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<document id="726020E679B021D5DD115780B844D99A" ID-CLB-Dataset="24989" ID-DOI="10.11646/zootaxa.4691.5.3" ID-GBIF-Dataset="64b46fbc-ecdd-4dde-96cf-548d9d74b08c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3527522" ID-ZooBank="B2156154-5C5B-4BC8-815D-8C3BC26E5C2A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1572857992512" checkinUser="plazi" docAuthor="Benedetti, Alípio R. &amp; Pinto-Da-Rocha, Ricardo" docDate="2019" docId="03B6A1005C2DFFC3E4E2FB6AFC31B1F5" docLanguage="en" docName="zootaxa.4691.5.3.pdf" docOrigin="Zootaxa 4691 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Progonyleptoidellus picinguaba Benedetti &amp; Pinto-Da-Rocha 2019, sp. nov." docType="treatment" docVersion="11" lastPageNumber="482" masterDocId="FF8FD9785C3CFFD6E475FFA3FF99B662" masterDocTitle="Description of two new species of Progonyleptoidellus (Opiliones: Gonyleptidae), with a cladistic analysis of the genus, an overview of relationships in the K 92 group, and taxonomic notes on Deltaspidium" masterLastPageNumber="490" masterPageNumber="461" pageNumber="478" updateTime="1698761920771" updateUser="ExternalLinkService">
<mods:mods id="2D8EA7F52E85AC47111782D5B43EE843" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="347C7118D1E25C15BB0F7CC7E290A790">Description of two new species of Progonyleptoidellus (Opiliones: Gonyleptidae), with a cladistic analysis of the genus, an overview of relationships in the K 92 group, and taxonomic notes on Deltaspidium</mods:title>
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<mods:namePart id="F61073E37F4C86598F1212E36FD1B432">Benedetti, Alípio R.</mods:namePart>
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<mods:namePart id="39B4DAA97DF9D19AFFACBC78A6958AA6">Pinto-Da-Rocha, Ricardo</mods:namePart>
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<mods:title id="D74A852267FAAD677789AC47A23ABDC3">Zootaxa</mods:title>
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<mods:date id="2C077C7D38776B7BE3EA2C6021D29FBF">2019</mods:date>
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<mods:number id="172047D0A7492AAF346481B938524596">2019-11-04</mods:number>
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<mods:classification id="4DF99C9045B5590D463B49F02137132B">journal article</mods:classification>
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<mods:identifier id="A9D1BE80F16F629E00B79FDB4EFF539F" type="DOI">10.11646/zootaxa.4691.5.3</mods:identifier>
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<treatment id="03B6A1005C2DFFC3E4E2FB6AFC31B1F5" ID-DOI="http://doi.org/10.5281/zenodo.5586305" ID-GBIF-Taxon="160126970" ID-Zenodo-Dep="5586305" LSID="urn:lsid:plazi:treatment:03B6A1005C2DFFC3E4E2FB6AFC31B1F5" httpUri="http://treatment.plazi.org/id/03B6A1005C2DFFC3E4E2FB6AFC31B1F5" lastPageId="21" lastPageNumber="482" pageId="17" pageNumber="478">
<subSubSection id="C305439D5C2DFFC7E4E2FB6AFDE5B286" box="[151,636,1225,1252]" pageId="17" pageNumber="478" type="nomenclature">
<paragraph id="8BA010165C2DFFC7E4E2FB6AFDE5B286" blockId="17.[151,636,1225,1281]" box="[151,636,1225,1252]" pageId="17" pageNumber="478">
<heading id="D0E8A77A5C2DFFC7E4E2FB6AFDE5B286" bold="true" box="[151,636,1225,1252]" fontSize="11" level="1" pageId="17" pageNumber="478" reason="1">
<emphasis id="B96BCC045C2DFFC7E4E2FB6AFDE5B286" bold="true" box="[151,636,1225,1252]" pageId="17" pageNumber="478">
<taxonomicName id="4C1F6B955C2DFFC7E4E2FB6AFD8EB286" ID-CoL="7YGQN" authority="Benedetti &amp; Pinto-Da-Rocha, 2019" authorityName="Benedetti &amp; Pinto-Da-Rocha" authorityYear="2019" box="[151,535,1225,1252]" class="Arachnida" family="Gonyleptidae" genus="Progonyleptoidellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="478" phylum="Arthropoda" rank="species" species="picinguaba" status="sp. nov.">
<emphasis id="B96BCC045C2DFFC7E4E2FB6AFD8EB286" bold="true" box="[151,535,1225,1252]" italics="true" pageId="17" pageNumber="478">Progonyleptoidellus picinguaba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258717F5C2DFFC7E66BFB69FDE5B286" box="[542,636,1226,1252]" pageId="17" pageNumber="478" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C305439D5C2DFFC7E4E2FB44FED3B363" box="[151,330,1254,1281]" pageId="17" pageNumber="478" type="description">
<paragraph id="8BA010165C2DFFC7E4E2FB44FED3B363" blockId="17.[151,636,1225,1281]" box="[151,330,1254,1281]" pageId="17" pageNumber="478">
(
<figureCitation id="13240C935C2DFFC7E4EAFB44FF76B363" box="[159,239,1255,1281]" captionStart="FIGURE 4" captionStartId="18.[151,250,1800,1825]" captionTargetBox="[168,1425,388,1769]" captionTargetId="figure@18.[151,1436,373,1780]" captionTargetPageId="18" captionText="FIGURE 4. Progonyleptoidellus picinguaba sp. nov., male holotype (MZSP 73602): A, habitus, dorsal view; B, right trochanter–tibia IV, dorsal view; C, coxa, trochanter and basal part of femur IV, retrodorsal view. Scale bars: A, C: 2 mm, B: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3527530" httpUri="https://zenodo.org/record/3527530/files/figure.png" pageId="17" pageNumber="478">Figs. 4</figureCitation>
,
<figureCitation id="13240C935C2DFFC7E48EFB45FE92B363" box="[251,267,1254,1281]" captionStart="FIGURE 5" captionStartId="19.[151,250,1915,1940]" captionTargetBox="[157,1429,174,1887]" captionTargetId="figure@19.[151,1436,174,1896]" captionTargetPageId="19" captionText="FIGURE 5. Progonyleptoidellus picinguaba sp. nov., male paratype (MZSP 16009) and female paratype (MZSP 16257): A, male habitus, dorsal view; B, female habitus, dorsal view; C, male habitus, ventral view; D, male trochanter–tibia IV, dorsal view; E, female trochanter–patella IV, dorsal view; F, male habitus, left lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3527532" httpUri="https://zenodo.org/record/3527532/files/figure.png" pageId="17" pageNumber="478">5</figureCitation>
,
<figureCitation id="13240C935C2DFFC7E562FB44FEBEB363" box="[279,295,1255,1281]" captionStart="FIGURE 6" captionStartId="20.[151,250,1728,1753]" captionTargetBox="[151,1436,174,1709]" captionTargetId="figure@20.[151,1436,174,1709]" captionTargetPageId="20" captionText="FIGURE 6. Scanning electron micrographs showing the distal part of Progonyleptoidellus spp. penis: A–B, P. bocaina sp. nov. (MZSP 18939), A. dorsal, B. lateral; C–D, P. picinguaba sp. nov. (MZSP 16259), A. dorsal, B. lateral. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3527534" httpUri="https://zenodo.org/record/3527534/files/figure.png" pageId="17" pageNumber="478">6</figureCitation>
,
<figureCitation id="13240C935C2DFFC7E546FB44FEDBB363" box="[307,322,1255,1281]" captionStart="FIGURE 9" captionStartId="26.[151,250,1847,1872]" captionTargetBox="[156,1431,1034,1820]" captionTargetId="figure@26.[151,1436,1021,1827]" captionTargetPageId="26" captionText="FIGURE 9. Distribution of Progonyleptoidellus spp. in Coastal Atlantic Rain Forest of São Paulo and Rio de Janeiro States, Brazil. White symbols represent literature records." figureDoi="http://doi.org/10.5281/zenodo.3527538" httpUri="https://zenodo.org/record/3527538/files/figure.png" pageId="17" pageNumber="478">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C305439D5C2DFFC7E4E2FA88FC95B326" box="[151,780,1323,1348]" pageId="17" pageNumber="478" type="reference_group">
<paragraph id="8BA010165C2DFFC7E4E2FA88FC95B326" blockId="17.[151,780,1323,1348]" box="[151,780,1323,1348]" pageId="17" pageNumber="478">
<treatmentCitationGroup id="AB0F37385C2DFFC7E4E2FA88FC95B326" box="[151,780,1323,1348]" pageId="17" pageNumber="478">
<taxonomicName id="4C1F6B955C2DFFC7E4E2FA88FEFDB321" ID-CoL="7PVPZ" authorityName="Piza" authorityYear="1940" box="[151,356,1323,1347]" class="Arachnida" family="Gonyleptidae" genus="Progonyleptoidellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="478" phylum="Arthropoda" rank="genus">
<emphasis id="B96BCC045C2DFFC7E4E2FA88FEFDB321" box="[151,356,1323,1347]" italics="true" pageId="17" pageNumber="478">Progonyleptoidellus</emphasis>
</taxonomicName>
sp.:
<treatmentCitation id="0ABE36075C2DFFC7E5E6FA88FD50B326" author="Almeida-Neto, M. &amp; Machado, G. &amp; Pinto-da-Rocha, R. &amp; Giaretta, A. A." box="[403,713,1323,1348]" page="367" pageId="17" pageNumber="478" year="2006">
<bibRefCitation id="EF8E6DE75C2DFFC7E5E6FA88FD50B326" author="Almeida-Neto, M. &amp; Machado, G. &amp; Pinto-da-Rocha, R. &amp; Giaretta, A. A." box="[403,713,1323,1348]" pageId="17" pageNumber="478" pagination="361 - 375" refId="ref19632" refString="Almeida-Neto, M., Machado, G., Pinto-da-Rocha, R. &amp; Giaretta, A. A. (2006) Harvestman (Arachnida: Opiliones) species distribution along three Neotropical elevational gradients: an alternative rescue effect to explain Rapoport's rule? Journal of Biogeography, 33, 361 - 375." type="journal article" year="2006">
Almeida-Neto
<emphasis id="B96BCC045C2DFFC7E65FFA8FFDCEB321" box="[554,599,1323,1347]" italics="true" pageId="17" pageNumber="478">et al</emphasis>
, 2006: 367
</bibRefCitation>
</treatmentCitation>
(dist).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C305439D5C2DFFC7E4E2FAD1FF77B07F" pageId="17" pageNumber="478" type="materials_examined">
<paragraph id="8BA010165C2DFFC7E4E2FAD1FF77B07F" blockId="17.[151,1437,1393,2032]" pageId="17" pageNumber="478">
<emphasis id="B96BCC045C2DFFC7E4E2FAD1FEA4B3EE" bold="true" box="[151,317,1393,1420]" pageId="17" pageNumber="478">Type material</emphasis>
.
<materialsCitation id="3B771A4B5C2DFFC7E53EFAD1FE35B3D3" ID-GBIF-Occurrence="2446790892" collectingDate="1995-12-21" collectionCode="MZSP" collectorName="G. Machado" country="Brazil" county="Ubatuba" location="Morro do Cuscuzeiro" municipality="Picinguaba" pageId="17" pageNumber="478" specimenCode="MZSP 73601" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="holotype">
<collectingCountry id="F30850865C2DFFC7E53EFAD1FE2DB3EE" box="[331,436,1394,1420]" name="Brazil" pageId="17" pageNumber="478">BRAZIL</collectingCountry>
.
<emphasis id="B96BCC045C2DFFC7E5CBFAD1FDABB3EE" box="[446,562,1394,1420]" italics="true" pageId="17" pageNumber="478">
<collectingRegion id="49DBDEF45C2DFFC7E5CBFAD1FDABB3EE" box="[446,562,1394,1420]" country="Brazil" name="Sao Paulo" pageId="17" pageNumber="478">São Paulo</collectingRegion>
</emphasis>
.
<collectingCounty id="62C1689A5C2DFFC7E64AFAD1FD38B3EE" box="[575,673,1394,1420]" pageId="17" pageNumber="478">Ubatuba</collectingCounty>
,
<collectingMunicipality id="6BC48A6C5C2DFFC7E6D9FAD1FCB2B3EE" box="[684,811,1394,1420]" pageId="17" pageNumber="478">Picinguaba</collectingMunicipality>
,
<location id="8EC046CD5C2DFFC7E743FAD1FBB1B3EE" LSID="urn:lsid:plazi:treatment:03B6A1005C2DFFC3E4E2FB6AFC31B1F5:8EC046CD5C2DFFC7E743FAD1FBB1B3EE" box="[822,1064,1394,1420]" country="Brazil" county="Ubatuba" municipality="Picinguaba" name="Morro do Cuscuzeiro" pageId="17" pageNumber="478" stateProvince="Sao Paulo">Morro do Cuscuzeiro</location>
,
<date id="FFA136D65C2DFFC7E047FAD2FB2CB3EE" box="[1074,1205,1393,1420]" pageId="17" pageNumber="478" value="1995-12-21">
<collectingDate id="EFE5CF3E5C2DFFC7E047FAD2FB2CB3EE" box="[1074,1205,1393,1420]" pageId="17" pageNumber="478" value="1995-12-21">21.xii.1995</collectingDate>
</date>
,
<collectorName id="26EA75C05C2DFFC7E0B5FAD1FAD0B3EE" box="[1216,1353,1394,1420]" pageId="17" pageNumber="478">G. Machado</collectorName>
leg.,
<specimenCount id="9D19DB9F5C2DFFC7E1F2FAD1FA05B3EE" box="[1415,1436,1394,1420]" pageId="17" pageNumber="478" type="male">♂</specimenCount>
<typeStatus id="54A4AEB45C2DFFC7E4E2FA35FF60B3D2" box="[151,249,1430,1456]" pageId="17" pageNumber="478" type="holotype">holotype</typeStatus>
(
<specimenCode id="DBB9B86D5C2DFFC7E57DFA35FE3DB3D3" box="[264,420,1430,1457]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="17" pageNumber="478" type="Museum">MZSP 73601</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B771A4B5C2DFFC7E5CDFA35FC91B3D2" ID-GBIF-Occurrence="2446790890" box="[440,776,1429,1456]" collectingDate="1995-12-21" collectionCode="MZSP" collectorName="G. Machado" country="Brazil" county="Ubatuba" isTailEnumeration="true" location="Morro do Cuscuzeiro" municipality="Picinguaba" pageId="17" pageNumber="478" specimenCode="MZSP 16257" specimenCount="2" specimenCount-female="2" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D19DB9F5C2DFFC7E5CDFA35FE7BB3D2" box="[440,482,1430,1456]" pageId="17" pageNumber="478" type="female">2 ♀</specimenCount>
<typeStatus id="54A4AEB45C2DFFC7E59FFA35FDCCB3D2" box="[490,597,1430,1456]" pageId="17" pageNumber="478" type="paratype">paratypes</typeStatus>
(
<specimenCode id="DBB9B86D5C2DFFC7E611FA35FC98B3D2" box="[612,769,1429,1456]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="17" pageNumber="478" type="Museum">MZSP 16257</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B771A4B5C2DFFC7E760FA35FF70B3B6" ID-GBIF-Occurrence="2446790882" collectingDate="1995-12-20" collectionCode="MZSP" collectorName="G. Machado" country="Brazil" pageId="17" pageNumber="478" specimenCode="MZSP 16258" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Sao Paulo" typeStatus="paratype">
same loc.,
<date id="FFA136D65C2DFFC7E7FBFA36FB96B3D2" box="[910,1039,1429,1456]" pageId="17" pageNumber="478" value="1995-12-20">
<collectingDate id="EFE5CF3E5C2DFFC7E7FBFA36FB96B3D2" box="[910,1039,1429,1456]" pageId="17" pageNumber="478" value="1995-12-20">20.xii.1995</collectingDate>
</date>
,
<collectorName id="26EA75C05C2DFFC7E069FA35FB3EB3D2" box="[1052,1191,1430,1456]" pageId="17" pageNumber="478">G. Machado</collectorName>
leg.,
<specimenCount id="9D19DB9F5C2DFFC7E093FA35FA88B3D2" box="[1254,1297,1430,1456]" pageId="17" pageNumber="478" type="male">3 ♂</specimenCount>
<specimenCount id="9D19DB9F5C2DFFC7E16CFA35FADAB3D2" box="[1305,1347,1430,1456]" pageId="17" pageNumber="478" type="female">4 ♀</specimenCount>
(
<specimenCode id="DBB9B86D5C2DFFC7E127FA35FF78B3B6" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="17" pageNumber="478" type="Museum">MZSP 16258</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B771A4B5C2DFFC7E480FA19FD9CB3B6" ID-GBIF-Occurrence="2446790871" box="[245,517,1465,1492]" collectingDate="1995-12-20" collectionCode="MZSP" collectorName="G. Machado" country="Brazil" isTailEnumeration="true" pageId="17" pageNumber="478" specimenCode="MZSP 16259" specimenCount="10" specimenCount-female="7" specimenCount-male="3" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D19DB9F5C2DFFC7E480FA19FEB9B3B6" box="[245,288,1466,1492]" pageId="17" pageNumber="478" type="male">3 ♂</specimenCount>
<specimenCount id="9D19DB9F5C2DFFC7E552FA19FECBB3B6" box="[295,338,1466,1492]" pageId="17" pageNumber="478" type="female">7 ♀</specimenCount>
(
<specimenCode id="DBB9B86D5C2DFFC7E514FA19FE64B3B6" box="[353,509,1465,1492]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="17" pageNumber="478" type="Museum">MZSP 16259</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B771A4B5C2DFFC7E664FA19FB4CB3B7" ID-GBIF-Occurrence="2446790870" box="[529,1237,1465,1493]" collectingDate="1991-04-25" collectingDateMax="1991-04-29" collectingDateMin="1991-04-25" collectionCode="MZSP" collectorName="P. Gnaspini" country="Brazil" pageId="17" pageNumber="478" specimenCode="MZSP 16009" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
same loc.,
<date id="FFA136D65C2DFFC7E6FFFA1AFCB5B3B6" box="[650,812,1465,1492]" pageId="17" pageNumber="478" value="1991-04-25" valueMax="1991-04-29" valueMin="1991-04-25">
<collectingDate id="EFE5CF3E5C2DFFC7E6FFFA1AFCB5B3B6" box="[650,812,1465,1492]" pageId="17" pageNumber="478" value="1991-04-25" valueMax="1991-04-29" valueMin="1991-04-25">25–29.iv.1991</collectingDate>
</date>
,
<collectorName id="26EA75C05C2DFFC7E74DFA19FC21B3B6" box="[824,952,1466,1492]" pageId="17" pageNumber="478">P. Gnaspini</collectorName>
leg.,
<specimenCount id="9D19DB9F5C2DFFC7E782FA19FBBBB3B6" box="[1015,1058,1466,1492]" pageId="17" pageNumber="478" type="male">1 ♂</specimenCount>
(
<specimenCode id="DBB9B86D5C2DFFC7E044FA19FB54B3B7" box="[1073,1229,1466,1493]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="17" pageNumber="478" type="Museum">MZSP 16009</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B771A4B5C2DFFC7E094FA19FF73B07F" ID-GBIF-Occurrence="2446790872" collectingDate="2008-03-14" collectionCode="MZSP" collectorName="F. Esteves &amp; R. Feitosa" country="Brazil" county="Ubatuba" elevation="600" location="Picinguaba" municipality="Parque Estadual Serra do Mar" pageId="17" pageNumber="478" specimenCode="MZSP 30176" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectingCounty id="62C1689A5C2DFFC7E094FA19FADAB3B6" box="[1249,1347,1466,1492]" pageId="17" pageNumber="478">Ubatuba</collectingCounty>
,
<collectingMunicipality id="6BC48A6C5C2DFFC7E13AFA19FE00B39A" pageId="17" pageNumber="478">Parque Estadual Serra do Mar</collectingMunicipality>
, núcleo
<location id="8EC046CD5C2DFFC7E58FFA7DFDE0B39A" LSID="urn:lsid:plazi:treatment:03B6A1005C2DFFC3E4E2FB6AFC31B1F5:8EC046CD5C2DFFC7E58FFA7DFDE0B39A" box="[506,633,1502,1528]" country="Brazil" county="Ubatuba" municipality="Parque Estadual Serra do Mar" name="Picinguaba" pageId="17" pageNumber="478" stateProvince="Sao Paulo">Picinguaba</location>
,
<quantity id="4CE7BDF35C2DFFC7E6F3FA7DFD51B39A" box="[646,712,1502,1528]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="478" unit="m" value="600.0">
<elevation id="0032F7255C2DFFC7E6F3FA7DFD51B39A" box="[646,712,1502,1528]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="478" unit="m" value="600.0">600m</elevation>
</quantity>
,
<date id="FFA136D65C2DFFC7E6A0FA7DFCC8B39A" box="[725,849,1502,1528]" pageId="17" pageNumber="478" value="2008-03-14">
<collectingDate id="EFE5CF3E5C2DFFC7E6A0FA7DFCC8B39A" box="[725,849,1502,1528]" pageId="17" pageNumber="478" value="2008-03-14">14.iii.2008</collectingDate>
</date>
,
<collectorName id="26EA75C05C2DFFC7E72BFA7DFC48B39A" box="[862,977,1502,1528]" pageId="17" pageNumber="478">F. Esteves</collectorName>
&amp;
<collectorName id="26EA75C05C2DFFC7E78DFA7DFBF5B39A" box="[1016,1132,1502,1528]" pageId="17" pageNumber="478">R. Feitosa</collectorName>
leg.,
<specimenCount id="9D19DB9F5C2DFFC7E0D8FA7DFB40B39A" box="[1197,1241,1502,1528]" pageId="17" pageNumber="478" type="female">1 ♀</specimenCount>
<typeStatus id="54A4AEB45C2DFFC7E097FA7DFADBB39A" box="[1250,1346,1502,1528]" pageId="17" pageNumber="478" type="paratype">paratype</typeStatus>
(
<specimenCode id="DBB9B86D5C2DFFC7E127FA7DFF7BB07F" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="17" pageNumber="478" type="Museum">MZSP 30176</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C305439D5C2DFFC7E4B2F985FEFBB006" pageId="17" pageNumber="478" type="etymology">
<paragraph id="8BA010165C2DFFC7E4B2F985FEFBB006" blockId="17.[151,1437,1393,2032]" pageId="17" pageNumber="478">
<emphasis id="B96BCC045C2DFFC7E4B2F985FED7B022" bold="true" box="[199,334,1574,1600]" pageId="17" pageNumber="478">Etymology.</emphasis>
The name, a noun in apposition, refers to the
<typeStatus id="54A4AEB45C2DFFC7E73FF985FCE3B022" box="[842,890,1574,1600]" pageId="17" pageNumber="478">type</typeStatus>
locality, a region in the municipality of Ubatuba,
<collectingRegion id="49DBDEF45C2DFFC7E4E2F9E9FE92B006" box="[151,267,1610,1636]" country="Brazil" name="Sao Paulo" pageId="17" pageNumber="478">São Paulo</collectingRegion>
,
<collectingCountry id="F30850865C2DFFC7E563F9E9FEC7B006" box="[278,350,1610,1636]" name="Brazil" pageId="17" pageNumber="478">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C305439D5C2DFFC7E4B2F9CDFC6BB0CE" pageId="17" pageNumber="478" type="materials_examined">
<paragraph id="8BA010165C2DFFC7E4B2F9CDFBD0B0EA" blockId="17.[151,1437,1393,2032]" box="[199,1097,1645,1672]" pageId="17" pageNumber="478">
<materialsCitation id="3B771A4B5C2DFFC7E4B2F9CDFBD0B0EA" ID-GBIF-Occurrence="2446790891" box="[199,1097,1645,1672]" country="Brazil" county="Ubatuba" location="Morro do Cuscuzeiro" municipality="Picinguaba" pageId="17" pageNumber="478" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<emphasis id="B96BCC045C2DFFC7E4B2F9CDFEFFB0EA" bold="true" box="[199,358,1645,1672]" pageId="17" pageNumber="478">
<typeStatus id="54A4AEB45C2DFFC7E4B2F9CDFE9BB0EA" box="[199,258,1646,1672]" pageId="17" pageNumber="478">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F30850865C2DFFC7E519F9CDFE4CB0EA" box="[364,469,1646,1672]" name="Brazil" pageId="17" pageNumber="478">BRAZIL</collectingCountry>
.
<emphasis id="B96BCC045C2DFFC7E5AAF9CDFDCAB0EA" box="[479,595,1646,1672]" italics="true" pageId="17" pageNumber="478">
<collectingRegion id="49DBDEF45C2DFFC7E5AAF9CDFDCAB0EA" box="[479,595,1646,1672]" country="Brazil" name="Sao Paulo" pageId="17" pageNumber="478">São Paulo</collectingRegion>
</emphasis>
.
<collectingCounty id="62C1689A5C2DFFC7E614F9CDFD5DB0EA" box="[609,708,1646,1672]" pageId="17" pageNumber="478">Ubatuba</collectingCounty>
:
<collectingMunicipality id="6BC48A6C5C2DFFC7E6BAF9CDFCD7B0EA" box="[719,846,1646,1672]" pageId="17" pageNumber="478">Picinguaba</collectingMunicipality>
,
<location id="8EC046CD5C2DFFC7E72CF9CDFBD0B0EA" LSID="urn:lsid:plazi:treatment:03B6A1005C2DFFC3E4E2FB6AFC31B1F5:8EC046CD5C2DFFC7E72CF9CDFBD0B0EA" box="[857,1097,1646,1672]" country="Brazil" county="Ubatuba" municipality="Picinguaba" name="Morro do Cuscuzeiro" pageId="17" pageNumber="478" stateProvince="Sao Paulo">Morro do Cuscuzeiro</location>
</materialsCitation>
</paragraph>
<paragraph id="8BA010165C2DFFC7E4B2F932FC6BB0CE" blockId="17.[151,1437,1393,2032]" box="[199,1010,1681,1708]" pageId="17" pageNumber="478">
<emphasis id="B96BCC045C2DFFC7E4B2F932FE67B0CE" bold="true" box="[199,510,1681,1708]" pageId="17" pageNumber="478">Geographical distribution</emphasis>
(
<figureCitation id="13240C935C2DFFC7E678F931FDC8B0CE" box="[525,593,1682,1708]" captionStart="FIGURE 9" captionStartId="26.[151,250,1847,1872]" captionTargetBox="[156,1431,1034,1820]" captionTargetId="figure@26.[151,1436,1021,1827]" captionTargetPageId="26" captionText="FIGURE 9. Distribution of Progonyleptoidellus spp. in Coastal Atlantic Rain Forest of São Paulo and Rio de Janeiro States, Brazil. White symbols represent literature records." figureDoi="http://doi.org/10.5281/zenodo.3527538" httpUri="https://zenodo.org/record/3527538/files/figure.png" pageId="17" pageNumber="478">Fig. 9</figureCitation>
). Known only from the
<typeStatus id="54A4AEB45C2DFFC7E72AF931FC16B0CE" box="[863,911,1682,1708]" pageId="17" pageNumber="478">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C305439D5C2DFFC7E4B2F916FAF7B15E" pageId="17" pageNumber="478" type="diagnosis">
<paragraph id="8BA010165C2DFFC7E4B2F916FAF7B15E" blockId="17.[151,1437,1393,2032]" pageId="17" pageNumber="478">
<emphasis id="B96BCC045C2DFFC7E4B2F916FEDBB0B2" bold="true" box="[199,322,1717,1744]" pageId="17" pageNumber="478">Diagnosis.</emphasis>
Males.
<taxonomicName id="4C1F6B955C2DFFC7E5E9F915FC9AB0B2" authority="Benedetti &amp; Pinto-Da-Rocha, 2019" authorityName="Benedetti &amp; Pinto-Da-Rocha" authorityYear="2019" box="[412,771,1718,1744]" class="Arachnida" family="Gonyleptidae" genus="Progonyleptoidellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="478" phylum="Arthropoda" rank="species" species="picinguaba" status="sp. nov.">
<emphasis id="B96BCC045C2DFFC7E5E9F915FC9AB0B2" box="[412,771,1718,1744]" italics="true" pageId="17" pageNumber="478">Progonyleptoidellus picinguaba</emphasis>
</taxonomicName>
<emphasis id="B96BCC045C2DFFC7E77EF915FCFCB0B2" bold="true" box="[779,869,1718,1744]" pageId="17" pageNumber="478">
<taxonomicNameLabel id="A258717F5C2DFFC7E77EF915FCFCB0B2" box="[779,869,1718,1744]" pageId="17" pageNumber="478" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from the other species of the genus by the following combination of characters: Free tergite III armed with a long spine; proapical apophysis on coxa IV straight, without curved apex; femur III unarmed; femur IV without DBA. Females. This species differs from the other species of the genus in the absence of sexual dimorphism on armature of coxa IV and femur IV.
</paragraph>
</subSubSection>
<subSubSection id="C305439D5C2DFFC3E4B2F8E6FC31B1F5" lastPageId="21" lastPageNumber="482" pageId="17" pageNumber="478" type="description">
<paragraph id="8BA010165C2DFFC7E4B2F8E6FD28B103" blockId="17.[151,1437,1393,2032]" box="[199,689,1861,1889]" pageId="17" pageNumber="478">
<emphasis id="B96BCC045C2DFFC7E4B2F8E6FEC0B102" bold="true" box="[199,345,1861,1888]" pageId="17" pageNumber="478">Description.</emphasis>
Male
<typeStatus id="54A4AEB45C2DFFC7E5D5F8E5FD9BB102" box="[416,514,1862,1888]" pageId="17" pageNumber="478" type="holotype">holotype</typeStatus>
(MZSP 73601)
</paragraph>
<paragraph id="8BA010165C2DFFC7E4B2F8C8FB7BB1E6" blockId="17.[151,1437,1393,2032]" box="[199,1250,1897,1924]" pageId="17" pageNumber="478">
<emphasis id="B96BCC045C2DFFC7E4B2F8C8FEEAB1E6" box="[199,371,1899,1924]" italics="true" pageId="17" pageNumber="478">Measurements:</emphasis>
CL 2.2; CW 3.2; DSL 4.3; DSW 5.1; FeP 3.7; FI 4.5; FII 12; FIII 8; FIV 10.6.
</paragraph>
<paragraph id="8BA010165C2DFFC4E4B2F82CFC59B705" blockId="17.[151,1437,1393,2032]" lastBlockId="18.[151,1437,153,359]" lastPageId="18" lastPageNumber="479" pageId="17" pageNumber="478">
<emphasis id="B96BCC045C2DFFC7E4B2F82CFEB8B1CA" box="[199,289,1935,1960]" italics="true" pageId="17" pageNumber="478">Dorsum</emphasis>
(
<figureCitation id="13240C935C2DFFC7E546F82DFE03B1CA" box="[307,410,1934,1960]" captionStart="FIGURE 4" captionStartId="18.[151,250,1800,1825]" captionTargetBox="[168,1425,388,1769]" captionTargetId="figure@18.[151,1436,373,1780]" captionTargetPageId="18" captionText="FIGURE 4. Progonyleptoidellus picinguaba sp. nov., male holotype (MZSP 73602): A, habitus, dorsal view; B, right trochanter–tibia IV, dorsal view; C, coxa, trochanter and basal part of femur IV, retrodorsal view. Scale bars: A, C: 2 mm, B: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3527530" httpUri="https://zenodo.org/record/3527530/files/figure.png" pageId="17" pageNumber="478">Figs. 4A</figureCitation>
,
<figureCitation id="13240C935C2DFFC7E5DDF82EFE74B1CA" box="[424,493,1933,1960]" captionStart="FIGURE 5" captionStartId="19.[151,250,1915,1940]" captionTargetBox="[157,1429,174,1887]" captionTargetId="figure@19.[151,1436,174,1896]" captionTargetPageId="19" captionText="FIGURE 5. Progonyleptoidellus picinguaba sp. nov., male paratype (MZSP 16009) and female paratype (MZSP 16257): A, male habitus, dorsal view; B, female habitus, dorsal view; C, male habitus, ventral view; D, male trochanter–tibia IV, dorsal view; E, female trochanter–patella IV, dorsal view; F, male habitus, left lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3527532" httpUri="https://zenodo.org/record/3527532/files/figure.png" pageId="17" pageNumber="478">5A, F</figureCitation>
): DS shape
<typeStatus id="54A4AEB45C2DFFC7E60EF82DFD32B1CA" box="[635,683,1934,1960]" pageId="17" pageNumber="478">type</typeStatus>
gamma-P, with two median acuminate tubercles on the corner of its anterior margin. Frontal hump low, slightly projected anteriorly, with two upwards-pointing median acuminate tubercles. Ocularium narrow, low, with two parallel acuminate tubercles upwards and one pair of small tubercles posteriorly. Carapace with a pair of rounded tubercles posterior to the ocularium and a few scattered granules. Scutal areas I–II each with a paramedian pair of tiny tubercles; scutal area III with a paramedian pair of large spines directed posteriorly, and scattered granules. Lateral margins of DS with 2–3 small tubercles and one median spine on posterior 1/3. Two ozopores. Posterior margin of DS straight with prominent corners, with a row of small tubercles (two median ones larger). Free tergites I–III each with a row of varied-sized tubercles, predominantly small; free tergites II–III each with a large median spine. Anal operculum unarmed.
</paragraph>
<caption id="DF60409E5C2EFFC4E4E2F8ABFAEFB126" ID-DOI="http://doi.org/10.5281/zenodo.3527530" ID-Zenodo-Dep="3527530" httpUri="https://zenodo.org/record/3527530/files/figure.png" pageId="18" pageNumber="479" startId="18.[151,250,1800,1825]" targetBox="[168,1425,388,1769]" targetPageId="18">
<paragraph id="8BA010165C2EFFC4E4E2F8ABFAEFB126" blockId="18.[151,1436,1800,1861]" pageId="18" pageNumber="479">
<emphasis id="B96BCC045C2EFFC4E4E2F8ABFE8BB143" bold="true" box="[151,274,1800,1825]" pageId="18" pageNumber="479">FIGURE 4.</emphasis>
<taxonomicName id="4C1F6B955C2EFFC4E56DF8AAFDC4B143" authority="Benedetti &amp; Pinto-Da-Rocha, 2019" authorityName="Benedetti &amp; Pinto-Da-Rocha" authorityYear="2019" box="[280,605,1801,1825]" class="Arachnida" family="Gonyleptidae" genus="Progonyleptoidellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="479" phylum="Arthropoda" rank="species" species="picinguaba" status="sp. nov.">
<emphasis id="B96BCC045C2EFFC4E56DF8AAFDC4B143" box="[280,605,1801,1825]" italics="true" pageId="18" pageNumber="479">Progonyleptoidellus picinguaba</emphasis>
</taxonomicName>
<emphasis id="B96BCC045C2EFFC4E616F8AAFD2AB143" bold="true" box="[611,691,1801,1825]" pageId="18" pageNumber="479">
<taxonomicNameLabel id="A258717F5C2EFFC4E616F8AAFD2AB143" box="[611,691,1801,1825]" pageId="18" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (MZSP 73602): A, habitus, dorsal view; B, right trochant-er–tibia IV, dorsal view; C, coxa, trochanter and basal part of femur IV, retrodorsal view. Scale bars: A, C: 2 mm, B: 3 mm.
</paragraph>
</caption>
<caption id="DF60409E5C2FFFC5E4E2F8D8FB07B1B9" ID-DOI="http://doi.org/10.5281/zenodo.3527532" ID-Zenodo-Dep="3527532" httpUri="https://zenodo.org/record/3527532/files/figure.png" pageId="19" pageNumber="480" startId="19.[151,250,1915,1940]" targetBox="[157,1429,174,1887]" targetPageId="19">
<paragraph id="8BA010165C2FFFC5E4E2F8D8FB07B1B9" blockId="19.[151,1436,1915,2012]" pageId="19" pageNumber="480">
<emphasis id="B96BCC045C2FFFC5E4E2F8D8FE8DB1F6" bold="true" box="[151,276,1915,1940]" pageId="19" pageNumber="480">FIGURE 5.</emphasis>
<taxonomicName id="4C1F6B955C2FFFC5E569F8DFFDFBB1F6" authority="Benedetti &amp; Pinto-Da-Rocha, 2019" authorityName="Benedetti &amp; Pinto-Da-Rocha" authorityYear="2019" box="[284,610,1916,1940]" class="Arachnida" family="Gonyleptidae" genus="Progonyleptoidellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="19" pageNumber="480" phylum="Arthropoda" rank="species" species="picinguaba" status="sp. nov.">
<emphasis id="B96BCC045C2FFFC5E569F8DFFDFBB1F6" box="[284,610,1916,1940]" italics="true" pageId="19" pageNumber="480">Progonyleptoidellus picinguaba</emphasis>
</taxonomicName>
<emphasis id="B96BCC045C2FFFC5E61FF8DFFD25B1F6" bold="true" box="[618,700,1916,1940]" pageId="19" pageNumber="480">
<taxonomicNameLabel id="A258717F5C2FFFC5E61FF8DFFD25B1F6" box="[618,700,1916,1940]" pageId="19" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype (MZSP 16009) and female paratype (MZSP 16257): A, male habitus, dorsal view; B, female habitus, dorsal view; C, male habitus, ventral view; D, male trochanter–tibia IV, dorsal view; E, female trochanter–patella IV, dorsal view; F, male habitus, left lateral view. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="DF60409E5C28FFC2E4E2F963FAF4B09F" ID-DOI="http://doi.org/10.5281/zenodo.3527534" ID-Zenodo-Dep="3527534" httpUri="https://zenodo.org/record/3527534/files/figure.png" pageId="20" pageNumber="481" startId="20.[151,250,1728,1753]" targetBox="[151,1436,174,1709]" targetPageId="20">
<paragraph id="8BA010165C28FFC2E4E2F963FAF4B09F" blockId="20.[151,1436,1728,1789]" pageId="20" pageNumber="481">
<emphasis id="B96BCC045C28FFC2E4E2F963FE8BB0BB" bold="true" box="[151,274,1728,1753]" pageId="20" pageNumber="481">FIGURE 6.</emphasis>
Scanning electron micrographs showing the distal part of
<taxonomicName id="4C1F6B955C28FFC2E728F963FBB3B0BA" authorityName="Piza" authorityYear="1940" box="[861,1066,1728,1752]" class="Arachnida" family="Gonyleptidae" genus="Progonyleptoidellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="20" pageNumber="481" phylum="Arthropoda" rank="genus">
<emphasis id="B96BCC045C28FFC2E728F963FBB3B0BA" box="[861,1066,1728,1752]" italics="true" pageId="20" pageNumber="481">Progonyleptoidellus</emphasis>
</taxonomicName>
spp. penis: A–B,
<taxonomicName id="4C1F6B955C28FFC2E0A9F962FADFB0BA" authority="Benedetti &amp; Pinto-Da-Rocha, 2019" authorityName="Benedetti &amp; Pinto-Da-Rocha" authorityYear="2019" box="[1244,1350,1728,1752]" class="Arachnida" family="Gonyleptidae" genus="Progonyleptoidellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="20" pageNumber="481" phylum="Arthropoda" rank="species" species="bocaina" status="sp. nov.">
<emphasis id="B96BCC045C28FFC2E0A9F962FADFB0BA" box="[1244,1350,1728,1752]" italics="true" pageId="20" pageNumber="481">P. bocaina</emphasis>
</taxonomicName>
<emphasis id="B96BCC045C28FFC2E139F963FA05B0BA" bold="true" box="[1356,1436,1728,1752]" pageId="20" pageNumber="481">
<taxonomicNameLabel id="A258717F5C28FFC2E139F963FA05B0BA" box="[1356,1436,1728,1752]" pageId="20" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MZSP 18939), A. dorsal, B. lateral; C–D,
<taxonomicName id="4C1F6B955C28FFC2E63EF946FD4CB09E" authority="Benedetti &amp; Pinto-Da-Rocha, 2019" authorityName="Benedetti &amp; Pinto-Da-Rocha" authorityYear="2019" box="[587,725,1764,1788]" class="Arachnida" family="Gonyleptidae" genus="Progonyleptoidellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="20" pageNumber="481" phylum="Arthropoda" rank="species" species="picinguaba" status="sp. nov.">
<emphasis id="B96BCC045C28FFC2E63EF946FD4CB09E" box="[587,725,1764,1788]" italics="true" pageId="20" pageNumber="481">P. picinguaba</emphasis>
</taxonomicName>
<emphasis id="B96BCC045C28FFC2E6A9F947FCB5B09E" bold="true" box="[732,812,1764,1788]" pageId="20" pageNumber="481">
<taxonomicNameLabel id="A258717F5C28FFC2E6A9F947FCB5B09E" box="[732,812,1764,1788]" pageId="20" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MZSP 16259), A. dorsal, B. lateral. Scale bars: 20 µm.
</paragraph>
</caption>
<paragraph id="8BA010165C28FFC2E4B2F88FFB28B1EF" blockId="20.[151,1437,1834,2005]" pageId="20" pageNumber="481">
<emphasis id="B96BCC045C28FFC2E4B2F88FFE96B127" box="[199,271,1836,1861]" italics="true" pageId="20" pageNumber="481">Venter</emphasis>
(
<figureCitation id="13240C935C28FFC2E568F888FEEFB127" box="[285,374,1834,1861]" captionStart="FIGURE 5" captionStartId="19.[151,250,1915,1940]" captionTargetBox="[157,1429,174,1887]" captionTargetId="figure@19.[151,1436,174,1896]" captionTargetPageId="19" captionText="FIGURE 5. Progonyleptoidellus picinguaba sp. nov., male paratype (MZSP 16009) and female paratype (MZSP 16257): A, male habitus, dorsal view; B, female habitus, dorsal view; C, male habitus, ventral view; D, male trochanter–tibia IV, dorsal view; E, female trochanter–patella IV, dorsal view; F, male habitus, left lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3527532" httpUri="https://zenodo.org/record/3527532/files/figure.png" pageId="20" pageNumber="481">Fig. 5C</figureCitation>
): Coxa I with a median row of enlarged setiferous tubercles; coxa II with a median row of small tubercles; coxae III–IV with scattered tubercles. Coxae II–IV connected by a row of tubercles. Posterior margin of the stigmatic area convex, most lateral parts touching coxa IV. Stigmata elliptical and slanted.
</paragraph>
<paragraph id="8BA010165C28FFC2E4B2F834FD70B1B7" blockId="20.[151,1437,1834,2005]" pageId="20" pageNumber="481">
<emphasis id="B96BCC045C28FFC2E4B2F834FEDBB1D3" box="[199,322,1943,1969]" italics="true" pageId="20" pageNumber="481">Chelicerae</emphasis>
: Segment I with scattered granules; bulla weakly marked; segment II with scattered granules; fixed finger with 3–4 teeth; movable finger with 3–4 teeth.
</paragraph>
<paragraph id="8BA010165C29FFC3E4B2FF3AFDDCB77D" blockId="21.[151,1437,152,1944]" pageId="21" pageNumber="482">
<emphasis id="B96BCC045C29FFC3E4B2FF3AFEAEB6D1" box="[199,311,153,179]" italics="true" pageId="21" pageNumber="482">Pedipalps</emphasis>
(
<figureCitation id="13240C935C29FFC3E533FF3AFE05B6D1" box="[326,412,152,179]" captionStart="FIGURE 5" captionStartId="19.[151,250,1915,1940]" captionTargetBox="[157,1429,174,1887]" captionTargetId="figure@19.[151,1436,174,1896]" captionTargetPageId="19" captionText="FIGURE 5. Progonyleptoidellus picinguaba sp. nov., male paratype (MZSP 16009) and female paratype (MZSP 16257): A, male habitus, dorsal view; B, female habitus, dorsal view; C, male habitus, ventral view; D, male trochanter–tibia IV, dorsal view; E, female trochanter–patella IV, dorsal view; F, male habitus, left lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3527532" httpUri="https://zenodo.org/record/3527532/files/figure.png" pageId="21" pageNumber="482">Fig. 5F</figureCitation>
): Coxa smooth and enlarged, reaching the cheliceral bulla and trochanter I. Trochanter with a dorsoapical small elevation, with one apical and one basal ventro-median small setiferous tubercle. Femur and patella elongated, slender and smooth. Tibial setation: mesal IiIi; ectal IiIi. Tarsus with 2 ventral median row of setae; tarsal setation: mesal Ii; ectal II.
</paragraph>
<paragraph id="8BA010165C29FFC3E4B2FE89FEDBB4C9" blockId="21.[151,1437,152,1944]" pageId="21" pageNumber="482">
<emphasis id="B96BCC045C29FFC3E4B2FE89FF65B721" box="[199,252,298,323]" italics="true" pageId="21" pageNumber="482">Legs</emphasis>
: Coxae I–II with one prodorsal apophysis; coxa III with two retrolateral apophyses. Coxa IV with scattered granules; a strong, large and straight proapical apophysis, with an acuminate apex, obliquely directed, and a retroapical small, acuminate apophysis. Trochanters I–III unarmed, with some apical small tubercles; trochanter IV (
<figureCitation id="13240C935C29FFC3E4EAFE36FE9BB7CD" box="[159,258,405,431]" captionStart="FIGURE 4" captionStartId="18.[151,250,1800,1825]" captionTargetBox="[168,1425,388,1769]" captionTargetId="figure@18.[151,1436,373,1780]" captionTargetPageId="18" captionText="FIGURE 4. Progonyleptoidellus picinguaba sp. nov., male holotype (MZSP 73602): A, habitus, dorsal view; B, right trochanter–tibia IV, dorsal view; C, coxa, trochanter and basal part of femur IV, retrodorsal view. Scale bars: A, C: 2 mm, B: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3527530" httpUri="https://zenodo.org/record/3527530/files/figure.png" pageId="21" pageNumber="482">Figs. 4B</figureCitation>
,
<figureCitation id="13240C935C29FFC3E57AFE37FED4B7CD" box="[271,333,404,431]" captionStart="FIGURE 5" captionStartId="19.[151,250,1915,1940]" captionTargetBox="[157,1429,174,1887]" captionTargetId="figure@19.[151,1436,174,1896]" captionTargetPageId="19" captionText="FIGURE 5. Progonyleptoidellus picinguaba sp. nov., male paratype (MZSP 16009) and female paratype (MZSP 16257): A, male habitus, dorsal view; B, female habitus, dorsal view; C, male habitus, ventral view; D, male trochanter–tibia IV, dorsal view; E, female trochanter–patella IV, dorsal view; F, male habitus, left lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3527532" httpUri="https://zenodo.org/record/3527532/files/figure.png" pageId="21" pageNumber="482">5A,D</figureCitation>
) prolaterally with one sub-basal and one apical tubercle, these short and acuminate, the sub-basal one the largest; retrolaterally with one median and one apical tubercles, these short and acuminate, the apical largest; ventral face with scattered granules. Femora I–III unarmed, with scattered granules; femur IV (
<figureCitation id="13240C935C29FFC3E175FE7EFAFAB795" box="[1280,1379,477,503]" captionStart="FIGURE 4" captionStartId="18.[151,250,1800,1825]" captionTargetBox="[168,1425,388,1769]" captionTargetId="figure@18.[151,1436,373,1780]" captionTargetPageId="18" captionText="FIGURE 4. Progonyleptoidellus picinguaba sp. nov., male holotype (MZSP 73602): A, habitus, dorsal view; B, right trochanter–tibia IV, dorsal view; C, coxa, trochanter and basal part of femur IV, retrodorsal view. Scale bars: A, C: 2 mm, B: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3527530" httpUri="https://zenodo.org/record/3527530/files/figure.png" pageId="21" pageNumber="482">Figs. 4B</figureCitation>
,
<figureCitation id="13240C935C29FFC3E104FE7FFA0DB795" box="[1393,1428,476,503]" captionStart="FIGURE 5" captionStartId="19.[151,250,1915,1940]" captionTargetBox="[157,1429,174,1887]" captionTargetId="figure@19.[151,1436,174,1896]" captionTargetPageId="19" captionText="FIGURE 5. Progonyleptoidellus picinguaba sp. nov., male paratype (MZSP 16009) and female paratype (MZSP 16257): A, male habitus, dorsal view; B, female habitus, dorsal view; C, male habitus, ventral view; D, male trochanter–tibia IV, dorsal view; E, female trochanter–patella IV, dorsal view; F, male habitus, left lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3527532" httpUri="https://zenodo.org/record/3527532/files/figure.png" pageId="21" pageNumber="482">5D</figureCitation>
) straight and long; dorsobasally with a row of 4–6 median spines (those located in the middle of the row larger); retrodorsally with a row of intercalated tubercles and spines of irregular size (those located in the middle of the row larger); proventrally with a row of very short conical tubercles. Patellae and tibiae I–IV unarmed, with ventral scattered granules. Metatarsi I–IV smooth, astragalus swollen. Claws smooth. Tarsal process large. Tarsal counts:.7(3),?, 12, 15.
</paragraph>
<paragraph id="8BA010165C29FFC3E4B2FD15FC4FB58D" blockId="21.[151,1437,152,1944]" pageId="21" pageNumber="482">
<emphasis id="B96BCC045C29FFC3E4B2FD15FE9CB4AD" box="[199,261,694,719]" italics="true" pageId="21" pageNumber="482">Penis</emphasis>
(MZSP 16259, see
<figureCitation id="13240C935C29FFC3E590FD16FDA8B4B2" box="[485,561,693,720]" captionStart="FIGURE 6" captionStartId="20.[151,250,1728,1753]" captionTargetBox="[151,1436,174,1709]" captionTargetId="figure@20.[151,1436,174,1709]" captionTargetPageId="20" captionText="FIGURE 6. Scanning electron micrographs showing the distal part of Progonyleptoidellus spp. penis: A–B, P. bocaina sp. nov. (MZSP 18939), A. dorsal, B. lateral; C–D, P. picinguaba sp. nov. (MZSP 16259), A. dorsal, B. lateral. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3527534" httpUri="https://zenodo.org/record/3527534/files/figure.png" pageId="21" pageNumber="482">Figs. 6</figureCitation>
C–D): VP subrectangular, ventral face totally covered with microsetae of
<typeStatus id="54A4AEB45C29FFC3E119FD16FA05B4AD" box="[1388,1436,693,719]" pageId="21" pageNumber="482">type</typeStatus>
1; distal margin with a U shaped cleft, slightly curved in lateral view. Lateral lobe sub-basal, sub-rectangular shaped, dorsally directed, with microsetae
<typeStatus id="54A4AEB45C29FFC3E656FD5EFDCAB575" box="[547,595,765,791]" pageId="21" pageNumber="482">type</typeStatus>
3 on its distal part. Macrosetae (MS) A1–A3 well developed, lanceolate, forming a slanted row on the lateral lobe; MS B1 short, conical, separated from MS A; MS C1–C3 long, well developed, basally straight and distally helicoidal; MS D1 short and conical, placed between MS C and MS A; MS E1–E2 short and conical, placed near MS C1–C3. Glans stylus subcylindrical with inclined apex, ornate with microspines subapically. Glans ventral process with stem thicker than stylus, and with flabelliform apex. Flabellum fan-like, with multi-serrated apex. Glans sac short, multi-folded, heel-shaped, projected as a dorsal process. Stylus stem arising from the ventralmost part of glans sac, inserted as a candelabrum-like.
</paragraph>
<paragraph id="8BA010165C29FFC3E4B2FC5AFECBB2C1" blockId="21.[151,1437,152,1944]" pageId="21" pageNumber="482">
<emphasis id="B96BCC045C29FFC3E4B2FC5AFEDBB271" box="[199,322,1017,1043]" italics="true" pageId="21" pageNumber="482">Coloration</emphasis>
(in ethanol; see
<figureCitation id="13240C935C29FFC3E673FC5AFDD5B271" box="[518,588,1016,1043]" captionStart="FIGURE 5" captionStartId="19.[151,250,1915,1940]" captionTargetBox="[157,1429,174,1887]" captionTargetId="figure@19.[151,1436,174,1896]" captionTargetPageId="19" captionText="FIGURE 5. Progonyleptoidellus picinguaba sp. nov., male paratype (MZSP 16009) and female paratype (MZSP 16257): A, male habitus, dorsal view; B, female habitus, dorsal view; C, male habitus, ventral view; D, male trochanter–tibia IV, dorsal view; E, female trochanter–patella IV, dorsal view; F, male habitus, left lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3527532" httpUri="https://zenodo.org/record/3527532/files/figure.png" pageId="21" pageNumber="482">Fig. 5</figureCitation>
): Light yellow with small black spots scattered on anterior, posterior and lateral margins, as well as on scutal areas of DS, ocularium, free tergites I–III, pedipalps, chelicerae and legs. Predominantly black on the carapace around ocularium, on the scutal area III spines and on dorsoapical apophyses of coxae IV. Dry-marks on scutal areas sulci, anterior margin of DS, anterior half of carapace and sparsely on posterior half of carapace.
</paragraph>
<paragraph id="8BA010165C29FFC3E4B2FB0DFEAEB069" blockId="21.[151,1437,152,1944]" pageId="21" pageNumber="482">
<emphasis id="B96BCC045C29FFC3E4B2FB0DFE0EB2A5" box="[199,407,1197,1223]" italics="true" pageId="21" pageNumber="482">Variation in males</emphasis>
(n=7): Measurements: CL 2.1–2.4; CW 3–3.2; DSL 4.4–5; DSW 4.6–5.5; FeP 3.2–3.6; FI 4.2–4.7; FII 11.1–12.4; FIII 7.5–8.2; FIV 9.9–11.5. Anterior margin of DS with 3–4 median acuminate tubercles on the corners (if more than three, the fourth much smaller than the others). Posterior margin of DS slightly sinuous in some males; armed with one or more acuminate tubercles. Free tergite I–II armed with one or more spines; free tergite III with more than one large spine. The armature of posterior margin of DS and free tergites I–III extremely variable in number, size, position and could be absent in some specimens. Pedipalps: tibia setation: mesal, IiIi, IIiIi, IiII; ectal IiIi, II; tarsal setation: mesal Ii, II, IIII; ectal Ii, II; proapical apophysis with apex slightly slanted. Armature of femur IV weak, with spines like tubercles. Dry-mark weak (inconspicuous in some regions); in some specimens totally or partially absent (in the latter case, notably absent in scutal areas sulci). Tarsal counts: 7(3), 13–14(3–4), 11–13, 15–18.
</paragraph>
<paragraph id="8BA010165C29FFC3E4B2F9B6FE75B04D" blockId="21.[151,1437,152,1944]" box="[199,492,1556,1583]" pageId="21" pageNumber="482">
Female (n=9;
<figureCitation id="13240C935C29FFC3E510F9B6FE7AB04D" box="[357,483,1556,1583]" captionStart="FIGURE 5" captionStartId="19.[151,250,1915,1940]" captionTargetBox="[157,1429,174,1887]" captionTargetId="figure@19.[151,1436,174,1896]" captionTargetPageId="19" captionText="FIGURE 5. Progonyleptoidellus picinguaba sp. nov., male paratype (MZSP 16009) and female paratype (MZSP 16257): A, male habitus, dorsal view; B, female habitus, dorsal view; C, male habitus, ventral view; D, male trochanter–tibia IV, dorsal view; E, female trochanter–patella IV, dorsal view; F, male habitus, left lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3527532" httpUri="https://zenodo.org/record/3527532/files/figure.png" pageId="21" pageNumber="482">Figs. 5B, E</figureCitation>
)
</paragraph>
<paragraph id="8BA010165C29FFC3E4B2F999FE3CB015" blockId="21.[151,1437,152,1944]" pageId="21" pageNumber="482">
<emphasis id="B96BCC045C29FFC3E4B2F999FEF0B031" box="[199,361,1594,1619]" italics="true" pageId="21" pageNumber="482">Measurements</emphasis>
: CL 2.2–2.5; CW 3–3.4; DSL 5–5.4; DSW 5.1–5.6; FeP 3–3.6; FI 3.9–4.4; FII 11.2–12.3; FIII 7.5–8.7; FIV 10.3–12.6.
</paragraph>
<paragraph id="8BA010165C29FFC3E4B2F921FCA7B149" blockId="21.[151,1437,152,1944]" pageId="21" pageNumber="482">
<emphasis id="B96BCC045C29FFC3E4B2F921FEB8B0F9" box="[199,289,1666,1691]" italics="true" pageId="21" pageNumber="482">Dorsum</emphasis>
: Spines on scutal area III greater than in males in some specimens. Posterior margin of dorsal scutum sub straight (sinuous appearance); posterior margin of DS unarmed or armed with acuminate tubercles. Free tergites I–III armed. Like in males, the armature of posterior margin of DS and free tergites I–III extremely variable in number, size, position, usually with more than one spine; free tergites I–III armature generally greater than those on males (although some specimens have less robust armature).
</paragraph>
<paragraph id="8BA010165C29FFC3E4B2F896FB2AB12D" blockId="21.[151,1437,152,1944]" box="[199,1203,1845,1871]" pageId="21" pageNumber="482">
<emphasis id="B96BCC045C29FFC3E4B2F896FEAEB12D" box="[199,311,1845,1871]" italics="true" pageId="21" pageNumber="482">Pedipalps</emphasis>
: tibial setation: mesal, IiIi, IiII; ectal IiIi; tarsal setation: mesal Ii, II; ectal Ii, II.
</paragraph>
<paragraph id="8BA010165C29FFC3E4B2F8F9FC31B1F5" blockId="21.[151,1437,152,1944]" pageId="21" pageNumber="482">
<emphasis id="B96BCC045C29FFC3E4B2F8F9FF65B111" box="[199,252,1882,1907]" italics="true" pageId="21" pageNumber="482">Legs</emphasis>
: Proapical apophysis of coxa IV like in males in size; vary in stoutness (some specimens with apophysis stouter than in males). Tarsal counts: 6–8(3), 13–14(4), 12–13, 15–16.
</paragraph>
</subSubSection>
</treatment>
</document>
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