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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.307.4673" ID-GBIF-Dataset="1c534aa5-955a-49b7-9ce1-835ca6720c3d" ID-PMC="PMC3689063" ID-Pensoft-Pub="1313-2970-307-45" ID-PubMed="23794923" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-307-45" ModsDocOrigin="ZooKeys 307" ModsDocTitle="A taxonomic guide to the brittle-stars (Echinodermata, Ophiuroidea) from the State of Paraíba continental shelf, Northeastern Brazil" checkinTime="1451247231301" checkinUser="pensoft" docAuthor="Gondim, Anne I., Alonso, Carmen, Dias, Thelma L. P., Manso, Cynthia L. C. &amp; Christoffersen, Martin L." docDate="2013" docId="790F0102B26C399EFCDD9B45F62AB4E4" docLanguage="en" docName="ZooKeys 307: 45-96" docOrigin="ZooKeys 307" docSource="http://dx.doi.org/10.3897/zookeys.307.4673" docTitle="Ophiophragmus brachyactis H. L. Clark 1915" docType="treatment" docVersion="4" lastPageNumber="58" masterDocId="FF83FFA5FF98BD52FFD3FF85FF8FA509" masterDocTitle="A taxonomic guide to the brittle-stars (Echinodermata, Ophiuroidea) from the State of Paraiba continental shelf, Northeastern Brazil" masterLastPageNumber="96" masterPageNumber="45" pageNumber="58" updateTime="1668155933863" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A taxonomic guide to the brittle-stars (Echinodermata, Ophiuroidea) from the State of Paraiba continental shelf, Northeastern Brazil</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gondim, Anne I.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Alonso, Carmen</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dias, Thelma L. P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Manso, Cynthia L. C.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Christoffersen, Martin L.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>307</mods:number>
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<mods:start>45</mods:start>
<mods:end>96</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.307.4673</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.307.4673</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-307-45</mods:identifier>
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<treatment ID-GBIF-Taxon="152044877" LSID="urn:lsid:plazi:treatment:790F0102B26C399EFCDD9B45F62AB4E4" httpUri="http://treatment.plazi.org/id/790F0102B26C399EFCDD9B45F62AB4E4" lastPageNumber="58" pageId="13" pageNumber="58">
<subSubSection pageId="13" pageNumber="58" type="nomenclature">
<paragraph pageId="13" pageNumber="58">
<taxonomicName LSID="http://species-id.net/wiki/Ophiophragmus_brachyactis" authority="H. L. Clark, 1915" authorityName="H. L. Clark" authorityYear="1915" class="Ophiuroidea" family="Amphiuridae" genus="Ophiophragmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiophragmus brachyactis" order="Amphilepidida" pageId="13" pageNumber="58" phylum="Echinodermata" rank="species" species="brachyactis">
<pageBreakToken pageId="13" pageNumber="58" start="start">Ophiophragmus</pageBreakToken>
brachyactis H.L. Clark, 1915
</taxonomicName>
Figure 7
<normalizedToken originalValue="ae">a-e</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="58" type="description">
<paragraph pageId="13" pageNumber="58">Description.</paragraph>
<paragraph pageId="13" pageNumber="58">Disk circular (dd = 4.45 mm). Covered by imbricating scales of irregular shape (Fig. 7a). Margin of interradial field with 12-13 small, enlarged, blunt spines that decrease in size in the direction of the radial shields (Fig. 7a). Radial shields slightly longer than wide, united, except at proximal end, where a small triangular scale occurs between the pair of radial shields. Ventral interradius covered by scales similar to dorsal scales (Fig. 7b). Oral shields arrow-shaped (Fig. 7c). Adoral shields enlarged laterally, contiguous in proximal region. Two papillae on each side of jaw angle, the distal one longer and wider (Fig. 7c). Infradental papillae robust, and rectangular. Dorsal arm plate wider than long, rectangular, with rounded borders (Fig. 7d). Ventral arm plate pentagonal, with small notch on distal margin (Fig. 7e). Two small, perpendicular, tentacle scales, inner scale slightly longer than outer scale. Three small, compressed, blunt, arm spines, the ventral one largest (Fig. 7e).</paragraph>
<caption pageId="13" pageNumber="58">
<paragraph pageId="13" pageNumber="58">
Figure 7. Species of the family
<taxonomicName family="Amphiuridae" lsidName="" pageId="13" pageNumber="58" rank="family">Amphiuridae</taxonomicName>
.
<taxonomicName class="Ophiuroidea" family="Amphiuridae" genus="Ophiophragmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiophragmus brachyactis" order="Amphilepidida" pageId="13" pageNumber="58" phylum="Echinodermata" rank="species" species="brachyactis">Ophiophragmus brachyactis</taxonomicName>
A dorsal view, detail of the fence of papillae B ventral view C jaw D dorsal view of the arms E ventral view of the arms.
<taxonomicName class="Ophiuroidea" family="Amphiuridae" genus="Ophiostigma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiostigma isocanthum" order="Amphilepidida" pageId="13" pageNumber="58" phylum="Echinodermata" rank="species" species="isocanthum">Ophiostigma isocanthum</taxonomicName>
F dorsal view G ventral view H detail of the radial shields I jaw J dorsal view of the arms L ventral view of the arms. Scale bar = 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="13" pageNumber="58" type="distribution">
<paragraph pageId="13" pageNumber="58">Distribution.</paragraph>
<paragraph pageId="13" pageNumber="58">
Florida and Dry Tortugas, the Antilles, Gulf of Mexico, and Brazil (
<bibRefCitation author="Thomas, LP" journalOrPublisher="Bulletin of Marine Science of the Gulf and Caribbean" pageId="36" pageNumber="81" pagination="623 - 694" title="The shallow water Amphiurid brittle stars (Echinodermata, Ophiuroidea) of Florida." volume="12" year="1962">Thomas 1962</bibRefCitation>
,
<bibRefCitation author="Hendler, G" journalOrPublisher="Smithsonian Institution Press, Washington" pageId="33" pageNumber="78" title="Sea stars, sea urchins and allies: echinoderms of Florida and the Caribbean." year="1995">Hendler et al. 1995</bibRefCitation>
). In Brazil from
<normalizedToken originalValue="Paraíba">Paraiba</normalizedToken>
(
<bibRefCitation author="Gondim, AI" journalOrPublisher="Memoirs of Museum Victoria" pageId="32" pageNumber="77" title="Echinodermata da Praia do Cabo Branco, Joao Pessoa, Paraiba, Brasil. Biota Neotropica 8: 151 - 159. http: // www. biotaneotropica. org. br / v 8 n 2 / pt / abstract? inventory + bn 02408022008," url="10.1590/S1676-06032008000200016" year="2008">Gondim et al. 2008</bibRefCitation>
),
<normalizedToken originalValue="Espírito">Espirito</normalizedToken>
Santo, and Rio de Janeiro (
<bibRefCitation author="Manso, CLC" journalOrPublisher="Revista Brasileira de Biologia" pageId="34" pageNumber="79" pagination="965 - 968" title="Sobre a ocorrencia de Ophionema intricata Luetken, 1869 e Ophiophragmus brachyactis H. L. Clark, 1915 no Brasil (Echinodermata, Ophiuroidea, Amphiurudae)." volume="48" year="1988">Manso 1988</bibRefCitation>
,
<bibRefCitation author="Manso, CLC" journalOrPublisher="Revista Brasileira de Biologia" pageId="34" pageNumber="79" pagination="189 - 195" title="Ofiuroides da Plataforma Brasileira. Parte II: Norte do Estado do Rio de Janeiro, Estado do Espirito Santo, Sul do Estado da Bahia e Bancos Royal Charlotte, Hostpur e Davis (Echinodermata: Ophiuroidea)." volume="35" year="1993">1993</bibRefCitation>
). Depth 22 to 87 m. Recorded from 30 m in this study.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="58" type="remarks">
<paragraph pageId="13" pageNumber="58">Remarks.</paragraph>
<paragraph pageId="13" pageNumber="58">
Species known from bottoms with sand, in which bryozoans predominate (
<bibRefCitation author="Manso, CLC" journalOrPublisher="Revista Brasileira de Biologia" pageId="34" pageNumber="79" pagination="965 - 968" title="Sobre a ocorrencia de Ophionema intricata Luetken, 1869 e Ophiophragmus brachyactis H. L. Clark, 1915 no Brasil (Echinodermata, Ophiuroidea, Amphiurudae)." volume="48" year="1988">Manso 1988</bibRefCitation>
). We consider
<taxonomicName class="Ophiuroidea" family="Amphiuridae" genus="Amphiodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphiodia riisei" order="Amphilepidida" pageId="13" pageNumber="58" phylum="Echinodermata" rank="species" species="riisei">Amphiodia riisei</taxonomicName>
distinct from
<taxonomicName class="Ophiuroidea" family="Amphiuridae" genus="Ophiophragmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiophragmus brachyactis" order="Amphilepidida" pageId="13" pageNumber="58" phylum="Echinodermata" rank="species" species="brachyactis">Ophiophragmus brachyactis</taxonomicName>
and use the name
<taxonomicName class="Ophiuroidea" family="Amphiuridae" genus="Amphiodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphiodia riisei" order="Amphilepidida" pageId="13" pageNumber="58" phylum="Echinodermata" rank="species" species="riisei">Amphiodia riisei</taxonomicName>
for the former taxon (see discussion under this species).
</paragraph>
</subSubSection>
</treatment>
</document>