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<document ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" approvalRequired="59" approvalRequired_for_taxonomicNames="59" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. &amp; Jansa, Sharon A." docDate="2022" docId="03EFDD5DF7046969D977FDF01BCCFDC5" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.8:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="8" docTitle="Evolestes undetermined" docType="treatment" docVersion="5" lastPageNumber="319" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="319" updateTime="1659987007554" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Beck, Robin M. D.</mods:namePart>
<mods:affiliation>School of Science, Engineering and Environment University of Salford, U. K. &amp; School of Biological, Earth &amp; Environmental Sciences University of New South Wales, Australia &amp; Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Voss, Robert S.</mods:namePart>
<mods:affiliation>Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jansa, Sharon A.</mods:namePart>
<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-28</mods:number>
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<mods:detail type="volume">
<mods:number>2022</mods:number>
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<mods:detail type="issue">
<mods:number>457</mods:number>
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<mods:end>353</mods:end>
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<mods:url>https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
<mods:identifier type="ISSN">0003-0090</mods:identifier>
<mods:identifier type="Zenodo-Dep">6971356</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6974442" ID-Zenodo-Dep="6974442" LSID="urn:lsid:plazi:treatment:03EFDD5DF7046969D977FDF01BCCFDC5" httpUri="http://treatment.plazi.org/id/03EFDD5DF7046969D977FDF01BCCFDC5" lastPageId="320" lastPageNumber="319" pageId="319" pageNumber="319">
<subSubSection box="[892,1005,605,630]" pageId="319" pageNumber="319" type="nomenclature">
<paragraph blockId="319.[892,1005,605,630]" box="[892,1005,605,630]" pageId="319" pageNumber="319">
<heading box="[892,1005,605,630]" centered="true" fontSize="10" level="2" pageId="319" pageNumber="319" reason="4">
<emphasis bold="true" box="[892,1005,605,630]" pageId="319" pageNumber="319">
<taxonomicName box="[906,1005,605,629]" class="Mammalia" family="Caenolestidae" genus="Evolestes" kingdom="Animalia" order="Paucituberculata" pageId="319" pageNumber="319" phylum="Chordata" rank="species">
<emphasis bold="true" box="[906,1005,605,629]" italics="true" pageId="319" pageNumber="319">Evolestes</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="319" pageNumber="319" type="synonymic_list">
<paragraph blockId="319.[683,1213,655,1738]" pageId="319" pageNumber="319">
SPECIES SCORED: †
<taxonomicName box="[947,1212,655,679]" class="Mammalia" family="Caenolestidae" genus="Evolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="319" pageNumber="319" phylum="Chordata" rank="species">
<emphasis box="[947,1212,655,679]" italics="true" pageId="319" pageNumber="319">Evolestes hadrommatos</emphasis>
</taxonomicName>
(
<typeStatus box="[692,741,689,713]" pageId="319" pageNumber="319">type</typeStatus>
and only described species), †
<taxonomicName box="[1076,1207,688,712]" class="Mammalia" family="Caenolestidae" genus="Evolestes" kingdom="Animalia" order="Paucituberculata" pageId="319" pageNumber="319" phylum="Chordata" rank="species">
<emphasis box="[1076,1169,688,712]" italics="true" pageId="319" pageNumber="319">Evolestes</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="319" pageNumber="319" type="materials_examined">
<paragraph blockId="319.[683,1213,655,1738]" pageId="319" pageNumber="319">
GEOLOGICAL PROVENANCE OF SCORED
<collectionCode box="[684,743,755,779]" pageId="319" pageNumber="319">SPECIMENS</collectionCode>
: Unit 6 of the Salla Beds,
<collectingRegion country="Bolivia" name="La Paz" pageId="319" pageNumber="319">La Paz Department</collectingRegion>
,
<collectingCountry box="[752,828,788,812]" name="Bolivia" pageId="319" pageNumber="319">Bolivia</collectingCountry>
(†
<taxonomicName box="[857,1103,788,812]" class="Mammalia" family="Caenolestidae" genus="Evolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="319" pageNumber="319" phylum="Chordata" rank="species">
<emphasis box="[857,1103,788,812]" italics="true" pageId="319" pageNumber="319">Evolestes hadrommatos</emphasis>
</taxonomicName>
); GBV-19 (“La Cantera” locality), Unit 3, Upper Puesto Almendra Member, Sarmiento Formation, Gran Barranca south of Lake Colhue-Huapi, Sarmiento Department,
<collectingRegion box="[854,1068,920,944]" country="Argentina" name="Chubut" pageId="319" pageNumber="319">Chubut Province</collectingRegion>
,
<collectingCountry box="[1095,1211,920,944]" name="Argentina" pageId="319" pageNumber="319">Argentina</collectingCountry>
(†
<taxonomicName box="[707,840,953,977]" class="Mammalia" family="Caenolestidae" genus="Evolestes" kingdom="Animalia" order="Paucituberculata" pageId="319" pageNumber="319" phylum="Chordata" rank="species">
<emphasis box="[707,800,953,977]" italics="true" pageId="319" pageNumber="319">Evolestes</emphasis>
sp.
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="319" pageNumber="319" type="description">
<paragraph blockId="319.[683,1213,655,1738]" pageId="319" pageNumber="319">AGE OF SCORED SPECIMENS: Unit 6 (= “the Upper White interval”) of the Salla Beds is 25.8225.89 Mya (i.e., late Oligocene) based on radiometric and magnetostratigraphic evidence (Kay et al., 1998: 191, fig. 2), whereas the “La Cantera” fossil locality is 29.531.1 Mya based on magnetostratigraphy (Ré et al., 2010: 57).</paragraph>
<paragraph blockId="319.[683,1213,655,1738]" box="[716,1197,1217,1242]" pageId="319" pageNumber="319">
ASSIGNED AGE
<collectionCode box="[887,964,1218,1242]" pageId="319" pageNumber="319">RANGE</collectionCode>
:
<geoCoordinate box="[978,1050,1217,1241]" degrees="31.100" direction="north" orientation="latitude" pageId="319" pageNumber="319" precision="55" value="31.1">31.100</geoCoordinate>
<geoCoordinate box="[1049,1135,1217,1241]" degrees="25.820" direction="west" orientation="longitude" pageId="319" pageNumber="319" precision="55" value="-25.82">25.820</geoCoordinate>
Mya.
</paragraph>
</subSubSection>
<subSubSection lastPageId="320" lastPageNumber="320" pageId="319" pageNumber="319" type="discussion">
<paragraph blockId="319.[683,1213,655,1738]" lastBlockId="320.[108,637,226,515]" lastPageId="320" lastPageNumber="320" pageId="319" pageNumber="319">
REMARKS: †
<taxonomicName box="[851,1100,1251,1275]" class="Mammalia" family="Caenolestidae" genus="Evolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="319" pageNumber="319" phylum="Chordata" rank="species">
<emphasis box="[851,1100,1251,1275]" italics="true" pageId="319" pageNumber="319">Evolestes hadrommatos</emphasis>
</taxonomicName>
is known from a single partial cranium (the
<typeStatus box="[1114,1212,1284,1308]" pageId="319" pageNumber="319">holotype</typeStatus>
MNHN-BOL-V-004017 [previously MNHN- BOL-96-400]; Goin et al., 2007). Of particular interest for this study, this specimen exhibits a molar dentition that is markedly more plesiomorphic than that of any extant paucituberculatan (Abello, 2007; Goin et al., 2007, 2010; Abello, 2013). Subsequently, additional dental material assignable to †
<taxonomicName box="[961,1056,1548,1572]" class="Mammalia" family="Caenolestidae" genus="Evolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
<emphasis box="[961,1056,1548,1572]" italics="true" pageId="319" pageNumber="319">Evolestes</emphasis>
</taxonomicName>
and possibly representing a new species was described from the “La Cantera” locality at the Gran Barranca in
<collectingCountry box="[684,793,1647,1671]" name="Argentina" pageId="319" pageNumber="319">Argentina</collectingCountry>
(Goin et al., 2010; Abello, 2013). The La Cantera material is important because it includes lower molars, which have not been described for †
<taxonomicName box="[293,482,226,250]" class="Mammalia" family="Caenolestidae" genus="Evolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="320" pageNumber="320" phylum="Chordata" rank="species">
<emphasis box="[293,482,226,250]" italics="true" pageId="320" pageNumber="320">E. hadrommatos</emphasis>
</taxonomicName>
. Goin et al. (2010) thought that †
<taxonomicName box="[353,447,259,283]" class="Mammalia" family="Caenolestidae" genus="Evolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="320" pageNumber="320" phylum="Chordata" rank="genus">
<emphasis box="[353,447,259,283]" italics="true" pageId="320" pageNumber="320">Evolestes</emphasis>
</taxonomicName>
might be a caenolestid, but such a familial assignment is not supported by published phylogenetic analyses (Goin et al., 2007, 2009a; Abello, 2013; Forasiepi et al., 2013; Rincón et al., 2015; Engelman et al., 2016; Abello et al., 2020), which instead suggest that †
<taxonomicName box="[170,263,457,481]" class="Mammalia" family="Caenolestidae" genus="Evolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="320" pageNumber="320" phylum="Chordata" rank="genus">
<emphasis box="[170,263,457,481]" italics="true" pageId="320" pageNumber="320">Evolestes</emphasis>
</taxonomicName>
represents an early-branching lineage within
<taxonomicName authorityName="Ameghino" authorityYear="1894" box="[241,426,490,514]" class="Mammalia" kingdom="Animalia" order="Paucituberculata" pageId="320" pageNumber="320" phylum="Chordata" rank="order">Paucituberculata</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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