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<mods:title id="AB1959F5F938D599FE2A69C3998EAFD9">The Pristimantis trachyblepharis species group, a clade of miniaturized frogs: description of four new species and insights into the evolution of body size in the genus</mods:title>
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<taxonomicName id="4C504D4FFFC05C17FC68FE6AFB71FE16" authority="Zumel &amp; Buckley &amp; Ron, 2022" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[922,1206,433,456]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="22" pageNumber="344" phylum="Chordata" rank="species" species="ujucami" status="sp. nov.">PRISTIMANTIS UJUCAMI</taxonomicName>
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<paragraph id="8BEF36CCFFC05C17FC43FE0CFACFFE21" blockId="22.[945,1288,471,496]" box="[945,1288,471,496]" pageId="22" pageNumber="337">
(
<figureCitation id="136B2A49FFC05C17FC4BFE0CFBFBFE21" box="[953,1084,471,496]" captionStart-0="Figure 15" captionStart-1="Figure 16" captionStart-2="Figure 17" captionStart-3="Figure 18" captionStart-4="Figure 19" captionStartId-0="23.[163,243,938,960]" captionStartId-1="24.[144,222,1392,1414]" captionStartId-2="25.[163,243,1483,1505]" captionStartId-3="26.[145,224,1675,1697]" captionStartId-4="27.[163,242,1664,1686]" captionTargetBox-0="[331,1277,200,891]" captionTargetBox-1="[305,1265,195,1352]" captionTargetBox-2="[166,1437,198,1435]" captionTargetBox-3="[180,1391,203,1632]" captionTargetBox-4="[173,1431,195,1635]" captionTargetId-0="figure-495@23.[323,1283,195,898]" captionTargetId-1="figure-304@24.[305,1265,195,1352]" captionTargetId-2="figure-122@25.[163,1446,195,1443]" captionTargetId-3="figure-27@26.[178,1391,195,1635]" captionTargetId-4="figure-26@27.[173,1431,195,1635]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="25" captionTargetPageId-3="26" captionTargetPageId-4="27" captionText-0="Figure 15. Photographs in live of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm). A, dorsolateral view; B, ventral view; C, dorsolateral and inguinal region view." captionText-1="Figure 16. Photographs of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm) in preservative. A, dorsal view; B, ventral view; C, ventral view of the left hand; D, ventral view of the right foot." captionText-2="Figure 17. Colour variation in life individuals of Pristimantis ujucami in dorsolateral view. A, QCAZ 49031 (subadult female, SVL 16.19 mm); B, QCAZ 57002 (adult male, SVL 14.39 mm); C, QCAZ 57010 (subadult female, 18.95 mm); D, QCAZ 57003 (adult male, SVL 15.47 mm); E, QCAZ 57004 (subadult female, SVL 18.91 mm); F, QCAZ 57646 (adult male, SVL 14.2 mm); G, QCAZ 60212 (adult female, SVL 22.33 mm); H, QCAZ 58865 (adult female, SVL 20.62 mm); I, QCAZ 60214 (adult female, SVL 23.76 mm); J, QCAZ 60194 (adult female, SVL 22.43 mm); K, QCAZ 60196 (adult female, SVL 20.45 mm); L, QCAZ 58874 (subadult female, SVL 16.74 mm)." captionText-3="Figure 18. Colour variation in life individuals of Pristimantis ujucami in ventral view. A, QCAZ 49031 (subadult female, SVL 16.19 mm); B, QCAZ 57002 (adult male, SVL 14.39 mm); C, QCAZ 57010 (subadult female, 18.95 mm); D, QCAZ 57003 (adult male, SVL 15.47 mm); E, QCAZ 57004 (subadult female, SVL 18.91 mm); F, QCAZ 57646 (adult male, SVL 14.2 mm); G, QCAZ 60212 (adult female, SVL 22.33 mm); H, QCAZ 58865 (adult female, SVL 20.62 mm); I, QCAZ 60214 (adult female, SVL 23.76 mm); J, QCAZ 60194 (adult female, SVL 22.43 mm); K, QCAZ 60196 (adult female, SVL 20.45 mm); L, QCAZ 58874 (subadult female, SVL 16.74 mm)." captionText-4="Figure 19. Colour variation in preserved individuals of Pristimantis ujucami: 1, dorsal views; 2, ventral views. A, QCAZ 49026 (adult female, SVL 22.62 mm); B, QCAZ 60194 (adult female, SVL 22.43 mm); C, QCAZ 60212 (adult female, SVL 22.33 mm); D, QCAZ 60214 (adult female, SVL 23.76 mm); E, QCAZ 58874 (subadult female, SVL 16.74 mm); F, QCAZ 60206 (subadult female, SVL 16.61 mm); G, QCAZ 57010 (subadult female, SVL 18.95 mm); H, QCAZ 57003 (adult male, SVL 15.47 mm); I, QCAZ 60621 (adult male, SVL 16.42 mm); J, QCAZ 57002 (adult male, 14.39 mm)." figureDoi-0="http://doi.org/10.5281/zenodo.6530605" figureDoi-1="http://doi.org/10.5281/zenodo.6530607" figureDoi-2="http://doi.org/10.5281/zenodo.6530609" figureDoi-3="http://doi.org/10.5281/zenodo.6530611" figureDoi-4="http://doi.org/10.5281/zenodo.6530613" httpUri-0="https://zenodo.org/record/6530605/files/figure.png" httpUri-1="https://zenodo.org/record/6530607/files/figure.png" httpUri-2="https://zenodo.org/record/6530609/files/figure.png" httpUri-3="https://zenodo.org/record/6530611/files/figure.png" httpUri-4="https://zenodo.org/record/6530613/files/figure.png" pageId="22" pageNumber="337">FIGS 15–19</figureCitation>
; 
<tableCitation id="C6D20377FFC05C17FBBBFE0CFB72FE21" box="[1097,1205,471,496]" captionStart="Table 2" captionStartId="7.[144,208,553,575]" captionTargetPageId="7" captionText="Table 2. Snout–vent length (SVL) and qualitative morphological traits in the Pristimantis trachyblepharis species group. Average SVL (in mm), is followed" httpUri="http://table.plazi.org/id/DF2F6644FFD15C06FF62FDF2FAFFFDAB" pageId="22" pageNumber="337" tableUuid="DF2F6644FFD15C06FF62FDF2FAFFFDAB">TABLES 2</tableCitation>
AND 
<tableCitation id="C6D20377FFC05C17FB00FE0CFAC7FE21" box="[1266,1280,471,496]" captionStart="Table 3" captionStartId="8.[145,209,196,217]" captionTargetPageId="8" captionText="Table 3. Qualitative morphological traits in the Pristimantis trachyblepharis species group. Coloration corresponds to live individuals. +: present; -: absent. Data from examined specimens of the QCAZ collection (listed in the Supporting Information, Table S3) except those of P. minimus, which are from Terán-Valdez &amp; Guayasamin (2010)" httpUri="http://table.plazi.org/id/DF2F6644FFDE5C09FF63FF1FFB79FEC5" pageId="22" pageNumber="337" tableUuid="DF2F6644FFDE5C09FF63FF1FFB79FEC5">3</tableCitation>
)
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<emphasis id="B924EADEFFC05C17FCDBFDDAFBB0FDC7" box="[809,1143,512,534]" italics="true" pageId="22" pageNumber="337">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act: 
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<typeStatus id="54EB886EFFC05C17FCDBFD86FC49FDA3" box="[809,910,605,626]" pageId="22" pageNumber="337" type="holotype">Holotype</typeStatus>
(
<figureCitation id="136B2A49FFC05C17FC6FFD85FC35FDA3" box="[925,1010,605,627]" captionStart="Figure 15" captionStartId="23.[163,243,938,960]" captionTargetBox="[331,1277,200,891]" captionTargetId="figure-495@23.[323,1283,195,898]" captionTargetPageId="23" captionText="Figure 15. Photographs in live of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm). A, dorsolateral view; B, ventral view; C, dorsolateral and inguinal region view." figureDoi="http://doi.org/10.5281/zenodo.6530605" httpUri="https://zenodo.org/record/6530605/files/figure.png" pageId="22" pageNumber="337">Figs 15</figureCitation>
, 
<figureCitation id="136B2A49FFC05C17FC0DFD86FBDCFDA3" box="[1023,1051,605,626]" captionStart="Figure 16" captionStartId="24.[144,222,1392,1414]" captionTargetBox="[305,1265,195,1352]" captionTargetId="figure-304@24.[305,1265,195,1352]" captionTargetPageId="24" captionText="Figure 16. Photographs of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm) in preservative. A, dorsal view; B, ventral view; C, ventral view of the left hand; D, ventral view of the right foot." figureDoi="http://doi.org/10.5281/zenodo.6530607" httpUri="https://zenodo.org/record/6530607/files/figure.png" pageId="22" pageNumber="337">16</figureCitation>
):
</emphasis>
<specimenCode id="DBF69EB7FFC05C17FBCFFD86FB08FDA3" box="[1085,1231,605,626]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="22" pageNumber="337" type="Museum">QCAZ 49030</specimenCode>
(field no. YSNA-130), adult female from 
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, 
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, 
<collectingMunicipality id="6B8BACB6FFC05C17FC52FD40FB9DFD61" box="[928,1114,667,689]" pageId="22" pageNumber="337">Cantón Morona</collectingMunicipality>
, 
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, on the eastern border of 
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, near the 
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(
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, 
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), 
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a.s.l.
</elevation>
, collected by 
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and 
<collectorName id="26A5531AFFC05C17FCA9FCCEFC34FCFA" box="[859,1011,789,811]" pageId="22" pageNumber="337">Rubén Jarrín</collectorName>
on 
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<collectingDate id="EFAAE9E4FFC05C17FBEEFCCEFB77FCFB" box="[1052,1200,789,811]" pageId="22" pageNumber="337" value="2010-07-25">25 July 2010</collectingDate>
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.
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</paragraph>
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<typeStatus id="54EB886EFFC05C17FCDBFC8FFC65FCB8" box="[809,930,852,873]" pageId="22" pageNumber="337" type="paratype">Paratypes</typeStatus>
(30: 
<specimenCount id="9D56FD45FFC05C17FC14FC8FFBA8FCB8" box="[998,1135,852,873]" pageId="22" pageNumber="337" type="male">nine males</specimenCount>
, 
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):
</emphasis>
<collectingCountry id="F347765CFFC05C17FAD1FC8FFA4BFCBB" box="[1315,1420,852,874]" name="Ecuador" pageId="22" pageNumber="337">Ecuador</collectingCountry>
: Provincia 
<collectingRegion id="4994F82EFFC05C17FC6EFCA8FBA4FC59" box="[924,1123,882,904]" country="Ecuador" name="Morona-Santiago" pageId="22" pageNumber="337">Morona Santiago</collectingRegion>
: Abanico Protected Forest, QCAZ 49026, adult female, and QCAZ 49031, subadult female (
<geoCoordinate id="EE64500BFFC05C17FC74FC6BFBC5FC14" box="[902,1026,944,966]" degrees="2.25901" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="1" value="-2.25901">2.25901º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FBFDFC6BFB53FC14" box="[1039,1172,944,966]" degrees="78.1950" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.195">78.1950º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FB5AFC6BFB39FC14" box="[1192,1278,944,965]" metricMagnitude="3" metricUnit="m" metricValue="1.72" pageId="22" pageNumber="337" unit="m" value="1720.0">1720 m</quantity>
a.s.l.; QCAZ 56998, adult male (
<geoCoordinate id="EE64500BFFC05C17FBE4FC15FB4FFC35" box="[1046,1160,974,996]" degrees="2.2576" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-2.2576">2.2576º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FB65FC15FAE5FC35" box="[1175,1314,974,996]" degrees="78.1986" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.1986">78.1986º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FACAFC15FA57FC35" box="[1336,1424,974,996]" metricMagnitude="3" metricUnit="m" metricValue="1.5419999999999998" pageId="22" pageNumber="337" unit="m" value="1542.0">1542 m</quantity>
a.s.l.; QCAZ 57001–57002, adult males (
<geoCoordinate id="EE64500BFFC05C17FAEAFC36FA4CFBD3" box="[1304,1419,1005,1027]" degrees="2.2579" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-2.2579">2.2579º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FCDBFBD7FC76FBF0" box="[809,945,1036,1058]" degrees="78.1985" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.1985">78.1985º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FC37FBD7FBDBFBF0" box="[965,1052,1036,1058]" metricMagnitude="3" metricUnit="m" metricValue="1.555" pageId="22" pageNumber="337" unit="m" value="1555.0">1555 m</quantity>
a.s.l.; QCAZ 57003, adult male (
<geoCoordinate id="EE64500BFFC05C17FCC2FBF1FC67FB91" box="[816,928,1066,1088]" degrees="2.2581" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-2.2581">2.2581º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FC5FFBF1FBF3FB91" box="[941,1076,1066,1088]" degrees="78.1986" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.1986">78.1986º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FBBAFBF1FB58FB91" box="[1096,1183,1066,1088]" metricMagnitude="3" metricUnit="m" metricValue="1.526" pageId="22" pageNumber="337" unit="m" value="1526.0">1526 m</quantity>
a.s.l.; QCAZ 57004– 57005, subadult females (
<geoCoordinate id="EE64500BFFC05C17FB9CFB92FB22FB8F" box="[1134,1253,1097,1119]" degrees="2.2582" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-2.2582">2.2582º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FB07FB92FA43FB8E" box="[1269,1412,1097,1119]" degrees="78.1984" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.1984">78.1984º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FCDBFBB3FC47FBAC" box="[809,896,1128,1150]" metricMagnitude="3" metricUnit="m" metricValue="1.564" pageId="22" pageNumber="337" unit="m" value="1564.0">1564 m</quantity>
a.s.l.; QCAZ 57008, adult female (
<geoCoordinate id="EE64500BFFC05C17FAEEFBB3FA4BFBAC" box="[1308,1420,1128,1150]" degrees="2.2582" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-2.2582">2.2582º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FCDBFB5CFC72FB4D" box="[809,949,1159,1180]" degrees="78.1984" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.1984">78.1984º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FC3EFB5DFBE2FB4D" box="[972,1061,1158,1180]" metricMagnitude="3" metricUnit="m" metricValue="1.573" pageId="22" pageNumber="337" unit="m" value="1573.0">1573 m</quantity>
a.s.l.; QCAZ 57010, subadult female (
<geoCoordinate id="EE64500BFFC05C17FC76FB7EFC28FB6B" box="[900,1007,1189,1211]" degrees="2.2583" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-2.2583">2.2583º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FC09FB7EFBB9FB6A" box="[1019,1150,1189,1211]" degrees="78.1984" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.1984">78.1984º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FB62FB7EFB23FB6A" box="[1168,1252,1189,1211]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="22" pageNumber="337" unit="m" value="1550.0">1550 m</quantity>
a.s.l. Collected by Y. Sagredo, R. Jarrín, F. Ayala, S. Arroyo, S. Valverde and L. Cedeño on 
<date id="FFEE100CFFC05C17FC0AFB39FB74FB29" box="[1016,1203,1250,1272]" pageId="22" pageNumber="337" value="2010-07-23" valueMax="2010-07-25" valueMin="2010-07-23">23–25 July 2010</date>
and 
<date id="FFEE100CFFC05C17FB1FFB38FA4AFB29" box="[1261,1421,1250,1272]" pageId="22" pageNumber="337" value="2014-03-02">2 March 2014</date>
; Volcano River, Sangay National Park. QCAZ 58865, adult female, and QCAZ 58874, subadult female (
<geoCoordinate id="EE64500BFFC05C17FCC2FAE4FC66FA85" box="[816,929,1343,1364]" degrees="2.0983" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-2.0983">2.0983º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FC5DFAE5FBFFFA85" box="[943,1080,1342,1364]" degrees="78.5554" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.5554">78.5554º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FBBEFAE5FB64FA85" box="[1100,1187,1342,1364]" metricMagnitude="3" metricUnit="m" metricValue="1.406" pageId="22" pageNumber="337" unit="m" value="1406.0">1406 m</quantity>
a.s.l.; QCAZ 58866, subadult female (
<geoCoordinate id="EE64500BFFC05C17FC02FA86FB9CFAA3" box="[1008,1115,1373,1394]" degrees="2.0987" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-2.0987">2.0987º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FB94FA86FB2FFAA2" box="[1126,1256,1373,1395]" degrees="78.1560" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.156">78.1560º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FB09FA86FA88FAA2" box="[1275,1359,1373,1395]" metricMagnitude="3" metricUnit="m" metricValue="1.403" pageId="22" pageNumber="337" unit="m" value="1403.0">1403 m</quantity>
a.s.l.; QCAZ 58868, subadult female (
<geoCoordinate id="EE64500BFFC05C17FB5AFAA7FADCFA40" box="[1192,1307,1404,1425]" degrees="2.0903" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-2.0903">2.0903º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FAD8FAA7FC83FA61" degrees="78.1897" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.1897">78.1897º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FCA5FA41FC6CFA61" box="[855,939,1434,1456]" metricMagnitude="3" metricUnit="m" metricValue="1.551" pageId="22" pageNumber="337" unit="m" value="1551.0">1551 m</quantity>
a.s.l.; QCAZ 58870, adult male (
<geoCoordinate id="EE64500BFFC05C17FAD2FA40FA4BFA61" box="[1312,1420,1435,1456]" degrees="2.0971" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-2.0971">2.0971º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FCDBFA62FC6DFA1F" box="[809,938,1465,1487]" degrees="78.1553" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.1553">78.1553º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FC4EFA62FBC8FA1F" box="[956,1039,1465,1486]" metricMagnitude="3" metricUnit="m" metricValue="1.411" pageId="22" pageNumber="337" unit="m" value="1411.0">1411 m</quantity>
a.s.l. Collected by D. Rivadeneira, F. Mora, J.C. Sánchez, and A. Correa, on 
<date id="FFEE100CFFC05C17FAFCFA03FCA4F9DD" pageId="22" pageNumber="337" value="2015-02-03">3 February 2015</date>
. Provincia 
<collectingRegion id="4994F82EFFC05C17FC2FFA2CFB77F9DD" box="[989,1200,1526,1548]" country="Ecuador" name="Zamora-Chinchipe" pageId="22" pageNumber="337">Zamora Chinchipe</collectingRegion>
: Numbami Reserve, QCAZ 57646, adult male (
<geoCoordinate id="EE64500BFFC05C17FBA2F9CEFB7CF9FB" box="[1104,1211,1557,1579]" degrees="4.1672" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-4.1672">4.1672º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FB34F9CEFA8FF9FB" box="[1222,1352,1557,1578]" degrees="78.9489" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.9489">78.9489º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FAA8F9CEFCF8F998" metricMagnitude="3" metricUnit="m" metricValue="1.458" pageId="22" pageNumber="337" unit="m" value="1458.0">1458 m</quantity>
a.s.l. Collected by S. Ron, D. Paucar, P. Venegas, D. Almeida, D. Velalcázar, M.J. Navarrete, S. Arroyo, N. Páez and Z. Lange, on 
<date id="FFEE100CFFC05C17FB93F9AAFB37F957" box="[1121,1264,1649,1671]" pageId="22" pageNumber="337" value="2014-07-08">8 July 2014</date>
; Bombuscaro River, Podocarpus National Park. QCAZ 60194, adult female (
<geoCoordinate id="EE64500BFFC05C17FC78F975FC3AF912" box="[906,1021,1710,1732]" degrees="4.1146" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-4.1146">4.1146º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FBFEF975FB50F915" box="[1036,1175,1710,1732]" degrees="78.9670" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.967">78.9670º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FB5EF975FB30F915" box="[1196,1271,1710,1732]" metricMagnitude="2" metricUnit="m" metricValue="9.809999999999999" pageId="22" pageNumber="337" unit="m" value="981.0">981 m</quantity>
a.s.l.; QCAZ 60195–60196, adult females (
<geoCoordinate id="EE64500BFFC05C17FB71F916FB36F933" box="[1155,1265,1741,1763]" degrees="4.1250" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-4.125">4.1250º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FB0DF916FA42F933" box="[1279,1413,1741,1762]" degrees="78.9788" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.9788">78.9788º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FCDBF937FCBAF8D0" box="[809,893,1772,1794]" metricMagnitude="3" metricUnit="m" metricValue="1.164" pageId="22" pageNumber="337" unit="m" value="1164.0">1164 m</quantity>
a.s.l.; QCAZ 60200, subadult male, and QCAZ 60202, adult female (
<geoCoordinate id="EE64500BFFC05C17FBD1F8D1FB55F8CE" box="[1059,1170,1802,1824]" degrees="4.1146" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-4.1146">4.1146º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FB6DF8D1FAE1F8F1" box="[1183,1318,1802,1824]" degrees="78.9633" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.9633">78.9633º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FAC8F8D1FA57F8F1" box="[1338,1424,1802,1824]" metricMagnitude="3" metricUnit="m" metricValue="1.112" pageId="22" pageNumber="337" unit="m" value="1112.0">1112 m</quantity>
a.s.l.; QCAZ 60205, QCAZ 60612 and QCAZ 60214, adult females (
<geoCoordinate id="EE64500BFFC05C17FC28F893FB8DF88C" box="[986,1098,1864,1885]" degrees="4.1344" direction="south" orientation="latitude" pageId="22" pageNumber="337" precision="5" value="-4.1344">4.1344º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC05C17FBAAF893FB18F88C" box="[1112,1247,1864,1885]" degrees="78.9938" direction="west" orientation="longitude" pageId="22" pageNumber="337" precision="5" value="-78.9938">78.9938º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC05C17FB01F893FA8DF88C" box="[1267,1354,1864,1885]" metricMagnitude="3" metricUnit="m" metricValue="1.443" pageId="22" pageNumber="337" unit="m" value="1443.0">1443 m</quantity>
a.s.l.; QCAZ 60206–60207, subadult females (
<geoCoordinate id="EE64500BFFC15C16FD62FC24FCC2FBC5" box="[656,773,1023,1044]" degrees="4.1344" direction="south" orientation="latitude" pageId="23" pageNumber="338" precision="5" value="-4.1344">4.1344º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC15C16FF51FBC5FEE1FBE2" box="[163,294,1054,1075]" degrees="78.9938" direction="west" orientation="longitude" pageId="23" pageNumber="338" precision="5" value="-78.9938">78.9938º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC15C16FECBFBC5FE49FBE2" box="[313,398,1054,1075]" metricMagnitude="3" metricUnit="m" metricValue="1.443" pageId="23" pageNumber="338" unit="m" value="1443.0">1443 m</quantity>
a.s.l.; QCAZ 60216, adult female (
<geoCoordinate id="EE64500BFFC15C16FF58FBE7FEDCFB80" box="[170,283,1084,1105]" degrees="4.1333" direction="south" orientation="latitude" pageId="23" pageNumber="338" precision="5" value="-4.1333">4.1333º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC15C16FEDBFBE7FE76FB83" box="[297,433,1084,1106]" degrees="78.9854" direction="west" orientation="longitude" pageId="23" pageNumber="338" precision="5" value="-78.9854">78.9854º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC15C16FE34FBE7FDDAFB83" box="[454,541,1084,1106]" metricMagnitude="3" metricUnit="m" metricValue="1.281" pageId="23" pageNumber="338" unit="m" value="1281.0">1281 m</quantity>
a.s.l.; QCAZ 60218, subadult female (
<geoCoordinate id="EE64500BFFC15C16FE9BFB80FE12FBA1" box="[361,469,1115,1136]" degrees="4.1333" direction="south" orientation="latitude" pageId="23" pageNumber="338" precision="5" value="-4.1333">4.1333º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC15C16FE12FB80FDA5FBA1" box="[480,610,1115,1137]" degrees="78.9854" direction="west" orientation="longitude" pageId="23" pageNumber="338" precision="5" value="-78.9854">78.9854º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC15C16FD87FB80FD0EFBA1" box="[629,713,1115,1136]" metricMagnitude="3" metricUnit="m" metricValue="1.281" pageId="23" pageNumber="338" unit="m" value="1281.0">1281 m</quantity>
a.s.l.; QCAZ 60621 and QCAZ 60629, adult males (
<geoCoordinate id="EE64500BFFC15C16FD69FBA1FCC1FB5E" box="[667,774,1146,1167]" degrees="4.1333" direction="south" orientation="latitude" pageId="23" pageNumber="338" precision="5" value="-4.1333">4.1333º S</geoCoordinate>
, 
<geoCoordinate id="EE64500BFFC15C16FF51FB43FEE3FB7F" box="[163,292,1176,1198]" degrees="78.9854" direction="west" orientation="longitude" pageId="23" pageNumber="338" precision="5" value="-78.9854">78.9854º W</geoCoordinate>
), 
<quantity id="4CA89B29FFC15C16FEC4FB43FE4EFB7F" box="[310,393,1176,1198]" metricMagnitude="3" metricUnit="m" metricValue="1.281" pageId="23" pageNumber="338" unit="m" value="1281.0">1281 m</quantity>
a.s.l. Collected by D. Rivadeneira, F. Mora, J.C. Sánchez, D. Velalcázar, D. Núñez, J. Pinto, K. Cruz and L. Tipantiza on 8 April, 2015.
</paragraph>
<caption id="DF2F6644FFC15C16FF51FC71FC73FC0C" ID-DOI="http://doi.org/10.5281/zenodo.6530605" ID-Zenodo-Dep="6530605" httpUri="https://zenodo.org/record/6530605/files/figure.png" pageId="23" pageNumber="338" startId="23.[163,243,938,960]" targetBox="[331,1277,200,891]" targetPageId="23">
<paragraph id="8BEF36CCFFC15C16FF51FC71FC73FC0C" blockId="23.[163,1443,938,989]" pageId="23" pageNumber="338">
<emphasis id="B924EADEFFC15C16FF51FC71FEDCFC6E" bold="true" box="[163,283,938,960]" pageId="23" pageNumber="338">Figure 15.</emphasis>
Photographs in live of the holotype of 
<taxonomicName id="4C504D4FFFC15C16FD4DFC71FC64FC11" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[703,931,938,960]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="23" pageNumber="338" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFC15C16FD4DFC71FC64FC11" box="[703,931,938,960]" italics="true" pageId="23" pageNumber="338">Pristimantis ujucami</emphasis>
</taxonomicName>
(QCAZ 49030; adult female, SVL 21.16 mm). A, dorsolateral view; B, ventral view; C, dorsolateral and inguinal region view.
</paragraph>
</caption>
<paragraph id="8BEF36CCFFC15C16FF51FACFFE15FA99" blockId="23.[163,779,1300,1353]" pageId="23" pageNumber="338">
<emphasis id="B924EADEFFC15C16FF51FACFFE0EFAFB" box="[163,457,1300,1322]" italics="true" pageId="23" pageNumber="338">Suggested common name:</emphasis>
English: Ujukam rain frog. Spanish: 
<emphasis id="B924EADEFFC15C16FEFEFAE8FE0CFA99" box="[268,459,1331,1352]" italics="true" pageId="23" pageNumber="338">cutín de Ujukam</emphasis>
.
</paragraph>
<paragraph id="8BEF36CCFFC15C16FF51FAA9FBF0F8BE" blockId="23.[163,779,1394,1906]" lastBlockId="23.[827,1443,1023,1903]" pageId="23" pageNumber="338">
<emphasis id="B924EADEFFC15C16FF51FAA9FE05FA56" box="[163,450,1394,1415]" italics="true" pageId="23" pageNumber="338">
Diagnosis (
<figureCitation id="136B2A49FFC15C16FEDFFAA9FE76FA56" box="[301,433,1394,1415]" captionStart-0="Figure 15" captionStart-1="Figure 16" captionStart-2="Figure 17" captionStart-3="Figure 18" captionStart-4="Figure 19" captionStartId-0="23.[163,243,938,960]" captionStartId-1="24.[144,222,1392,1414]" captionStartId-2="25.[163,243,1483,1505]" captionStartId-3="26.[145,224,1675,1697]" captionStartId-4="27.[163,242,1664,1686]" captionTargetBox-0="[331,1277,200,891]" captionTargetBox-1="[305,1265,195,1352]" captionTargetBox-2="[166,1437,198,1435]" captionTargetBox-3="[180,1391,203,1632]" captionTargetBox-4="[173,1431,195,1635]" captionTargetId-0="figure-495@23.[323,1283,195,898]" captionTargetId-1="figure-304@24.[305,1265,195,1352]" captionTargetId-2="figure-122@25.[163,1446,195,1443]" captionTargetId-3="figure-27@26.[178,1391,195,1635]" captionTargetId-4="figure-26@27.[173,1431,195,1635]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="25" captionTargetPageId-3="26" captionTargetPageId-4="27" captionText-0="Figure 15. Photographs in live of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm). A, dorsolateral view; B, ventral view; C, dorsolateral and inguinal region view." captionText-1="Figure 16. Photographs of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm) in preservative. A, dorsal view; B, ventral view; C, ventral view of the left hand; D, ventral view of the right foot." captionText-2="Figure 17. Colour variation in life individuals of Pristimantis ujucami in dorsolateral view. A, QCAZ 49031 (subadult female, SVL 16.19 mm); B, QCAZ 57002 (adult male, SVL 14.39 mm); C, QCAZ 57010 (subadult female, 18.95 mm); D, QCAZ 57003 (adult male, SVL 15.47 mm); E, QCAZ 57004 (subadult female, SVL 18.91 mm); F, QCAZ 57646 (adult male, SVL 14.2 mm); G, QCAZ 60212 (adult female, SVL 22.33 mm); H, QCAZ 58865 (adult female, SVL 20.62 mm); I, QCAZ 60214 (adult female, SVL 23.76 mm); J, QCAZ 60194 (adult female, SVL 22.43 mm); K, QCAZ 60196 (adult female, SVL 20.45 mm); L, QCAZ 58874 (subadult female, SVL 16.74 mm)." captionText-3="Figure 18. Colour variation in life individuals of Pristimantis ujucami in ventral view. A, QCAZ 49031 (subadult female, SVL 16.19 mm); B, QCAZ 57002 (adult male, SVL 14.39 mm); C, QCAZ 57010 (subadult female, 18.95 mm); D, QCAZ 57003 (adult male, SVL 15.47 mm); E, QCAZ 57004 (subadult female, SVL 18.91 mm); F, QCAZ 57646 (adult male, SVL 14.2 mm); G, QCAZ 60212 (adult female, SVL 22.33 mm); H, QCAZ 58865 (adult female, SVL 20.62 mm); I, QCAZ 60214 (adult female, SVL 23.76 mm); J, QCAZ 60194 (adult female, SVL 22.43 mm); K, QCAZ 60196 (adult female, SVL 20.45 mm); L, QCAZ 58874 (subadult female, SVL 16.74 mm)." captionText-4="Figure 19. Colour variation in preserved individuals of Pristimantis ujucami: 1, dorsal views; 2, ventral views. A, QCAZ 49026 (adult female, SVL 22.62 mm); B, QCAZ 60194 (adult female, SVL 22.43 mm); C, QCAZ 60212 (adult female, SVL 22.33 mm); D, QCAZ 60214 (adult female, SVL 23.76 mm); E, QCAZ 58874 (subadult female, SVL 16.74 mm); F, QCAZ 60206 (subadult female, SVL 16.61 mm); G, QCAZ 57010 (subadult female, SVL 18.95 mm); H, QCAZ 57003 (adult male, SVL 15.47 mm); I, QCAZ 60621 (adult male, SVL 16.42 mm); J, QCAZ 57002 (adult male, 14.39 mm)." figureDoi-0="http://doi.org/10.5281/zenodo.6530605" figureDoi-1="http://doi.org/10.5281/zenodo.6530607" figureDoi-2="http://doi.org/10.5281/zenodo.6530609" figureDoi-3="http://doi.org/10.5281/zenodo.6530611" figureDoi-4="http://doi.org/10.5281/zenodo.6530613" httpUri-0="https://zenodo.org/record/6530605/files/figure.png" httpUri-1="https://zenodo.org/record/6530607/files/figure.png" httpUri-2="https://zenodo.org/record/6530609/files/figure.png" httpUri-3="https://zenodo.org/record/6530611/files/figure.png" httpUri-4="https://zenodo.org/record/6530613/files/figure.png" pageId="23" pageNumber="338">Figs 15–19</figureCitation>
):
</emphasis>
A species of 
<taxonomicName id="4C504D4FFFC15C16FD81FAA9FCCCFA56" authorityName="Jimenez de la Espada" authorityYear="1870" box="[627,779,1394,1415]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="23" pageNumber="338" phylum="Chordata" rank="genus">
<emphasis id="B924EADEFFC15C16FD81FAA9FCCCFA56" box="[627,779,1394,1415]" italics="true" pageId="23" pageNumber="338">Pristimantis</emphasis>
</taxonomicName>
characterized by the following combination of characters:(1) dorsal skin slightly shagreen to shagreen, bedecked with pustules and/or spicules; scapular folds distinct,&gt; &lt;, \ / or) (-shaped, with scapular subconical or conical tubercles along them; scapular folds may join posteriorly and extend as diffuse paravertebral folds; ventral skin weakly areolate on throat and chest, areolate on belly and coarsely areolate on posteroventral surface of thighs; discoidal fold distinct; (2) tympanic annulus distinct and externally visible through skin; tympanic membrane present but hidden by skin; upper and posterior margins of tympanic annulus covered by low supratympanic fold; two distinct postrictal subconical tubercles on each side of the head; (3) snout moderate in length with a small rostral papilla at tip, subacuminate to acuminate in dorsal view, rounded in lateral view; 
<emphasis id="B924EADEFFC15C16FB13FC24FA65FBC5" box="[1249,1442,1023,1044]" italics="true" pageId="23" pageNumber="338">canthus rostralis</emphasis>
very distinct, rounded in lateral view and concave or straight in dorsal view (4) upper eyelid with several subconical tubercles; (5) cranial crests absent; (6) vocal slits and nuptial pads absent; (7) Finger I shorter than Finger II; discs of digits expanded, rounded to truncate; (8) narrow lateral fringes on fingers; (9) low, rounded and subconical ulnar tubercles; (10) heel bearing one distinct subconical tubercle and a few nearby lower pustules; inner tarsal fold distinct; low and subconical outer tarsal tubercles; (12) narrow lateral fringes on toes; basal toe webbing, most distinctive between Toe IV and Toe V; Toe V slightly longer than Toe III; toe discs nearly as large as those on fingers; (13) coloration in life highly variable, dorsum generally pale brown or yellowish; scapular and supratympanic folds, and scapular tubercles darker than background; dark labial bars on both sides of head; flanks and dorsal surfaces of limbs usually with the same colour of the dorsum, showing several darker diagonal stripes; slightly translucent venter with dark flecks distributed on throat, chest, belly and ventral surfaces of limbs, but usually more numerous on throat, chest and ventral surfaces of hindlimbs; in males, groins and venter with same colour; in females groins are salmon reddish; bright golden iris; and (14) SVL in adult females from 
<quantity id="4CA89B29FFC15C16FCC9F8C7FBDFF8E0" box="[827,1048,1820,1842]" metricMagnitude="-2" metricUnit="m" metricValue="2.1340000000000003" metricValueMax="2.3760000000000003" metricValueMin="1.8920000000000001" pageId="23" pageNumber="338" unit="mm" value="21.34" valueMax="23.76" valueMin="18.92">18.92 to 23.76 mm</quantity>
(mean = 21.46; 
<emphasis id="B924EADEFFC15C16FB29F8C6FB28F8E3" box="[1243,1263,1821,1842]" italics="true" pageId="23" pageNumber="338">N</emphasis>
= 12), in adult males SVL from 
<quantity id="4CA89B29FFC15C16FBF5F8E0FB21F881" box="[1031,1254,1851,1873]" metricMagnitude="-2" metricUnit="m" metricValue="1.5265" metricValueMax="1.642" metricValueMin="1.411" pageId="23" pageNumber="338" unit="mm" value="15.265" valueMax="16.42" valueMin="14.11">14.11 to 16.42 mm</quantity>
(mean = 15.19; 
<emphasis id="B924EADEFFC15C16FCC9F881FC88F8BE" box="[827,847,1882,1903]" italics="true" pageId="23" pageNumber="338">N</emphasis>
= 8; 
<tableCitation id="C6D20377FFC15C16FC76F882FC25F8BF" box="[900,994,1881,1903]" captionStart="Table 2" captionStartId="7.[144,208,553,575]" captionTargetPageId="7" captionText="Table 2. Snout–vent length (SVL) and qualitative morphological traits in the Pristimantis trachyblepharis species group. Average SVL (in mm), is followed" httpUri="http://table.plazi.org/id/DF2F6644FFD15C06FF62FDF2FAFFFDAB" pageId="23" pageNumber="338" tableUuid="DF2F6644FFD15C06FF62FDF2FAFFFDAB">Tables 2</tableCitation>
and 
<tableCitation id="C6D20377FFC15C16FBE8F882FBEFF8BF" box="[1050,1064,1881,1902]" captionStart="Table 3" captionStartId="8.[145,209,196,217]" captionTargetPageId="8" captionText="Table 3. Qualitative morphological traits in the Pristimantis trachyblepharis species group. Coloration corresponds to live individuals. +: present; -: absent. Data from examined specimens of the QCAZ collection (listed in the Supporting Information, Table S3) except those of P. minimus, which are from Terán-Valdez &amp; Guayasamin (2010)" httpUri="http://table.plazi.org/id/DF2F6644FFDE5C09FF63FF1FFB79FEC5" pageId="23" pageNumber="338" tableUuid="DF2F6644FFDE5C09FF63FF1FFB79FEC5">3</tableCitation>
).
</paragraph>
<caption id="DF2F6644FFCE5C19FF62FAABFBBEFA72" ID-DOI="http://doi.org/10.5281/zenodo.6530607" ID-Zenodo-Dep="6530607" httpUri="https://zenodo.org/record/6530607/files/figure.png" pageId="24" pageNumber="339" startId="24.[144,222,1392,1414]" targetBox="[305,1265,195,1352]" targetPageId="24">
<paragraph id="8BEF36CCFFCE5C19FF62FAABFBBEFA72" blockId="24.[144,1426,1392,1443]" pageId="24" pageNumber="339">
<emphasis id="B924EADEFFCE5C19FF62FAABFEC3FA54" bold="true" box="[144,260,1392,1414]" pageId="24" pageNumber="339">Figure 16.</emphasis>
Photographs of the holotype of 
<taxonomicName id="4C504D4FFFCE5C19FDB9FAABFCEFFA57" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[587,808,1392,1414]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCE5C19FDB9FAABFCEFFA57" box="[587,808,1392,1414]" italics="true" pageId="24" pageNumber="339">Pristimantis ujucami</emphasis>
</taxonomicName>
(QCAZ 49030; adult female, SVL 21.16 mm) in preservative. A, dorsal view; B, ventral view; C, ventral view of the left hand; D, ventral view of the right foot.
</paragraph>
</caption>
<paragraph id="8BEF36CCFFCE5C19FF63FA0BFE15F8A5" blockId="24.[145,761,1488,1908]" pageId="24" pageNumber="339">
<emphasis id="B924EADEFFCE5C19FF63FA0BFDE4FA34" box="[145,547,1488,1509]" italics="true" pageId="24" pageNumber="339">Comparisons with other species:</emphasis>
In this section, coloration refers to live individuals unless otherwise noticed. Among the species of the 
<taxonomicName id="4C504D4FFFCE5C19FDD6F9D5FD3FF9F3" baseAuthorityName="Boulenger" baseAuthorityYear="1918" box="[548,760,1549,1571]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="trachyblepharis">
<emphasis id="B924EADEFFCE5C19FDD6F9D5FD3FF9F3" box="[548,760,1549,1571]" italics="true" pageId="24" pageNumber="339">P. trachyblepharis</emphasis>
</taxonomicName>
species group, 
<taxonomicName id="4C504D4FFFCE5C19FE92F9F7FE37F990" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[352,496,1580,1601]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCE5C19FE92F9F7FE37F990" box="[352,496,1580,1601]" italics="true" pageId="24" pageNumber="339">P. ujucami</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C504D4FFFCE5C19FF63F990FEFCF98E" box="[145,315,1610,1632]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="aquilonaris">
<emphasis id="B924EADEFFCE5C19FF63F990FEFCF98E" box="[145,315,1610,1632]" italics="true" pageId="24" pageNumber="339">P. aquilonaris</emphasis>
</taxonomicName>
by having a moderately long snout, usually subacuminate in dorsal view and rounded in profile. However, 
<taxonomicName id="4C504D4FFFCE5C19FE78F953FDF3F94C" box="[394,564,1672,1693]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="aquilonaris">
<emphasis id="B924EADEFFCE5C19FE78F953FDF3F94C" box="[394,564,1672,1693]" italics="true" pageId="24" pageNumber="339">P. aquilonaris</emphasis>
</taxonomicName>
has an angular 
<emphasis id="B924EADEFFCE5C19FF63F97DFE97F96A" box="[145,336,1702,1723]" italics="true" pageId="24" pageNumber="339">canthus rostralis</emphasis>
in lateral view, which is rounded and more distinct in 
<taxonomicName id="4C504D4FFFCE5C19FEA9F91DFE1EF90A" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[347,473,1734,1755]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCE5C19FEA9F91DFE1EF90A" box="[347,473,1734,1755]" italics="true" pageId="24" pageNumber="339">P. ujucami</emphasis>
</taxonomicName>
(
<figureCitation id="136B2A49FFCE5C19FE1EF91EFDFAF90B" box="[492,573,1733,1755]" captionStart="Figure 17" captionStartId="25.[163,243,1483,1505]" captionTargetBox="[166,1437,198,1435]" captionTargetId="figure-122@25.[163,1446,195,1443]" captionTargetPageId="25" captionText="Figure 17. Colour variation in life individuals of Pristimantis ujucami in dorsolateral view. A, QCAZ 49031 (subadult female, SVL 16.19 mm); B, QCAZ 57002 (adult male, SVL 14.39 mm); C, QCAZ 57010 (subadult female, 18.95 mm); D, QCAZ 57003 (adult male, SVL 15.47 mm); E, QCAZ 57004 (subadult female, SVL 18.91 mm); F, QCAZ 57646 (adult male, SVL 14.2 mm); G, QCAZ 60212 (adult female, SVL 22.33 mm); H, QCAZ 58865 (adult female, SVL 20.62 mm); I, QCAZ 60214 (adult female, SVL 23.76 mm); J, QCAZ 60194 (adult female, SVL 22.43 mm); K, QCAZ 60196 (adult female, SVL 20.45 mm); L, QCAZ 58874 (subadult female, SVL 16.74 mm)." figureDoi="http://doi.org/10.5281/zenodo.6530609" httpUri="https://zenodo.org/record/6530609/files/figure.png" pageId="24" pageNumber="339">Fig. 17</figureCitation>
). They are also similar because both show red groins and a golden iris. Nevertheless, in 
<taxonomicName id="4C504D4FFFCE5C19FEAEF8D8FDC5F8C6" box="[348,514,1794,1816]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="aquilonaris">
<emphasis id="B924EADEFFCE5C19FEAEF8D8FDC5F8C6" box="[348,514,1794,1816]" italics="true" pageId="24" pageNumber="339">P. aquilonaris</emphasis>
</taxonomicName>
, the reddish colours are usually arranged in rounded blotches, which are absent in 
<taxonomicName id="4C504D4FFFCE5C19FEF4F89BFE47F884" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[262,384,1856,1877]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCE5C19FEF4F89BFE47F884" box="[262,384,1856,1877]" italics="true" pageId="24" pageNumber="339">P. ujucami</emphasis>
</taxonomicName>
. Additionally, the iris is usually more brilliant in 
<taxonomicName id="4C504D4FFFCE5C19FEA6F884FE0CF8A5" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[340,459,1887,1908]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCE5C19FEA6F884FE0CF8A5" box="[340,459,1887,1908]" italics="true" pageId="24" pageNumber="339">P. ujucami</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6547FFCE5C18FCB3FA0BFEDBF905" lastPageId="25" lastPageNumber="340" pageId="24" pageNumber="339" type="reference_group">
<paragraph id="8BEF36CCFFCE5C18FCB3FA0BFEDBF905" blockId="24.[809,1426,1488,1908]" lastBlockId="25.[163,779,1696,1748]" lastPageId="25" lastPageNumber="340" pageId="24" pageNumber="339">
<taxonomicName id="4C504D4FFFCE5C19FCB3FA0BFB82FA34" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[833,1093,1488,1509]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCE5C19FCB3FA0BFB82FA34" box="[833,1093,1488,1509]" italics="true" pageId="24" pageNumber="339">Pristimantis ujucami</emphasis>
</taxonomicName>
is similar to a few species outside the 
<taxonomicName id="4C504D4FFFCE5C19FC5BFA34FBB6F9D2" ID-CoL="4MJQL" baseAuthorityName="Boulenger" baseAuthorityYear="1918" box="[937,1137,1518,1540]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="trachyblepharis">
<emphasis id="B924EADEFFCE5C19FC5BFA34FBB6F9D2" box="[937,1137,1518,1540]" italics="true" pageId="24" pageNumber="339">P. trachyblepharis</emphasis>
</taxonomicName>
group, such as 
<taxonomicName id="4C504D4FFFCE5C19FAEBFA34FB41F9F3" ID-CoL="77WHD" authority="(Guayasamin &amp; Funk, 2009)" baseAuthorityName="Guayasamin &amp; Funk" baseAuthorityYear="2009" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="bicantus">
<emphasis id="B924EADEFFCE5C19FAEBFA34FA56F9D2" box="[1305,1425,1518,1540]" italics="true" pageId="24" pageNumber="339">P. bicantus</emphasis>
(
<bibRefCitation id="EFC14B3DFFCE5C19FCC0F9D6FBBCF9F3" author="Guayasamin JM &amp; Funk WC" box="[818,1147,1549,1571]" pageId="24" pageNumber="339" pagination="41 - 66" refId="ref23372" refString="Guayasamin JM, Funk WC. 2009. The amphibian community at Yanayacu Biological Station, Ecuador, with a comparison of vertical microhabitat use among Pristimantis species and the description of a new species of the Pristimantis myersi group. Zootaxa 2220: 41 - 66." type="journal article" year="2009">Guayasamin &amp; Funk, 2009</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C504D4FFFCE5C19FB67F9D5FC2CF990" ID-CoL="4MJLT" authority="(Lynch &amp; Duellman, 1980)" baseAuthorityName="Lynch &amp; Duellman" baseAuthorityYear="1980" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="prolatus">
<emphasis id="B924EADEFFCE5C19FB67F9D5FAD0F9F3" box="[1173,1303,1549,1571]" italics="true" pageId="24" pageNumber="339">P. prolatus</emphasis>
(
<bibRefCitation id="EFC14B3DFFCE5C19FAD8F9D6FC24F990" author="Lynch JD &amp; Duellman WE" pageId="24" pageNumber="339" pagination="1 - 86" refId="ref24102" refString="Lynch JD, Duellman WE. 1980. The Eleutherodactylus of the Amazonian slopes of the Ecuadorian Andes (Anura: Leptodactylidae). Miscellaneous Publications - University of Kansas, Museum of Natural History 69: 1 - 86." type="journal article" year="1980">Lynch &amp; Duellman, 1980</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C504D4FFFCE5C19FBDBF9F7FB15F991" authorityName="Valencia, Valladares-Suntasig, Tipantiza-Tuguminago &amp; Duenas" authorityYear="2019" box="[1065,1234,1579,1601]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="nelsongalloi">
<emphasis id="B924EADEFFCE5C19FBDBF9F7FB15F991" box="[1065,1234,1579,1601]" italics="true" pageId="24" pageNumber="339">P. nelsongalloi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC14B3DFFCE5C19FB11F9F0FCA6F98E" author="Valencia JH &amp; Valladares-Suntasig F &amp; Tuguminago L &amp; Duenas MR" pageId="24" pageNumber="339" pagination="509 - 525" refId="ref25245" refString="Valencia JH, Valladares-Suntasig F, Tipantiza- Tuguminago L, Duenas MR. 2019. A new species of terrestrial-breeding frog of the genus Pristimantis (Anura: Terrarana: Craugastoridae) from the eastern Andean slopes of the southern Ecuador. Zootaxa 4658: 509 - 525." type="journal article" year="2019">
<collectingRegion id="4994F82EFFCE5C19FB11F9F0FA80F990" box="[1251,1351,1579,1601]" country="Spain" name="Comunitat Valenciana" pageId="24" pageNumber="339">Valencia</collectingRegion>
<emphasis id="B924EADEFFCE5C19FAA2F9F7FA4CF991" box="[1360,1419,1579,1601]" italics="true" pageId="24" pageNumber="339">et al.</emphasis>
, 2019
</bibRefCitation>
), as they also have a distinct 
<emphasis id="B924EADEFFCE5C19FB3AF991FA56F98E" box="[1224,1425,1610,1631]" italics="true" pageId="24" pageNumber="339">canthus rostralis</emphasis>
and some individuals have similar snouts in profile. In addition, 
<taxonomicName id="4C504D4FFFCE5C19FC64F953FB4BF94C" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[918,1164,1672,1693]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCE5C19FC64F953FB4BF94C" box="[918,1164,1672,1693]" italics="true" pageId="24" pageNumber="339">Pristimantis ujucami</emphasis>
</taxonomicName>
is like 
<taxonomicName id="4C504D4FFFCE5C19FB13F953FA99F94D" ID-CoL="77WHD" baseAuthorityName="Guayasamin &amp; Funk" baseAuthorityYear="2009" box="[1249,1374,1671,1693]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="bicantus">
<emphasis id="B924EADEFFCE5C19FB13F953FA99F94D" box="[1249,1374,1671,1693]" italics="true" pageId="24" pageNumber="339">P. bicantus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C504D4FFFCE5C19FCDBF97CFC02F96A" authorityName="Valencia, Valladares-Suntasig, Tipantiza-Tuguminago &amp; Duenas" authorityYear="2019" box="[809,965,1702,1724]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="nelsongalloi">
<emphasis id="B924EADEFFCE5C19FCDBF97CFC02F96A" box="[809,965,1702,1724]" italics="true" pageId="24" pageNumber="339">P.nelsongalloi</emphasis>
</taxonomicName>
in having red groins, and like 
<taxonomicName id="4C504D4FFFCE5C19FAF6F97CFAB0F96A" ID-CoL="4MJLT" baseAuthorityName="Lynch &amp; Duellman" baseAuthorityYear="1980" box="[1284,1399,1702,1724]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="prolatus">
<emphasis id="B924EADEFFCE5C19FAF6F97CFAB0F96A" box="[1284,1399,1702,1724]" italics="true" pageId="24" pageNumber="339">P.prolatus</emphasis>
</taxonomicName>
in having a golden iris. However, 
<taxonomicName id="4C504D4FFFCE5C19FB8AF91DFB28F90B" ID-CoL="77WHD" baseAuthorityName="Guayasamin &amp; Funk" baseAuthorityYear="2009" box="[1144,1263,1733,1755]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="bicantus">
<emphasis id="B924EADEFFCE5C19FB8AF91DFB28F90B" box="[1144,1263,1733,1755]" italics="true" pageId="24" pageNumber="339">P. bicantus</emphasis>
</taxonomicName>
has a rounded snout in dorsal view (subacuminate in 
<taxonomicName id="4C504D4FFFCE5C19FB26F93FFA8FF928" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[1236,1352,1764,1785]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCE5C19FB26F93FFA8FF928" box="[1236,1352,1764,1785]" italics="true" pageId="24" pageNumber="339">P. ujucami</emphasis>
</taxonomicName>
), have vocal slits and lacks ulnar, outer tarsal and heel tubercles (unlike 
<taxonomicName id="4C504D4FFFCE5C19FC8FF8FAFC37F8E7" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[893,1008,1825,1846]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCE5C19FC8FF8FAFC37F8E7" box="[893,1008,1825,1846]" italics="true" pageId="24" pageNumber="339">P. ujucami</emphasis>
</taxonomicName>
). 
<taxonomicName id="4C504D4FFFCE5C19FBF6F8FAFB35F8E7" ID-CoL="4MJLT" baseAuthorityName="Lynch &amp; Duellman" baseAuthorityYear="1980" box="[1028,1266,1825,1846]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="prolatus">
<emphasis id="B924EADEFFCE5C19FBF6F8FAFB35F8E7" box="[1028,1266,1825,1846]" italics="true" pageId="24" pageNumber="339">Pristimantis prolatus</emphasis>
</taxonomicName>
has vocal slits too and does not show red groins and hindlimbs. Finally, 
<taxonomicName id="4C504D4FFFCE5C19FCDBF884FC0FF8A2" authorityName="Valencia, Valladares-Suntasig, Tipantiza-Tuguminago &amp; Duenas" authorityYear="2019" box="[809,968,1886,1908]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="nelsongalloi">
<emphasis id="B924EADEFFCE5C19FCDBF884FC0FF8A2" box="[809,968,1886,1908]" italics="true" pageId="24" pageNumber="339">P. nelsongalloi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C504D4FFFCE5C19FBA5F884FB0BF8A5" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[1111,1228,1887,1908]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="24" pageNumber="339" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCE5C19FBA5F884FB0BF8A5" box="[1111,1228,1887,1908]" italics="true" pageId="24" pageNumber="339">P. ujucami</emphasis>
</taxonomicName>
by having clearly distinguishable dorsolateral folds and by lacking inner tarsal fold.
</paragraph>
</subSubSection>
<caption id="DF2F6644FFCF5C18FF51FA10FD68F9A5" ID-DOI="http://doi.org/10.5281/zenodo.6530609" ID-Zenodo-Dep="6530609" httpUri="https://zenodo.org/record/6530609/files/figure.png" pageId="25" pageNumber="340" startId="25.[163,243,1483,1505]" targetBox="[166,1437,198,1435]" targetPageId="25">
<paragraph id="8BEF36CCFFCF5C18FF51FA10FD68F9A5" blockId="25.[163,1443,1483,1652]" pageId="25" pageNumber="340">
<emphasis id="B924EADEFFCF5C18FF51FA10FEDBFA31" bold="true" box="[163,284,1483,1505]" pageId="25" pageNumber="340">Figure 17.</emphasis>
Colour variation in life individuals of 
<taxonomicName id="4C504D4FFFCF5C18FD33FA10FC60FA30" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[705,935,1483,1505]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="25" pageNumber="340" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCF5C18FD33FA10FC60FA30" box="[705,935,1483,1505]" italics="true" pageId="25" pageNumber="340">Pristimantis ujucami</emphasis>
</taxonomicName>
in dorsolateral view. A, QCAZ 49031 (subadult female, SVL 16.19 mm); B, QCAZ 57002 (adult male, SVL 14.39 mm); C, QCAZ 57010 (subadult female, 18.95 mm); D, QCAZ 57003 (adult male, SVL 15.47 mm); E, QCAZ 57004 (subadult female, SVL 18.91 mm); F, QCAZ 57646 (adult male, SVL 14.2 mm); G, QCAZ 60212 (adult female, SVL 22.33 mm); H, QCAZ 58865 (adult female, SVL 20.62 mm); I, QCAZ 60214 (adult female, SVL 23.76 mm); J, QCAZ 60194 (adult female, SVL 22.43 mm); K, QCAZ 60196 (adult female, SVL 20.45 mm); L, QCAZ 58874 (subadult female, SVL 16.74 mm).
</paragraph>
</caption>
<subSubSection id="C34A6547FFCF5C1CFF51F925FD3EF9B3" lastPageId="29" lastPageNumber="344" pageId="25" pageNumber="340" type="description">
<paragraph id="8BEF36CCFFCF5C18FF51F925FAC8F8C3" blockId="25.[163,779,1789,1903]" lastBlockId="25.[827,1443,1696,1810]" pageId="25" pageNumber="340">
<emphasis id="B924EADEFFCF5C18FF51F925FE31F8C3" box="[163,502,1789,1811]" italics="true" pageId="25" pageNumber="340">
Description of the 
<typeStatus id="54EB886EFFCF5C18FE7BF926FE37F8C3" box="[393,496,1789,1810]" pageId="25" pageNumber="340" type="holotype">holotype</typeStatus>
:
</emphasis>
Adult female (QCAZ 49030). Measurements (in mm): SVL 21.10; tibia length 11.37; foot length 9.54; head length 6.55; head width 7.74; interorbital distance 2.13; width of upper eyelid 2.13; internarial distance 1.93; eye–nostril distance 2.17; eye diameter 2.72; tympanum diameter 1.05. Colour of the 
<typeStatus id="54EB886EFFCF5C18FBE7F906FBB1F922" box="[1045,1142,1757,1779]" pageId="25" pageNumber="340" type="holotype">holotype</typeStatus>
in life and preservative is shown in 
<figureCitation id="136B2A49FFCF5C18FC5AF927FBE6F8C3" box="[936,1057,1788,1810]" captionStart="Figure 15" captionStartId="23.[163,243,938,960]" captionTargetBox="[331,1277,200,891]" captionTargetId="figure-495@23.[323,1283,195,898]" captionTargetPageId="23" captionText="Figure 15. Photographs in live of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm). A, dorsolateral view; B, ventral view; C, dorsolateral and inguinal region view." figureDoi="http://doi.org/10.5281/zenodo.6530605" httpUri="https://zenodo.org/record/6530605/files/figure.png" pageId="25" pageNumber="340">Figures 15</figureCitation>
and 
<figureCitation id="136B2A49FFCF5C18FBA8F927FBB2F8C3" box="[1114,1141,1788,1810]" captionStart="Figure 16" captionStartId="24.[144,222,1392,1414]" captionTargetBox="[305,1265,195,1352]" captionTargetId="figure-304@24.[305,1265,195,1352]" captionTargetPageId="24" captionText="Figure 16. Photographs of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm) in preservative. A, dorsal view; B, ventral view; C, ventral view of the left hand; D, ventral view of the right foot." figureDoi="http://doi.org/10.5281/zenodo.6530607" httpUri="https://zenodo.org/record/6530607/files/figure.png" pageId="25" pageNumber="340">16</figureCitation>
, respectively.
</paragraph>
<paragraph id="8BEF36CCFFCF5C1DFCC9F8E1FD01FDBB" blockId="25.[827,1443,1850,1903]" lastBlockId="28.[145,762,197,618]" lastPageId="28" lastPageNumber="343" pageId="25" pageNumber="340">
<emphasis id="B924EADEFFCF5C18FCC9F8E1FB96F881" box="[827,1105,1850,1872]" italics="true" pageId="25" pageNumber="340">
Head (
<figureCitation id="136B2A49FFCF5C18FC79F8E0FC25F881" box="[907,994,1851,1872]" captionStart="Figure 15" captionStartId="23.[163,243,938,960]" captionTargetBox="[331,1277,200,891]" captionTargetId="figure-495@23.[323,1283,195,898]" captionTargetPageId="23" captionText="Figure 15. Photographs in live of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm). A, dorsolateral view; B, ventral view; C, dorsolateral and inguinal region view." figureDoi="http://doi.org/10.5281/zenodo.6530605" httpUri="https://zenodo.org/record/6530605/files/figure.png" pageId="25" pageNumber="340">Figs 15</figureCitation>
, 
<figureCitation id="136B2A49FFCF5C18FC00F8E1FB86F881" box="[1010,1089,1850,1872]" captionStart="Figure 16" captionStartId="24.[144,222,1392,1414]" captionTargetBox="[305,1265,195,1352]" captionTargetId="figure-304@24.[305,1265,195,1352]" captionTargetPageId="24" captionText="Figure 16. Photographs of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm) in preservative. A, dorsal view; B, ventral view; C, ventral view of the left hand; D, ventral view of the right foot." figureDoi="http://doi.org/10.5281/zenodo.6530607" httpUri="https://zenodo.org/record/6530607/files/figure.png" pageId="25" pageNumber="340">16A, B</figureCitation>
):
</emphasis>
Wider than long, less wide than body; snout with a small rostral papilla at the tip, subacuminate in dorsal view, rounded in profile; 
<emphasis id="B924EADEFFCA5C1DFD50FF1EFF34FF28" italics="true" pageId="28" pageNumber="343">canthus rostralis</emphasis>
distinct, concave in dorsal view, angular in lateral view; loreal region concave; tympanic annulus distinct and externally visible through skin; tympanic membrane present but hidden by skin; upper margin of tympanic annulus covered by low supratympanic fold; two subconical postrictal tubercles on each side of head; four subconical tubercles and several pustules and spicules on upper eyelid; skin on throat weakly areolate; dentigerous processes of vomer rounded, slightly oblique, each vomer bearing four or five teeth, widely separated and posterior to choanae; choanae not concealed by palatal shelf of maxilla; tongue longer than wide, not notched posteriorly, posterior two thirds not adherent to the mouth floor.
</paragraph>
<caption id="DF2F6644FFCC5C1BFF63F950FDFBF8E5" ID-DOI="http://doi.org/10.5281/zenodo.6530611" ID-Zenodo-Dep="6530611" httpUri="https://zenodo.org/record/6530611/files/figure.png" pageId="26" pageNumber="341" startId="26.[145,224,1675,1697]" targetBox="[180,1391,203,1632]" targetPageId="26">
<paragraph id="8BEF36CCFFCC5C1BFF63F950FDFBF8E5" blockId="26.[145,1425,1675,1844]" pageId="26" pageNumber="341">
<emphasis id="B924EADEFFCC5C1BFF63F950FEC0F971" bold="true" box="[145,263,1675,1697]" pageId="26" pageNumber="341">Figure 18.</emphasis>
Colour variation in life individuals of 
<taxonomicName id="4C504D4FFFCC5C1BFD53F950FC46F970" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[673,897,1675,1697]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="26" pageNumber="341" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCC5C1BFD53F950FC46F970" box="[673,897,1675,1697]" italics="true" pageId="26" pageNumber="341">Pristimantis ujucami</emphasis>
</taxonomicName>
in ventral view. A, QCAZ 49031 (subadult female, SVL 16.19 mm); B, QCAZ 57002 (adult male, SVL 14.39 mm); C, QCAZ 57010 (subadult female, 18.95 mm); D, QCAZ 57003 (adult male, SVL 15.47 mm); E, QCAZ 57004 (subadult female, SVL 18.91 mm); F, QCAZ 57646 (adult male, SVL 14.2 mm); G, QCAZ 60212 (adult female, SVL 22.33 mm); H, QCAZ 58865 (adult female, SVL 20.62 mm); I, QCAZ 60214 (adult female, SVL 23.76 mm); J, QCAZ 60194 (adult female, SVL 22.43 mm); K, QCAZ 60196 (adult female, SVL 20.45 mm); L, QCAZ 58874 (subadult female, SVL 16.74 mm).
</paragraph>
</caption>
<caption id="DF2F6644FFCD5C1AFF51F95BFB4CF8DD" ID-DOI="http://doi.org/10.5281/zenodo.6530613" ID-Zenodo-Dep="6530613" httpUri="https://zenodo.org/record/6530613/files/figure.png" pageId="27" pageNumber="342" startId="27.[163,242,1664,1686]" targetBox="[173,1431,195,1635]" targetPageId="27">
<paragraph id="8BEF36CCFFCD5C1AFF51F95BFB4CF8DD" blockId="27.[163,1443,1664,1804]" pageId="27" pageNumber="342">
<emphasis id="B924EADEFFCD5C1AFF51F95BFEDEF944" bold="true" box="[163,281,1664,1686]" pageId="27" pageNumber="342">Figure 19.</emphasis>
Colour variation in preserved individuals of 
<taxonomicName id="4C504D4FFFCD5C1AFD09F95BFC1AF947" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[763,989,1664,1686]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="27" pageNumber="342" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCD5C1AFD09F95BFC1AF947" box="[763,989,1664,1686]" italics="true" pageId="27" pageNumber="342">Pristimantis ujucami</emphasis>
</taxonomicName>
: 1, dorsal views; 2, ventral views. A, QCAZ 49026 (adult female, SVL 22.62 mm); B, QCAZ 60194 (adult female, SVL 22.43 mm); C, QCAZ 60212 (adult female, SVL 22.33 mm); D, QCAZ 60214 (adult female, SVL 23.76 mm); E, QCAZ 58874 (subadult female, SVL 16.74 mm); F, QCAZ 60206 (subadult female, SVL 16.61 mm); G, QCAZ 57010 (subadult female, SVL 18.95 mm); H, QCAZ 57003 (adult male, SVL 15.47 mm); I, QCAZ 60621 (adult male, SVL 16.42 mm); J, QCAZ 57002 (adult male, 14.39 mm).
</paragraph>
</caption>
<paragraph id="8BEF36CCFFCA5C1DFF63FD48FD5CFC0A" blockId="28.[145,761,658,987]" pageId="28" pageNumber="343">
<emphasis id="B924EADEFFCA5C1DFF63FD48FD90FD79" box="[145,599,658,680]" italics="true" pageId="28" pageNumber="343">
Dorsum and venter (
<figureCitation id="136B2A49FFCA5C1DFE7EFD48FE22FD79" box="[396,485,659,680]" captionStart="Figure 15" captionStartId="23.[163,243,938,960]" captionTargetBox="[331,1277,200,891]" captionTargetId="figure-495@23.[323,1283,195,898]" captionTargetPageId="23" captionText="Figure 15. Photographs in live of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm). A, dorsolateral view; B, ventral view; C, dorsolateral and inguinal region view." figureDoi="http://doi.org/10.5281/zenodo.6530605" httpUri="https://zenodo.org/record/6530605/files/figure.png" pageId="28" pageNumber="343">Figs 15</figureCitation>
, 
<figureCitation id="136B2A49FFCA5C1DFE07FD49FD81FD79" box="[501,582,658,680]" captionStart="Figure 16" captionStartId="24.[144,222,1392,1414]" captionTargetBox="[305,1265,195,1352]" captionTargetId="figure-304@24.[305,1265,195,1352]" captionTargetPageId="24" captionText="Figure 16. Photographs of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm) in preservative. A, dorsal view; B, ventral view; C, ventral view of the left hand; D, ventral view of the right foot." figureDoi="http://doi.org/10.5281/zenodo.6530607" httpUri="https://zenodo.org/record/6530607/files/figure.png" pageId="28" pageNumber="343">16A, B</figureCitation>
):
</emphasis>
Dorsal skin shagreen bedecked with some pustules; scapular folds distinct, \ /-shaped, with a pair of subconical scapular tubercles towards its posterior end, and another pair outside the fold and posterolateral to it; small pustules grouped in rows to form very low, almost negligible paravertebral and dorsolateral folds on dorsum, and lateral folds on flanks; one pair of subconical sacral tubercles; several subconical tubercles and pustules on dorsal surface of cloacal sheath; skin on chest weakly areolate, belly areolate; discoidal fold distinct.
</paragraph>
<paragraph id="8BEF36CCFFCA5C1DFF63FC27FDD6FAF7" blockId="28.[145,761,1020,1318]" pageId="28" pageNumber="343">
<emphasis id="B924EADEFFCA5C1DFF63FC27FE48FBC0" box="[145,399,1020,1041]" italics="true" pageId="28" pageNumber="343">
Forelimbs (
<figureCitation id="136B2A49FFCA5C1DFEE8FC27FEB9FBC0" box="[282,382,1020,1041]" captionStart="Figure 16" captionStartId="24.[144,222,1392,1414]" captionTargetBox="[305,1265,195,1352]" captionTargetId="figure-304@24.[305,1265,195,1352]" captionTargetPageId="24" captionText="Figure 16. Photographs of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm) in preservative. A, dorsal view; B, ventral view; C, ventral view of the left hand; D, ventral view of the right foot." figureDoi="http://doi.org/10.5281/zenodo.6530607" httpUri="https://zenodo.org/record/6530607/files/figure.png" pageId="28" pageNumber="343">Fig. 16C</figureCitation>
):
</emphasis>
Low ulnar tubercles; Finger I shorter than Finger II; discs expanded, rounded; all fingers having ventral pads surrounded by circumferential grooves; palmar tubercles well defined, outer palmar tubercle bifid, approximately 1.5 times the size of the ovoid thenar tubercle; subarticular tubercles prominent; hyperdistal subarticular tubercles low; distinct supernumerary tubercles at base of fingers; fingers bearing narrow lateral fringes; basal webbing slightly noticeable.
</paragraph>
<paragraph id="8BEF36CCFFCA5C1DFF63FA92FDD4F8DD" blockId="28.[145,761,1353,1804]" pageId="28" pageNumber="343">
<emphasis id="B924EADEFFCA5C1DFF63FA92FE4AFA8F" box="[145,397,1353,1375]" italics="true" pageId="28" pageNumber="343">
Hindlimbs (
<figureCitation id="136B2A49FFCA5C1DFEEEFA91FEB9FA8F" box="[284,382,1353,1375]" captionStart="Figure 16" captionStartId="24.[144,222,1392,1414]" captionTargetBox="[305,1265,195,1352]" captionTargetId="figure-304@24.[305,1265,195,1352]" captionTargetPageId="24" captionText="Figure 16. Photographs of the holotype of Pristimantis ujucami (QCAZ 49030; adult female, SVL 21.16 mm) in preservative. A, dorsal view; B, ventral view; C, ventral view of the left hand; D, ventral view of the right foot." figureDoi="http://doi.org/10.5281/zenodo.6530607" httpUri="https://zenodo.org/record/6530607/files/figure.png" pageId="28" pageNumber="343">Fig. 16D</figureCitation>
):
</emphasis>
Heel bearing several pustules and one low subconical tubercle; three subconical outer tarsal tubercles; inner tarsal fold distinct; Toe V slightly longer than Toe III; discs expanded, rounded; all toes having ventral pads surrounded by circumferential grooves; inner metatarsal tubercle enlarged, approximately five times the size of the rounded outer metatarsal tubercle; subarticular tubercles prominent, well defined; hyperdistal subarticular tubercles low; distinct supernumerary tubercles at base of toes; toes bearing broad lateral fringes; basal webbing, most distinctive between Toes IV and V; discs nearly as large as those on fingers; skin on chest and anteroventral surfaces of thighs weakly areolate, posteroventral surface of thighs coarsely areolate.
</paragraph>
<paragraph id="8BEF36CCFFCA5C1DFF63F8EDFAB8FE1E" blockId="28.[145,761,1845,1898]" lastBlockId="28.[809,1425,197,464]" pageId="28" pageNumber="343">
<emphasis id="B924EADEFFCA5C1DFF63F8EDFECBF89A" box="[145,268,1846,1867]" italics="true" pageId="28" pageNumber="343">Variation:</emphasis>
The 14 individuals of 
<taxonomicName id="4C504D4FFFCA5C1DFDC4F8EDFD7FF89A" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[566,696,1846,1867]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="28" pageNumber="343" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCA5C1DFDC4F8EDFD7FF89A" box="[566,696,1846,1867]" italics="true" pageId="28" pageNumber="343">P. ujucami</emphasis>
</taxonomicName>
with available genetic material show a mean intraspecific uncorrected genetic 
<emphasis id="B924EADEFFCA5C1DFBFCFF1DFBE3FF0A" box="[1038,1060,198,219]" italics="true" pageId="28" pageNumber="343">p-</emphasis>
distance of 1.6% in the 16S gene sequence. The highest distances are those between populations in 
<collectingRegion id="4994F82EFFCA5C1DFC20FED8FB53FEC9" box="[978,1172,258,280]" country="Ecuador" name="Morona-Santiago" pageId="28" pageNumber="343">Morona Santiago</collectingRegion>
vs. 
<collectingRegion id="4994F82EFFCA5C1DFB4DFED8FA56FEC9" box="[1215,1425,258,280]" country="Ecuador" name="Zamora-Chinchipe" pageId="28" pageNumber="343">Zamora Chinchipe</collectingRegion>
Provinces. In the type series, adult males (SVL = 
<quantity id="4CA89B29FFCA5C1DFAB7FEFAFC5EFE84" metricMagnitude="-2" metricUnit="m" metricValue="1.5265" metricValueMax="1.642" metricValueMin="1.411" pageId="28" pageNumber="343" unit="mm" value="15.265" valueMax="16.42" valueMin="14.11">14.11– 16.42 mm</quantity>
) are between 13.22 and 40.52% smaller than adult females (SVL = 
<quantity id="4CA89B29FFCA5C1DFBD7FE85FB22FEA5" box="[1061,1253,350,372]" metricMagnitude="-2" metricUnit="m" metricValue="2.1340000000000003" metricValueMax="2.3760000000000003" metricValueMin="1.8920000000000001" pageId="28" pageNumber="343" unit="mm" value="21.34" valueMax="23.76" valueMin="18.92">18.92–23.76 mm</quantity>
). Below we list character states distinct from those described in the 
<typeStatus id="54EB886EFFCA5C1DFCDBFE40FC4AFE60" box="[809,909,411,433]" pageId="28" pageNumber="343" type="holotype">holotype</typeStatus>
, followed by an example. Colour variation in life and in preservative is shown in 
<figureCitation id="136B2A49FFCA5C1DFB32FE61FAFEFE1E" box="[1216,1337,442,464]" captionStart="Figure 17" captionStartId="25.[163,243,1483,1505]" captionTargetBox="[166,1437,198,1435]" captionTargetId="figure-122@25.[163,1446,195,1443]" captionTargetPageId="25" captionText="Figure 17. Colour variation in life individuals of Pristimantis ujucami in dorsolateral view. A, QCAZ 49031 (subadult female, SVL 16.19 mm); B, QCAZ 57002 (adult male, SVL 14.39 mm); C, QCAZ 57010 (subadult female, 18.95 mm); D, QCAZ 57003 (adult male, SVL 15.47 mm); E, QCAZ 57004 (subadult female, SVL 18.91 mm); F, QCAZ 57646 (adult male, SVL 14.2 mm); G, QCAZ 60212 (adult female, SVL 22.33 mm); H, QCAZ 58865 (adult female, SVL 20.62 mm); I, QCAZ 60214 (adult female, SVL 23.76 mm); J, QCAZ 60194 (adult female, SVL 22.43 mm); K, QCAZ 60196 (adult female, SVL 20.45 mm); L, QCAZ 58874 (subadult female, SVL 16.74 mm)." figureDoi="http://doi.org/10.5281/zenodo.6530609" httpUri="https://zenodo.org/record/6530609/files/figure.png" pageId="28" pageNumber="343">Figures 17</figureCitation>
to 19.
</paragraph>
<paragraph id="8BEF36CCFFCA5C1DFCDBFE22FAEAFC0A" blockId="28.[809,1425,505,987]" pageId="28" pageNumber="343">
<emphasis id="B924EADEFFCA5C1DFCDBFE22FBCBFDDF" box="[809,1036,505,526]" italics="true" pageId="28" pageNumber="343">
Head (
<figureCitation id="136B2A49FFCA5C1DFC8BFE22FC3CFDDF" box="[889,1019,505,526]" captionStart-0="Figure 17" captionStart-1="Figure 18" captionStart-2="Figure 19" captionStartId-0="25.[163,243,1483,1505]" captionStartId-1="26.[145,224,1675,1697]" captionStartId-2="27.[163,242,1664,1686]" captionTargetBox-0="[166,1437,198,1435]" captionTargetBox-1="[180,1391,203,1632]" captionTargetBox-2="[173,1431,195,1635]" captionTargetId-0="figure-122@25.[163,1446,195,1443]" captionTargetId-1="figure-27@26.[178,1391,195,1635]" captionTargetId-2="figure-26@27.[173,1431,195,1635]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionText-0="Figure 17. Colour variation in life individuals of Pristimantis ujucami in dorsolateral view. A, QCAZ 49031 (subadult female, SVL 16.19 mm); B, QCAZ 57002 (adult male, SVL 14.39 mm); C, QCAZ 57010 (subadult female, 18.95 mm); D, QCAZ 57003 (adult male, SVL 15.47 mm); E, QCAZ 57004 (subadult female, SVL 18.91 mm); F, QCAZ 57646 (adult male, SVL 14.2 mm); G, QCAZ 60212 (adult female, SVL 22.33 mm); H, QCAZ 58865 (adult female, SVL 20.62 mm); I, QCAZ 60214 (adult female, SVL 23.76 mm); J, QCAZ 60194 (adult female, SVL 22.43 mm); K, QCAZ 60196 (adult female, SVL 20.45 mm); L, QCAZ 58874 (subadult female, SVL 16.74 mm)." captionText-1="Figure 18. Colour variation in life individuals of Pristimantis ujucami in ventral view. A, QCAZ 49031 (subadult female, SVL 16.19 mm); B, QCAZ 57002 (adult male, SVL 14.39 mm); C, QCAZ 57010 (subadult female, 18.95 mm); D, QCAZ 57003 (adult male, SVL 15.47 mm); E, QCAZ 57004 (subadult female, SVL 18.91 mm); F, QCAZ 57646 (adult male, SVL 14.2 mm); G, QCAZ 60212 (adult female, SVL 22.33 mm); H, QCAZ 58865 (adult female, SVL 20.62 mm); I, QCAZ 60214 (adult female, SVL 23.76 mm); J, QCAZ 60194 (adult female, SVL 22.43 mm); K, QCAZ 60196 (adult female, SVL 20.45 mm); L, QCAZ 58874 (subadult female, SVL 16.74 mm)." captionText-2="Figure 19. Colour variation in preserved individuals of Pristimantis ujucami: 1, dorsal views; 2, ventral views. A, QCAZ 49026 (adult female, SVL 22.62 mm); B, QCAZ 60194 (adult female, SVL 22.43 mm); C, QCAZ 60212 (adult female, SVL 22.33 mm); D, QCAZ 60214 (adult female, SVL 23.76 mm); E, QCAZ 58874 (subadult female, SVL 16.74 mm); F, QCAZ 60206 (subadult female, SVL 16.61 mm); G, QCAZ 57010 (subadult female, SVL 18.95 mm); H, QCAZ 57003 (adult male, SVL 15.47 mm); I, QCAZ 60621 (adult male, SVL 16.42 mm); J, QCAZ 57002 (adult male, 14.39 mm)." figureDoi-0="http://doi.org/10.5281/zenodo.6530609" figureDoi-1="http://doi.org/10.5281/zenodo.6530611" figureDoi-2="http://doi.org/10.5281/zenodo.6530613" httpUri-0="https://zenodo.org/record/6530609/files/figure.png" httpUri-1="https://zenodo.org/record/6530611/files/figure.png" httpUri-2="https://zenodo.org/record/6530613/files/figure.png" pageId="28" pageNumber="343">Figs 17–19</figureCitation>
):
</emphasis>
Shagreen and highly spiculate (QCAZ 58865); smooth with scattered pustules (QCAZ 60196). Head wider than body (QCAZ 60621). Snout acuminate in dorsal view (QCAZ 60212). 
<emphasis id="B924EADEFFCA5C1DFAD8FD8EFC53FD59" italics="true" pageId="28" pageNumber="343">Canthus rostralis</emphasis>
straight in dorsal view (QCAZ 60194); rounded in profile (QCAZ 60612). Postrictal tubercles on each side of head: two conical (QCAZ 60196); one conical and two subconical (QCAZ 60195); one conical and one subconical (QCAZ 60212). Tubercles on upper eyelid: two subconical (QCAZ 58874); two conical and several subconical (QCAZ 49031). Interorbital tubercle: one conical (QCAZ 58866); two subconical (QCAZ 60207). Low interocular fold (QCAZ 60195). Subconical occipital tubercle (QCAZ 56998). Dentigerous processes of vomer narrowly separated (QCAZ 60194). Throat smooth (QCAZ 60196).
</paragraph>
<paragraph id="8BEF36CCFFCA5C1DFCDBFBD7FBD4F932" blockId="28.[809,1426,1035,1763]" pageId="28" pageNumber="343">
<emphasis id="B924EADEFFCA5C1DFCDBFBD7FB64FBF0" box="[809,1187,1035,1057]" italics="true" pageId="28" pageNumber="343">
Dorsum and venter (
<figureCitation id="136B2A49FFCA5C1DFBE4FBD7FB54FBF0" box="[1046,1171,1036,1057]" captionStart-0="Figure 17" captionStart-1="Figure 18" captionStart-2="Figure 19" captionStartId-0="25.[163,243,1483,1505]" captionStartId-1="26.[145,224,1675,1697]" captionStartId-2="27.[163,242,1664,1686]" captionTargetBox-0="[166,1437,198,1435]" captionTargetBox-1="[180,1391,203,1632]" captionTargetBox-2="[173,1431,195,1635]" captionTargetId-0="figure-122@25.[163,1446,195,1443]" captionTargetId-1="figure-27@26.[178,1391,195,1635]" captionTargetId-2="figure-26@27.[173,1431,195,1635]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionText-0="Figure 17. Colour variation in life individuals of Pristimantis ujucami in dorsolateral view. A, QCAZ 49031 (subadult female, SVL 16.19 mm); B, QCAZ 57002 (adult male, SVL 14.39 mm); C, QCAZ 57010 (subadult female, 18.95 mm); D, QCAZ 57003 (adult male, SVL 15.47 mm); E, QCAZ 57004 (subadult female, SVL 18.91 mm); F, QCAZ 57646 (adult male, SVL 14.2 mm); G, QCAZ 60212 (adult female, SVL 22.33 mm); H, QCAZ 58865 (adult female, SVL 20.62 mm); I, QCAZ 60214 (adult female, SVL 23.76 mm); J, QCAZ 60194 (adult female, SVL 22.43 mm); K, QCAZ 60196 (adult female, SVL 20.45 mm); L, QCAZ 58874 (subadult female, SVL 16.74 mm)." captionText-1="Figure 18. Colour variation in life individuals of Pristimantis ujucami in ventral view. A, QCAZ 49031 (subadult female, SVL 16.19 mm); B, QCAZ 57002 (adult male, SVL 14.39 mm); C, QCAZ 57010 (subadult female, 18.95 mm); D, QCAZ 57003 (adult male, SVL 15.47 mm); E, QCAZ 57004 (subadult female, SVL 18.91 mm); F, QCAZ 57646 (adult male, SVL 14.2 mm); G, QCAZ 60212 (adult female, SVL 22.33 mm); H, QCAZ 58865 (adult female, SVL 20.62 mm); I, QCAZ 60214 (adult female, SVL 23.76 mm); J, QCAZ 60194 (adult female, SVL 22.43 mm); K, QCAZ 60196 (adult female, SVL 20.45 mm); L, QCAZ 58874 (subadult female, SVL 16.74 mm)." captionText-2="Figure 19. Colour variation in preserved individuals of Pristimantis ujucami: 1, dorsal views; 2, ventral views. A, QCAZ 49026 (adult female, SVL 22.62 mm); B, QCAZ 60194 (adult female, SVL 22.43 mm); C, QCAZ 60212 (adult female, SVL 22.33 mm); D, QCAZ 60214 (adult female, SVL 23.76 mm); E, QCAZ 58874 (subadult female, SVL 16.74 mm); F, QCAZ 60206 (subadult female, SVL 16.61 mm); G, QCAZ 57010 (subadult female, SVL 18.95 mm); H, QCAZ 57003 (adult male, SVL 15.47 mm); I, QCAZ 60621 (adult male, SVL 16.42 mm); J, QCAZ 57002 (adult male, 14.39 mm)." figureDoi-0="http://doi.org/10.5281/zenodo.6530609" figureDoi-1="http://doi.org/10.5281/zenodo.6530611" figureDoi-2="http://doi.org/10.5281/zenodo.6530613" httpUri-0="https://zenodo.org/record/6530609/files/figure.png" httpUri-1="https://zenodo.org/record/6530611/files/figure.png" httpUri-2="https://zenodo.org/record/6530613/files/figure.png" pageId="28" pageNumber="343">Figs 17–19</figureCitation>
):
</emphasis>
<collectorName id="26A5531AFFCA5C1DFB44FBD0FAFEFBF0" box="[1206,1337,1035,1057]" pageId="28" pageNumber="343">Individuals</collectorName>
of both 
<collectingRegion id="4994F82EFFCA5C1DFCDBFBF0FB24FB91" box="[809,1251,1066,1088]" isEnumeration="true" pageId="28" pageNumber="343">Morona Santiago and Zamora Chinchipe</collectingRegion>
<collectorName id="26A5531AFFCA5C1DFB1BFBF1FA93FB91" box="[1257,1364,1066,1088]" pageId="28" pageNumber="343">Provinces</collectorName>
show a similar morphology despite being at a distance of 
<quantity id="4CA89B29FFCA5C1DFA9AFB92FC97FBAC" metricMagnitude="5" metricUnit="m" metricValue="2.2" pageId="28" pageNumber="343" unit="km" value="220.0">220 km</quantity>
. A frequent polymorphism (16% of the 
<typeStatus id="54EB886EFFCA5C1DFAEBFBB3FA4AFBAC" box="[1305,1421,1128,1149]" pageId="28" pageNumber="343" type="paratype">paratypes</typeStatus>
, 
<emphasis id="B924EADEFFCA5C1DFCDBFB5CFCFAFB4D" box="[809,829,1159,1180]" italics="true" pageId="28" pageNumber="343">N</emphasis>
= 5) that appears only on individuals of 
<collectingRegion id="4994F82EFFCA5C1DFAC4FB5CFC34FB6A" country="Ecuador" name="Morona-Santiago" pageId="28" pageNumber="343">Morona Santiago Province</collectingRegion>
is the presence of a pale broad stripe that starts at the back of the head and narrows posteriorly down to the cloacal region (
<specimenCode id="DBF69EB7FFCA5C1DFB9BFB39FAC5FB29" box="[1129,1282,1250,1272]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="28" pageNumber="343" type="Museum">QCAZ 57010</specimenCode>
). 
<collectingMunicipality id="6B8BACB6FFCA5C1DFAE6FB38FA83FB29" box="[1300,1348,1251,1272]" pageId="28" pageNumber="343">Two</collectingMunicipality>
males (
<specimenCode id="DBF69EB7FFCA5C1DFCDDFADAFC78FAC6" box="[815,959,1281,1303]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="28" pageNumber="343" type="Museum">QCAZ 56998</specimenCode>
, 
<specimenCode id="DBF69EB7FFCA5C1DFC38FADAFB9EFAC6" box="[970,1113,1281,1303]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="28" pageNumber="343" type="Museum">QCAZ 57001</specimenCode>
), out of 
<specimenCount id="9D56FD45FFCA5C1DFB42FADAFAD6FAC6" box="[1200,1297,1281,1303]" pageId="28" pageNumber="343" type="adult">six adult</specimenCount>
males with photographs in life, show a small diffuse reddish blotch on groins. 
<location id="8E8F6017FFCA5C1DFC88FAE5FC7BFA85" LSID="urn:lsid:plazi:treatment:03F987DAFFC05C1CFC68FE6AFD3EF9B3:8E8F6017FFCA5C1DFC88FAE5FC7BFA85" box="[890,956,1342,1364]" municipality="Two" name="Other" pageId="28" pageNumber="343" stateProvince="Morona Santiago and Zamora Chinchipe">Other</location>
males do not show reddish colours on any part of their body. Among females with photographs in life (
<specimenCount id="9D56FD45FFCA5C1DFCABFAA7FC74FA40" box="[857,947,1403,1425]" pageId="28" pageNumber="343" type="adult">11 adult</specimenCount>
females; 
<specimenCount id="9D56FD45FFCA5C1DFBEAFAA0FB68FA40" box="[1048,1199,1403,1425]" pageId="28" pageNumber="343" type="subadult">nine subadult</specimenCount>
females), all of them show reddish colours on groins. 
<collectorName id="26A5531AFFCA5C1DFB7DFA41FB39FA61" box="[1167,1278,1434,1456]" pageId="28" pageNumber="343">Therefore</collectorName>
, there seems to be a certain sexual dimorphism in colour. 
<collectorName id="26A5531AFFCA5C1DFAC7FA62FA56FA1F" box="[1333,1425,1465,1486]" pageId="28" pageNumber="343">Dorsum</collectorName>
smooth with scattered pustules (
<specimenCode id="DBF69EB7FFCA5C1DFB7AFA0CFAD0FA3C" box="[1160,1303,1495,1517]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="28" pageNumber="343" type="Museum">QCAZ 60196</specimenCode>
); shagreen, highly spiculated with scattered conical and subconical tubercles (
<specimenCode id="DBF69EB7FFCA5C1DFC55F9CEFB82F9FA" box="[935,1093,1557,1579]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="28" pageNumber="343" type="Museum">QCAZ 60207</specimenCode>
). 
<location id="8E8F6017FFCA5C1DFBA9F9CEFB02F9FA" LSID="urn:lsid:plazi:treatment:03F987DAFFC05C1CFC68FE6AFD3EF9B3:8E8F6017FFCA5C1DFBA9F9CEFB02F9FA" box="[1115,1221,1557,1579]" name="Scapular" pageId="28" pageNumber="343" stateProvince="Morona Santiago and Zamora Chinchipe">Scapular</location>
folds:) (-shaped (
<specimenCode id="DBF69EB7FFCA5C1DFCDDF9E8FC79F998" box="[815,958,1587,1609]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="28" pageNumber="343" type="Museum">QCAZ 60194</specimenCode>
);&gt; &lt;-shaped (
<specimenCode id="DBF69EB7FFCA5C1DFBA2F9E8FB1AF998" box="[1104,1245,1587,1609]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="28" pageNumber="343" type="Museum">QCAZ 49031</specimenCode>
), low and diffuse (
<specimenCode id="DBF69EB7FFCA5C1DFCC2F989FC0FF9B9" box="[816,968,1618,1640]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="28" pageNumber="343" type="Museum">QCAZ 57646</specimenCode>
); several subconical scapular tubercles along the scapular fold (
<specimenCode id="DBF69EB7FFCA5C1DFBDCF9AAFB79F957" box="[1070,1214,1648,1670]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="28" pageNumber="343" type="Museum">QCAZ 57010</specimenCode>
). 
<location id="8E8F6017FFCA5C1DFB3CF9AAFA90F956" LSID="urn:lsid:plazi:treatment:03F987DAFFC05C1CFC68FE6AFD3EF9B3:8E8F6017FFCA5C1DFB3CF9AAFA90F956" box="[1230,1367,1649,1671]" name="Dorsolateral" pageId="28" pageNumber="343" stateProvince="Morona Santiago and Zamora Chinchipe">Dorsolateral</location>
folds formed by rows of tubercles and pustules (
<specimenCode id="DBF69EB7FFCA5C1DFB04F954FA40F974" box="[1270,1415,1679,1701]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="28" pageNumber="343" type="Museum">QCAZ 58874</specimenCode>
). 
<location id="8E8F6017FFCA5C1DFCDBF975FC43F915" LSID="urn:lsid:plazi:treatment:03F987DAFFC05C1CFC68FE6AFD3EF9B3:8E8F6017FFCA5C1DFCDBF975FC43F915" box="[809,900,1710,1732]" name="Distinct" pageId="28" pageNumber="343" stateProvince="Morona Santiago and Zamora Chinchipe">Distinct</location>
paravertebral folds (
<specimenCode id="DBF69EB7FFCA5C1DFB86F975FACDF915" box="[1140,1290,1710,1732]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="28" pageNumber="343" type="Museum">QCAZ 58868</specimenCode>
). 
<location id="8E8F6017FFCA5C1DFAEEF975FA99F915" LSID="urn:lsid:plazi:treatment:03F987DAFFC05C1CFC68FE6AFD3EF9B3:8E8F6017FFCA5C1DFAEEF975FA99F915" box="[1308,1374,1710,1732]" name="Chest" pageId="28" pageNumber="343" stateProvince="Morona Santiago and Zamora Chinchipe">Chest</location>
and belly smooth (60196).
</paragraph>
<paragraph id="8BEF36CCFFCA5C1CFCDBF8CDFCCBFF2B" blockId="28.[809,1424,1814,1897]" lastBlockId="29.[163,780,197,250]" lastPageId="29" lastPageNumber="344" pageId="28" pageNumber="343">
<emphasis id="B924EADEFFCA5C1DFCDBF8CDFBD5F8FA" box="[809,1042,1814,1836]" italics="true" pageId="28" pageNumber="343">
Forelimbs (
<figureCitation id="136B2A49FFCA5C1DFC42F8CCFBC6F8FA" box="[944,1025,1814,1836]" captionStart="Figure 19" captionStartId="27.[163,242,1664,1686]" captionTargetBox="[173,1431,195,1635]" captionTargetId="figure-26@27.[173,1431,195,1635]" captionTargetPageId="27" captionText="Figure 19. Colour variation in preserved individuals of Pristimantis ujucami: 1, dorsal views; 2, ventral views. A, QCAZ 49026 (adult female, SVL 22.62 mm); B, QCAZ 60194 (adult female, SVL 22.43 mm); C, QCAZ 60212 (adult female, SVL 22.33 mm); D, QCAZ 60214 (adult female, SVL 23.76 mm); E, QCAZ 58874 (subadult female, SVL 16.74 mm); F, QCAZ 60206 (subadult female, SVL 16.61 mm); G, QCAZ 57010 (subadult female, SVL 18.95 mm); H, QCAZ 57003 (adult male, SVL 15.47 mm); I, QCAZ 60621 (adult male, SVL 16.42 mm); J, QCAZ 57002 (adult male, 14.39 mm)." figureDoi="http://doi.org/10.5281/zenodo.6530613" httpUri="https://zenodo.org/record/6530613/files/figure.png" pageId="28" pageNumber="343">Fig. 19</figureCitation>
):
</emphasis>
In females, reddish colours on shoulders (QCAZ 58865). Fingers without lateral fringes (QCAZ 60205). Basal webbing absent (QCAZ 49026). Posteroventral surface of thighs areolate (QCAZ 60196). Discs on fingers truncate (QCAZ 57010).
</paragraph>
<paragraph id="8BEF36CCFFCB5C1CFF51FEF9FDCDFDFF" blockId="29.[163,779,290,558]" pageId="29" pageNumber="344">
<emphasis id="B924EADEFFCB5C1CFF51FEF9FE4DFEE9" box="[163,394,290,312]" italics="true" pageId="29" pageNumber="344">
Hindlimbs (
<figureCitation id="136B2A49FFCB5C1CFEDFFEF8FEBCFEE9" box="[301,379,291,312]" captionStart="Figure 19" captionStartId="27.[163,242,1664,1686]" captionTargetBox="[173,1431,195,1635]" captionTargetId="figure-26@27.[173,1431,195,1635]" captionTargetPageId="27" captionText="Figure 19. Colour variation in preserved individuals of Pristimantis ujucami: 1, dorsal views; 2, ventral views. A, QCAZ 49026 (adult female, SVL 22.62 mm); B, QCAZ 60194 (adult female, SVL 22.43 mm); C, QCAZ 60212 (adult female, SVL 22.33 mm); D, QCAZ 60214 (adult female, SVL 23.76 mm); E, QCAZ 58874 (subadult female, SVL 16.74 mm); F, QCAZ 60206 (subadult female, SVL 16.61 mm); G, QCAZ 57010 (subadult female, SVL 18.95 mm); H, QCAZ 57003 (adult male, SVL 15.47 mm); I, QCAZ 60621 (adult male, SVL 16.42 mm); J, QCAZ 57002 (adult male, 14.39 mm)." figureDoi="http://doi.org/10.5281/zenodo.6530613" httpUri="https://zenodo.org/record/6530613/files/figure.png" pageId="29" pageNumber="344">Fig. 19</figureCitation>
):
</emphasis>
In females, reddish colours may appear on inner surface of shanks, anterior, posterior and dorsal surfaces of thighs, and inner surfaces of tarsus (QCAZ 58865). One conical and one subconical tubercle on dorsal surface of shanks (QCAZ 58865). Low outer tarsal subconical tubercles (QCAZ 58870). Toes bearing narrow lateral fringes (QCAZ 60205). High inner metatarsal tubercle (QCAZ 60205). Discs on toes truncate (QCAZ 60194).
</paragraph>
<paragraph id="8BEF36CCFFCB5C1CFF51FD8DFD1FFABA" blockId="29.[163,779,598,1387]" pageId="29" pageNumber="344">
<emphasis id="B924EADEFFCB5C1CFF51FD8DFE5FFD5A" italics="true" pageId="29" pageNumber="344">
Distribution, natural history and conservation status: 
<taxonomicName id="4C504D4FFFCB5C1CFF51FDADFE5FFD5A" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[163,408,630,651]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="29" pageNumber="344" phylum="Chordata" rank="species" species="ujucami">Pristimantis ujucami</taxonomicName>
</emphasis>
is known from four localities in 
<collectingRegion id="4994F82EFFCB5C1CFF51FD4FFEA1FD7B" box="[163,358,660,682]" country="Ecuador" name="Morona-Santiago" pageId="29" pageNumber="344">Morona Santiago</collectingRegion>
and 
<collectingRegion id="4994F82EFFCB5C1CFE6DFD4FFDB6FD7B" box="[415,625,660,682]" country="Ecuador" name="Zamora-Chinchipe" pageId="29" pageNumber="344">Zamora Chinchipe</collectingRegion>
Provinces, in 
<collectingCountry id="F347765CFFCB5C1CFF51FD69FEC4FD19" box="[163,259,690,712]" name="Ecuador" pageId="29" pageNumber="344">Ecuador</collectingCountry>
(
<figureCitation id="136B2A49FFCB5C1CFEE0FD69FE94FD19" box="[274,339,690,712]" captionStart="Figure 3" captionStartId="11.[163,243,1676,1698]" captionTargetBox="[193,1411,198,1633]" captionTargetId="figure-95@11.[191,1415,195,1637]" captionTargetPageId="11" captionText="Figure 3. Distribution of Pristimantis trachyblepharis species group. Locality information for examined specimens is shown in Supporting Information, Table S2. UCS: Unconfirmed Candidate Species. Elevations are indicated in metres above sea level." figureDoi="http://doi.org/10.5281/zenodo.6530579" httpUri="https://zenodo.org/record/6530579/files/figure.png" pageId="29" pageNumber="344">Fig. 3</figureCitation>
). In 
<collectingRegion id="4994F82EFFCB5C1CFE78FD68FD96FD19" box="[394,593,690,712]" country="Ecuador" name="Morona-Santiago" pageId="29" pageNumber="344">Morona Santiago</collectingRegion>
, it was found at the Abanico Protected Forest near the Hidroabanico Hydroelectric Power Plant, and near the Volcano River in Sangay National Park. In 
<collectingRegion id="4994F82EFFCB5C1CFE1BFCD4FD7CFCF5" box="[489,699,782,804]" country="Ecuador" name="Zamora-Chinchipe" pageId="29" pageNumber="344">Zamora Chinchipe</collectingRegion>
, it was reported near the Bombuscaro River, in Podocarpus National Park, and at the Numbami Reserve. Elevation range is 
<quantity id="4CA89B29FFCB5C1CFEF1FCB0FE66FC51" box="[259,417,875,896]" metricMagnitude="3" metricUnit="m" metricValue="1.3505" metricValueMax="1.72" metricValueMin="0.9809999999999999" pageId="29" pageNumber="344" unit="m" value="1350.5" valueMax="1720.0" valueMin="981.0">981 to 1720 m</quantity>
a.s.l. Frogs were found perching from 
<quantity id="4CA89B29FFCB5C1CFF13FC52FEB1FC4E" box="[225,374,905,927]" metricMagnitude="-1" metricUnit="m" metricValue="9.5" metricValueMax="18.0" metricValueMin="1.0" pageId="29" pageNumber="344" unit="cm" value="95.0" valueMax="180.0" valueMin="10.0">10 to 180 cm</quantity>
above the ground level, mainly on leaves and ferns by both day and night. Their natural region is Eastern Montane Forest and Eastern Foothills. Occurrences were near the forest edge in areas covered by native woodland, shrubby vegetation or even pasturelands (
<figureCitation id="136B2A49FFCB5C1CFE6FFBF9FE1BFBE9" box="[413,476,1058,1080]" captionStart="Figure 4" captionStartId="12.[146,225,1344,1366]" captionTargetBox="[160,1421,205,1286]" captionTargetId="figure-243@12.[145,1429,195,1304]" captionTargetPageId="12" captionText="Figure 4. Land coverage in the Ecuadorian localities where individuals of the eight described species of the Pristimantis trachyblepharis species group were found. Land coverage shapefile was downloaded from Ministerio del Ambiente y Agua of Ecuador, at http://ide.ambiente.gob.ec/mapainteractivo. Locality information for examined specimens is shown in Supporting Information, Table S2." figureDoi="http://doi.org/10.5281/zenodo.6530581" httpUri="https://zenodo.org/record/6530581/files/figure.png" pageId="29" pageNumber="344">Fig. 4</figureCitation>
). Proximity to pastureland suggests a certain tolerance to anthropogenic habitat disturbances. Extent of Occurrence of 
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and Area of Occupancy of 
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. 
<collectingRegion id="4994F82EFFCB5C1CFDC6FBA4FCCDFB45" box="[564,778,1150,1172]" country="Ecuador" name="Zamora-Chinchipe" pageId="29" pageNumber="344">Zamora Chinchipe</collectingRegion>
populations were found within Podocarpus National Park, Bombuscaro sector, and a private protected area, Reserva Numbami.Airline distance between populations from 
<collectingRegion id="4994F82EFFCB5C1CFF2DFB21FDDCFADE" box="[223,539,1273,1295]" country="Ecuador" name="Zamora-Chinchipe" pageId="29" pageNumber="344">Zamora Chinchipe Province</collectingRegion>
and those of 
<collectingRegion id="4994F82EFFCB5C1CFD41FB21FECDFAFF" country="Ecuador" name="Morona-Santiago" pageId="29" pageNumber="344">Morona Santiago</collectingRegion>
is 
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. Therefore, because intermediate areas are largely unexplored, we assign 
<taxonomicName id="4C504D4FFFCB5C1CFD86FAECFD2AFA9D" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[628,749,1335,1356]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="29" pageNumber="344" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCB5C1CFD86FAECFD2AFA9D" box="[628,749,1335,1356]" italics="true" pageId="29" pageNumber="344">P. ujucami</emphasis>
</taxonomicName>
to the Data Deficient Red List Category (
<bibRefCitation id="EFC14B3DFFCB5C1CFDB9FA8EFD0EFABB" author="IUCN" box="[587,713,1365,1386]" pageId="29" pageNumber="344" refId="ref23602" refString="IUCN. 2021. The IUCN Red List of Threatened Species. Available from: https: // www. iucnredlist. org (accessed 18 May 2021)." type="url" year="2021">IUCN 2021</bibRefCitation>
).
</paragraph>
<paragraph id="8BEF36CCFFCB5C1CFF51FA4FFD3EF9B3" blockId="29.[163,780,1428,1634]" pageId="29" pageNumber="344">
<emphasis id="B924EADEFFCB5C1CFF51FA4FFEE6FA78" box="[163,289,1428,1449]" italics="true" pageId="29" pageNumber="344">Etymology:</emphasis>
The specific epithet 
<taxonomicName id="4C504D4FFFCB5C1CFDEFFA4FFDBCFA78" authorityName="Zumel &amp; Buckley &amp; Ron" authorityYear="2022" box="[541,635,1428,1449]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="29" pageNumber="344" phylum="Chordata" rank="species" species="ujucami">
<emphasis id="B924EADEFFCB5C1CFDEFFA4FFDBCFA78" box="[541,635,1428,1449]" italics="true" pageId="29" pageNumber="344">ujucami</emphasis>
</taxonomicName>
is a noun in the genitive case and is a patronym for Martín Ujukam, known as the last Shuar warrior of Zamora, who led the struggles to prevent the Shuar people, a native American group, from being displaced from its territory. He lived at the confluence of the Zamora and Jamboé rivers. The Jamboé valley is inhabited by this species.
</paragraph>
</subSubSection>
</treatment>
</document>