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<document id="A00856B0A7B5CB3A8712CA13D60BDE49" ID-CLB-Dataset="281520" ID-DOI="10.5252/zoosystema2023v45a23" ID-GBIF-Dataset="3f666a8b-4246-43ae-9afe-ce53163f4dc5" ID-ISSN="1638-9387" ID-Zenodo-Dep="10275978" ID-ZooBank="9115B736-A081-44B8-A5D6-DF5D45185BC3" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1701851624808" checkinUser="plazi" docAuthor="Gérard, Sylvain, Marchán, Daniel Fernández, Navarro, Alejandro Martinez, Hedde, Mickaël &amp; Decaëns, Thibaud" docDate="2023" docId="03F987A6FFF4AA12B1B5771CFD71FA6E" docLanguage="en" docName="zoosystema.2023.45.23.pdf" docOrigin="Zoosystema 45 (23)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a23.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Vignysa Bouche 1970" docType="treatment" docVersion="5" lastPageNumber="759" masterDocId="FFC0FFDEFFFFAA1EB2657348FF9AFF84" masterDocTitle="Resampling Bouché’s historical localities reveals three new species and helps identifying a new genus of earthworms (Oligochaeta, Hormogastridae and Lumbricidae) in Southeastern France" masterLastPageNumber="768" masterPageNumber="749" pageNumber="758" updateTime="1702288701709" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="046E44F7AAB2589028DE83B4A114FEF4">Resampling Bouché’s historical localities reveals three new species and helps identifying a new genus of earthworms (Oligochaeta, Hormogastridae and Lumbricidae) in Southeastern France</mods:title>
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<mods:namePart id="D712EA472DF129CC6FE31AAD8A456820">Gérard, Sylvain</mods:namePart>
<mods:affiliation id="694C30C71499685E0901D1F03C126353">Eco &amp; Sols, INRAE, IRD, CIRAD, SupAgro Montpellier, Campus SupAgro, Bâtiment 12, 2 place Viala, F- 34060 Montpellier cedex 2 (France) Université de Montpellier, CNRS, EPHE, IRD, 1919, route de Mende, F- 34293 Montpellier cedex 5 (France) sylvain. gerard @ ens-paris-saclay. fr, sylvain. gerard 57 @ gmail. com (corresponding author)</mods:affiliation>
<mods:nameIdentifier id="CAEBA21896032C1D3F82918246387465" type="email">sylvain.gerard@ens-paris-saclay.fr &amp; sylvain.gerard57@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AA62ABA51DB35FDFDBBE58CE4394E3DF">Marchán, Daniel Fernández</mods:namePart>
<mods:affiliation id="F5A9CF345AB8AA066B7A3DCD6C5FE175">CEFE, Université de Montpellier, CNRS, EPHE, IRD, 1919, route de Mende, F- 34293 Montpellier cedex 5 (France) &amp; Biodiversity, Ecology and Evolution Department, Faculty of Biology, Universidad Complutense de Madrid, Avenida de Séneca, 2 Ciudad Universitaria E- 28040 Madrid (Spain)</mods:affiliation>
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<mods:namePart id="610F9CBD7017C0E5781D0C3E55A252F2">Navarro, Alejandro Martinez</mods:namePart>
<mods:affiliation id="AC89086578872D141E3DD8456E788A8C">Biodiversity, Ecology and Evolution Department, Faculty of Biology, Universidad Complutense de Madrid, Avenida de Séneca, 2 Ciudad Universitaria E- 28040 Madrid (Spain)</mods:affiliation>
<mods:nameIdentifier id="82FBC033C3F51DCCFA358F4DD62192DC" type="email">alejma21@ucm.es</mods:nameIdentifier>
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<mods:namePart id="50DE71A3E7BE48E6EFC98485BE47DAED">Hedde, Mickaël</mods:namePart>
<mods:affiliation id="356E88FCC786E4AEAB560B516CED85D1">Eco &amp; Sols, INRAE, IRD, CIRAD, SupAgro Montpellier, SupAgro, Bâtiment 12, 2 place Viala, F- 34060 Montpellier cedex 2 (France)</mods:affiliation>
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<mods:namePart id="4269FB77ADB8881FFB5ECDB116C5F6F1">Decaëns, Thibaud</mods:namePart>
<mods:affiliation id="772240EA8826A949B2BD53BF91BCBE75">CEFE, Université de Montpellier, CNRS, EPHE, IRD, 1919, route de Mende, F- 34293 Montpellier cedex 5 (France)</mods:affiliation>
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<mods:title id="E2610C0994C67D6CC162A26DB65293AC">Zoosystema</mods:title>
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<mods:date id="30C5C41003B1F8B9B4A9798D90805C9E">2023</mods:date>
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Genus
<taxonomicName id="4C504D33FFF4AA15B678771DFA91FBEA" ID-CoL="84VPL" authority="Bouche, 1970" authorityName="Bouche" authorityYear="1970" box="[1053,1291,1108,1134]" class="Clitellata" family="Hormogastridae" genus="Vignysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="genus">
<emphasis id="B924EAA2FFF4AA15B678771DFBF7FBEA" box="[1053,1133,1109,1134]" italics="true" pageId="11" pageNumber="758">Vignysa</emphasis>
<bibRefCitation id="EFC14B41FFF4AA15B611771CFA91FBEA" author="BOUCHE M. B." box="[1140,1291,1108,1134]" firstAuthor="Bouche" journalOrPublisher="Pedobiologia" pageId="11" pageNumber="758" pagination="246 - 256" part="10" refId="ref12055" refString="BOUCHE M. B. 1970. - Remarques sur quelques Lumbricina de France et consequences de la decouverte des noveaux taxons Vignysinae (Subfam. nov) et Diporodrilidae (Fam. nov.). Pedobiologia 10: 246 - 256." title="Remarques sur quelques Lumbricina de France et consequences de la decouverte des noveaux taxons Vignysinae (Subfam. nov) et Diporodrilidae (Fam. nov.)" type="journal article" year="1970">Bouché, 1970</bibRefCitation>
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. —
<taxonomicName id="4C504D33FFF4AA15B1B277C5FAB8FB21" authority="Duges, 1828: 289" authorityName="Duges" authorityPageNumber="289" authorityYear="1828" box="[983,1314,1165,1189]" class="Clitellata" family="Lumbricidae" genus="Lumbricus" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="species" species="teres">
<emphasis id="B924EAA2FFF4AA15B1B277C5FBF2FB21" box="[983,1128,1165,1189]" italics="true" pageId="11" pageNumber="758">Lumbricus teres</emphasis>
<bibRefCitation id="EFC14B41FFF4AA15B60A77C5FAB8FB21" author="DUGES A. - L." box="[1135,1314,1165,1189]" firstAuthor="Duges" journalOrPublisher="Annales des Sciences naturelles, Paris" page="289" pageId="11" pageNumber="758" pagination="284 - 337" part="15" refId="ref12691" refString="DUGES A. - L. 1828. - Recherches sur la circulation, la respiration et la reproduction des Annelides abranches. Annales des Sciences naturelles, Paris Serie 1 15: 284 - 337." title="Recherches sur la circulation, la respiration et la reproduction des Annelides abranches" type="journal article" year="1828">Dugès, 1828: 289</bibRefCitation>
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.
</paragraph>
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<subSubSection id="C34A653BFFF4AA12B14877F5FEF6FB71" lastPageId="12" lastPageNumber="759" pageId="11" pageNumber="758" type="diagnosis">
<paragraph id="8BEF36B0FFF4AA12B14877F5FEF6FB71" blockId="11.[813,1456,1213,1290]" lastBlockId="12.[132,776,1112,1269]" lastPageId="12" lastPageNumber="759" pageId="11" pageNumber="758">
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. — Small-to-medium-sized species without cutaneous pigmentation. Prostomium prolobous. Transversal furrows on segments I-II. Setae closely paired. Male pores in 15/16 with well-developed porophores. Spermathecal pores in 9/10, 10/11, simple. Nephridial pores aligned. Clitellum between XIV and XXIX (1/2 XXX). Two gizzards in VI-VII. Typhlosole lyre- or Y-shaped. Two pairs of spermathecae, disc-shaped or tubular. Two pairs of seminal vesicles in XI-XII. Nephridial bladders fish-hook shaped with caeca from the first segments.
</paragraph>
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<paragraph id="8BEF36B0FFF4AA15B2E47601FB74FA0C" blockId="11.[129,1455,1352,1416]" pageId="11" pageNumber="758">
<smallCapsWord id="8D09A06CFFF4AA15B2E47601FF35FADE" baselines="1366,1367" box="[129,175,1353,1371]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Table" pageId="11" pageNumber="758">TABLE</smallCapsWord>
4. — Main morphological characters distinguishing the newly described species
<taxonomicName id="4C504D33FFF4AA15B13C7601FAB6FADE" authority="Gerard, Decaens &amp; Marchan" authorityName="Gerard, Decaens &amp; Marchan" authorityYear="2023" box="[857,1324,1352,1371]" class="Clitellata" family="Lumbricidae" genus="Flabellodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="species" species="luberonensis" status="n. gen., n. sp.">
<emphasis id="B924EAA2FFF4AA15B13C7601FBACFADF" box="[857,1078,1353,1371]" italics="true" pageId="11" pageNumber="758">Flabellodrilus luberonensis</emphasis>
Gérard, Decaëns &amp; Marchán
</taxonomicName>
,
<taxonomicNameLabel id="A21757D9FFF4AA15B7517601FA36FADF" box="[1332,1452,1353,1371]" pageId="11" pageNumber="758" rank="species">n. gen., n. sp.</taxonomicNameLabel>
,
<taxonomicName id="4C504D33FFF4AA15B2E47617FDC8FAF5" authority="Gerard, Decaens &amp; Marchan" authorityName="Gerard, Decaens &amp; Marchan" authorityYear="2023" box="[129,594,1375,1393]" class="Clitellata" family="Lumbricidae" genus="Allolobophora" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="species" species="delitescens" status="sp. nov.">
<emphasis id="B924EAA2FFF4AA15B2E47617FEC0FAF5" box="[129,346,1375,1393]" italics="true" pageId="11" pageNumber="758">Allolobophora delitescens</emphasis>
Gérard, Decaëns &amp; Marchán
</taxonomicName>
,
<taxonomicNameLabel id="A21757D9FFF4AA15B03F7617FD11FAF5" box="[602,651,1375,1393]" pageId="11" pageNumber="758" rank="species">n. sp.</taxonomicNameLabel>
and their closest relatives
<taxonomicName id="4C504D33FFF4AA15B1087617FB46FAF5" authority="(Bouche, 1970)" authorityName="Gérard &amp; Marchán &amp; Navarro &amp; Hedde &amp; Decaëns" authorityYear="2023" baseAuthorityName="Bouche" baseAuthorityYear="1970" box="[877,1244,1375,1393]" class="Clitellata" family="Lumbricidae" genus="Flabellodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="subSpecies" species="bartolii" status="n. comb." subSpecies="bartolii">
<emphasis id="B924EAA2FFF4AA15B1087617FBCDFAF5" box="[877,1111,1375,1393]" italics="true" pageId="11" pageNumber="758">Flabellodrilus bartolii bartolii</emphasis>
(
<bibRefCitation id="EFC14B41FFF4AA15B6077617FB42FAF5" author="BOUCHE M. B." box="[1122,1240,1375,1393]" firstAuthor="Bouche" journalOrPublisher="Pedobiologia" pageId="11" pageNumber="758" pagination="246 - 256" part="10" refId="ref12055" refString="BOUCHE M. B. 1970. - Remarques sur quelques Lumbricina de France et consequences de la decouverte des noveaux taxons Vignysinae (Subfam. nov) et Diporodrilidae (Fam. nov.). Pedobiologia 10: 246 - 256." title="Remarques sur quelques Lumbricina de France et consequences de la decouverte des noveaux taxons Vignysinae (Subfam. nov) et Diporodrilidae (Fam. nov.)" type="journal article" year="1970">Bouché, 1970</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A21757D9FFF4AA15B6867617FAA8FAF5" box="[1251,1330,1375,1393]" pageId="11" pageNumber="758" rank="species">n. comb.</taxonomicNameLabel>
,
<taxonomicName id="4C504D33FFF4AA15B75C7617FEDEFA0C" authority="(Savigny, 1826)" baseAuthorityName="Savigny" baseAuthorityYear="1826" class="Clitellata" family="Lumbricidae" genus="Allolobophora" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="species" species="icterica">
<emphasis id="B924EAA2FFF4AA15B75C7617FF27FA0C" italics="true" pageId="11" pageNumber="758">Allolobophora icterica</emphasis>
(
<bibRefCitation id="EFC14B41FFF4AA15B2A2763EFEA5FA0C" author="SAVIGNY J. - C. &amp; CUVIER G." box="[199,319,1398,1416]" firstAuthor="Savigny" journalOrPublisher="Memoires de l'Academie (royale) des sciences de l'Institut (imperial) de France" pageId="11" pageNumber="758" pagination="176 - 184" refId="ref15576" refString="SAVIGNY J. - C. &amp; CUVIER G. 1826. - Analyse des travaux de l'Academie royale des Sciences pendant l'annee 1821, partie physique [Zoology from p. 172 ̿. Memoires de l'Academie (royale) des sciences de l'Institut (imperial) de France: 176 - 184." title="Analyse des travaux de l'Academie royale des Sciences pendant l'annee 1821, partie physique" type="journal article" year="1826">Savigny, 1826</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C504D33FFF4AA15B32E763EFDEDFA0C" authority="(Orley, 1881)" baseAuthorityName="Orley" baseAuthorityYear="1881" box="[331,631,1395,1416]" class="Clitellata" family="Lumbricidae" genus="Allolobophora" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="species" species="dubiosa">
<emphasis id="B924EAA2FFF4AA15B32E763EFD9DFA0C" box="[331,519,1398,1416]" italics="true" pageId="11" pageNumber="758">Allolobophora dubiosa</emphasis>
(
<bibRefCitation id="EFC14B41FFF4AA15B075763BFDEBFA0C" author="ORLEY L." box="[528,625,1395,1416]" firstAuthor="Orley" journalOrPublisher="Mathematikai es Termeszettudomanyi Kozlemenyek" pageId="11" pageNumber="758" pagination="562 - 611" part="16" refId="ref14465" refString="ORLEY L. 1881. - A magyarorszagi Oligochaetak faunaja. I. Terricolae. Mathematikai es Termeszettudomanyi Kozlemenyek 16: 562 - 611." title="A magyarorszagi Oligochaetak faunaja. I. Terricolae" type="journal article" year="1881">Örley, 1881</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C504D33FFF4AA15B01B763EFB70FA0C" authority="Trigo, Mascates, Briones and Diaz Cosin, 1990" authorityName="Trigo, Mascates, Briones and Diaz Cosin" authorityYear="1990" box="[638,1258,1398,1416]" class="Clitellata" family="Lumbricidae" genus="Allolobophora" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="species" species="monchicana">
<emphasis id="B924EAA2FFF4AA15B01B763EFCC7FA0C" box="[638,861,1398,1416]" italics="true" pageId="11" pageNumber="758">Allolobophora monchicana</emphasis>
Trigo, Mascates, Briones and Diaz Cosin, 1990
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BEF36B0FFF4AA15B34F768AFB9FF864" pageId="11" pageNumber="758">
<table id="F950C410FFF455E1B2E6768AFA29F864" box="[131,1459,1474,2016]" gridcols="7" gridrows="23" pageId="11" pageNumber="758">
<tr id="356034F2FFF455E1B2E6768AFA29FA52" box="[131,1459,1474,1494]" gridrow="0" pageId="11" pageNumber="758" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B15D8EFFF455E1B34F768AFE48FA52" box="[298,466,1474,1494]" gridcol="1" gridrow="0" pageId="11" pageNumber="758">
<taxonomicName id="4C504D33FFF4AA15B34F768AFE28FA52" authorityName="Gerard, Decaens &amp; Marchan" authorityYear="2023" box="[298,434,1474,1494]" class="Clitellata" family="Lumbricidae" genus="Flabellodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="genus">
<emphasis id="B924EAA2FFF4AA15B34F768AFE28FA52" bold="true" box="[298,434,1474,1494]" italics="true" pageId="11" pageNumber="758">Flabellodrilus</emphasis>
</taxonomicName>
</th>
<th id="76B15D8EFFF455E1B3B9768AFD19FA52" box="[476,643,1474,1494]" gridcol="2" gridrow="0" pageId="11" pageNumber="758">
<taxonomicName id="4C504D33FFF4AA15B3B9768AFDFEFA52" authorityName="Gerard, Decaens &amp; Marchan" authorityYear="2023" box="[476,612,1474,1494]" class="Clitellata" family="Lumbricidae" genus="Flabellodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="species" species="luberonensis">
<emphasis id="B924EAA2FFF4AA15B3B9768AFDFEFA52" bold="true" box="[476,612,1474,1494]" italics="true" pageId="11" pageNumber="758">Flabellodrilus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="356034F2FFF455E1B2E67693FA29FA6B" box="[131,1459,1499,1519]" gridrow="1" pageId="11" pageNumber="758" rowspan-0="1">
<td id="76B15D8EFFF455E1B34F7693FE48FA6B" box="[298,466,1499,1519]" gridcol="1" gridrow="1" pageId="11" pageNumber="758">
<emphasis id="B924EAA2FFF4AA15B35C7693FE21FA6B" bold="true" box="[313,443,1499,1519]" italics="true" pageId="11" pageNumber="758">luberonensis</emphasis>
</td>
<td id="76B15D8EFFF455E1B3B97693FD19FA6B" box="[476,643,1499,1519]" gridcol="2" gridrow="1" pageId="11" pageNumber="758">
<emphasis id="B924EAA2FFF4AA15B38E7693FDE5FA6B" bold="true" box="[491,639,1499,1519]" italics="true" pageId="11" pageNumber="758">
<taxonomicName id="4C504D33FFF4AA15B38E7693FDA8FA6B" authorityName="Gérard &amp; Marchán &amp; Navarro &amp; Hedde &amp; Decaëns" authorityYear="2023" baseAuthorityName="Bouche" baseAuthorityYear="1970" box="[491,562,1499,1519]" class="Clitellata" family="Lumbricidae" genus="Flabellodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="species" species="bartolii">bartolii</taxonomicName>
<taxonomicName id="4C504D33FFF4AA15B05D7693FDE5FA6B" authorityName="Gérard &amp; Marchán &amp; Navarro &amp; Hedde &amp; Decaëns" authorityYear="2023" baseAuthorityName="Bouche" baseAuthorityYear="1970" box="[568,639,1499,1519]" class="Clitellata" family="Lumbricidae" genus="Flabellodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="species" species="bartolii">bartolii</taxonomicName>
</emphasis>
</td>
<td id="76B15D8EFFF455E1B0F77693FCC8FA6B" box="[658,850,1499,1519]" gridcol="3" gridrow="1" pageId="11" pageNumber="758">
<taxonomicName id="4C504D33FFF4AA15B0F77693FC85FA6B" authorityName="Eisen" authorityYear="1874" box="[658,799,1499,1519]" class="Clitellata" family="Lumbricidae" genus="Allolobophora" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="genus">
<emphasis id="B924EAA2FFF4AA15B0F77693FC85FA6B" bold="true" box="[658,799,1499,1519]" italics="true" pageId="11" pageNumber="758">Allolobophora</emphasis>
</taxonomicName>
</td>
<td id="76B15D8EFFF455E1B1047693FB8EFA6B" box="[865,1044,1499,1519]" gridcol="4" gridrow="1" pageId="11" pageNumber="758">
<taxonomicName id="4C504D33FFF4AA15B1047693FC74FA6B" authorityName="Eisen" authorityYear="1874" box="[865,1006,1499,1519]" class="Clitellata" family="Lumbricidae" genus="Allolobophora" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="genus">
<emphasis id="B924EAA2FFF4AA15B1047693FC74FA6B" bold="true" box="[865,1006,1499,1519]" italics="true" pageId="11" pageNumber="758">Allolobophora</emphasis>
</taxonomicName>
</td>
<td id="76B15D8EFFF455E1B6797693FB24FA6B" box="[1052,1214,1499,1519]" gridcol="5" gridrow="1" pageId="11" pageNumber="758">
<taxonomicName id="4C504D33FFF4AA15B6797693FB33FA6B" authorityName="Eisen" authorityYear="1874" box="[1052,1193,1499,1519]" class="Clitellata" family="Lumbricidae" genus="Allolobophora" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="genus">
<emphasis id="B924EAA2FFF4AA15B6797693FB33FA6B" bold="true" box="[1052,1193,1499,1519]" italics="true" pageId="11" pageNumber="758">Allolobophora</emphasis>
</taxonomicName>
</td>
<td id="76B15D8EFFF455E1B6AE7693FA29FA6B" box="[1227,1459,1499,1519]" gridcol="6" gridrow="1" pageId="11" pageNumber="758">
<taxonomicName id="4C504D33FFF4AA15B6AE7693FAC2FA6B" authorityName="Eisen" authorityYear="1874" box="[1227,1368,1499,1519]" class="Clitellata" family="Lumbricidae" genus="Allolobophora" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="genus">
<emphasis id="B924EAA2FFF4AA15B6AE7693FAC2FA6B" bold="true" box="[1227,1368,1499,1519]" italics="true" pageId="11" pageNumber="758">Allolobophora</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="356034F2FFF455E1B2E676BCFA29F98D" box="[131,1459,1524,1545]" gridrow="2" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E676BCFEBBF98D" box="[131,289,1524,1545]" gridcol="0" gridrow="2" pageId="11" pageNumber="758">
<emphasis id="B924EAA2FFF4AA15B2E676BCFF6EF98C" bold="true" box="[131,244,1524,1544]" pageId="11" pageNumber="758">Characters</emphasis>
</th>
<td id="76B15D8EFFF455E1B34F76BCFE48F98D" box="[298,466,1524,1545]" gridcol="1" gridrow="2" pageId="11" pageNumber="758">
<emphasis id="B924EAA2FFF4AA15B35C76BDFEEBF98D" bold="true" box="[313,369,1525,1545]" pageId="11" pageNumber="758">
<taxonomicNameLabel id="A21757D9FFF4AA15B35C76BDFEEBF98D" box="[313,369,1525,1545]" pageId="11" pageNumber="758" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</td>
<td id="76B15D8EFFF455E1B3B976BCFD19F98D" box="[476,643,1524,1545]" gridcol="2" gridrow="2" pageId="11" pageNumber="758">
<emphasis id="B924EAA2FFF4AA15B38E76BDFDD9F98D" bold="true" box="[491,579,1525,1545]" pageId="11" pageNumber="758">n. comb.</emphasis>
</td>
<td id="76B15D8EFFF455E1B0F776BCFCC8F98D" box="[658,850,1524,1545]" gridcol="3" gridrow="2" pageId="11" pageNumber="758">
<emphasis id="B924EAA2FFF4AA15B0C476BDFCC8F98D" bold="true" box="[673,850,1525,1545]" italics="true" pageId="11" pageNumber="758">
<taxonomicName id="4C504D33FFF4AA15B0C476BDFC8FF98D" authority="Gérard &amp; Marchán &amp; Navarro &amp; Hedde &amp; Decaëns, 2023" authorityName="Gérard &amp; Marchán &amp; Navarro &amp; Hedde &amp; Decaëns" authorityYear="2023" box="[673,789,1525,1545]" class="Clitellata" family="Lumbricidae" genus="Allolobophora" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="species" species="delitescens" status="sp. nov.">delitescens</taxonomicName>
<taxonomicNameLabel id="A21757D9FFF4AA15B17E76BDFCC8F98D" box="[795,850,1525,1545]" pageId="11" pageNumber="758" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</td>
<td id="76B15D8EFFF455E1B10476BCFB8EF98D" box="[865,1044,1524,1545]" gridcol="4" gridrow="2" pageId="11" pageNumber="758">
<taxonomicName id="4C504D33FFF4AA15B11576BDFC21F98D" baseAuthorityName="Savigny" baseAuthorityYear="1826" box="[880,955,1525,1545]" class="Clitellata" family="Lumbricidae" genus="Allolobophora" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="species" species="icterica">
<emphasis id="B924EAA2FFF4AA15B11576BDFC21F98D" bold="true" box="[880,955,1525,1545]" italics="true" pageId="11" pageNumber="758">icterica</emphasis>
</taxonomicName>
</td>
<td id="76B15D8EFFF455E1B67976BCFB24F98D" box="[1052,1214,1524,1545]" gridcol="5" gridrow="2" pageId="11" pageNumber="758">
<taxonomicName id="4C504D33FFF4AA15B64E76BDFBE6F98D" baseAuthorityName="Orley" baseAuthorityYear="1881" box="[1067,1148,1525,1545]" class="Clitellata" family="Lumbricidae" genus="Allolobophora" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="species" species="dubiosa">
<emphasis id="B924EAA2FFF4AA15B64E76BDFBE6F98D" bold="true" box="[1067,1148,1525,1545]" italics="true" pageId="11" pageNumber="758">dubiosa</emphasis>
</taxonomicName>
</td>
<td id="76B15D8EFFF455E1B6AE76BCFA29F98D" box="[1227,1459,1524,1545]" gridcol="6" gridrow="2" pageId="11" pageNumber="758">
<taxonomicName id="4C504D33FFF4AA15B6BF76BDFACDF98D" authorityName="Trigo, Mascates, Briones and Diaz Cosin" authorityYear="1990" box="[1242,1367,1525,1545]" class="Clitellata" family="Lumbricidae" genus="Allolobophora" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="758" phylum="Annelida" rank="species" species="monchicana">
<emphasis id="B924EAA2FFF4AA15B6BF76BDFACDF98D" bold="true" box="[1242,1367,1525,1545]" italics="true" pageId="11" pageNumber="758">monchicana</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="356034F2FFF455E1B2E67550FA29F9A8" box="[131,1459,1560,1580]" gridrow="3" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E67550FEBBF9A8" box="[131,289,1560,1580]" gridcol="0" gridrow="3" pageId="11" pageNumber="758">Length</th>
<td id="76B15D8EFFF455E1B34F7550FE48F9A8" box="[298,466,1560,1580]" gridcol="1" gridrow="3" pageId="11" pageNumber="758">70-100 mm</td>
<td id="76B15D8EFFF455E1B3B97550FD19F9A8" box="[476,643,1560,1580]" gridcol="2" gridrow="3" pageId="11" pageNumber="758">60-120 mm</td>
<td id="76B15D8EFFF455E1B0F77550FCC8F9A8" box="[658,850,1560,1580]" gridcol="3" gridrow="3" pageId="11" pageNumber="758">70-98 mm</td>
<td id="76B15D8EFFF455E1B1047550FB8EF9A8" box="[865,1044,1560,1580]" gridcol="4" gridrow="3" pageId="11" pageNumber="758">70-90 mm</td>
<td id="76B15D8EFFF455E1B6797550FB24F9A8" box="[1052,1214,1560,1580]" gridcol="5" gridrow="3" pageId="11" pageNumber="758">120-250 mm</td>
<td id="76B15D8EFFF455E1B6AE7550FA29F9A8" box="[1227,1459,1560,1580]" gridcol="6" gridrow="3" pageId="11" pageNumber="758">50-117 mm</td>
</tr>
<tr id="356034F2FFF455E1B2E67578FA29F9C0" box="[131,1459,1584,1604]" gridrow="4" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E67578FEBBF9C0" box="[131,289,1584,1604]" gridcol="0" gridrow="4" pageId="11" pageNumber="758">N. segments</th>
<td id="76B15D8EFFF455E1B34F7578FE48F9C0" box="[298,466,1584,1604]" gridcol="1" gridrow="4" pageId="11" pageNumber="758">187-353</td>
<td id="76B15D8EFFF455E1B3B97578FD19F9C0" box="[476,643,1584,1604]" gridcol="2" gridrow="4" pageId="11" pageNumber="758">280-394</td>
<td id="76B15D8EFFF455E1B0F77578FCC8F9C0" box="[658,850,1584,1604]" gridcol="3" gridrow="4" pageId="11" pageNumber="758">200-211</td>
<td id="76B15D8EFFF455E1B1047578FB8EF9C0" box="[865,1044,1584,1604]" gridcol="4" gridrow="4" pageId="11" pageNumber="758">140-200</td>
<td id="76B15D8EFFF455E1B6797578FB24F9C0" box="[1052,1214,1584,1604]" gridcol="5" gridrow="4" pageId="11" pageNumber="758">120-303</td>
<td id="76B15D8EFFF455E1B6AE7578FA29F9C0" box="[1227,1459,1584,1604]" gridcol="6" gridrow="4" pageId="11" pageNumber="758">130-158</td>
</tr>
<tr id="356034F2FFF455E1B2E67500FA29F9D8" box="[131,1459,1608,1628]" gridrow="5" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E67500FEBBF9D8" box="[131,289,1608,1628]" gridcol="0" gridrow="5" pageId="11" pageNumber="758">Weight</th>
<td id="76B15D8EFFF455E1B34F7500FE48F9D8" box="[298,466,1608,1628]" gridcol="1" gridrow="5" pageId="11" pageNumber="758">0.85-1.24 g</td>
<td id="76B15D8EFFF455E1B3B97500FD19F9D8" box="[476,643,1608,1628]" gridcol="2" gridrow="5" pageId="11" pageNumber="758">0.85-1.49 g</td>
<td id="76B15D8EFFF455E1B0F77500FCC8F9D8" box="[658,850,1608,1628]" gridcol="3" gridrow="5" pageId="11" pageNumber="758">0.64-1.2 g</td>
<td id="76B15D8EFFF455E1B1047500FB8EF9D8" box="[865,1044,1608,1628]" gridcol="4" gridrow="5" pageId="11" pageNumber="758">0.3-1.33 g</td>
<td id="76B15D8EFFF455E1B6797500FB24F9D8" box="[1052,1214,1608,1628]" gridcol="5" gridrow="5" pageId="11" pageNumber="758">2.96-3.47 g</td>
<td id="76B15D8EFFF455E1B6AE7500FA29F9D8" box="[1227,1459,1608,1628]" gridcol="6" gridrow="5" pageId="11" pageNumber="758">0.74-0.96 g</td>
</tr>
<tr id="356034F2FFF455E1B2E67528FA29F9F0" box="[131,1459,1632,1652]" gridrow="6" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E67528FEBBF9F0" box="[131,289,1632,1652]" gridcol="0" gridrow="6" pageId="11" pageNumber="758">Pigmentation</th>
<td id="76B15D8EFFF455E1B34F7528FE48F9F0" box="[298,466,1632,1652]" gridcol="1" gridrow="6" pageId="11" pageNumber="758">Absent,</td>
<td id="76B15D8EFFF455E1B3B97528FD19F9F0" box="[476,643,1632,1652]" gridcol="2" gridrow="6" pageId="11" pageNumber="758">Brown-yellow,</td>
<td id="76B15D8EFFF455E1B0F77528FCC8F9F0" box="[658,850,1632,1652]" gridcol="3" gridrow="6" pageId="11" pageNumber="758">Absent</td>
<td id="76B15D8EFFF455E1B1047528FB8EF9F0" box="[865,1044,1632,1652]" gridcol="4" gridrow="6" pageId="11" pageNumber="758">Absent</td>
<td id="76B15D8EFFF455E1B6797528FB24F9F0" box="[1052,1214,1632,1652]" gridcol="5" gridrow="6" pageId="11" pageNumber="758">Dark green to</td>
<td id="76B15D8EFFF455E1B6AE7528FA29F9F0" box="[1227,1459,1632,1652]" gridcol="6" gridrow="6" pageId="11" pageNumber="758">Absent</td>
</tr>
<tr id="356034F2FFF455E1B2E6753EFA29F90E" box="[131,1459,1654,1674]" gridrow="7" pageId="11" pageNumber="758" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<td id="76B15D8EFFF455E1B34F753EFE48F90E" box="[298,466,1654,1674]" gridcol="1" gridrow="7" pageId="11" pageNumber="758">dark spots</td>
<td id="76B15D8EFFF455E1B3B9753EFD19F90E" box="[476,643,1654,1674]" gridcol="2" gridrow="7" pageId="11" pageNumber="758">dark spots</td>
<td id="76B15D8EFFF455E1B679753EFB24F90E" box="[1052,1214,1654,1674]" gridcol="5" gridrow="7" pageId="11" pageNumber="758">greenish brown</td>
</tr>
<tr id="356034F2FFF455E1B2E675C6FA29F926" box="[131,1459,1678,1698]" gridrow="8" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E675C6FEBBF926" box="[131,289,1678,1698]" gridcol="0" gridrow="8" pageId="11" pageNumber="758">Papillae</th>
<td id="76B15D8EFFF455E1B34F75C6FE48F926" box="[298,466,1678,1698]" gridcol="1" gridrow="8" pageId="11" pageNumber="758">XI-XII, XXVI-</td>
<td id="76B15D8EFFF455E1B3B975C6FD19F926" box="[476,643,1678,1698]" gridcol="2" gridrow="8" pageId="11" pageNumber="758">X, XI, XXXIII-XLIV</td>
<td id="76B15D8EFFF455E1B0F775C6FCC8F926" box="[658,850,1678,1698]" gridcol="3" gridrow="8" pageId="11" pageNumber="758">XXXVII, (XXXVIII),</td>
<td id="76B15D8EFFF455E1B10475C6FB8EF926" box="[865,1044,1678,1698]" gridcol="4" gridrow="8" pageId="11" pageNumber="758">XI, XLIV, XLV</td>
<td id="76B15D8EFFF455E1B67975C6FB24F926" box="[1052,1214,1678,1698]" gridcol="5" gridrow="8" pageId="11" pageNumber="758">Unknown</td>
<td id="76B15D8EFFF455E1B6AE75C6FA29F926" box="[1227,1459,1678,1698]" gridcol="6" gridrow="8" pageId="11" pageNumber="758">XXII, XLVII, XLVIII</td>
</tr>
<tr id="356034F2FFF455E1B2E675EBFA29F933" box="[131,1459,1699,1719]" gridrow="9" pageId="11" pageNumber="758" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="76B15D8EFFF455E1B34F75EBFE48F933" box="[298,466,1699,1719]" gridcol="1" gridrow="9" pageId="11" pageNumber="758">XXVIII, XXXIV</td>
<td id="76B15D8EFFF455E1B0F775EBFCC8F933" box="[658,850,1699,1719]" gridcol="3" gridrow="9" pageId="11" pageNumber="758">XXXIX</td>
</tr>
<tr id="356034F2FFF455E1B2E675F3FA29F94B" box="[131,1459,1723,1743]" gridrow="10" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E675F3FEBBF94B" box="[131,289,1723,1743]" gridcol="0" gridrow="10" pageId="11" pageNumber="758">Spermathecal</th>
<td id="76B15D8EFFF455E1B34F75F3FE48F94B" box="[298,466,1723,1743]" gridcol="1" gridrow="10" pageId="11" pageNumber="758">9/10, 10/11</td>
<td id="76B15D8EFFF455E1B3B975F3FD19F94B" box="[476,643,1723,1743]" gridcol="2" gridrow="10" pageId="11" pageNumber="758">9/10, 10/11</td>
<td id="76B15D8EFFF455E1B0F775F3FCC8F94B" box="[658,850,1723,1743]" gridcol="3" gridrow="10" pageId="11" pageNumber="758">8/9, 9/10, 10/11</td>
<td id="76B15D8EFFF455E1B10475F3FB8EF94B" box="[865,1044,1723,1743]" gridcol="4" gridrow="10" pageId="11" pageNumber="758">(7/8)8/9, 9/10,</td>
<td id="76B15D8EFFF455E1B67975F3FB24F94B" box="[1052,1214,1723,1743]" gridcol="5" gridrow="10" pageId="11" pageNumber="758">8/9, 9/10, 10/11</td>
<td id="76B15D8EFFF455E1B6AE75F3FA29F94B" box="[1227,1459,1723,1743]" gridcol="6" gridrow="10" pageId="11" pageNumber="758">(8/9) 9/10, 10/11</td>
</tr>
<tr id="356034F2FFF455E1B2E67599FA29F961" box="[131,1459,1745,1765]" gridrow="11" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E67599FEBBF961" box="[131,289,1745,1765]" gridcol="0" gridrow="11" pageId="11" pageNumber="758">pores</th>
<td id="76B15D8EFFF455E1B34F7599FE48F961" box="[298,466,1745,1765]" gridcol="1" gridrow="11" pageId="11" pageNumber="758">double</td>
<td id="76B15D8EFFF455E1B3B97599FD19F961" box="[476,643,1745,1765]" gridcol="2" gridrow="11" pageId="11" pageNumber="758">double</td>
<td id="76B15D8EFFF455E1B0F77599FCC8F961" box="[658,850,1745,1765]" gridcol="3" gridrow="11" pageId="11" pageNumber="758">simple in c</td>
<td id="76B15D8EFFF455E1B1047599FB8EF961" box="[865,1044,1745,1765]" gridcol="4" gridrow="11" pageId="11" pageNumber="758">10/11 simple in c</td>
<td id="76B15D8EFFF455E1B6797599FB24F961" box="[1052,1214,1745,1765]" gridcol="5" gridrow="11" pageId="11" pageNumber="758">simple in c</td>
<td id="76B15D8EFFF455E1B6AE7599FA29F961" box="[1227,1459,1745,1765]" gridcol="6" gridrow="11" pageId="11" pageNumber="758">simple in c</td>
</tr>
<tr id="356034F2FFF455E1B2E675A0FA29F979" box="[131,1459,1768,1789]" gridrow="12" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E675A0FEBBF979" box="[131,289,1768,1789]" gridcol="0" gridrow="12" pageId="11" pageNumber="758">Clitellum</th>
<td id="76B15D8EFFF455E1B34F75A0FE48F979" box="[298,466,1768,1789]" gridcol="1" gridrow="12" pageId="11" pageNumber="758">XXXII-XLII (XLIII)</td>
<td id="76B15D8EFFF455E1B3B975A0FCC8F979" box="[476,850,1768,1789]" colspan="2" colspanRight="1" gridcol="2" gridrow="12" pageId="11" pageNumber="758">(XXXIII) XXXIV-XLIII (XXXV) XXXVI–(XLVI)</td>
<td id="76B15D8EFFF455E1B10475A0FB8EF979" box="[865,1044,1768,1789]" gridcol="4" gridrow="12" pageId="11" pageNumber="758">(XXXIII) XXXIV-XLIII</td>
<td id="76B15D8EFFF455E1B67975A0FB24F979" box="[1052,1214,1768,1789]" gridcol="5" gridrow="12" pageId="11" pageNumber="758">(XXXVI) XXXVII-</td>
<td id="76B15D8EFFF455E1B6AE75A0FA29F979" box="[1227,1459,1768,1789]" gridcol="6" gridrow="12" pageId="11" pageNumber="758">(XXXVII)XXXVIII(XXXIX)-</td>
</tr>
<tr id="356034F2FFF455E1B2E675B6FA29F896" box="[131,1459,1790,1810]" gridrow="13" pageId="11" pageNumber="758" rowspan-0="1" rowspan-1="1">
<td id="76B15D8EFFF455E1B3B975B6FD19F896" box="[476,643,1790,1810]" gridcol="2" gridrow="13" pageId="11" pageNumber="758">(XLIV)</td>
<td id="76B15D8EFFF455E1B0F775B6FCC8F896" box="[658,850,1790,1810]" gridcol="3" gridrow="13" pageId="11" pageNumber="758">XLVII</td>
<td id="76B15D8EFFF455E1B10475B6FB8EF896" box="[865,1044,1790,1810]" gridcol="4" gridrow="13" pageId="11" pageNumber="758">(XLIV)</td>
<td id="76B15D8EFFF455E1B67975B6FB24F896" box="[1052,1214,1790,1810]" gridcol="5" gridrow="13" pageId="11" pageNumber="758">XLVII (XLVIII)</td>
<td id="76B15D8EFFF455E1B6AE75B6FA29F896" box="[1227,1459,1790,1810]" gridcol="6" gridrow="13" pageId="11" pageNumber="758">XLVII(XLVIII,XLIX,L)</td>
</tr>
<tr id="356034F2FFF455E1B2E6745EFA29F8AE" box="[131,1459,1814,1834]" gridrow="14" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E6745EFEBBF8AE" box="[131,289,1814,1834]" gridcol="0" gridrow="14" pageId="11" pageNumber="758">Tubercula</th>
<td id="76B15D8EFFF455E1B34F745EFE48F8AE" box="[298,466,1814,1834]" gridcol="1" gridrow="14" pageId="11" pageNumber="758">XXXV-XL</td>
<td id="76B15D8EFFF455E1B3B9745EFD19F8AE" box="[476,643,1814,1834]" gridcol="2" gridrow="14" pageId="11" pageNumber="758">(XXXV) XXXVI-XLI</td>
<td id="76B15D8EFFF455E1B0F7745EFCC8F8AE" box="[658,850,1814,1834]" gridcol="3" gridrow="14" pageId="11" pageNumber="758">XL-XLVI</td>
<td id="76B15D8EFFF455E1B104745EFB8EF8AE" box="[865,1044,1814,1834]" gridcol="4" gridrow="14" pageId="11" pageNumber="758">(XXXV) XXXVI-XLII</td>
<td id="76B15D8EFFF455E1B679745EFB24F8AE" box="[1052,1214,1814,1834]" gridcol="5" gridrow="14" pageId="11" pageNumber="758">(XLIII) XLIV-XLVII</td>
<td id="76B15D8EFFF455E1B6AE745EFA29F8AE" box="[1227,1459,1814,1834]" gridcol="6" gridrow="14" pageId="11" pageNumber="758">(XXXIX)XL(XLI)-</td>
</tr>
<tr id="356034F2FFF455E1B2E67463FA29F8BB" box="[131,1459,1835,1855]" gridrow="15" pageId="11" pageNumber="758" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76B15D8EFFF455E1B2E67463FEBBF8BB" box="[131,289,1835,1855]" gridcol="0" gridrow="15" pageId="11" pageNumber="758">pubertatis</th>
<td id="76B15D8EFFF455E1B1047463FB8EF8BB" box="[865,1044,1835,1855]" gridcol="4" gridrow="15" pageId="11" pageNumber="758">(XLIII)</td>
<td id="76B15D8EFFF455E1B6797463FB24F8BB" box="[1052,1214,1835,1855]" gridcol="5" gridrow="15" pageId="11" pageNumber="758">(XVIII)</td>
<td id="76B15D8EFFF455E1B6AE7463FA29F8BB" box="[1227,1459,1835,1855]" gridcol="6" gridrow="15" pageId="11" pageNumber="758">XLVI(XLVII,XLVIII,XLIX,L)</td>
</tr>
<tr id="356034F2FFF455E1B2E6740BFA29F8DC" box="[131,1459,1859,1880]" gridrow="16" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E6740BFEBBF8DC" box="[131,289,1859,1880]" gridcol="0" gridrow="16" pageId="11" pageNumber="758">Typhlosole</th>
<td id="76B15D8EFFF455E1B34F740BFE48F8DC" box="[298,466,1859,1880]" gridcol="1" gridrow="16" pageId="11" pageNumber="758">Four lamellae</td>
<td id="76B15D8EFFF455E1B3B9740BFD19F8DC" box="[476,643,1859,1880]" gridcol="2" gridrow="16" pageId="11" pageNumber="758">Four lamellae</td>
<td id="76B15D8EFFF455E1B0F7740BFCC8F8DC" box="[658,850,1859,1880]" gridcol="3" gridrow="16" pageId="11" pageNumber="758">Five lamellae</td>
<td id="76B15D8EFFF455E1B104740BFB8EF8DC" box="[865,1044,1859,1880]" gridcol="4" gridrow="16" pageId="11" pageNumber="758">Bifid</td>
<td id="76B15D8EFFF455E1B679740BFB24F8DC" box="[1052,1214,1859,1880]" gridcol="5" gridrow="16" pageId="11" pageNumber="758">Bifid</td>
<td id="76B15D8EFFF455E1B6AE740BFA29F8DC" box="[1227,1459,1859,1880]" gridcol="6" gridrow="16" pageId="11" pageNumber="758">Bifid</td>
</tr>
<tr id="356034F2FFF455E1B2E67413FA29F8F4" box="[131,1459,1883,1904]" gridrow="17" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E67413FEBBF8F4" box="[131,289,1883,1904]" gridcol="0" gridrow="17" pageId="11" pageNumber="758">Nephridial</th>
<td id="76B15D8EFFF455E1B34F7413FE48F8F4" box="[298,466,1883,1904]" gridcol="1" gridrow="17" pageId="11" pageNumber="758">Sigmoid, bag-like</td>
<td id="76B15D8EFFF455E1B3B97413FD19F8F4" box="[476,643,1883,1904]" gridcol="2" gridrow="17" pageId="11" pageNumber="758">Sigmoid, bag-like</td>
<td id="76B15D8EFFF455E1B0F77413FCC8F8F4" box="[658,850,1883,1904]" gridcol="3" gridrow="17" pageId="11" pageNumber="758">U-shaped</td>
<td id="76B15D8EFFF455E1B1047413FB8EF8F4" box="[865,1044,1883,1904]" gridcol="4" gridrow="17" pageId="11" pageNumber="758">U-shaped</td>
<td id="76B15D8EFFF455E1B6797413FB24F8F4" box="[1052,1214,1883,1904]" gridcol="5" gridrow="17" pageId="11" pageNumber="758">U-shaped</td>
<td id="76B15D8EFFF455E1B6AE7413FA29F8F4" box="[1227,1459,1883,1904]" gridcol="6" gridrow="17" pageId="11" pageNumber="758">U-shaped</td>
</tr>
<tr id="356034F2FFF455E1B2E67439FA29F801" box="[131,1459,1905,1925]" gridrow="18" pageId="11" pageNumber="758" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B15D8EFFF455E1B2E67439FEBBF801" box="[131,289,1905,1925]" gridcol="0" gridrow="18" pageId="11" pageNumber="758">bladders</th>
<td id="76B15D8EFFF455E1B34F7439FE48F801" box="[298,466,1905,1925]" gridcol="1" gridrow="18" pageId="11" pageNumber="758">distension</td>
<td id="76B15D8EFFF455E1B3B97439FD19F801" box="[476,643,1905,1925]" gridcol="2" gridrow="18" pageId="11" pageNumber="758">distension</td>
</tr>
<tr id="356034F2FFF455E1B2E674C1FA29F819" box="[131,1459,1929,1949]" gridrow="19" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E674C1FEBBF819" box="[131,289,1929,1949]" gridcol="0" gridrow="19" pageId="11" pageNumber="758">Seminal vesicles</th>
<td id="76B15D8EFFF455E1B34F74C1FE48F819" box="[298,466,1929,1949]" gridcol="1" gridrow="19" pageId="11" pageNumber="758">XI, XII</td>
<td id="76B15D8EFFF455E1B3B974C1FD19F819" box="[476,643,1929,1949]" gridcol="2" gridrow="19" pageId="11" pageNumber="758">XI, XII</td>
<td id="76B15D8EFFF455E1B0F774C1FCC8F819" box="[658,850,1929,1949]" gridcol="3" gridrow="19" pageId="11" pageNumber="758">IX-XII</td>
<td id="76B15D8EFFF455E1B10474C1FB8EF819" box="[865,1044,1929,1949]" gridcol="4" gridrow="19" pageId="11" pageNumber="758">IX-XII</td>
<td id="76B15D8EFFF455E1B67974C1FB24F819" box="[1052,1214,1929,1949]" gridcol="5" gridrow="19" pageId="11" pageNumber="758">IX-XII</td>
<td id="76B15D8EFFF455E1B6AE74C1FA29F819" box="[1227,1459,1929,1949]" gridcol="6" gridrow="19" pageId="11" pageNumber="758">IX-XII</td>
</tr>
<tr id="356034F2FFF455E1B2E674E9FA29F831" box="[131,1459,1953,1973]" gridrow="20" pageId="11" pageNumber="758">
<th id="76B15D8EFFF455E1B2E674E9FEBBF831" box="[131,289,1953,1973]" gridcol="0" gridrow="20" pageId="11" pageNumber="758">Spermathecae</th>
<td id="76B15D8EFFF455E1B34F74E9FE48F831" box="[298,466,1953,1973]" gridcol="1" gridrow="20" pageId="11" pageNumber="758">Double, oval in</td>
<td id="76B15D8EFFF455E1B3B974E9FD19F831" box="[476,643,1953,1973]" gridcol="2" gridrow="20" pageId="11" pageNumber="758">Double, oval in</td>
<td id="76B15D8EFFF455E1B0F774E9FCC8F831" box="[658,850,1953,1973]" gridcol="3" gridrow="20" pageId="11" pageNumber="758">Simple, intraseptal</td>
<td id="76B15D8EFFF455E1B10474E9FB8EF831" box="[865,1044,1953,1973]" gridcol="4" gridrow="20" pageId="11" pageNumber="758">Simple, intraseptal</td>
<td id="76B15D8EFFF455E1B67974E9FB24F831" box="[1052,1214,1953,1973]" gridcol="5" gridrow="20" pageId="11" pageNumber="758">Simple in 8/9,</td>
<td id="76B15D8EFFF455E1B6AE74E9FA29F831" box="[1227,1459,1953,1973]" gridcol="6" gridrow="20" pageId="11" pageNumber="758">Simple in (8/9), 9/10,</td>
</tr>
<tr id="356034F2FFF455E1B2E674FEFA29F84E" box="[131,1459,1974,1994]" gridrow="21" pageId="11" pageNumber="758" rowspan-0="1">
<td id="76B15D8EFFF455E1B34F74FEFE48F84E" box="[298,466,1974,1994]" gridcol="1" gridrow="21" pageId="11" pageNumber="758">X, XI</td>
<td id="76B15D8EFFF455E1B3B974FEFD19F84E" box="[476,643,1974,1994]" gridcol="2" gridrow="21" pageId="11" pageNumber="758">X, XI</td>
<td id="76B15D8EFFF455E1B0F774FEFCC8F84E" box="[658,850,1974,1994]" gridcol="3" gridrow="21" pageId="11" pageNumber="758">in 8/9, 9/10, 10/11</td>
<td id="76B15D8EFFF455E1B10474FEFB8EF84E" box="[865,1044,1974,1994]" gridcol="4" gridrow="21" pageId="11" pageNumber="758">to intracoelomic,</td>
<td id="76B15D8EFFF455E1B67974FEFB24F84E" box="[1052,1214,1974,1994]" gridcol="5" gridrow="21" pageId="11" pageNumber="758">9/10, 10/11</td>
<td id="76B15D8EFFF455E1B6AE74FEFA29F84E" box="[1227,1459,1974,1994]" gridcol="6" gridrow="21" pageId="11" pageNumber="758">10/11</td>
</tr>
<tr id="356034F2FFF455E1B2E67484FA29F864" box="[131,1459,1996,2016]" gridrow="22" pageId="11" pageNumber="758" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<td id="76B15D8EFFF455E1B1047484FB8EF864" box="[865,1044,1996,2016]" gridcol="4" gridrow="22" pageId="11" pageNumber="758">in (VIII), IX, X, XI</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEF36B0FFF3AA12B2B57019FE5BFCE1" blockId="12.[207,1355,849,955]" box="[208,449,849,869]" pageId="12" pageNumber="759">
<taxonomicName id="4C504D33FFF3AA12B2B57019FE1EFCE1" authority="Gérard &amp; Marchán &amp; Navarro &amp; Hedde &amp; Decaëns, 2023" authorityName="Gérard &amp; Marchán &amp; Navarro &amp; Hedde &amp; Decaëns" authorityYear="2023" box="[208,388,849,869]" class="Clitellata" family="Hormogastridae" genus="Vignysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="759" phylum="Annelida" rank="species" species="callasensis" status="sp. nov.">
<emphasis id="B924EAA2FFF3AA12B2B57019FE1EFCE1" box="[208,388,849,869]" italics="true" pageId="12" pageNumber="759">Vignysa callasensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757D9FFF3AA12B3EF7019FE5BFCE1" box="[394,449,849,869]" pageId="12" pageNumber="759" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BEF36B0FFF3AA12B2AA7033FED7FC0B" blockId="12.[207,1355,849,955]" box="[207,333,891,911]" pageId="12" pageNumber="759">
<taxonomicName id="4C504D33FFF3AA12B2AA7033FED7FC0B" baseAuthorityName="Duges" baseAuthorityYear="1828" box="[207,333,891,911]" class="Clitellata" family="Hormogastridae" genus="Vignysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="759" phylum="Annelida" rank="species" species="teres">
<emphasis id="B924EAA2FFF3AA12B2AA7033FED7FC0B" box="[207,333,891,911]" italics="true" pageId="12" pageNumber="759">Vignysa teres</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BEF36B0FFF3AA12B77270D4FAD1FC2A" blockId="12.[207,1355,849,955]" box="[1303,1355,924,942]" pageId="12" pageNumber="759">50 km</paragraph>
<paragraph id="8BEF36B0FFF3AA12B2AA70EFFEEBFC3F" blockId="12.[207,1355,849,955]" box="[207,369,935,955]" pageId="12" pageNumber="759">
<taxonomicName id="4C504D33FFF3AA12B2AA70EFFEEBFC3F" authorityName="Rota" authorityYear="1994" box="[207,369,935,955]" class="Clitellata" family="Hormogastridae" genus="Vignysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="759" phylum="Annelida" rank="species" species="vedovinii">
<emphasis id="B924EAA2FFF3AA12B2AA70EFFEEBFC3F" box="[207,369,935,955]" italics="true" pageId="12" pageNumber="759">Vignysa vedovinii</emphasis>
</taxonomicName>
</paragraph>
<caption id="DF2F6638FFF3AA12B2E070B7FD5BFBAC" ID-DOI="http://doi.org/10.5281/zenodo.10351731" ID-Zenodo-Dep="10351731" httpUri="https://zenodo.org/record/10351731/files/figure.png" pageId="12" pageNumber="759" startId="12.[133,144,1023,1041]" targetBox="[132,1453,214,981]" targetPageId="12" targetType="figure">
<paragraph id="8BEF36B0FFF3AA12B2E070B7FD5BFBAC" blockId="12.[133,1433,1023,1064]" pageId="12" pageNumber="759">
<smallCapsWord id="8D09A06CFFF3AA12B2E070B7FF07FB94" baselines="1036,1037" box="[133,157,1023,1041]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="12" pageNumber="759">FIG</smallCapsWord>
. 8. — Distribution of
<taxonomicName id="4C504D33FFF3AA12B32A70B7FD78FB95" authority="Gerard, Decaens &amp; Marchan" authorityName="Gerard, Characters Decaens &amp; Marchan" authorityYear="2023" box="[335,738,1023,1041]" class="Clitellata" family="Hormogastridae" genus="Vignysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="759" phylum="Annelida" rank="species" species="callasensis" status="sp. nov.">
<emphasis id="B924EAA2FFF3AA12B32A70B7FE77FB95" box="[335,493,1023,1041]" italics="true" pageId="12" pageNumber="759">Vignysa callasensis</emphasis>
Gérard, Decaëns &amp; Marchán
</taxonomicName>
,
<taxonomicNameLabel id="A21757D9FFF3AA12B08F70B7FC80FB95" box="[746,794,1023,1041]" pageId="12" pageNumber="759" rank="species">n. sp.</taxonomicNameLabel>
(
<emphasis id="B924EAA2FFF3AA12B14770B7FCDFFB95" bold="true" box="[802,837,1023,1041]" pageId="12" pageNumber="759">star</emphasis>
) and its closest relatives (
<emphasis id="B924EAA2FFF3AA12B67970B7FBCDFB95" bold="true" box="[1052,1111,1023,1041]" pageId="12" pageNumber="759">circles</emphasis>
) in Southern France. All species of the genus
<taxonomicName id="4C504D33FFF3AA12B2D8775EFEECFBAC" authority="Bouche, 1970" authorityName="Bouche" authorityYear="1970" box="[189,374,1045,1064]" class="Clitellata" family="Hormogastridae" genus="Vignysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="759" phylum="Annelida" rank="genus">
<emphasis id="B924EAA2FFF3AA12B2D8775EFF67FBAC" box="[189,253,1046,1064]" italics="true" pageId="12" pageNumber="759">Vignysa</emphasis>
<bibRefCitation id="EFC14B41FFF3AA12B367775DFEECFBAC" author="BOUCHE M. B." box="[258,374,1045,1064]" firstAuthor="Bouche" journalOrPublisher="Pedobiologia" pageId="12" pageNumber="759" pagination="246 - 256" part="10" refId="ref12055" refString="BOUCHE M. B. 1970. - Remarques sur quelques Lumbricina de France et consequences de la decouverte des noveaux taxons Vignysinae (Subfam. nov) et Diporodrilidae (Fam. nov.). Pedobiologia 10: 246 - 256." title="Remarques sur quelques Lumbricina de France et consequences de la decouverte des noveaux taxons Vignysinae (Subfam. nov) et Diporodrilidae (Fam. nov.)" type="journal article" year="1970">Bouché, 1970</bibRefCitation>
</taxonomicName>
are represented. Credits:
<bibRefCitation id="EFC14B41FFF3AA12B02B775DFD27FBAC" accessDate="2022-12-13" author="GOOGLE" box="[590,701,1045,1064]" firstAuthor="Google" pageId="12" pageNumber="759" publicationUrl="http://google.com/maps" refId="ref12848" refString="GOOGLE 2022 - Available at: http: // google. com / maps [Accessed: 13 December 2022 ̿." type="url" year="2022">Google 2022</bibRefCitation>
.
</paragraph>
</caption>
<subSubSection id="C34A653BFFF3AA12B2E1765EFEBAFAE6" pageId="12" pageNumber="759" type="taxon_list">
<paragraph id="8BEF36B0FFF3AA12B2E1765EFEBAFAE6" blockId="12.[132,775,1301,1378]" pageId="12" pageNumber="759">
<smallCapsWord id="8D09A06CFFF3AA12B2E1765EFF71FAA8" baselines="1319,1320" box="[132,235,1302,1325]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Included" pageId="12" pageNumber="759">INCLUDED</smallCapsWord>
<smallCapsWord id="8D09A06CFFF3AA12B2967653FEA0FAA8" baselines="1320" box="[243,314,1307,1324]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="species" pageId="12" pageNumber="759">SPECIES</smallCapsWord>
. —
<taxonomicName id="4C504D33FFF3AA12B30F765EFD2BFAA9" authority="(Duges, 1828: 289)" baseAuthorityName="Duges" baseAuthorityPageNumber="289" baseAuthorityYear="1828" box="[362,689,1301,1325]" class="Clitellata" family="Hormogastridae" genus="Vignysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="759" phylum="Annelida" rank="species" species="teres">
<emphasis id="B924EAA2FFF3AA12B30F765EFE79FAA9" box="[362,483,1302,1325]" italics="true" pageId="12" pageNumber="759">Vignysa teres</emphasis>
(
<bibRefCitation id="EFC14B41FFF3AA12B397765DFD30FAA9" author="DUGES A. - L." box="[498,682,1301,1325]" firstAuthor="Duges" journalOrPublisher="Annales des Sciences naturelles, Paris" page="289" pageId="12" pageNumber="759" pagination="284 - 337" part="15" refId="ref12691" refString="DUGES A. - L. 1828. - Recherches sur la circulation, la respiration et la reproduction des Annelides abranches. Annales des Sciences naturelles, Paris Serie 1 15: 284 - 337." title="Recherches sur la circulation, la respiration et la reproduction des Annelides abranches" type="journal article" year="1828">Dugès, 1828: 289</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C504D33FFF3AA12B0DB765EFE1AFACC" authority="Rota, 1994: 27" authorityName="Rota" authorityPageNumber="27" authorityYear="1994" class="Clitellata" family="Hormogastridae" genus="Vignysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="759" phylum="Annelida" rank="species" species="vedovinii">
<emphasis id="B924EAA2FFF3AA12B0DB765EFF41FACC" italics="true" pageId="12" pageNumber="759">Vignysa vedovinii</emphasis>
<bibRefCitation id="EFC14B41FFF3AA12B2817678FE1AFACC" author="ROTA E." box="[228,384,1328,1352]" firstAuthor="Rota" journalOrPublisher="Journal of Natural History" page="27" pageId="12" pageNumber="759" pagination="27 - 36" part="28" refId="ref15472" refString="ROTA E. 1994. - Contribution to the taxonomy of the Hormogastridae (Annelida: Oligochaeta) with description of two new species from southern France. Journal of Natural History 28 (1): 27 - 36. https: // doi. org / 10.1080 / 00222939400770031" title="Contribution to the taxonomy of the Hormogastridae (Annelida: Oligochaeta) with description of two new species from southern France" type="journal article" year="1994">Rota, 1994: 27</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C504D33FFF3AA12B3E97678FF7BFAE6" authority="Gerard, Decaens &amp; Marchan" authorityName="Gerard, Characters Decaens &amp; Marchan" authorityYear="2023" class="Clitellata" family="Hormogastridae" genus="Vignysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="759" phylum="Annelida" rank="species" species="callasensis" status="sp. nov.">
<emphasis id="B924EAA2FFF3AA12B3E97678FDA1FACC" box="[396,571,1328,1352]" italics="true" pageId="12" pageNumber="759">Vignysa callasensis</emphasis>
Gérard, Decaëns &amp; Marchán
</taxonomicName>
,
<taxonomicNameLabel id="A21757D9FFF3AA12B28E7603FEBAFAE6" box="[235,288,1355,1378]" pageId="12" pageNumber="759" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C34A653BFFF3AA12B2E176CBFD71FA6E" pageId="12" pageNumber="759" type="distribution">
<paragraph id="8BEF36B0FFF3AA12B2E176CBFD71FA6E" blockId="12.[132,776,1411,1514]" pageId="12" pageNumber="759">
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<smallCapsWord id="8D09A06CFFF3AA12B34476C0FE33FA1D" baselines="1429" box="[289,425,1416,1433]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="distribution" pageId="12" pageNumber="759">DISTRIBUTION</smallCapsWord>
. — This genus currently comprises three species occurring in Southeastern
<collectingCountry id="F3477620FFF3AA12B06976D6FDCAFA31" box="[524,592,1438,1461]" name="France" pageId="12" pageNumber="759">France</collectingCountry>
, from Montpellier (Hérault, Occitanie) to the north of Fréjus (Var, Occitanie), and from the Mediterranean Sea to the 44th north parallel (
<figureCitation id="136B2A35FFF3AA12B0C6769AFD44FA6E" box="[675,734,1490,1514]" captionStart="FIG" captionStartId="12.[133,144,1023,1041]" captionTargetId="figure-502@12.[131,793,213,789]" captionTargetPageId="12" captionText="FIG. 8. — Distribution of Vignysa callasensis Gérard, Decaëns &amp; Marchán, n. sp. (star) and its closest relatives (circles) in Southern France. All species of the genus Vignysa Bouché, 1970 are represented. Credits: Google 2022." figureDoi="http://doi.org/10.5281/zenodo.10351731" httpUri="https://zenodo.org/record/10351731/files/figure.png" pageId="12" pageNumber="759">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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