB24F9A9FAEBF023" box="[1204,1311,1612,1631]" pageId="1" pageNumber="44" value="1998-03-04">
<collectingDate id="EFCEEE3BAC229D4CFB24F9A9FAEBF023" box="[1204,1311,1612,1631]" pageId="1" pageNumber="44" value="1998-03-04">4 Mar 1998</collectingDate>
</date>
,
<collectorName id="26C154C5AC229D4CFAB9F9A9FA67F022" box="[1321,1427,1611,1631]" pageId="1" pageNumber="44">N. Forteath</collectorName>
; 1 MI, 2FI (all reared from nymphs),
<location id="8EEB67C8AC229D4CFBFAF983FB2AF005" LSID="urn:lsid:plazi:treatment:039D8005AC229D49FCA1FC45FD55F2A7:8EEB67C8AC229D4CFBFAF983FB2AF005" box="[1130,1246,1638,1657]" county="Lake Pedder" municipality="Piney Creek" name="Lake Pedder" pageId="1" pageNumber="44" stateProvince="Tasmania">Lake Pedder</location>
,
<collectorName id="26C154C5AC229D4CFB77F983FA8FF005" box="[1255,1403,1638,1657]" pageId="1" pageNumber="44">Serpentine Arm</collectorName>
,
<date id="FF8A17D3AC229D4CFA14F983FC95F0EF" pageId="1" pageNumber="44" value="2005-04">
<collectingDate id="EFCEEE3BAC229D4CFA14F983FC95F0EF" pageId="1" pageNumber="44" value="2005-04">Apr 2005</collectingDate>
</date>
,
<collectorName id="26C154C5AC229D4CFCFAF965FC3AF0EF" box="[874,974,1664,1683]" pageId="1" pageNumber="44">N.Forteath</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8B3113AC229D4EFCA1F94EFB5EF3F2" blockId="1.[817,1449,1707,1903]" lastBlockId="3.[817,1448,1310,1903]" lastPageId="3" lastPageNumber="46" pageId="1" pageNumber="44">
<emphasis id="B940ED01AC229D4CFCA1F94EFBBEF0BC" box="[817,1098,1707,1728]" italics="true" pageId="1" pageNumber="44">Description. Male imago</emphasis>
: Length: body
<quantity id="4CCC9CF6AC229D4CFA9BF94EFA55F0BC" box="[1291,1441,1707,1728]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.2" metricValueMin="8.8" pageId="1" pageNumber="44" unit="mm" value="9.5" valueMax="10.2" valueMin="8.8">8.810.2 mm</quantity>
., forewing
<quantity id="4CCC9CF6AC229D4CFC02F92DFBEDF0A1" box="[914,1049,1736,1757]" metricMagnitude="-3" metricUnit="m" metricValue="9.45" metricValueMax="10.1" metricValueMin="8.8" pageId="1" pageNumber="44" unit="mm" value="9.45" valueMax="10.1" valueMin="8.8">8.810.1 mm</quantity>
., caudal filaments
<quantity id="4CCC9CF6AC229D4CFB44F92DFA9CF0A1" box="[1236,1384,1736,1757]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" metricValueMax="1.42" metricValueMin="1.24" pageId="1" pageNumber="44" unit="mm" value="13.3" valueMax="14.2" valueMin="12.4">12.414.2 mm</quantity>
. Eyes: upper lobes pale pink/brown, separated mesally by a distance approximately 1/8 of the diameter of the upper lobe in dorsal view; lower lobes grey. Thorax: medium-dark brown; pronotum with narrow raised carina along median line, dark brown markings medially on posterior half of segment, dark brown along lateral margins and curved dark brown band parallel to lateral margin approximately midway between margin and medial carina. Legs: predominantly medium brown, tending to paler in apical half of tibiae and tarsi; all femora with two broad dark brown bands, one just beyond midlength and the other sub-apical; tarsal claws similar, each claw with terminal sclerotised hook and opposing ventral process. Forewing (fig. 1): membrane hyaline, stigmatic region opaque, white; costal crossveins in basal half of wing heavily suffused with dark brown, subcostal and
<collectionCode id="ED25A9D6AC209D4EFEE7F9C8FE73F03E" box="[375,391,1581,1602]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="46">R</collectionCode>
<subScript id="17B03356AC209D4EFE18F9D9FE7BF034" attach="right" box="[392,399,1596,1608]" fontSize="5" pageId="3" pageNumber="46">1</subScript>
<collectionCode id="ED25A9D6AC209D4EFE0CF9C8FE59F03E" box="[412,429,1581,1602]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="46">R</collectionCode>
<subScript id="17B03356AC209D4EFE3DF9D9FE40F034" attach="left" box="[429,436,1596,1608]" fontSize="5" pageId="3" pageNumber="46">2</subScript>
crossveins in basal half of wing less strongly suffused with dark brown, crossveins near MA fork not suffused with brown. Abdomen: terga generally reddish-brown with dark brown markings, tergum 6 with four conspicuous dark brown longitudinal bands extending over most of the segment (fig. 5), segment 7 with similar bands but restricted to anterior half of the segment, segment 8 with dark markings only adjacent to the anterior margin; sterna reddish brown, inconspicuous paler markings on segments 2-5 (fig. 6), darker brown markings poorly defined. Genitalia (figs 7, 8): penes lobes fused basally, widely separated apically; each lobe relatively broad and approximately parallel-sided, apically truncate, outer margin somewhat angular just before apex; three moderate sized ventral spines close to inner margin posterior to where the lobes separate.
</paragraph>
<caption id="DF4B619BAC219D4FFF1CFC53FB40F5B5" ID-DOI="http://doi.org/10.5281/zenodo.10665903" ID-Zenodo-Dep="10665903" box="[140,1204,950,970]" httpUri="https://zenodo.org/record/10665903/files/figure.png" pageId="2" pageNumber="45" startId="2.[140,208,950,969]" targetBox="[149,1426,223,903]" targetPageId="2" targetType="figure">
<paragraph id="8B8B3113AC219D4FFF1CFC53FB40F5B5" blockId="2.[140,1204,950,970]" box="[140,1204,950,970]" pageId="2" pageNumber="45">
Figures 14.
<taxonomicName id="4C344A90AC219D4FFE93FC52FE3FF5B5" authorityName="Dean &amp; Forteath &amp; Osborn" authorityYear="2008" box="[259,459,950,969]" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="B940ED01AC219D4FFE93FC52FE3FF5B5" box="[259,459,950,969]" italics="true" pageId="2" pageNumber="45">Marmenuera tillyardi</emphasis>
</taxonomicName>
. Male imago: 1, forewing; 2, hind wing. Female imago: 3, forewing; 4, hind wing.
</paragraph>
</caption>
<caption id="DF4B619BAC219D4FFF1CF8A7FDAFF113" ID-DOI="http://doi.org/10.5281/zenodo.10665907" ID-Zenodo-Dep="10665907" httpUri="https://zenodo.org/record/10665907/files/figure.png" pageId="2" pageNumber="45" startId="2.[140,208,1858,1877]" targetBox="[140,1415,1032,1832]" targetPageId="2" targetType="figure">
<paragraph id="8B8B3113AC219D4FFF1CF8A7FDAFF113" blockId="2.[140,1441,1858,1904]" pageId="2" pageNumber="45">
Figures 510.
<taxonomicName id="4C344A90AC219D4FFE9DF8A6FE20F129" authorityName="Dean &amp; Forteath &amp; Osborn" authorityYear="2008" box="[269,468,1858,1877]" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="B940ED01AC219D4FFE9DF8A6FE20F129" box="[269,468,1858,1877]" italics="true" pageId="2" pageNumber="45">Marmenuera tillyardi</emphasis>
</taxonomicName>
. Male imago: 5, abdomen, dorsal; 6, abdomen, ventral; 7, penes, dorsal; 8, penes, ventral, slide preparation; 9, foretarsal claw. Female imago: 10, sternum nine.
</paragraph>
</caption>
<caption id="DF4B619BAC209D4EFF04FB66FDB7F2CC" ID-DOI="http://doi.org/10.5281/zenodo.10665909" ID-Zenodo-Dep="10665909" httpUri="https://zenodo.org/record/10665909/files/figure.png" pageId="3" pageNumber="46" startId="3.[148,216,1155,1174]" targetBox="[147,1424,227,1127]" targetPageId="3" targetType="figure">
<paragraph id="8B8B3113AC209D4EFF04FB66FDB7F2CC" blockId="3.[147,1448,1155,1201]" pageId="3" pageNumber="46">
Figures 1116.
<taxonomicName id="4C344A90AC209D4EFE8CFB61FE17F2EA" authorityName="Dean &amp; Forteath &amp; Osborn" authorityYear="2008" box="[284,483,1155,1174]" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="B940ED01AC209D4EFE8CFB61FE17F2EA" box="[284,483,1155,1174]" italics="true" pageId="3" pageNumber="46">Marmenuera tillyardi</emphasis>
</taxonomicName>
. Nymph: 11, abdomen, dorsal; 12, labrum; 13, forefemur; 14, foretarsus; 15, posterior margin of abdominal segment four; 16, gill, abdominal segment four.
</paragraph>
</caption>
<paragraph id="8B8B3113AC209D49FCC7FA73FDD6F4B1" blockId="3.[817,1448,1310,1903]" lastBlockId="4.[140,771,229,717]" lastPageId="4" lastPageNumber="47" pageId="3" pageNumber="46">
<emphasis id="B940ED01AC209D4EFCC7FA73FC04F3D0" box="[855,1008,1430,1452]" italics="true" pageId="3" pageNumber="46">Female imago</emphasis>
. Length: body
<quantity id="4CCC9CF6AC209D4EFB32FA73FACFF3D0" box="[1186,1339,1430,1452]" metricMagnitude="-2" metricUnit="m" metricValue="1.1149999999999998" metricValueMax="1.22" metricValueMin="1.01" pageId="3" pageNumber="46" unit="mm" value="11.149999999999999" valueMax="12.2" valueMin="10.1">10.112.2 mm</quantity>
; forewing 10.0
<quantity id="4CCC9CF6AC209D4EFCF9FA50FC37F3B6" box="[873,963,1461,1482]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="3" pageNumber="46" unit="mm" value="11.8">11.8 mm</quantity>
; caudal filaments
<quantity id="4CCC9CF6AC209D4EFBE6FA51FAFCF3B6" box="[1142,1288,1460,1482]" metricMagnitude="-2" metricUnit="m" metricValue="1.4899999999999998" metricValueMax="1.64" metricValueMin="1.34" pageId="3" pageNumber="46" unit="mm" value="14.899999999999999" valueMax="16.4" valueMin="13.4">13.416.4 mm</quantity>
. General colour similar to male imago. Forewing: membrane slightly opaque; costal, subcostal and
<collectionCode id="ED25A9D6AC209D4EFBB2FA14FBC6F07A" box="[1058,1074,1521,1542]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="46">R</collectionCode>
1
<collectionCode id="ED25A9D6AC209D4EFBD6FA14FBA3F07A" box="[1094,1111,1521,1542]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="46">R</collectionCode>
2 crossveins all surrounded by suffusion of dark brown, more strongly developed than in male. Abdomen: medium reddish-brown, terga with darker brown markings, sterna medium reddish-brown, no strong markings; sternum nine deeply cleft (fig. 10).
<emphasis id="B940ED01AC209D4EFB65F98CFAAFF002" box="[1269,1371,1641,1662]" italics="true" pageId="3" pageNumber="46">Subimago</emphasis>
. Wings pale fawn/yellow, brown blotches and/or suffusions similar in distribution and intensity to corresponding imago.
<emphasis id="B940ED01AC209D4EFACCF940FC81F0A4" italics="true" pageId="3" pageNumber="46">Mature nymph</emphasis>
. Body length ranging from about
<quantity id="4CCC9CF6AC209D4EFA92F926FA54F0A4" box="[1282,1440,1731,1752]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="3" pageNumber="46" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10 to 13 mm</quantity>
.; antennae about half length of body; cerci about 1½ times length of body, terminal filament a little longer. General colour yellow with dark brown markings; abdomen with conspicuous pattern of dark brown and yellow, abdominal segments 6 and 9 darker than remaining segments (fig. 11); all legs with dark banding. Mouthparts: labrum (fig. 12) a little broader than clypeus; maximum width 1.71.9 times length along median line; maximum width at about 3/4 labrum length, basal to this the lateral margins relatively straight and diverging; anterior margin shallowly concave, with five well developed medial denticles; single row of setae close to frontal edge of labrum, sub-apical setal fringe also consisting of a single row set back from anterior margin at about 0.9 labrum length. Legs relatively slender; forefemora length 3½4 times width (
<figureCitation id="130F2D96AC279D49FDFFFE2BFD4DF798" box="[623,697,462,484]" captionStart="Figures 1116" captionStartId="3.[148,216,1155,1174]" captionTargetBox="[147,1424,227,1127]" captionTargetId="figure-384@3.[147,1446,227,1138]" captionTargetPageId="3" captionText="Figures 1116. Marmenuera tillyardi. Nymph: 11, abdomen, dorsal; 12, labrum; 13, forefemur; 14, foretarsus; 15, posterior margin of abdominal segment four; 16, gill, abdominal segment four." figureDoi="http://doi.org/10.5281/zenodo.10665909" httpUri="https://zenodo.org/record/10665909/files/figure.png" pageId="4" pageNumber="47">Fig. 13</figureCitation>
), outer margin with moderate number of short, spine-like setae and also longer, hair-like setae; foretarsus (fig. 14) with 3040 ventral spines; tarsal claws smooth, without ventral teeth. Postero-lateral spines on abdominal segments 7 (small) and 8-9 (relatively large); posterior margins of abdominal terga with series of large, conspicuous spines interspersed with the occasional shorter spine (fig. 15). Gills lanceolate, lateral tracheae moderately developed (fig. 16).
</paragraph>
<paragraph id="8B8B3113AC279D49FF1CFD01FF21F548" blockId="4.[140,770,740,820]" pageId="4" pageNumber="47">
<emphasis id="B940ED01AC279D49FF1CFD01FF0EF485" box="[140,250,740,761]" italics="true" pageId="4" pageNumber="47">Etymology</emphasis>
: The species is named for
<collectionCode id="ED25A9D6AC279D49FD8FFD01FDC5F486" box="[543,561,740,762]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="47">R</collectionCode>
.J.Tillyard, who first recognised that this was probably an undescribed species (see below).
</paragraph>
<paragraph id="8B8B3113AC279D49FF1CFCAFFE44F273" blockId="4.[140,772,842,1243]" pageId="4" pageNumber="47">
<emphasis id="B940ED01AC279D49FF1CFCAFFF18F523" box="[140,236,842,863]" italics="true" pageId="4" pageNumber="47">Remarks.</emphasis>
When describing
<taxonomicName id="4C344A90AC279D49FE3CFCAFFCF6F51C" authority=", Tillyard (1936)" authorityName=", Tillyard" authorityYear="1936" box="[428,770,842,864]" class="Insecta" family="Leptophlebiidae" genus="Atalophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="ida">
<emphasis id="B940ED01AC279D49FE3CFCAFFDADF523" box="[428,601,842,863]" italics="true" pageId="4" pageNumber="47">Atalophlebia ida</emphasis>
,
<bibRefCitation id="EFA54CE2AC279D49FDF8FCAFFCF6F51C" author="Tillyard, R. J." box="[616,770,842,864]" pageId="4" pageNumber="47" pagination="23 - 59" refId="ref4883" refString="Tillyard, R. J. 1936. The trout-food insects of Tasmania. Part II: A monograph of the mayflies of Tasmania. Papers and Proceedings of the Royal Society of Tasmania. 1935: 23 - 59." type="journal article" year="1936">Tillyard (1936)</bibRefCitation>
</taxonomicName>
referred to specimens from Cradle Mountain which were smaller and less spotted than the
<typeStatus id="548F8FB1AC279D49FE65FC60FDD5F5E6" box="[501,545,901,922]" pageId="4" pageNumber="47">type</typeStatus>
series. It now seems likely that these belong to the species herein described as
<taxonomicName id="4C344A90AC279D49FD77FC47FF2AF5A8" authorityName="Dean &amp; Forteath &amp; Osborn" authorityYear="2008" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="B940ED01AC279D49FD77FC47FF2AF5A8" italics="true" pageId="4" pageNumber="47">M. tillyardi</emphasis>
</taxonomicName>
, and Tillyards decision not to describe the male imago of
<taxonomicName id="4C344A90AC279D49FF3BFC39FF19F58D" authorityName=", Tillyard" authorityYear="1936" box="[171,237,988,1009]" class="Insecta" family="Leptophlebiidae" genus="Atalophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="ida">
<emphasis id="B940ED01AC279D49FF3BFC39FF19F58D" box="[171,237,988,1009]" italics="true" pageId="4" pageNumber="47">A. ida</emphasis>
</taxonomicName>
based on the Cradle Mountain material has been proven taxonomically astute.
</paragraph>
<paragraph id="8B8B3113AC279D49FF22FBF3FD55F2A7" blockId="4.[140,772,842,1243]" pageId="4" pageNumber="47">
<bibRefCitation id="EFA54CE2AC279D49FF22FBF3FEB7F250" author="Scholes, D." box="[178,323,1046,1068]" pageId="4" pageNumber="47" refId="ref4862" refString="Scholes, D. 1961. Fly-fisher in Tasmania. Melbourne University Press: Carlton. Pp xiv + 208." type="book" year="1961">Scholes (1961)</bibRefCitation>
recommended that the imago and subimago of
<taxonomicName id="4C344A90AC279D49FF35FBD1FF15F234" authorityName=", Tillyard" authorityYear="1936" box="[165,225,1075,1097]" class="Insecta" family="Leptophlebiidae" genus="Atalophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="ida">
<emphasis id="B940ED01AC279D49FF35FBD1FF15F234" box="[165,225,1075,1097]" italics="true" pageId="4" pageNumber="47">A. ida</emphasis>
</taxonomicName>
should be referred to as the “Large Speckled Spinner” and “Large Speckled Dun” respectively, and we suggest that “Lesser Speckled Spinner” and “Lesser Speckled Dun” are appropriate common names for the imago and subimago of
<taxonomicName id="4C344A90AC279D49FF1CFB4CFE9AF2C1" authorityName="Dean &amp; Forteath &amp; Osborn" authorityYear="2008" box="[140,366,1192,1214]" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="B940ED01AC279D49FF1CFB4CFE9AF2C1" box="[140,366,1192,1214]" italics="true" pageId="4" pageNumber="47">Marmenuera tillyardi</emphasis>
</taxonomicName>
. This reflects both the smaller size of the new species and the reduced wing pigmentation.
</paragraph>
</subSubSection>
</treatment>
</document>