257 lines
32 KiB
XML
257 lines
32 KiB
XML
<document id="46F734352E098B6902CBF4B15863F5E3" ID-CLB-Dataset="8407" ID-DOI="10.11646/zootaxa.4852.4.3" ID-GBIF-Dataset="4b35f4f2-e111-4162-997b-f607da4bbb44" ID-ISSN="1175-5326" ID-Zenodo-Dep="4410079" ID-ZooBank="D21984B5-3BC7-4175-AA15-3290FEC3E9DB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609596865468" checkinUser="plazi" docAuthor="Hutchinson, Paul M. & Allsopp, Peter G." docDate="2020" docId="03F52C0EFF8CAB7CFF12FCE2FC33FCC8" docLanguage="en" docName="zootaxa.4852.4.3.pdf" docOrigin="Zootaxa 4852 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cavonus fovealacus Hutchinson & Allsopp 2020, new species" docType="treatment" docUuid="27E8992D-0EF2-4729-BE2A-EBD5EEE97910" docUuidSource="ZooBank" docVersion="7" lastPageNumber="458" masterDocId="FFCC5476FF8AAB75FF85FFFDFF8DFFF1" masterDocTitle="Cavonus Sharp, 1875 (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini Pseudoryctina) in Western Australia with description of a new species and the female of Cavonus sculpturatus Blackburn, 1888" masterLastPageNumber="460" masterPageNumber="449" pageNumber="455" updateTime="1698851062327" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="98FFF0E158EC98AA2DE6769323056BB9">Cavonus Sharp, 1875 (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini Pseudoryctina) in Western Australia with description of a new species and the female of Cavonus sculpturatus Blackburn, 1888</mods:title>
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<mods:name id="1C76113B7C1C6A9A5448B3E177305739" type="personal">
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<mods:roleTerm id="9032E7F398DDA509D46A9A177621580D">Author</mods:roleTerm>
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<mods:namePart id="89A9FA39B801D5C38B2402106995DDF7">Hutchinson, Paul M.</mods:namePart>
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<mods:affiliation id="DF04050C2925D8DBA1F157033E8A9834">Department of Agriculture and Food WA (Quarantine WA), Locked Bag 69, Welshpool DC, Western Australia 6986, Australia. paul. hutchinson @ dpird. wa. gov. au; https: // orcid. org / 0000 - 0001 - 8747 - 1320</mods:affiliation>
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<mods:nameIdentifier id="3E70806BF572C20B3F243CB67B4BB49A" type="email">paul.hutchinson@dpird.wa.gov.au</mods:nameIdentifier>
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<mods:namePart id="65469377075ABFA7F1BA0D33AD50A6C9">Allsopp, Peter G.</mods:namePart>
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<mods:affiliation id="31C34A17DB7394F3A25A8D01E61EE5F6">Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia.</mods:affiliation>
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<mods:title id="51C2D206536E6BE29CD69942CAFC85EA">Zootaxa</mods:title>
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<mods:date id="465E052B3623BA1C8053CBE4904131DB">2020</mods:date>
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<mods:number id="81E05C44C76B1BAA10A164FF6A945471">2020-09-17</mods:number>
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<treatment id="03F52C0EFF8CAB7CFF12FCE2FC33FCC8" ID-DOI="http://doi.org/10.5281/zenodo.4410077" ID-GBIF-Taxon="177309746" ID-Zenodo-Dep="4410077" LSID="urn:lsid:zoobank.org:act:27E8992D-0EF2-4729-BE2A-EBD5EEE97910" httpUri="http://treatment.plazi.org/id/03F52C0EFF8CAB7CFF12FCE2FC33FCC8" lastPageId="9" lastPageNumber="458" pageId="6" pageNumber="455">
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<subSubSection id="C346CE93FF8CAB73FF12FCE2FCCBFCC9" box="[151,838,798,825]" pageId="6" pageNumber="455" type="nomenclature">
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<heading id="D0AB2A74FF8CAB73FF12FCE2FCCBFCC9" bold="true" box="[151,838,798,825]" fontSize="11" level="1" pageId="6" pageNumber="455" reason="1">
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<emphasis id="B928410AFF8CAB73FF12FCE2FCCBFCC9" bold="true" box="[151,838,798,825]" pageId="6" pageNumber="455">
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<taxonomicName id="4C5CE69BFF8CAB73FF12FCE2FD24FCC9" authority="Hutchinson & Allsopp" authorityName="Hutchinson & Allsopp" authorityYear="2020" box="[151,681,798,825]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="species" species="fovealacus" status="sp. nov.">
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<emphasis id="B928410AFF8CAB73FF12FCE2FE04FCC8" bold="true" box="[151,393,798,825]" italics="true" pageId="6" pageNumber="455">Cavonus fovealacus</emphasis>
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Hutchinson & Allsopp
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21BFC71FF8CAB73FD33FCE2FCCBFCC9" box="[694,838,798,825]" pageId="6" pageNumber="455" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C346CE93FF8CAB7CFF12FCBEFE2FFF08" lastPageId="9" lastPageNumber="458" pageId="6" pageNumber="455" type="description">
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<paragraph id="8BE39D18FF8CAB73FF12FCBEFEA3FCAC" blockId="6.[151,851,798,889]" box="[151,302,835,861]" pageId="6" pageNumber="455">
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(
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<figureCitation id="1367819DFF8CAB73FF1AFCBEFEABFCAC" box="[159,294,835,861]" captionStart-0="FIGURES 17–25" captionStart-1="FIGURES 26–27" captionStart-2="FIGURE 28" captionStartId-0="7.[151,264,1960,1984]" captionStartId-1="8.[151,264,1943,1967]" captionStartId-2="9.[151,250,1556,1580]" captionTargetBox-0="[205,1382,193,1929]" captionTargetBox-1="[155,1433,181,1920]" captionTargetBox-2="[151,1435,849,1533]" captionTargetId-0="figure-19@7.[205,1382,181,1937]" captionTargetId-1="figure-25@8.[155,1433,181,1920]" captionTargetId-2="figure-292@9.[151,1435,849,1533]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURES 17–25. Cavonus fovealacus new species, holotype male. 17, Dorsum; 18, venter; 19, lateral; 20, head, lateral; 21, head, dorsal; 22, pygidium; 23, parameres; 24, aedeagus, lateral; 25, labels." captionText-1="FIGURES 26–27. Collection areas of Cavonus fovealacus new species. 26, Lake Pallarup (holotype); 27, Lake Carmody (paratype)." captionText-2="FIGURE 28. Collection localities of Cavonus fovealacus new species and C. sculpturatus Blackburn, 1888 in Western Australia and South Australia." figureDoi-0="http://doi.org/10.5281/zenodo.4410087" figureDoi-1="http://doi.org/10.5281/zenodo.4410089" figureDoi-2="http://doi.org/10.5281/zenodo.4410091" httpUri-0="https://zenodo.org/record/4410087/files/figure.png" httpUri-1="https://zenodo.org/record/4410089/files/figure.png" httpUri-2="https://zenodo.org/record/4410091/files/figure.png" pageId="6" pageNumber="455">Figs. 17–28</figureCitation>
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)
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</paragraph>
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<paragraph id="8BE39D18FF8CAB73FF12FC9EFCDEFC88" blockId="6.[151,851,798,889]" box="[151,851,867,889]" pageId="6" pageNumber="455">
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urn:lsid:zoobank.org:act:
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<paragraph id="8BE39D18FF8CAB73FF12FC55FE8DFBFB" blockId="6.[151,1437,936,2043]" pageId="6" pageNumber="455">
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<emphasis id="B928410AFF8CAB73FF12FC55FF69FC33" bold="true" box="[151,228,936,962]" pageId="6" pageNumber="455">Types.</emphasis>
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<materialsCitation id="3B349745FF8CAB73FF6EFC55FE8DFBFB" ID-GBIF-Occurrence="3032720305" collectingDate="2008-02-12" collectionCode="LAKE, PALLARUP, WAM" country="Australia" location="WESTERN AUSTRALIA" pageId="6" pageNumber="455" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia" typeStatus="holotype">
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<collectingCountry id="F34BDD88FF8CAB73FF6EFC55FE00FC33" box="[235,397,936,962]" name="Australia" pageId="6" pageNumber="455">AUSTRALIA</collectingCountry>
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.
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<collectingRegion id="499853FAFF8CAB73FE12FC55FD4FFC33" box="[407,706,936,962]" country="Australia" name="Western Australia" pageId="6" pageNumber="455">WESTERN AUSTRALIA</collectingRegion>
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.
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<typeStatus id="54E723BAFF8CAB73FD48FC55FCB8FC33" box="[717,821,936,962]" pageId="6" pageNumber="455" type="holotype">Holotype</typeStatus>
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male:
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<specimenCount id="9D5A5691FF8CAB73FC04FC54FC1BFC32" box="[897,918,937,963]" pageId="6" pageNumber="455" type="male">♂</specimenCount>
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<collectionCode id="ED4D05DDFF8CAB73FC18FC55FBF6FC33" box="[925,1147,936,962]" pageId="6" pageNumber="455">LAKE PALLARUP</collectionCode>
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(STRANDLINE)
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<collectingRegion id="499853FAFF8CAB73FACEFC55FA14FC33" box="[1355,1433,936,962]" country="Australia" name="WEST" pageId="6" pageNumber="455">WEST</collectingRegion>
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.
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<collectingRegion id="499853FAFF8CAB73FF12FC31FF6FFC17" box="[151,226,972,998]" country="Australia" name="AUST" pageId="6" pageNumber="455">AUST</collectingRegion>
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.
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<date id="FFE2BBD8FF8CAB73FF6BFC31FEF7FC17" box="[238,378,972,998]" pageId="6" pageNumber="455" value="2008-02-12">
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<collectingDate id="EFA64230FF8CAB73FF6BFC31FEF7FC17" box="[238,378,972,998]" pageId="6" pageNumber="455" value="2008-02-12">12.Feb.2008</collectingDate>
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</date>
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<collectingRegion id="499853FAFF8CAB73FE06FC30FE1EFC16" box="[387,403,973,999]" country="Australia" name="P" pageId="6" pageNumber="455">P</collectingRegion>
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. HUTCHINSON | lake strandline // PMH Coll. # Dyn 0473 // genitalia on card [in
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<collectionCode id="ED4D05DDFF8CAB73FACBFC36FA1BFC17" box="[1358,1430,971,998]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="6" pageNumber="455" type="Museum">WAM</collectionCode>
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] (
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<figureCitation id="1367819DFF8CAB73FF25FC0DFF7EFBFB" box="[160,243,1008,1034]" captionStart="FIGURES 17–25" captionStartId="7.[151,264,1960,1984]" captionTargetBox="[205,1382,193,1929]" captionTargetId="figure-19@7.[205,1382,181,1937]" captionTargetPageId="7" captionText="FIGURES 17–25. Cavonus fovealacus new species, holotype male. 17, Dorsum; 18, venter; 19, lateral; 20, head, lateral; 21, head, dorsal; 22, pygidium; 23, parameres; 24, aedeagus, lateral; 25, labels." figureDoi="http://doi.org/10.5281/zenodo.4410087" httpUri="https://zenodo.org/record/4410087/files/figure.png" pageId="6" pageNumber="455">Fig. 17</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BE39D18FF8CAB73FF42FBE8FC8AFBA3" blockId="6.[151,1437,936,2043]" pageId="6" pageNumber="455">
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<typeStatus id="54E723BAFF8CAB73FF42FBE8FEA4FBDE" box="[199,297,1045,1071]" pageId="6" pageNumber="455" type="paratype">Paratype</typeStatus>
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<specimenCount id="9D5A5691FF8CAB73FEB4FBE8FEC4FBDE" box="[305,329,1045,1071]" pageId="6" pageNumber="455" type="male">♂</specimenCount>
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:
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<materialsCitation id="3B349745FF8CAB73FED0FBE8FC8EFBA3" ID-GBIF-Occurrence="3032720306" collectingDate="2001-10-23" collectionCode="PMH" collectorName="PMH" location="Lake Carmody West. Aust. Dead" pageId="6" pageNumber="455" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<specimenCount id="9D5A5691FF8CAB73FED0FBE8FEE7FBDE" box="[341,362,1045,1071]" pageId="6" pageNumber="455" type="male">♂</specimenCount>
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<location id="8E83CBC3FF8CAB73FEF7FBE9FD6EFBDF" LSID="urn:lsid:plazi:treatment:03F52C0EFF8CAB7CFF12FCE2FC33FCC8:8E83CBC3FF8CAB73FEF7FBE9FD6EFBDF" box="[370,739,1044,1071]" name="Lake Carmody West. Aust. Dead" pageId="6" pageNumber="455">Lake Carmody West. Aust. Dead</location>
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<date id="FFE2BBD8FF8CAB73FD6EFBE9FCF8FBDF" box="[747,885,1044,1070]" pageId="6" pageNumber="455" value="2001-10-23">
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<collectingDate id="EFA64230FF8CAB73FD6EFBE9FCF8FBDF" box="[747,885,1044,1070]" pageId="6" pageNumber="455" value="2001-10-23">23.Oct.2001</collectingDate>
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||
</date>
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edge saltlake
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<collectorName id="26A9F8CEFF8CAB73FB93FBE8FB3FFBDF" box="[1046,1202,1044,1071]" pageId="6" pageNumber="455">P. Hutchinson</collectorName>
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| dead. edge saltlake // PMH
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<collectorName id="26A9F8CEFF8CAB73FF74FBC5FEA4FBA3" box="[241,297,1080,1106]" pageId="6" pageNumber="455">Coll.</collectorName>
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#
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<collectorName id="26A9F8CEFF8CAB73FEC0FBC4FEF8FBA2" box="[325,373,1081,1107]" pageId="6" pageNumber="455">Dyn</collectorName>
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0474 // genitalia on card [in
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<collectionCode id="ED4D05DDFF8CAB73FD3CFBCAFD71FBA3" box="[697,764,1079,1106]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14877" name="City Museum and Records Office" pageId="6" pageNumber="455" type="Herbarium">PMH</collectionCode>
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||
]
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</materialsCitation>
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||
.
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</paragraph>
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||
<paragraph id="8BE39D18FF8CAB7CFF42FBA1FDCDFF24" blockId="6.[151,1437,936,2043]" lastBlockId="9.[151,1437,150,825]" lastPageId="9" lastPageNumber="458" pageId="6" pageNumber="455">
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||
<emphasis id="B928410AFF8CAB73FF42FBA1FE12FB87" bold="true" box="[199,415,1116,1142]" pageId="6" pageNumber="455">Description. Male</emphasis>
|
||
(
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<figureCitation id="1367819DFF8CAB73FE35FBA1FDB5FB86" box="[432,568,1116,1143]" captionStart="FIGURES 17–25" captionStartId="7.[151,264,1960,1984]" captionTargetBox="[205,1382,193,1929]" captionTargetId="figure-19@7.[205,1382,181,1937]" captionTargetPageId="7" captionText="FIGURES 17–25. Cavonus fovealacus new species, holotype male. 17, Dorsum; 18, venter; 19, lateral; 20, head, lateral; 21, head, dorsal; 22, pygidium; 23, parameres; 24, aedeagus, lateral; 25, labels." figureDoi="http://doi.org/10.5281/zenodo.4410087" httpUri="https://zenodo.org/record/4410087/files/figure.png" pageId="6" pageNumber="455">Figs. 17–25</figureCitation>
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). Body black, palps brown, antennomeres 2–10 black with dark-brown suffusion,
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<quantity id="4CA430FDFF8CAB73FF69FB7DFEDDFB6A" box="[236,336,1152,1179]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" pageId="6" pageNumber="455" unit="mm" value="15.7">15.7 mm</quantity>
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long,
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<quantity id="4CA430FDFF8CAB73FE1DFB7DFE62FB6A" box="[408,495,1152,1179]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="6" pageNumber="455" unit="mm" value="9.3">9.3 mm</quantity>
|
||
wide. Mentum wide at base, deflexed, narrowed to globular apex, reflexed at acute angle and constricted to ligula exposing labial palp insertions, surface setose; ligula margins straight, apex with narrowly expanded arcuate ridge; mandibular palps distinctly exposed beyond clypeus, palpomeres 2 and 3 subequal, palpomere 4 conical and widest at truncate apex, bearing slight dorsolateral fovea in basal half. Antenna with 10 antennomeres, scape and pedicel setose, club long, subequal to length of stalk, inner face setose, outer face unitexturous. Clypeus at 45° to plane of frons, transverse, subcircular in outline with apex broadly arcuate, anterior margin reflexed, lateral margins raised, surface with medial third slightly convex, finely and sparsely punctate, anterior face deep, oblique, glabrous; clypeofrontal ridge strongly raised, transverse, slightly flattened in middle, declivous to frons. Frons medially flattened, concentrically rugulose; ocular canthi short, coarsely punctate, anterior margin at slight angle to line of clypeus, straight, bearing short stout setae, apex rounded and not surpassing lateral margin of eye. Pronotum with anterior margin membranous, anterolateral angle at 90°, posterolateral angle distinct, obtuse, lateral margins evenly arcuate, widest at midlength, anteromedian tubercle stout, erect, slightly recurved, preceding a Y-shaped moderately deep impression, impression anteriorly highly transverse in anterior half reaching lateral margins, with broad moderately deep longitudinal depression interrupting basal convexity, laterally borders rounded, surface with fine, sparse punctation, coarser and denser along lateral and posterior margins, impression with incomplete punctures becoming denser and rugulose anteriorly, posterior ridge continuous. Scutellum with disc coarsely punctate. Each elytron with epipleural setae confined to humeral calli, sutural costa linear-punctate, disc with moderately to deeply impressed coarse punctures in well-defined striae, striae 3 and 4 finer anterior to apical calli, intervals impunctate except for a cluster at base of stria 6. Pygidium strongly convex, disc sparsely punctate, denser across base, setose basolaterally, scarcer across middle, apical ridge moderately broadened and sparsely setose. Postcoxal prosternal process long, narrow, evenly arcuate posteriorly, setose, apex tapering with surface glabrous. Abdominal sternites 2–6 punctate across posterior margin, medially narrowly glabrous. Protibiae quite stout, tridentate, denticles equidistant; protarsomere 5 longer than protarsomere 1; metafemora stout, unisulcate; metatibiae moderately stout with length about 2.5x apical width, bicarinate, basal carina bearing long setae, medial carina bearing long, acute cilia set 1 width apart, apical spurs tapering, smaller slightly curved, larger straight; metatarsomere 5 longer than metatarsomere 1. Genitalia with parameres curving inwards from about half length and apices at right angles to basal portion.
|
||
</paragraph>
|
||
<caption id="DF23CD90FF8DAB72FF12F855FC1EF814" ID-DOI="http://doi.org/10.5281/zenodo.4410087" ID-Zenodo-Dep="4410087" httpUri="https://zenodo.org/record/4410087/files/figure.png" pageId="7" pageNumber="456" startId="7.[151,264,1960,1984]" targetBox="[205,1382,193,1929]" targetPageId="7">
|
||
<paragraph id="8BE39D18FF8DAB72FF12F855FC1EF814" blockId="7.[151,1436,1960,2021]" pageId="7" pageNumber="456">
|
||
<emphasis id="B928410AFF8DAB72FF12F855FED8F830" bold="true" box="[151,341,1960,1985]" pageId="7" pageNumber="456">FIGURES 17–25.</emphasis>
|
||
<taxonomicName id="4C5CE69BFF8DAB72FED9F854FDA5F830" authority="Hutchinson & Allsopp, 2020" authorityName="Hutchinson & Allsopp" authorityYear="2020" box="[348,552,1961,1985]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="fovealacus" status="sp. nov.">
|
||
<emphasis id="B928410AFF8DAB72FED9F854FDA5F830" box="[348,552,1961,1985]" italics="true" pageId="7" pageNumber="456">Cavonus fovealacus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928410AFF8DAB72FDB5F854FD23F830" bold="true" box="[560,686,1961,1985]" pageId="7" pageNumber="456">
|
||
<taxonomicNameLabel id="A21BFC71FF8DAB72FDB5F854FD23F830" box="[560,686,1961,1985]" pageId="7" pageNumber="456" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male.
|
||
<emphasis id="B928410AFF8DAB72FCDFF854FCFEF830" bold="true" box="[858,883,1961,1985]" pageId="7" pageNumber="456">17</emphasis>
|
||
, Dorsum;
|
||
<emphasis id="B928410AFF8DAB72FC64F854FC77F830" bold="true" box="[993,1018,1961,1985]" pageId="7" pageNumber="456">18</emphasis>
|
||
, venter;
|
||
<emphasis id="B928410AFF8DAB72FBD3F854FBE2F830" bold="true" box="[1110,1135,1961,1985]" pageId="7" pageNumber="456">19</emphasis>
|
||
, lateral;
|
||
<emphasis id="B928410AFF8DAB72FB4FF854FB6EF830" bold="true" box="[1226,1251,1961,1985]" pageId="7" pageNumber="456">20</emphasis>
|
||
, head, lateral;
|
||
<emphasis id="B928410AFF8DAB72FAF8F854FA1BF830" bold="true" box="[1405,1430,1961,1985]" pageId="7" pageNumber="456">21</emphasis>
|
||
, head, dorsal;
|
||
<emphasis id="B928410AFF8DAB72FE9AF830FEB5F814" bold="true" box="[287,312,1997,2021]" pageId="7" pageNumber="456">22</emphasis>
|
||
, pygidium;
|
||
<emphasis id="B928410AFF8DAB72FE36F830FE41F814" bold="true" box="[435,460,1997,2021]" pageId="7" pageNumber="456">23</emphasis>
|
||
, parameres;
|
||
<emphasis id="B928410AFF8DAB72FDCAF830FDE5F814" bold="true" box="[591,616,1997,2021]" pageId="7" pageNumber="456">24</emphasis>
|
||
, aedeagus, lateral;
|
||
<emphasis id="B928410AFF8DAB72FCAEF830FCC9F814" bold="true" box="[811,836,1997,2021]" pageId="7" pageNumber="456">25</emphasis>
|
||
, labels.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF23CD90FF82AB7DFF12F86AFE8AF825" ID-DOI="http://doi.org/10.5281/zenodo.4410089" ID-Zenodo-Dep="4410089" httpUri="https://zenodo.org/record/4410089/files/figure.png" pageId="8" pageNumber="457" startId="8.[151,264,1943,1967]" targetBox="[155,1433,181,1920]" targetPageId="8">
|
||
<paragraph id="8BE39D18FF82AB7DFF12F86AFE8AF825" blockId="8.[151,1436,1943,2004]" pageId="8" pageNumber="457">
|
||
<emphasis id="B928410AFF82AB7DFF12F86AFED5F85E" bold="true" box="[151,344,1943,1967]" pageId="8" pageNumber="457">FIGURES 26–27.</emphasis>
|
||
Collection areas of
|
||
<taxonomicName id="4C5CE69BFF82AB7DFDB4F865FD72F841" authority="Hutchinson & Allsopp, 2020" authorityName="Hutchinson & Allsopp" authorityYear="2020" box="[561,767,1944,1968]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="fovealacus" status="sp. nov.">
|
||
<emphasis id="B928410AFF82AB7DFDB4F865FD72F841" box="[561,767,1944,1968]" italics="true" pageId="8" pageNumber="457">Cavonus fovealacus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928410AFF82AB7DFC8FF865FC3EF85E" bold="true" box="[778,947,1943,1968]" pageId="8" pageNumber="457">
|
||
<taxonomicNameLabel id="A21BFC71FF82AB7DFC8FF865FC06F85E" box="[778,907,1943,1968]" pageId="8" pageNumber="457" rank="species">new species</taxonomicNameLabel>
|
||
. 26
|
||
</emphasis>
|
||
, Lake Pallarup (holotype);
|
||
<emphasis id="B928410AFF82AB7DFB5DF86AFB7CF85E" bold="true" box="[1240,1265,1943,1967]" pageId="8" pageNumber="457">27</emphasis>
|
||
, Lake Carmody (paratype).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE39D18FF83AB7CFF42FF22FE2FFF08" blockId="9.[151,1437,150,825]" box="[199,418,223,249]" pageId="9" pageNumber="458">
|
||
<emphasis id="B928410AFF83AB7CFF42FF22FEA8FF08" bold="true" box="[199,293,223,249]" pageId="9" pageNumber="458">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346CE93FF83AB7CFF42FEFEFAF1FE97" pageId="9" pageNumber="458" type="etymology">
|
||
<paragraph id="8BE39D18FF83AB7CFF42FEFEFAF1FE97" blockId="9.[151,1437,150,825]" pageId="9" pageNumber="458">
|
||
<emphasis id="B928410AFF83AB7CFF42FEFEFEC3FEEC" bold="true" box="[199,334,259,285]" pageId="9" pageNumber="458">Etymology.</emphasis>
|
||
The specific name is a combination of fovea (Latin for pitfall) and lacus (Latin for lake), in recognition of the many naturally occurring salt lakes that dot the Western Australian landscape and that act as ‘Nature’s pitfall traps’. Both known specimens of the new species were collected from salt-lake strandlines (
|
||
<figureCitation id="1367819DFF83AB7CFB63FEB6FAE2FE97" box="[1254,1391,331,358]" captionStart="FIGURES 26–27" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[155,1433,181,1920]" captionTargetId="figure-25@8.[155,1433,181,1920]" captionTargetPageId="8" captionText="FIGURES 26–27. Collection areas of Cavonus fovealacus new species. 26, Lake Pallarup (holotype); 27, Lake Carmody (paratype)." figureDoi="http://doi.org/10.5281/zenodo.4410089" httpUri="https://zenodo.org/record/4410089/files/figure.png" pageId="9" pageNumber="458">Figs. 26–27</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346CE93FF83AB7CFF42FE93FC33FCC8" pageId="9" pageNumber="458" type="diagnosis">
|
||
<paragraph id="8BE39D18FF83AB7CFF42FE93FEA9FE04" blockId="9.[151,1437,150,825]" pageId="9" pageNumber="458">
|
||
<emphasis id="B928410AFF83AB7CFF42FE93FECFFE79" bold="true" box="[199,322,366,392]" pageId="9" pageNumber="458">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C5CE69BFF83AB7CFECFFE92FDA0FE78" authorityName="Hutchinson & Allsopp" authorityYear="2020" box="[330,557,367,393]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="fovealacus">
|
||
<emphasis id="B928410AFF83AB7CFECFFE92FDA0FE78" box="[330,557,367,393]" italics="true" pageId="9" pageNumber="458">Cavonus fovealacus</emphasis>
|
||
</taxonomicName>
|
||
males can be distinguished from its congeners by the following combination of characters: clypeus rounded; mandibular palps with apical palpomere conical, widest and truncate at apex; pronotum with single tubercle preceding a deep Y-shaped depression and basal ridge continuous; pygidium with apical ridge setose.
|
||
</paragraph>
|
||
<paragraph id="8BE39D18FF83AB7CFF42FE03FC33FCC8" blockId="9.[151,1437,150,825]" pageId="9" pageNumber="458">
|
||
<emphasis id="B928410AFF83AB7CFF42FE03FEA7FDE9" bold="true" box="[199,298,510,536]" pageId="9" pageNumber="458">Habitat.</emphasis>
|
||
Lake Carmody and Lake Pallarup are amongst a series of lakes that are arranged in a north-south orientation between Hyden and Ravensthorpe,
|
||
<collectingRegion id="499853FAFF83AB7CFD2EFDDEFCF9FDCC" box="[683,884,547,573]" country="Australia" name="Western Australia" pageId="9" pageNumber="458">Western Australia</collectingRegion>
|
||
(
|
||
<figureCitation id="1367819DFF83AB7CFC01FDDEFC54FDCC" box="[900,985,547,573]" captionStart="FIGURE 28" captionStartId="9.[151,250,1556,1580]" captionTargetBox="[151,1435,849,1533]" captionTargetId="figure-292@9.[151,1435,849,1533]" captionTargetPageId="9" captionText="FIGURE 28. Collection localities of Cavonus fovealacus new species and C. sculpturatus Blackburn, 1888 in Western Australia and South Australia." figureDoi="http://doi.org/10.5281/zenodo.4410091" httpUri="https://zenodo.org/record/4410091/files/figure.png" pageId="9" pageNumber="458">Fig. 28</figureCitation>
|
||
). Both natural salt lakes are surrounded by mallee (
|
||
<taxonomicName id="4C5CE69BFF83AB7CFE91FDBAFE1DFD90" box="[276,400,583,609]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="458" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B928410AFF83AB7CFE91FDBAFE1DFD90" box="[276,400,583,609]" italics="true" pageId="9" pageNumber="458">Eucalyptus</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<taxonomicName id="4C5CE69BFF83AB7CFE2CFDBAFDA8FD90" box="[425,549,583,609]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="458" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
|
||
), with an understorey composed of
|
||
<taxonomicName id="4C5CE69BFF83AB7CFC37FDBAFB36FD90" box="[946,1211,583,609]" class="Agaricomycetes" family="Tricholomataceae" genus="Melaleuca" higherTaxonomySource="CoL" kingdom="Fungi" order="Agaricales" pageId="9" pageNumber="458" phylum="Basidiomycota" rank="genus">
|
||
<emphasis id="B928410AFF83AB7CFC37FDBAFBA5FD90" box="[946,1064,583,609]" italics="true" pageId="9" pageNumber="458">Melaleuca</emphasis>
|
||
(Myrtaceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5CE69BFF83AB7CFB40FDBAFEABFD77" class="Magnoliopsida" family="Casuarinaceae" genus="Allocasuarina" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="458" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B928410AFF83AB7CFB40FDBAFAE9FD90" box="[1221,1380,583,609]" italics="true" pageId="9" pageNumber="458">Allocasuarina</emphasis>
|
||
(Casuarinaceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5CE69BFF83AB7CFEB1FD96FD87FD77" box="[308,522,619,646]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="458" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B928410AFF83AB7CFEB1FD96FE0FFD74" box="[308,386,619,645]" italics="true" pageId="9" pageNumber="458">Acacia</emphasis>
|
||
(Fabaceae)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5CE69BFF83AB7CFDCFFD96FCB4FD74" box="[586,825,619,645]" class="Magnoliopsida" family="Proteaceae" genus="Banksia" kingdom="Plantae" order="Proteales" pageId="9" pageNumber="458" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B928410AFF83AB7CFDCFFD96FD29FD74" box="[586,676,619,645]" italics="true" pageId="9" pageNumber="458">Banksia</emphasis>
|
||
(Proteaceae)
|
||
</taxonomicName>
|
||
over white sand (
|
||
<figureCitation id="1367819DFF83AB7CFB88FD96FB17FD77" box="[1037,1178,619,646]" captionStart="FIGURES 26–27" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[155,1433,181,1920]" captionTargetId="figure-25@8.[155,1433,181,1920]" captionTargetPageId="8" captionText="FIGURES 26–27. Collection areas of Cavonus fovealacus new species. 26, Lake Pallarup (holotype); 27, Lake Carmody (paratype)." figureDoi="http://doi.org/10.5281/zenodo.4410089" httpUri="https://zenodo.org/record/4410089/files/figure.png" pageId="9" pageNumber="458">Figs. 26–27</figureCitation>
|
||
). The area has a Köppen-Geiger climate of BSk (cold, semi-arid,
|
||
<bibRefCitation id="EFCDE0E9FF83AB7CFD1EFD72FCD1FD58" author="Beck, H. E. & Zimmermann, N. E. & McVicar, T. R. & Vergopolan, N. & Berg, A. & Wood, E. F." box="[667,860,655,681]" pageId="9" pageNumber="458" pagination="180214" refId="ref6347" refString="Beck, H. E., Zimmermann, N. E., McVicar, T. R., Vergopolan, N., Berg, A. & Wood, E. F. (2018) Present and future Koppen-Geiger climate classification maps at 1 - km resolution. Scientific Data, 5, 180214. https: // doi. org / 10.1038 / sdata. 2018.214" type="journal article" year="2018">
|
||
Beck
|
||
<emphasis id="B928410AFF83AB7CFD5BFD72FC94FD58" box="[734,793,655,681]" italics="true" pageId="9" pageNumber="458">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
) with an average annual rainfall of
|
||
<quantity id="4CA430FDFF83AB7CFB7BFD72FA14FD58" box="[1278,1433,655,681]" metricMagnitude="-1" metricUnit="m" metricValue="3.85" metricValueMax="4.3" metricValueMin="3.4" pageId="9" pageNumber="458" unit="mm" value="385.0" valueMax="430.0" valueMin="340.0">340–430 mm</quantity>
|
||
. Most adult specimens of
|
||
<taxonomicName id="4C5CE69BFF83AB7CFE32FD4EFD94FD3C" authorityName="Sharp" authorityYear="1875" box="[439,537,691,717]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928410AFF83AB7CFE32FD4EFD94FD3C" box="[439,537,691,717]" italics="true" pageId="9" pageNumber="458">Cavonus</emphasis>
|
||
</taxonomicName>
|
||
were recorded from September to November and March (
|
||
<bibRefCitation id="EFCDE0E9FF83AB7CFB37FD4EFAB5FD3C" author="Carne, P. B." box="[1202,1336,690,717]" pageId="9" pageNumber="458" refId="ref6492" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne 1957</bibRefCitation>
|
||
) and the condition of the specimens with quite minor deterioration of colour by sun bleaching and mostly intact appendages is of similar stage of decomposition to other strandline coleopteran specimens where emergence times thus age of specimens are known and suggests it emerges in September–November.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |