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<document id="67A37A741F9C79B1AE22840F6ACA4C8A" ID-DOI="10.1649/0010-065x-69.4.773" ID-ISSN="1938-4394" ID-Zenodo-Dep="10106588" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630178626354" checkinUser="diego" docAuthor="Ratcliffe, Brett C." docDate="2015" docId="03EC0108C137FFF3A88AFBCAFDEDF985" docLanguage="en" docName="0010-065X-69.4.773.pdf.imf" docOrigin="The Coleopterists Bulletin 69 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-69.4.773" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Stenocrates hastatus Ratcliffe 2015, new species" docType="treatment" docVersion="3" lastPageNumber="775" masterDocId="FFD57970C135FFF0A842FFCEFFEDFF8A" masterDocTitle="A Revised Catalog of the Species ofStenocratesBurmeister (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini), with Descriptions of Three New Species from Peru and Brazil andStenocrates inpaiRatcliffe, 1978 Placed in Junior Synonymy withStenocrates popeiEndrödi, 1971" masterLastPageNumber="779" masterPageNumber="773" pageNumber="775" updateTime="1699638037398" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="ACC795B7679E6A91EF91B257321D7C27">
<mods:title id="419D16E05429E8EF4F601C6D44D4377F">A Revised Catalog of the Species ofStenocratesBurmeister (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini), with Descriptions of Three New Species from Peru and Brazil andStenocrates inpaiRatcliffe, 1978 Placed in Junior Synonymy withStenocrates popeiEndrödi, 1971</mods:title>
</mods:titleInfo>
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<mods:role id="BECD6C271BB085414CB36029DC2DC38E">
<mods:roleTerm id="67EBF8AEDFCE1C973F564B45C1B47A2D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="19F310766F98B701934F7276F9F8C7EF">Ratcliffe, Brett C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="E92CAC6AC7284500A3FAE3B41D566409">text</mods:typeOfResource>
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<mods:title id="C7FCBAAAA73188442A0149624330315C">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="6FCF954EFEE9CB665509DF92FC84FC99">
<mods:date id="1927FB46673910E5BEAEA9FE4DEE4283">2015</mods:date>
<mods:detail id="AC1FADBCD248C84113E39701FAC35CED" type="pubDate">
<mods:number id="E81DD2CE3625EBB2417C31569BCB33FA">2015-12-31</mods:number>
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<mods:number id="86DDE3D265FE652517CAF9E30555F930">69</mods:number>
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<mods:number id="B223EEC6C89C73C76404920F3CCB6122">4</mods:number>
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<mods:url id="DB32C327842D041EE95D6376ADBD6AC2">http://dx.doi.org/10.1649/0010-065x-69.4.773</mods:url>
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<mods:classification id="0C7B6E31520A0630AC4B5A298FC44D67">journal article</mods:classification>
<mods:identifier id="C56A4A2B48029B3EF62B6C01CB069B3C" type="DOI">10.1649/0010-065x-69.4.773</mods:identifier>
<mods:identifier id="264103B0E45C47BBCF462791FA183C6A" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="1F7A58D2AD098BD17CDF56F5D969089B" type="Zenodo-Dep">10106588</mods:identifier>
</mods:mods>
<treatment id="03EC0108C137FFF3A88AFBCAFDEDF985" LSID="urn:lsid:plazi:treatment:03EC0108C137FFF3A88AFBCAFDEDF985" httpUri="http://treatment.plazi.org/id/03EC0108C137FFF3A88AFBCAFDEDF985" lastPageId="3" lastPageNumber="775" pageId="2" pageNumber="775">
<subSubSection id="C35FE395C137FFF2A88AFBCAFE38FBBE" pageId="2" pageNumber="775" type="nomenclature">
<paragraph id="8BFAB01EC137FFF2A88AFBCAFE38FBBE" blockId="2.[200,631,1028,1076]" pageId="2" pageNumber="775">
<emphasis id="B9316C0CC137FFF2A88AFBCAFD9AFB93" bold="true" box="[200,631,1028,1049]" pageId="2" pageNumber="775">
<taxonomicName id="4C45CB9DC137FFF2A88AFBCAFE19FB93" ID-CoL="4ZXQP" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2015" box="[200,500,1028,1049]" class="Insecta" family="Scarabaeidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="775" phylum="Arthropoda" rank="species" species="hastatus" status="sp. nov.">
<emphasis id="B9316C0CC137FFF2A88AFBCAFE7EFB93" bold="true" box="[200,403,1028,1049]" italics="true" pageId="2" pageNumber="775">Stenocrates hastatus</emphasis>
Ratcliffe
</taxonomicName>
,
<taxonomicNameLabel id="A202D177C137FFF2AA43FBCAFD9AFB93" box="[513,631,1028,1049]" pageId="2" pageNumber="775" rank="species">new species</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="137EAC9BC137FFF2A933FBD1FE23FBBE" box="[369,462,1054,1076]" captionStart="Figs" captionStartId="2.[197,234,830,849]" captionTargetBox="[181,654,192,806]" captionTargetId="figure-614@2.[173,660,183,825]" captionTargetPageId="2" captionText="Figs.36. Stenocrates species, parameres. 3) S. howdeni; 4) S. laborator; 56) S. hastatus. Illustrations in Figs. 34 from Dechambre and Hardy (2004)." figureDoi="http://doi.org/10.5281/zenodo.10106592" httpUri="https://zenodo.org/record/10106592/files/figure.png" pageId="2" pageNumber="775">Figs. 56</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35FE395C137FFF2A887FB87FDD9F9B6" pageId="2" pageNumber="775" type="materials_examined">
<paragraph id="8BFAB01EC137FFF2A887FB87FD79FAE2" blockId="2.[173,660,1097,1729]" pageId="2" pageNumber="775">
<emphasis id="B9316C0CC137FFF2A887FB87FEB1FBD4" bold="true" box="[197,348,1097,1118]" pageId="2" pageNumber="775">Type Material.</emphasis>
<materialsCitation id="3B2DBA43C137FFF2A92BFB87FE14FB19" collectingDate="2011-03" collectorName="E. J. Grossi" country="Brazil" elevation="100" location="BRAZIL" pageId="2" pageNumber="775" specimenCount="2" typeStatus="holotype">
<typeStatus id="54FE0EBCC137FFF2A92BFB87FE25FBD4" box="[361,456,1097,1118]" pageId="2" pageNumber="775" type="holotype">Holotype</typeStatus>
labeled “
<collectingCountry id="F352F08EC137FFF2AA73FB87FD62FBD4" box="[561,655,1097,1118]" name="Brazil" pageId="2" pageNumber="775">BRAZIL</collectingCountry>
: RJ/Rio das Ostras/
<date id="FFFB96DEC137FFF2A92FFBAAFE50FBF3" box="[365,445,1124,1145]" pageId="2" pageNumber="775" value="2011-03">
<collectingDate id="EFBF6F36C137FFF2A92FFBAAFE50FBF3" box="[365,445,1124,1145]" pageId="2" pageNumber="775" value="2011-03">III-2011</collectingDate>
</date>
<quantity id="4CBD1DFBC137FFF2A985FBAAFDE4FBF3" box="[455,521,1124,1145]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="775" unit="m" value="100.0">
<elevation id="0068572DC137FFF2A985FBAAFDE4FBF3" box="[455,521,1124,1145]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="775" unit="m" value="100.0">100 m</elevation>
</quantity>
/
<collectorName id="26B0D5C8C137FFF2AA53FBAAFD6AFBF3" box="[529,647,1124,1145]" pageId="2" pageNumber="775">E. J. Grossi</collectorName>
” and with my red
<typeStatus id="54FE0EBCC137FFF2A921FBB0FE56FB19" box="[355,443,1150,1171]" pageId="2" pageNumber="775" type="holotype">holotype</typeStatus>
label
</materialsCitation>
.
<materialsCitation id="3B2DBA43C137FFF2AA44FBB0FD7DFAE2" collectingDate="2011-03" collectorName="Dr. L. Gomez &amp; P. Sandig" country="Brazil" elevation="100" location="BRAZIL" pageId="2" pageNumber="775" specimenCount="3" specimenCount-female="3" typeStatus="allotype">
<typeStatus id="54FE0EBCC137FFF2AA44FBB0FDB0FB19" box="[518,605,1150,1171]" pageId="2" pageNumber="775" type="allotype">Allotype</typeStatus>
with same data and my red
<typeStatus id="54FE0EBCC137FFF2A9DBFB57FE05FB24" box="[409,488,1177,1198]" pageId="2" pageNumber="775" type="allotype">allotype</typeStatus>
label. One male and
<specimenCount id="9D437B97C137FFF2A895FB7DFEACFB42" box="[215,321,1203,1224]" pageId="2" pageNumber="775" type="female">one female</specimenCount>
<typeStatus id="54FE0EBCC137FFF2A908FB7DFE76FB42" box="[330,411,1203,1224]" pageId="2" pageNumber="775" type="paratype">paratype</typeStatus>
with same data and each with my yellow
<typeStatus id="54FE0EBCC137FFF2A90FFB00FE73FB69" box="[333,414,1230,1251]" pageId="2" pageNumber="775" type="paratype">paratype</typeStatus>
label. One male and
<specimenCount id="9D437B97C137FFF2AA33FB00FF00FB74" pageId="2" pageNumber="775" type="female">one female</specimenCount>
<typeStatus id="54FE0EBCC137FFF2A8B0FB27FEAEFB74" box="[242,323,1257,1278]" pageId="2" pageNumber="775" type="paratype">paratype</typeStatus>
labeled “
<collectingCountry id="F352F08EC137FFF2A9DFFB27FE18FB74" box="[413,501,1257,1278]" name="Brazil" pageId="2" pageNumber="775">BRAZIL</collectingCountry>
, Esp. Santo/40 k no Linhares/17-19-X-82/Col:
<collectorName id="26B0D5C8C137FFF2A996FACDFDB0FA92" box="[468,605,1283,1304]" pageId="2" pageNumber="775">Dr. L. Gomez</collectorName>
” and each with my yellow
<typeStatus id="54FE0EBCC137FFF2A9C8FAD0FE32FAB9" box="[394,479,1310,1331]" pageId="2" pageNumber="775" type="paratype">paratype</typeStatus>
label. Two males and
<specimenCount id="9D437B97C137FFF2A895FAF6FEAFFAC7" box="[215,322,1336,1357]" pageId="2" pageNumber="775" type="female">one female</specimenCount>
labeled “Guapi, Braz/Mar.3.1935/
<collectorName id="26B0D5C8C137FFF2A8EFFA9DFEE8FAE2" box="[173,261,1363,1384]" pageId="2" pageNumber="775">P. Sandig</collectorName>
” and each with my yellow
<typeStatus id="54FE0EBCC137FFF2AA4AFA9DFDB4FAE2" box="[520,601,1363,1384]" pageId="2" pageNumber="775" type="paratype">paratype</typeStatus>
label
</materialsCitation>
.
</paragraph>
<paragraph id="8BFAB01EC137FFF2A887FAA3FDD9F9B6" blockId="2.[173,660,1097,1729]" pageId="2" pageNumber="775">
<materialsCitation id="3B2DBA43C137FFF2A887FAA3FF0BFA3D" collectorName="Lincoln, NE" country="United States of America" location="University" municipality="Museum" pageId="2" pageNumber="775" specimenCount="2" stateProvince="Nebraska" typeStatus="holotype">
<typeStatus id="54FE0EBCC137FFF2A887FAA3FECEFA08" box="[197,291,1389,1410]" pageId="2" pageNumber="775" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54FE0EBCC137FFF2A918FAA3FE46FA08" box="[346,427,1389,1410]" pageId="2" pageNumber="775" type="allotype">allotype</typeStatus>
deposited at the
<location id="8E9AE6C5C137FFF2AA27FAA3FF03FA17" LSID="urn:lsid:plazi:treatment:03EC0108C137FFF3A88AFBCAFDEDF985:8E9AE6C5C137FFF2AA27FAA3FF03FA17" country="United States of America" municipality="Museum" name="University" pageId="2" pageNumber="775" stateProvince="Nebraska">University</location>
of
<collectingRegion id="49817EFCC137FFF2A951FA46FE48FA17" box="[275,421,1416,1437]" country="United States of America" name="Nebraska" pageId="2" pageNumber="775">Nebraska State</collectingRegion>
<collectingMunicipality id="6B9E2A64C137FFF2A9EDFA46FDEEFA17" box="[431,515,1416,1437]" pageId="2" pageNumber="775">Museum</collectingMunicipality>
(
<collectorName id="26B0D5C8C137FFF2AA50FA46FD62FA17" box="[530,655,1416,1437]" pageId="2" pageNumber="775">Lincoln, NE</collectorName>
,
<collectingCountry id="F352F08EC137FFF2A8EFFA6CFF0DFA3D" box="[173,224,1442,1463]" name="United States of America" pageId="2" pageNumber="775">USA</collectingCountry>
)
</materialsCitation>
.
<materialsCitation id="3B2DBA43C137FFF2A8ACFA6CFEB6F9A8" collectionCode="US, USNM, DC" country="Brazil" county="Museu Nacional" location="National Museum" municipality="University" pageId="2" pageNumber="775" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
Three
<typeStatus id="54FE0EBCC137FFF2A968FA6CFE6FFA3D" box="[298,386,1442,1463]" pageId="2" pageNumber="775" type="paratype">paratypes</typeStatus>
deposited in the
<collectionCode id="ED5428DBC137FFF2AA62FA6CFDD3FA3D" box="[544,574,1442,1463]" country="South Africa" name="University of Stellenbosch" pageId="2" pageNumber="775">US</collectionCode>
<location id="8E9AE6C5C137FFF2AA06FA6CFEECFA58" LSID="urn:lsid:plazi:treatment:03EC0108C137FFF3A88AFBCAFDEDF985:8E9AE6C5C137FFF2AA06FA6CFEECFA58" country="Brazil" county="Museu Nacional" municipality="University" name="National Museum" pageId="2" pageNumber="775" stateProvince="Rio de Janeiro">National Museum</location>
(
<collectionCode id="ED5428DBC137FFF2A952FA73FEB5FA58" box="[272,344,1469,1490]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="775" type="Museum">USNM</collectionCode>
) (Washington,
<collectionCode id="ED5428DBC137FFF2A9B0FA73FDFBFA58" box="[498,534,1469,1490]" pageId="2" pageNumber="775">DC</collectionCode>
, currently at the
<collectingMunicipality id="6B9E2A64C137FFF2A88CFA16FEDDFA67" box="[206,304,1496,1517]" pageId="2" pageNumber="775">University</collectingMunicipality>
of Nebraska for offsite enhancement),
<specimenCount id="9D437B97C137FFF2A8EFFA3CFEC8F98D" box="[173,293,1522,1543]" pageId="2" pageNumber="775" type="generic" typeStatus="paratype">one paratype</specimenCount>
deposited in the
<collectingCounty id="629BC892C137FFF2A98AFA3CFD8EF98D" box="[456,611,1522,1543]" pageId="2" pageNumber="775">Museu Naçional</collectingCounty>
(
<collectingRegion id="49817EFCC137FFF2AA32FA3CFEE2F9A8" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="775">Rio de Janeiro</collectingRegion>
,
<collectingCountry id="F352F08EC137FFF2A958F9C3FEB8F9A8" box="[282,341,1549,1570]" name="Brazil" pageId="2" pageNumber="775">Brazil</collectingCountry>
)
</materialsCitation>
,
<materialsCitation id="3B2DBA43C137FFF2A924F9C3FDDDF9B6" collectorName="B. C. Ratcliffe Collection &amp; Lincoln, NE" country="United States of America" location="USA" pageId="2" pageNumber="775" specimenCount="1" typeStatus="holotype">
and
<specimenCount id="9D437B97C137FFF2A9CDF9C3FDF0F9A8" box="[399,541,1549,1570]" pageId="2" pageNumber="775" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
in the
<collectorName id="26B0D5C8C137FFF2AA23F9C3FE89F9B6" pageId="2" pageNumber="775">B. C. Ratcliffe Collection</collectorName>
(
<collectorName id="26B0D5C8C137FFF2A931F9E9FE01F9B6" box="[371,492,1575,1596]" pageId="2" pageNumber="775">Lincoln, NE</collectorName>
,
<collectingCountry id="F352F08EC137FFF2A9B4F9E9FDC4F9B6" box="[502,553,1575,1596]" name="United States of America" pageId="2" pageNumber="775">USA</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35FE395C137FFF2A887F98CFBE8FAE2" pageId="2" pageNumber="775" type="description">
<paragraph id="8BFAB01EC137FFF2A887F98CFC91FCE5" blockId="2.[173,660,1097,1729]" lastBlockId="2.[707,1194,194,1729]" pageId="2" pageNumber="775">
<emphasis id="B9316C0CC137FFF2A887F98CFE5CF9DD" bold="true" box="[197,433,1602,1623]" pageId="2" pageNumber="775">
Description of
<typeStatus id="54FE0EBCC137FFF2A913F98CFE40F9DD" box="[337,429,1602,1623]" pageId="2" pageNumber="775" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male. Length
<quantity id="4CBD1DFBC137FFF2AA7EF98CFD62F9DD" box="[572,655,1602,1623]" metricMagnitude="-2" metricUnit="m" metricValue="2.08" pageId="2" pageNumber="775" unit="mm" value="20.8">20.8 mm</quantity>
; width
<quantity id="4CBD1DFBC137FFF2A8AEF992FEAAF9FB" box="[236,327,1628,1649]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="2" pageNumber="775" unit="mm" value="10.1">10.1 mm</quantity>
. Color black.
<emphasis id="B9316C0CC137FFF2A996F992FDFFF9FB" bold="true" box="[468,530,1628,1649]" pageId="2" pageNumber="775">Head:</emphasis>
Frons shiny, with sparse micropunctures. Frontoclypeal suture impressed, distinctly broader at middle, ridge in front of suture distinctly carinulate either side of depressed center. Clypeus transversely rugose; apex weakly emarginate, narrowly reflexed, anterior face thickened. Interocular width equals 3.0 transverse eye diameters. Antenna with 10 antennomeres, club subequal in length to antennomeres 27.
<emphasis id="B9316C0CC137FFF2AC7FFEE2FB44FECB" bold="true" box="[1085,1193,300,321]" pageId="2" pageNumber="775">Pronotum:</emphasis>
Surface shiny, nearly smooth, except for a few small punctures on disc and a field of large, moderately dense punctures in both posterior and anterior angles. Lateral margins with thick marginal bead, base without marginal bead.
<emphasis id="B9316C0CC137FFF2AB97FE7FFBF1FE4C" bold="true" box="[981,1052,433,454]" pageId="2" pageNumber="775">Elytra:</emphasis>
Surface shiny, with punctate sutural stria and 2 pairs of distinct discal striae and 1 pair of striae behind humerus; each stria comprised of large, deep, umbilicate, closely adjacent punctures. First broad interval with an irregular row of similar punctures, second broad interval with punctures only on apical third.
<emphasis id="B9316C0CC137FFF2AA81FDA5FCC7FD0A" bold="true" box="[707,810,619,640]" pageId="2" pageNumber="775">Pygidium:</emphasis>
Surface shiny, completely and densely punctate, punctures moderately large, deep, glabrous. In lateral view, surface regularly convex.
<emphasis id="B9316C0CC137FFF2AA81FD75FD13FD5A" bold="true" box="[707,766,699,720]" pageId="2" pageNumber="775">Legs:</emphasis>
Protibia tridentate, basal tooth slightly removed from other teeth. Metatarsus incomplete but, if complete, shorter than metatibia.
<emphasis id="B9316C0CC137FFF2AC1CFD3EFB44FC8F" bold="true" box="[1118,1193,752,773]" pageId="2" pageNumber="775">Venter:</emphasis>
Prosternal process columnar, long, thick, apex flattened into longitudinally oval disc.
<emphasis id="B9316C0CC137FFF2AC70FCEBFB44FCB0" bold="true" box="[1074,1193,805,826]" pageId="2" pageNumber="775">Parameres:</emphasis>
Form widest at about middle, apices elongate, lanceolate (
<figureCitation id="137EAC9BC137FFF2AB51FC94FC9DFCE5" box="[787,880,858,880]" captionStart="Figs" captionStartId="2.[197,234,830,849]" captionTargetBox="[181,654,192,806]" captionTargetId="figure-614@2.[173,660,183,825]" captionTargetPageId="2" captionText="Figs.36. Stenocrates species, parameres. 3) S. howdeni; 4) S. laborator; 56) S. hastatus. Illustrations in Figs. 34 from Dechambre and Hardy (2004)." figureDoi="http://doi.org/10.5281/zenodo.10106592" httpUri="https://zenodo.org/record/10106592/files/figure.png" pageId="2" pageNumber="775">Figs. 56</figureCitation>
).
</paragraph>
<paragraph id="8BFAB01EC137FFF2AA99FCBBFBCAFC53" blockId="2.[707,1194,194,1729]" pageId="2" pageNumber="775">
<emphasis id="B9316C0CC137FFF2AA99FCBBFCD4FC00" bold="true" box="[731,825,885,906]" pageId="2" pageNumber="775">
<typeStatus id="54FE0EBCC137FFF2AA99FCBBFCD8FC00" box="[731,821,885,906]" pageId="2" pageNumber="775" type="allotype">Allotype</typeStatus>
.
</emphasis>
Female. Length
<quantity id="4CBD1DFBC137FFF2ABB9FCBBFBB3FC00" box="[1019,1118,885,906]" metricMagnitude="-2" metricUnit="m" metricValue="2.19" pageId="2" pageNumber="775" unit="mm" value="21.9">21.9 mm</quantity>
; width across humeri
<quantity id="4CBD1DFBC137FFF2AB0BFC41FC4DFC2E" box="[841,928,911,932]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="2" pageNumber="775" unit="mm" value="10.4">10.4 mm</quantity>
. The
<typeStatus id="54FE0EBCC137FFF2AB90FC41FBF0FC2E" box="[978,1053,911,932]" pageId="2" pageNumber="775" type="allotype">allotype</typeStatus>
does not differ significantly from the
<typeStatus id="54FE0EBCC137FFF2ABD7FC64FC04FC35" box="[917,1001,938,959]" pageId="2" pageNumber="775" type="holotype">holotype</typeStatus>
. The prosternal process is tapered into an elongate cone.
</paragraph>
<paragraph id="8BFAB01EC137FFF2AA99FC11FCBDFB24" blockId="2.[707,1194,194,1729]" pageId="2" pageNumber="775">
<emphasis id="B9316C0CC137FFF2AA99FC11FCD2FC7E" bold="true" box="[731,831,991,1012]" pageId="2" pageNumber="775">Variation.</emphasis>
<materialsCitation id="3B2DBA43C137FFF2AB0EFC11FCA1FB24" location="All" pageId="2" pageNumber="775" specimenCount="6" stateProvince="Espirito Santo" typeStatus="paratype">
Males (
<specimenCount id="9D437B97C137FFF2ABDBFC11FBFDFC7E" box="[921,1040,991,1012]" pageId="2" pageNumber="775" type="generic" typeStatus="paratypes">5 paratypes</specimenCount>
). Length
<quantity id="4CBD1DFBC137FFF2AC30FC11FCF0FB85" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.05" metricValueMin="1.95" pageId="2" pageNumber="775" unit="mm" value="20.0" valueMax="20.5" valueMin="19.5">19.5 20.5 mm</quantity>
; width across humeri
<quantity id="4CBD1DFBC137FFF2ABB7FC34FB94FB85" box="[1013,1145,1017,1039]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" metricValueMax="1.06" metricValueMin="0.98" pageId="2" pageNumber="775" unit="mm" value="10.2" valueMax="10.6" valueMin="9.8">9.810.6 mm</quantity>
. The male
<typeStatus id="54FE0EBCC137FFF2AABAFBDAFCBEFBA3" box="[760,851,1044,1065]" pageId="2" pageNumber="775" type="paratype">paratypes</typeStatus>
do not differ significantly from the
<typeStatus id="54FE0EBCC137FFF2AA81FBE1FCF4FBCE" box="[707,793,1071,1092]" pageId="2" pageNumber="775" type="holotype">holotype</typeStatus>
. The
<specimenCount id="9D437B97C137FFF2AB10FBE1FC0CFBCE" box="[850,993,1071,1092]" pageId="2" pageNumber="775" type="generic">two specimens</specimenCount>
from
<collectingRegion id="49817EFCC137FFF2AC62FBE1FB44FBCE" box="[1056,1193,1071,1092]" country="Brazil" name="Espirito Santo" pageId="2" pageNumber="775">Espírito Santo</collectingRegion>
have the pronotum and first broad elytral interval slightly more densely punctate.
<location id="8E9AE6C5C137FFF2AC67FBAAFBA9FBF3" LSID="urn:lsid:plazi:treatment:03EC0108C137FFF3A88AFBCAFDEDF985:8E9AE6C5C137FFF2AC67FBAAFBA9FBF3" box="[1061,1092,1124,1145]" name="All" pageId="2" pageNumber="775" stateProvince="Espirito Santo">All</location>
<typeStatus id="54FE0EBCC137FFF2AC0EFBAAFB45FBF3" box="[1100,1192,1124,1145]" pageId="2" pageNumber="775" type="paratype">paratypes</typeStatus>
have the second broad interval punctate on the posterior half
</materialsCitation>
.
</paragraph>
<paragraph id="8BFAB01EC137FFF2AA99FB7AFBE8FAE2" blockId="2.[707,1194,194,1729]" pageId="2" pageNumber="775">
<materialsCitation id="3B2DBA43C137FFF2AA99FB7AFBECFAE2" location="Rio de Janeiro" pageId="2" pageNumber="775" specimenCount="2" stateProvince="Rio de Janeiro" typeStatus="paratype">
Females (
<specimenCount id="9D437B97C137FFF2AB79FB7AFC5FFB43" box="[827,946,1204,1225]" pageId="2" pageNumber="775" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
). Length
<quantity id="4CBD1DFBC137FFF2AC50FB7AFB48FB43" box="[1042,1189,1203,1225]" metricMagnitude="-2" metricUnit="m" metricValue="2.1399999999999997" metricValueMax="2.16" metricValueMin="2.12" pageId="2" pageNumber="775" unit="mm" value="21.4" valueMax="21.6" valueMin="21.2">21.221.6 mm</quantity>
; width across humeri
<quantity id="4CBD1DFBC137FFF2ABDAFB00FBC1FB69" box="[920,1068,1230,1252]" metricMagnitude="-2" metricUnit="m" metricValue="1.035" metricValueMax="1.04" metricValueMin="1.03" pageId="2" pageNumber="775" unit="mm" value="10.35" valueMax="10.4" valueMin="10.3">10.310.4 mm</quantity>
. The female
<typeStatus id="54FE0EBCC137FFF2AA81FB27FCF3FB74" box="[707,798,1257,1278]" pageId="2" pageNumber="775" type="paratype">paratypes</typeStatus>
do not differ significantly from the
<typeStatus id="54FE0EBCC137FFF2AC3CFB27FD01FA92" pageId="2" pageNumber="775" type="allotype">allotype</typeStatus>
except that both have the prosternal process similar to that of the
<typeStatus id="54FE0EBCC137FFF2ABE7FAD0FBEDFAB9" box="[933,1024,1310,1331]" pageId="2" pageNumber="775" type="holotype">holotype</typeStatus>
, and the
<collectingRegion id="49817EFCC137FFF2AC26FAD0FCEBFAC7" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="775">Rio de Janeiro</collectingRegion>
specimen has the second broad elytral interval punctate on the posterior half
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35FE395C137FFF2AA99FAA0FCFFFA58" pageId="2" pageNumber="775" type="etymology">
<paragraph id="8BFAB01EC137FFF2AA99FAA0FCFFFA58" blockId="2.[707,1194,194,1729]" pageId="2" pageNumber="775">
<emphasis id="B9316C0CC137FFF2AA99FAA0FCA7FA09" bold="true" box="[731,842,1390,1411]" pageId="2" pageNumber="775">Etymology.</emphasis>
The specific epithet is derived from the Latin,
<emphasis id="B9316C0CC137FFF2AB67FA46FCB4FA17" box="[805,857,1416,1437]" italics="true" pageId="2" pageNumber="775">hasta</emphasis>
, meaning a spear, and is used here as
<taxonomicName id="4C45CB9DC137FFF2AA9CFA6CFCC0FA3D" authorityName="Ratcliffe" authorityYear="2015" box="[734,813,1442,1463]" class="Insecta" family="Scarabaeidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="775" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B9316C0CC137FFF2AA9CFA6CFCC0FA3D" box="[734,813,1442,1463]" italics="true" pageId="2" pageNumber="775">hastatus</emphasis>
</taxonomicName>
to indicate the spear-shaped parameres (
<figureCitation id="137EAC9BC137FFF2AA88FA73FCE8FA58" box="[714,773,1469,1490]" captionStart="Figs" captionStartId="2.[197,234,830,849]" captionTargetBox="[181,654,192,806]" captionTargetId="figure-614@2.[173,660,183,825]" captionTargetPageId="2" captionText="Figs.36. Stenocrates species, parameres. 3) S. howdeni; 4) S. laborator; 56) S. hastatus. Illustrations in Figs. 34 from Dechambre and Hardy (2004)." figureDoi="http://doi.org/10.5281/zenodo.10106592" httpUri="https://zenodo.org/record/10106592/files/figure.png" pageId="2" pageNumber="775">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35FE395C137FFF2AA99FA16FB47F9F8" pageId="2" pageNumber="775" type="distribution">
<paragraph id="8BFAB01EC137FFF2AA99FA16FC90F98D" blockId="2.[707,1194,194,1729]" pageId="2" pageNumber="775">
<emphasis id="B9316C0CC137FFF2AA99FA16FCB5FA67" bold="true" box="[731,856,1496,1517]" pageId="2" pageNumber="775">Distribution.</emphasis>
<taxonomicName id="4C45CB9DC137FFF2AB27FA19FBF2FA66" authorityName="Ratcliffe" authorityYear="2015" box="[869,1055,1495,1516]" class="Insecta" family="Scarabaeidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="775" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B9316C0CC137FFF2AB27FA19FBF2FA66" box="[869,1055,1495,1516]" italics="true" pageId="2" pageNumber="775">Stenocrates hastatus</emphasis>
</taxonomicName>
is known from southeastern
<collectingCountry id="F352F08EC137FFF2AB7FFA3CFC95F98D" box="[829,888,1522,1543]" name="Brazil" pageId="2" pageNumber="775">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="8BFAB01EC137FFF2AA99F9C3FB6EF9DD" blockId="2.[707,1194,194,1729]" pageId="2" pageNumber="775">
<emphasis id="B9316C0CC137FFF2AA99F9C3FC7DF9A8" bold="true" box="[731,912,1549,1570]" pageId="2" pageNumber="775">Locality Records.</emphasis>
<materialsCitation id="3B2DBA43C137FFF2ABDFF9C3FBAEF9B7" country="Brazil" location="ESPIRITO SANTO" pageId="2" pageNumber="775" specimenCount="1" stateProvince="Espirito Santo">
<emphasis id="B9316C0CC137FFF2ABDFF9C3FBDCF9A8" bold="true" box="[925,1073,1549,1570]" pageId="2" pageNumber="775">
<collectingCountry id="F352F08EC137FFF2ABDFF9C3FC13F9A8" box="[925,1022,1549,1570]" name="Brazil" pageId="2" pageNumber="775">BRAZIL</collectingCountry>
(9):
</emphasis>
<collectingRegion id="49817EFCC137FFF2AC7EF9C3FCFBF9B7" country="Brazil" name="Espirito Santo" pageId="2" pageNumber="775">ESPÍRITO SANTO</collectingRegion>
(2): Linhares (
<quantity id="4CBD1DFBC137FFF2ABFFF9E6FBEFF9B7" box="[957,1026,1576,1597]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="775" unit="km" value="40.0">40 km</quantity>
NW)
</materialsCitation>
.
<materialsCitation id="3B2DBA43C137FFF2AC10F9E6FB6DF9DD" collectorName="Rio das Ostras" country="Brazil" location="RIO DE JANEIRO" pageId="2" pageNumber="775" specimenCount="1" stateProvince="Rio de Janeiro">
<collectingRegion id="49817EFCC137FFF2AC10F9E6FCC5F9DD" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="775">RIO DE JANEIRO</collectingRegion>
(7): Guapimirim,
<collectorName id="26B0D5C8C137FFF2ABA4F98CFB6DF9DD" box="[998,1152,1602,1623]" pageId="2" pageNumber="775">Rio das Ostras</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFAB01EC137FFF2AA99F993FB47F9F8" blockId="2.[707,1194,194,1729]" box="[731,1194,1629,1650]" pageId="2" pageNumber="775">
<emphasis id="B9316C0CC137FFF2AA99F993FC54F9F8" bold="true" box="[731,953,1629,1650]" pageId="2" pageNumber="775">Temporal Distribution.</emphasis>
March (7), November (2).
</paragraph>
</subSubSection>
<subSubSection id="C35FE395C137FFF3AA99F9B9FDEDF985" lastPageId="3" lastPageNumber="776" pageId="2" pageNumber="775" type="diagnosis">
<paragraph id="8BFAB01EC137FFF3AA99F9B9FDEDF985" blockId="2.[707,1194,194,1729]" lastBlockId="3.[151,638,1291,1551]" lastPageId="3" lastPageNumber="776" pageId="2" pageNumber="775">
<emphasis id="B9316C0CC137FFF2AA99F9B9FCABF906" bold="true" box="[731,838,1655,1676]" pageId="2" pageNumber="775">Diagnosis.</emphasis>
<taxonomicName id="4C45CB9DC137FFF2AB11F9B9FBC7F906" authorityName="Ratcliffe" authorityYear="2015" box="[851,1066,1655,1676]" class="Insecta" family="Scarabaeidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="775" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B9316C0CC137FFF2AB11F9B9FBC7F906" box="[851,1066,1655,1676]" italics="true" pageId="2" pageNumber="775">Stenocrates hastatus</emphasis>
</taxonomicName>
will key to
<taxonomicName id="4C45CB9DC137FFF2AA81F95FFC55F92D" authority="Prell" authorityName="Prell" authorityYear="1937" box="[707,952,1681,1703]" class="Insecta" family="Scarabaeidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="775" phylum="Arthropoda" rank="species" species="celatus">
<emphasis id="B9316C0CC137FFF2AA81F95FFC6CF92C" box="[707,897,1681,1702]" italics="true" pageId="2" pageNumber="775">Stenocrates celatus</emphasis>
Prell
</taxonomicName>
in Endrödi (1985), and
<taxonomicName id="4C45CB9DC137FFF2AA81F962FCCEF94B" authorityName="Prell" authorityYear="1937" box="[707,803,1708,1729]" class="Insecta" family="Scarabaeidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="775" phylum="Arthropoda" rank="species" species="celatus">
<emphasis id="B9316C0CC137FFF2AA81F962FCCEF94B" box="[707,803,1708,1729]" italics="true" pageId="2" pageNumber="775">S. celatus</emphasis>
</taxonomicName>
is the only other species in the genus with a lanceolate apex of the parameres. The two species differ in that
<taxonomicName id="4C45CB9DC136FFF3A902FAEBFE48FAB0" authorityName="Ratcliffe" authorityYear="2015" box="[320,421,1317,1338]" class="Insecta" family="Scarabaeidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="776" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B9316C0CC136FFF3A902FAEBFE48FAB0" box="[320,421,1317,1338]" italics="true" pageId="3" pageNumber="776">S. hastatus</emphasis>
</taxonomicName>
has sparse micropunctures on the frons (with small, dense punctures in
<taxonomicName id="4C45CB9DC136FFF3A8D5FA94FF02FAE5" authorityName="Prell" authorityYear="1937" box="[151,239,1370,1391]" class="Insecta" family="Scarabaeidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="776" phylum="Arthropoda" rank="species" species="celatus">
<emphasis id="B9316C0CC136FFF3A8D5FA94FF02FAE5" box="[151,239,1370,1391]" italics="true" pageId="3" pageNumber="776">S. celatus</emphasis>
</taxonomicName>
), the pronotum has punctures mostly in the posterior and anterior angles (with large, moderately dense punctures on each lateral third in
<emphasis id="B9316C0CC136FFF3AA56FA5EFD93FA2F" box="[532,638,1424,1445]" italics="true" pageId="3" pageNumber="776">
<taxonomicName id="4C45CB9DC136FFF3AA56FA5EFD9FFA2F" authorityName="Prell" authorityYear="1937" box="[532,626,1424,1445]" class="Insecta" family="Scarabaeidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="776" phylum="Arthropoda" rank="species" species="celatus">S. celatus</taxonomicName>
),
</emphasis>
and the parameres are widest at about the middle (widest just below the middle in
<taxonomicName id="4C45CB9DC136FFF3A985FA0BFDF2FA50" authorityName="Prell" authorityYear="1937" box="[455,543,1477,1498]" class="Insecta" family="Scarabaeidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="776" phylum="Arthropoda" rank="species" species="celatus">
<emphasis id="B9316C0CC136FFF3A985FA0BFDF2FA50" box="[455,543,1477,1498]" italics="true" pageId="3" pageNumber="776">S. celatus</emphasis>
</taxonomicName>
) and with their apices narrowly lanceolate (broadly lanceolate in
<taxonomicName id="4C45CB9DC136FFF3A8F2FA34FEE7F985" authorityName="Prell" authorityYear="1937" box="[176,266,1530,1551]" class="Insecta" family="Scarabaeidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="776" phylum="Arthropoda" rank="species" species="celatus">
<emphasis id="B9316C0CC136FFF3A8F2FA34FEE7F985" box="[176,266,1530,1551]" italics="true" pageId="3" pageNumber="776">S. celatus</emphasis>
</taxonomicName>
) (compare
<figureCitation id="137EAC9BC136FFF3A934FA34FE5AF985" box="[374,439,1530,1551]" captionStart="Figs" captionStartId="2.[197,234,830,849]" captionTargetBox="[181,654,192,806]" captionTargetId="figure-614@2.[173,660,183,825]" captionTargetPageId="2" captionText="Figs.36. Stenocrates species, parameres. 3) S. howdeni; 4) S. laborator; 56) S. hastatus. Illustrations in Figs. 34 from Dechambre and Hardy (2004)." figureDoi="http://doi.org/10.5281/zenodo.10106592" httpUri="https://zenodo.org/record/10106592/files/figure.png" pageId="3" pageNumber="776">Figs. 5</figureCitation>
and
<figureCitation id="137EAC9BC136FFF3A9A4FA34FE1EF985" box="[486,499,1530,1551]" captionStart="Fig" captionStartId="3.[175,207,1215,1234]" captionTargetBox="[151,621,194,1191]" captionTargetId="figure-279@3.[151,638,194,1196]" captionTargetPageId="3" captionText="Fig. 7. Stenocrates celatus, parameres." figureDoi="http://doi.org/10.5281/zenodo.10106598" httpUri="https://zenodo.org/record/10106598/files/figure.png" pageId="3" pageNumber="776">7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>