312 lines
43 KiB
XML
312 lines
43 KiB
XML
<document id="B57E76D9A66DFC236918A0B745DA5388" ID-CLB-Dataset="32767" ID-DOI="10.11646/zootaxa.4286.1.1" ID-GBIF-Dataset="a614f1d9-bda9-4df0-a486-aef5af7813fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="828206" ID-ZooBank="6A22EB50-0146-465A-8E2A-523E2F8FE470" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500108241184" checkinUser="plazi" docAuthor="Galvis, William" docDate="2017" docId="0386A719572F9078FF21FE5BFB29FAD8" docLanguage="en" docName="zootaxa.4286.1.1.pdf" docOrigin="Zootaxa 4286 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphidraus guaitipan Galvis, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="9" masterDocId="FFBFDF6157289070FFB6FFCFFF81FFE8" masterDocTitle="Nineteen new species of Amphidraus Simon, 1900 (Salticidae: Euophryini) from Colombia, with comments about their conservation" masterLastPageNumber="40" masterPageNumber="1" pageNumber="8" updateTime="1698465056562" updateUser="ExternalLinkService">
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<mods:title id="279DE244ADFD5F3240748C3DF9AB5755">Nineteen new species of Amphidraus Simon, 1900 (Salticidae: Euophryini) from Colombia, with comments about their conservation</mods:title>
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<mods:namePart id="7005DEB7C888EC6A88ED99FA5A18742D">Galvis, William</mods:namePart>
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<mods:date id="06A6339671222A37BC35DADFE9F5122D">2017</mods:date>
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<treatment id="0386A719572F9078FF21FE5BFB29FAD8" ID-DOI="http://doi.org/10.5281/zenodo.6045631" ID-GBIF-Taxon="132548507" ID-Zenodo-Dep="6045631" LSID="urn:lsid:plazi:treatment:0386A719572F9078FF21FE5BFB29FAD8" httpUri="http://treatment.plazi.org/id/0386A719572F9078FF21FE5BFB29FAD8" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<subSubSection id="C3354584572F9077FF21FE5BFDEDFE27" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="8B90160F572F9077FF21FE5BFD8DFE46" blockId="7.[151,620,404,463]" box="[151,524,404,430]" pageId="7" pageNumber="8">
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<emphasis id="B95BCA1D572F9077FF21FE5BFD8DFE46" bold="true" box="[151,524,404,430]" pageId="7" pageNumber="8">
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<taxonomicName id="4C2F6D8C572F9077FF21FE5BFE29FE46" ID-CoL="D2GB" box="[151,424,404,430]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="guaitipan" status="sp. nov.">
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<emphasis id="B95BCA1D572F9077FF21FE5BFE29FE46" bold="true" box="[151,424,404,430]" italics="true" pageId="7" pageNumber="8">Amphidraus guaitipan</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2687766572F9077FE18FE5BFD8DFE46" box="[430,524,404,430]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="8B90160F572F9077FF21FE79FDEDFE27" blockId="7.[151,620,404,463]" box="[151,620,438,463]" pageId="7" pageNumber="8">
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<figureCitation id="13140A8A572F9077FF21FE79FE86FE27" box="[151,263,438,463]" captionStart="FIGURES 4" captionStartId="7.[151,264,1904,1926]" captionTargetBox="[275,1321,710,1880]" captionTargetId="figure@7.[268,1321,690,1891]" captionTargetPageId="7" captionText="FIGURES 4 a – d. A. guaitipan sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.10 mm." httpUri="https://zenodo.org/record/828214/files/figure.png" pageId="7" pageNumber="8">Figs 4a–d</figureCitation>
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,
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<figureCitation id="13140A8A572F9077FEA5FE79FED6FE27" box="[275,343,438,463]" captionStart="FIGURES 16" captionStartId="27.[151,264,1766,1788]" captionTargetBox="[344,1247,526,1743]" captionTargetId="figure@27.[340,1247,517,1745]" captionTargetPageId="27" captionText="FIGURES 16 a – i. A. bochica sp. nov., a) male holotype (ICN – Ar 8276); b) female paratype (ICN – Ar 8269). A. chie sp. nov., c) male holotype (ICN – Ar 8273); d) female paratype (ICN – Ar 8274). A. colombianus sp. nov., e) male holotype (ICN – Ar 7292). A. guaitipan sp. nov., f) male holotype (ICN – Ar 8257); g) female paratype (ANDES – IN 3871). A. mae sp. nov., h) male holotype (ICN – Ar 8259); i) female paratype (ICN – Ar 8260). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828238/files/figure.png" pageId="7" pageNumber="8">16f–g</figureCitation>
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,
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<figureCitation id="13140A8A572F9077FED4FE79FE25FE27" box="[354,420,438,463]" captionStart="FIGURES 20" captionStartId="31.[151,264,1906,1928]" captionTargetBox="[248,1341,212,1891]" captionTargetId="figure@31.[228,1365,193,1894]" captionTargetPageId="31" captionText="FIGURES 20 a – k. Amphidraus spp, left male palp. A. bochica sp. nov., a) ventral view; b) same, retrolateral view. A. chie sp. nov., c) ventral view; d) same, retrolateral view. A. colombianus sp. nov., e) ventral view; f) same, retroventral view; g) same, retrolateral view. A. guaitipan sp. nov., h) ventral view (arrow shows the retrolateral macrosetae at the base of the RTA); i) same, retrolateral view. A. mae sp. nov., j) ventral view; k) same, retrolateral view." httpUri="https://zenodo.org/record/828246/files/figure.png" pageId="7" pageNumber="8">20h–i</figureCitation>
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,
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<figureCitation id="13140A8A572F9077FE06FE78FD8CFE27" box="[432,525,438,463]" captionStart="FIGURES 24" captionStartId="35.[151,264,1828,1850]" captionTargetBox="[155,1418,206,1799]" captionTargetId="figure@35.[151,1436,193,1807]" captionTargetPageId="35" captionText="FIGURES 24 a – l. Amphidraus spp, epigyne. A. bochica sp. nov., a) ventral view; g) disposition of the internal structures. A. chie sp. nov., b) ventral view; h) disposition of the internal structures. A. guaitipan sp. nov., c) ventral view; i) disposition of the internal structures. A. mae sp. nov., d) ventral view; j) disposition of the internal structures. A. quinini sp. nov., e) ventral view; k) disposition of the internal structures. A. sie sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828254/files/figure.png" pageId="7" pageNumber="8">
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24c,
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<date id="FF9130CF572F9077FE50FE79FD8CFE27" box="[486,525,438,463]" pageId="7" pageNumber="8" value="1924-01">24i</date>
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</figureCitation>
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,
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<figureCitation id="13140A8A572F9077FDAEFE79FDC4FE27" box="[536,581,438,463]" captionStart="FIGURES 26" captionStartId="37.[151,264,1181,1203]" captionTargetBox="[151,1436,193,1159]" captionTargetId="figure@37.[151,1436,193,1159]" captionTargetPageId="37" captionText="FIGURES 26 a – d. Different altitudinal ecosystems where some of the species of Amphidraus here described were found in Colombia. a) Páramo ecosystem, at Las Lagunas Encantadas Natural Reserve, Duitama, Colombia, habitat of A. tundama sp. nov. (~ 3800 m); b) Andean wet forest at Cueva de los Guácharos National Natural Park, Huila, Colombia, habitat of A. guaitipan sp. nov. (~ 2000 m); c) Andean Oak forest (Quercus humboldtii) at Quinini Natural Reserve, Cundinamarca, Colombia, habitat of A. quinini sp. nov. (~ 1700 m); d) Secondary lowland semi wet forest on the outskirts of the city of Villavicencio, Meta, Colombia, habitat of A. sikuani sp. nov. (~ 500 m)." httpUri="https://zenodo.org/record/828258/files/figure.png" pageId="7" pageNumber="8">26b</figureCitation>
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,
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<figureCitation id="13140A8A572F9077FDE6FE78FDEDFE27" box="[592,620,439,463]" captionStart="FIGURE 27" captionStartId="38.[151,250,1317,1339]" captionTargetBox="[138,1418,182,1273]" captionTargetId="figure@38.[137,1422,179,1282]" captionTargetPageId="38" captionText="FIGURE 27. Distribution records of the species of the genus Amphidraus Simon from Colombia treated here. Symbols with a dot inside represent species that have distributions in the Cundiboyacense high-Andean plateau, in the Boyacá and Cundinamarca departments." httpUri="https://zenodo.org/record/828260/files/figure.png" pageId="7" pageNumber="8">27</figureCitation>
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.
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:
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male from
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de los
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,
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,
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<collectorName id="26DA73D9572F9077FCDBFDECFBB3FDD3" box="[877,1074,546,571]" pageId="7" pageNumber="8">V. Muñoz-Charry</collectorName>
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(
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<collectionCode id="ED3E8ECA572F9077FBF4FDECFBF2FDD3" box="[1090,1139,547,571]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="7" pageNumber="8">ICN</collectionCode>
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–Ar 8257)
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.
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<materialsCitation id="3B471C52572F9077FB40FDEDFDF3FD6B" ID-GBIF-Occurrence="1572324835" collectionCode="ANDES" collectorName="Fonseca" country="Colombia" county="Palestina" elevation="2170" latitude="1.55" location="National Natural Park Cueva" longLatPrecision="785" longitude="-76.15" municipality="Guacharos" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Huila" typeStatus="paratype">
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<typeStatus id="5494A8AD572F9077FB40FDEDFAE6FDD3" box="[1270,1383,546,571]" pageId="7" pageNumber="8" type="paratype">Paratype</typeStatus>
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:
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</emphasis>
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one female from
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<location id="8EF040D4572F9077FE82FD89FD0AFDB7" LSID="urn:lsid:plazi:treatment:0386A719572F9078FF21FE5BFB29FAD8:8EF040D4572F9077FE82FD89FD0AFDB7" box="[308,651,582,607]" country="Colombia" county="Palestina" latitude="1.55" longLatPrecision="785" longitude="-76.15" municipality="Guacharos" name="National Natural Park Cueva" pageId="7" pageNumber="8" stateProvince="Huila">National Natural Park Cueva</location>
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de los
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<collectingMunicipality id="6BF48C75572F9077FD59FD89FCEBFDB7" box="[751,874,582,607]" pageId="7" pageNumber="8">Guácharos</collectingMunicipality>
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,
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,
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[
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],
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, M.
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<collectorName id="26DA73D9572F9077FE98FDA4FE0BFD6A" box="[302,394,619,642]" pageId="7" pageNumber="8">Fonseca</collectorName>
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(
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<collectionCode id="ED3E8ECA572F9077FE2EFDA4FE77FD6B" box="[408,502,619,643]" country="Colombia" httpUri="http://grbio.org/cool/zmn5-q6qj" name="La Universidad de Los Andes" pageId="7" pageNumber="8">ANDES</collectionCode>
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–IN 3871).
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|
||
<paragraph id="8B90160F572F9077FF21F8BFFC86F802" blockId="7.[151,1436,1904,2026]" pageId="7" pageNumber="8">
|
||
<emphasis id="B95BCA1D572F9077FF21F8BFFEC8F86E" bold="true" box="[151,329,1904,1926]" pageId="7" pageNumber="8">FIGURES 4a–d.</emphasis>
|
||
<taxonomicName id="4C2F6D8C572F9077FEF9F8BEFE4CF86E" box="[335,461,1905,1926]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="guaitipan" status="sp. nov.">
|
||
<emphasis id="B95BCA1D572F9077FEF9F8BEFE4CF86E" box="[335,461,1905,1926]" italics="true" pageId="7" pageNumber="8">A. guaitipan</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2687766572F9077FE62F8BFFDA4F86E" box="[468,549,1904,1926]" pageId="7" pageNumber="8" rank="species">
|
||
<emphasis id="B95BCA1D572F9077FE62F8BFFDA4F86E" bold="true" box="[468,549,1904,1926]" pageId="7" pageNumber="8">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B95BCA1D572F9077FD85F8BFFDC6F86E" bold="true" box="[563,583,1904,1926]" pageId="7" pageNumber="8">a)</emphasis>
|
||
left male palp, ventral view;
|
||
<emphasis id="B95BCA1D572F9077FCC2F8BFFC0AF86E" bold="true" box="[884,907,1904,1926]" pageId="7" pageNumber="8">b)</emphasis>
|
||
same, retrolateral view;
|
||
<emphasis id="B95BCA1D572F9077FB31F8BFFB1AF86E" bold="true" box="[1159,1179,1904,1926]" pageId="7" pageNumber="8">c)</emphasis>
|
||
epigyne, cleared, ventral view;
|
||
<emphasis id="B95BCA1D572F9077FF6AF85DFF72F840" bold="true" box="[220,243,1938,1960]" pageId="7" pageNumber="8">d)</emphasis>
|
||
same, dorsal view. Abbreviations: cd=copulatory duct, co=copulatory opening, e=embolus, ED=embolic disc, EF=embolic filament, fd=fertilization duct, po=epigynal pocket, RTA=retrolateral tibial apophysis of the male palp, SD=sperm duct, sp=spermatheca, TL=tegular lobe. Scale bars: 0.10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C335458457209078FF71FF58FC6DFEA8" pageId="8" pageNumber="9" type="etymology">
|
||
<paragraph id="8B90160F57209078FF71FF58FC6DFEA8" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="9">
|
||
<emphasis id="B95BCA1D57209078FF71FF58FECFFF58" bold="true" box="[199,334,151,176]" pageId="8" pageNumber="9">Etymology.</emphasis>
|
||
The epithet is a noun in apposition that honors the Cacique
|
||
<taxonomicName id="4C2F6D8C57209078FBA6FF58FBFFFF58" box="[1040,1150,151,176]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="38" phylum="Arthropoda" rank="species" species="guaitipan" status="sp. nov.">Guaitipán</taxonomicName>
|
||
(or better known by her Spanish name transliteration, Gaitana), one of the maximum leaders and lady of the indigenous people that inhabited the central and eastern region of the Central Mountain Range of the Colombian Andes, among which are the Andaquí, Avirama, Guanaca, Páez, Pinao, Pijao, Timaná and Yalcón people. Indigenous heroin, renowned by her brave and fierce defense of their territories against the Spanish invaders.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C335458457209078FF71FE83FF4AFDD5" pageId="8" pageNumber="9" type="diagnosis">
|
||
<paragraph id="8B90160F57209078FF71FE83FF4AFDD5" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="9">
|
||
<emphasis id="B95BCA1D57209078FF71FE83FEC0FE8D" bold="true" box="[199,321,332,357]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
|
||
Males of
|
||
<taxonomicName id="4C2F6D8C57209078FE02FE83FDC1FE8C" box="[436,576,332,356]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="guaitipan" status="sp. nov.">
|
||
<emphasis id="B95BCA1D57209078FE02FE83FDC1FE8C" box="[436,576,332,356]" italics="true" pageId="8" pageNumber="9">A. guaitipan</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A268776657209078FDF1FE83FD20FE8D" box="[583,673,332,357]" pageId="8" pageNumber="9" rank="species">
|
||
<emphasis id="B95BCA1D57209078FDF1FE83FD20FE8D" bold="true" box="[583,673,332,357]" pageId="8" pageNumber="9">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
are similar to those of
|
||
<taxonomicName id="4C2F6D8C57209078FC11FE83FB99FE8D" box="[935,1048,332,357]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sikuani" status="sp. nov.">
|
||
<emphasis id="B95BCA1D57209078FC11FE83FB99FE8D" box="[935,1048,332,357]" italics="true" pageId="8" pageNumber="9">A. sikuani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A268776657209078FB96FE83FBFBFE8D" box="[1056,1146,332,357]" pageId="8" pageNumber="9" rank="species">
|
||
<emphasis id="B95BCA1D57209078FB96FE83FBFBFE8D" bold="true" box="[1056,1146,332,357]" pageId="8" pageNumber="9">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C2F6D8C57209078FB04FE83FABBFE8C" box="[1202,1338,332,356]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tanimuca" status="sp. nov.">
|
||
<emphasis id="B95BCA1D57209078FB04FE83FABBFE8C" box="[1202,1338,332,356]" italics="true" pageId="8" pageNumber="9">A. tanimuca</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A268776657209078FAF4FE83FA1DFE8D" box="[1346,1436,332,357]" pageId="8" pageNumber="9" rank="species">
|
||
<emphasis id="B95BCA1D57209078FAF4FE83FA1DFE8D" bold="true" box="[1346,1436,332,357]" pageId="8" pageNumber="9">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by the shape of the RTA. However, they can be distinguished from those by their wider RTA, with the retro-dorsal lobe longer and wider, and the retro-ventral lobe turned prolaterally, with a retrolateral macroseta at its base (
|
||
<figureCitation id="13140A8A57209078FADCFE5BFF51FE38" captionStart="FIGURES 4" captionStartId="7.[151,264,1904,1926]" captionTargetBox="[275,1321,710,1880]" captionTargetId="figure@7.[268,1321,690,1891]" captionTargetPageId="7" captionText="FIGURES 4 a – d. A. guaitipan sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.10 mm." httpUri="https://zenodo.org/record/828214/files/figure.png" pageId="8" pageNumber="9">Figs 4a–b</figureCitation>
|
||
,
|
||
<figureCitation id="13140A8A57209078FF57FE78FEA6FE38" box="[225,295,439,464]" captionStart="FIGURES 20" captionStartId="31.[151,264,1906,1928]" captionTargetBox="[248,1341,212,1891]" captionTargetId="figure@31.[228,1365,193,1894]" captionTargetPageId="31" captionText="FIGURES 20 a – k. Amphidraus spp, left male palp. A. bochica sp. nov., a) ventral view; b) same, retrolateral view. A. chie sp. nov., c) ventral view; d) same, retrolateral view. A. colombianus sp. nov., e) ventral view; f) same, retroventral view; g) same, retrolateral view. A. guaitipan sp. nov., h) ventral view (arrow shows the retrolateral macrosetae at the base of the RTA); i) same, retrolateral view. A. mae sp. nov., j) ventral view; k) same, retrolateral view." httpUri="https://zenodo.org/record/828246/files/figure.png" pageId="8" pageNumber="9">20h–I</figureCitation>
|
||
). Females of
|
||
<taxonomicName id="4C2F6D8C57209078FE79FE77FDDEFE38" box="[463,607,440,464]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="guaitipan" status="sp. nov.">
|
||
<emphasis id="B95BCA1D57209078FE79FE77FDDEFE38" box="[463,607,440,464]" italics="true" pageId="8" pageNumber="9">A. guaitipan</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A268776657209078FDDAFE78FD4DFE38" box="[620,716,439,464]" pageId="8" pageNumber="9" rank="species">
|
||
<emphasis id="B95BCA1D57209078FDDAFE78FD4DFE38" bold="true" box="[620,716,439,464]" pageId="8" pageNumber="9">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
are similar to those of
|
||
<taxonomicName id="4C2F6D8C57209078FC44FE77FBE7FE38" box="[1010,1126,440,464]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="quinini" status="sp. nov.">
|
||
<emphasis id="B95BCA1D57209078FC44FE77FBE7FE38" box="[1010,1126,440,464]" italics="true" pageId="8" pageNumber="9">A. quinini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A268776657209078FBC2FE78FB52FE38" box="[1140,1235,439,464]" pageId="8" pageNumber="9" rank="species">
|
||
<emphasis id="B95BCA1D57209078FBC2FE78FB52FE38" bold="true" box="[1140,1235,439,464]" pageId="8" pageNumber="9">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by their general disposition of internal structures of the epigyne, but they can be distinguished from those by their shorter copulatory ducts, with the posterior epigynal border not concave, and a rounded epigynal pocket (
|
||
<figureCitation id="13140A8A57209078FB5BFE30FAE0FDF0" box="[1261,1377,511,536]" captionStart="FIGURES 4" captionStartId="7.[151,264,1904,1926]" captionTargetBox="[275,1321,710,1880]" captionTargetId="figure@7.[268,1321,690,1891]" captionTargetPageId="7" captionText="FIGURES 4 a – d. A. guaitipan sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.10 mm." httpUri="https://zenodo.org/record/828214/files/figure.png" pageId="8" pageNumber="9">Figs 4c–d</figureCitation>
|
||
,
|
||
<figureCitation id="13140A8A57209078FADBFDCFFF3FFDD5" captionStart="FIGURES 24" captionStartId="35.[151,264,1828,1850]" captionTargetBox="[155,1418,206,1799]" captionTargetId="figure@35.[151,1436,193,1807]" captionTargetPageId="35" captionText="FIGURES 24 a – l. Amphidraus spp, epigyne. A. bochica sp. nov., a) ventral view; g) disposition of the internal structures. A. chie sp. nov., b) ventral view; h) disposition of the internal structures. A. guaitipan sp. nov., c) ventral view; i) disposition of the internal structures. A. mae sp. nov., d) ventral view; j) disposition of the internal structures. A. quinini sp. nov., e) ventral view; k) disposition of the internal structures. A. sie sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828254/files/figure.png" pageId="8" pageNumber="9">
|
||
24c,
|
||
<date id="FF9130CF57209078FF21FDEBFF3FFDD5" box="[151,190,548,573]" pageId="8" pageNumber="9" value="1924-01">24i</date>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C335458457209078FF71FD88FB2AFB2D" pageId="8" pageNumber="9" type="description">
|
||
<paragraph id="8B90160F57209078FF71FD88FB2DFC4D" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="9">
|
||
<emphasis id="B95BCA1D57209078FF71FD88FDACFD88" bold="true" box="[199,557,583,608]" pageId="8" pageNumber="9">
|
||
Description. Male (
|
||
<typeStatus id="5494A8AD57209078FE03FD88FDA0FD88" box="[437,545,583,608]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
).
|
||
</emphasis>
|
||
Total length: 3.04. Carapace blackish brown with two posterior stripes of scarce white hairs, 1.55 long, 1.13 wide, 0.83 high (
|
||
<figureCitation id="13140A8A57209078FD6FFDA3FCB7FD6D" box="[729,822,620,645]" captionStart="FIGURES 16" captionStartId="27.[151,264,1766,1788]" captionTargetBox="[344,1247,526,1743]" captionTargetId="figure@27.[340,1247,517,1745]" captionTargetPageId="27" captionText="FIGURES 16 a – i. A. bochica sp. nov., a) male holotype (ICN – Ar 8276); b) female paratype (ICN – Ar 8269). A. chie sp. nov., c) male holotype (ICN – Ar 8273); d) female paratype (ICN – Ar 8274). A. colombianus sp. nov., e) male holotype (ICN – Ar 7292). A. guaitipan sp. nov., f) male holotype (ICN – Ar 8257); g) female paratype (ANDES – IN 3871). A. mae sp. nov., h) male holotype (ICN – Ar 8259); i) female paratype (ICN – Ar 8260). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828238/files/figure.png" pageId="8" pageNumber="9">Fig. 16f</figureCitation>
|
||
). OC black, 0.67 long. Anterior eye row 1.14 wide and posterior 1.04 wide. Sternum dark brown, 0.64 long, 0.53 wide. Labium dark brown, 0.11 long, 0.14 wide. Chelicerae dark brown with five retromarginal and one promarginal tooth. Palp dark brown with dorsal white hairs in the femur and patella, with a wide RTA with two lobes: the retro-dorsal one wide and pointed, and the retroventral one long and turned prolaterally with a macroseta at its base; rounded tegular lobe (TL) and slender embolus (
|
||
<figureCitation id="13140A8A57209078FEB3FCD0FEF6FCD0" box="[261,375,799,824]" captionStart="FIGURES 4" captionStartId="7.[151,264,1904,1926]" captionTargetBox="[275,1321,710,1880]" captionTargetId="figure@7.[268,1321,690,1891]" captionTargetPageId="7" captionText="FIGURES 4 a – d. A. guaitipan sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.10 mm." httpUri="https://zenodo.org/record/828214/files/figure.png" pageId="8" pageNumber="9">Figs 4a–b</figureCitation>
|
||
,
|
||
<figureCitation id="13140A8A57209078FE35FCD0FE49FCD0" box="[387,456,799,824]" captionStart="FIGURES 20" captionStartId="31.[151,264,1906,1928]" captionTargetBox="[248,1341,212,1891]" captionTargetId="figure@31.[228,1365,193,1894]" captionTargetPageId="31" captionText="FIGURES 20 a – k. Amphidraus spp, left male palp. A. bochica sp. nov., a) ventral view; b) same, retrolateral view. A. chie sp. nov., c) ventral view; d) same, retrolateral view. A. colombianus sp. nov., e) ventral view; f) same, retroventral view; g) same, retrolateral view. A. guaitipan sp. nov., h) ventral view (arrow shows the retrolateral macrosetae at the base of the RTA); i) same, retrolateral view. A. mae sp. nov., j) ventral view; k) same, retrolateral view." httpUri="https://zenodo.org/record/828246/files/figure.png" pageId="8" pageNumber="9">20h–i</figureCitation>
|
||
). Legs 4132, the first two pairs brown and the two posterior ones yellow, all with black rings. Leg macrosetae: femur, I–II d 1 di; III–IV d 2 di; tibia, I
|
||
<date id="FF9130CF57209078FCE2FC8AFC10FCB4" box="[852,913,836,860]" pageId="8" pageNumber="9">v 2-2</date>
|
||
-2; II
|
||
<date id="FF9130CF57209078FC78FC8AFB8BFCB4" box="[974,1034,836,860]" pageId="8" pageNumber="9">v 1-2</date>
|
||
-1, p 1 di; III–IV
|
||
<date id="FF9130CF57209078FB7EFC8AFB6AFCB4" box="[1224,1259,836,860]" pageId="8" pageNumber="9">v 1</date>
|
||
pr, p 0-1-1, r 0- 1-1; metatarsus, I
|
||
<date id="FF9130CF57209078FED4FCA7FE21FC68" box="[354,416,872,896]" pageId="8" pageNumber="9">v 2-2</date>
|
||
; II
|
||
<date id="FF9130CF57209078FE73FCA7FD82FC68" box="[453,515,872,896]" pageId="8" pageNumber="9">v 2-2</date>
|
||
, p 1 di; III
|
||
<date id="FF9130CF57209078FD37FCA7FD23FC68" box="[641,674,872,896]" pageId="8" pageNumber="9">v 1</date>
|
||
-0-1, p 1-0-2, r 1-0-2; IV p 2 di, r 1-0-2. Abdomen dark brown with lateral sides covered by some scarce white scales, dorsally with light brown marks (
|
||
<figureCitation id="13140A8A57209078FBF7FC43FB1EFC4D" box="[1089,1183,908,933]" captionStart="FIGURES 16" captionStartId="27.[151,264,1766,1788]" captionTargetBox="[344,1247,526,1743]" captionTargetId="figure@27.[340,1247,517,1745]" captionTargetPageId="27" captionText="FIGURES 16 a – i. A. bochica sp. nov., a) male holotype (ICN – Ar 8276); b) female paratype (ICN – Ar 8269). A. chie sp. nov., c) male holotype (ICN – Ar 8273); d) female paratype (ICN – Ar 8274). A. colombianus sp. nov., e) male holotype (ICN – Ar 7292). A. guaitipan sp. nov., f) male holotype (ICN – Ar 8257); g) female paratype (ANDES – IN 3871). A. mae sp. nov., h) male holotype (ICN – Ar 8259); i) female paratype (ICN – Ar 8260). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828238/files/figure.png" pageId="8" pageNumber="9">Fig. 16f</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B90160F57209078FF71FC60FB2AFB2D" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="9">
|
||
<emphasis id="B95BCA1D57209078FF71FC60FE2DFC20" bold="true" box="[199,428,943,968]" pageId="8" pageNumber="9">
|
||
Female (
|
||
<typeStatus id="5494A8AD57209078FE99FC60FE1EFC20" box="[303,415,943,968]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
|
||
).
|
||
</emphasis>
|
||
Total length 3.59. Carapace brown with a posterior lighter stripe, 1.69 long, 1.24 wide, 0.90 high (
|
||
<figureCitation id="13140A8A57209078FEA2FC1BFEF6FC05" box="[276,375,980,1005]" captionStart="FIGURES 16" captionStartId="27.[151,264,1766,1788]" captionTargetBox="[344,1247,526,1743]" captionTargetId="figure@27.[340,1247,517,1745]" captionTargetPageId="27" captionText="FIGURES 16 a – i. A. bochica sp. nov., a) male holotype (ICN – Ar 8276); b) female paratype (ICN – Ar 8269). A. chie sp. nov., c) male holotype (ICN – Ar 8273); d) female paratype (ICN – Ar 8274). A. colombianus sp. nov., e) male holotype (ICN – Ar 7292). A. guaitipan sp. nov., f) male holotype (ICN – Ar 8257); g) female paratype (ANDES – IN 3871). A. mae sp. nov., h) male holotype (ICN – Ar 8259); i) female paratype (ICN – Ar 8260). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828238/files/figure.png" pageId="8" pageNumber="9">
|
||
Fig.
|
||
<quantity id="4CD7BBEA57209078FEFCFC1BFEF6FC05" box="[330,375,980,1005]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="8" pageNumber="9" unit="g" value="16.0">16g</quantity>
|
||
</figureCitation>
|
||
). OC black, 0.71 long. Anterior eye row 1.18 wide and posterior 1.11 wide. Sternum brown, 0.67 long, 0.52 wide. Labium brown, 0.19 long, 0.29 wide. Chelicerae brown, with five retromarginal and two promarginal teeth. Legs 4312, all yellow with black marks. Leg macrosetae: femur, I–II, IV d 1 di; III d 2 di; patellae, III–IV r 1 me; tibiae, I
|
||
<date id="FF9130CF57209078FE4AFB8FFDB8FBB0" box="[508,569,1088,1112]" pageId="8" pageNumber="9">v 2-2</date>
|
||
-2; II
|
||
<date id="FF9130CF57209078FDC0FB8FFD30FBB0" box="[630,689,1088,1112]" pageId="8" pageNumber="9">v 2-2</date>
|
||
-2, p 1 di; III
|
||
<date id="FF9130CF57209078FCF0FB8FFCE9FBB0" box="[838,872,1088,1112]" pageId="8" pageNumber="9">v 1</date>
|
||
pr, p 1-1, r 1-1; IV
|
||
<date id="FF9130CF57209078FBF2FB8FFBE7FBB0" box="[1092,1126,1088,1112]" pageId="8" pageNumber="9">v 1</date>
|
||
pr, p 1-0, r 1-1; metatarsus, I
|
||
<date id="FF9130CF57209078FF1EFBAAFF66FB94" box="[168,231,1124,1148]" pageId="8" pageNumber="9">v 2-2</date>
|
||
;
|
||
<date id="FF9130CF57209078FF42FBAAFEB6FB94" box="[244,311,1124,1148]" pageId="8" pageNumber="9">II 2-2</date>
|
||
, p 1 di; III–IV
|
||
<date id="FF9130CF57209078FE53FBAAFD89FB94" box="[485,520,1124,1148]" pageId="8" pageNumber="9">v 1</date>
|
||
-0-1, p 1-0-2, r 1-0-2. Abdomen brown with a light gray reticulated pattern (
|
||
<figureCitation id="13140A8A57209078FAD9FBABFF45FB48" captionStart="FIGURES 16" captionStartId="27.[151,264,1766,1788]" captionTargetBox="[344,1247,526,1743]" captionTargetId="figure@27.[340,1247,517,1745]" captionTargetPageId="27" captionText="FIGURES 16 a – i. A. bochica sp. nov., a) male holotype (ICN – Ar 8276); b) female paratype (ICN – Ar 8269). A. chie sp. nov., c) male holotype (ICN – Ar 8273); d) female paratype (ICN – Ar 8274). A. colombianus sp. nov., e) male holotype (ICN – Ar 7292). A. guaitipan sp. nov., f) male holotype (ICN – Ar 8257); g) female paratype (ANDES – IN 3871). A. mae sp. nov., h) male holotype (ICN – Ar 8259); i) female paratype (ICN – Ar 8260). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828238/files/figure.png" pageId="8" pageNumber="9">
|
||
Fig.
|
||
<quantity id="4CD7BBEA57209078FF21FB48FF45FB48" box="[151,196,1159,1184]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="8" pageNumber="9" unit="g" value="16.0">16g</quantity>
|
||
</figureCitation>
|
||
). Epigyne (
|
||
<figureCitation id="13140A8A57209078FEF0FB48FE38FB48" box="[326,441,1159,1184]" captionStart="FIGURES 4" captionStartId="7.[151,264,1904,1926]" captionTargetBox="[275,1321,710,1880]" captionTargetId="figure@7.[268,1321,690,1891]" captionTargetPageId="7" captionText="FIGURES 4 a – d. A. guaitipan sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.10 mm." httpUri="https://zenodo.org/record/828214/files/figure.png" pageId="8" pageNumber="9">Figs 4c–d</figureCitation>
|
||
,
|
||
<figureCitation id="13140A8A57209078FE73FB47FDA0FB48" box="[453,545,1159,1184]" captionStart="FIGURES 24" captionStartId="35.[151,264,1828,1850]" captionTargetBox="[155,1418,206,1799]" captionTargetId="figure@35.[151,1436,193,1807]" captionTargetPageId="35" captionText="FIGURES 24 a – l. Amphidraus spp, epigyne. A. bochica sp. nov., a) ventral view; g) disposition of the internal structures. A. chie sp. nov., b) ventral view; h) disposition of the internal structures. A. guaitipan sp. nov., c) ventral view; i) disposition of the internal structures. A. mae sp. nov., d) ventral view; j) disposition of the internal structures. A. quinini sp. nov., e) ventral view; k) disposition of the internal structures. A. sie sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828254/files/figure.png" pageId="8" pageNumber="9">
|
||
24c,
|
||
<date id="FF9130CF57209078FE4BFB48FDA0FB48" box="[509,545,1159,1184]" pageId="8" pageNumber="9" value="1924-01">24i</date>
|
||
</figureCitation>
|
||
) with anterior copulatory openings, concave disposition of the anterior section of the copulatory ducts, oval anterior spermathecae, and posterior single and rounded pocket.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C335458457209078FF71FB00FF69FAE5" pageId="8" pageNumber="9" type="discussion">
|
||
<paragraph id="8B90160F57209078FF71FB00FF69FAE5" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="9">
|
||
<emphasis id="B95BCA1D57209078FF71FB00FECEFB00" bold="true" box="[199,335,1231,1256]" pageId="8" pageNumber="9">Comments.</emphasis>
|
||
The specimens were collected in a preserved Andean wet ecosystem (
|
||
<figureCitation id="13140A8A57209078FB23FB00FB7AFB00" box="[1173,1275,1231,1256]" captionStart="FIGURES 26" captionStartId="37.[151,264,1181,1203]" captionTargetBox="[151,1436,193,1159]" captionTargetId="figure@37.[151,1436,193,1159]" captionTargetPageId="37" captionText="FIGURES 26 a – d. Different altitudinal ecosystems where some of the species of Amphidraus here described were found in Colombia. a) Páramo ecosystem, at Las Lagunas Encantadas Natural Reserve, Duitama, Colombia, habitat of A. tundama sp. nov. (~ 3800 m); b) Andean wet forest at Cueva de los Guácharos National Natural Park, Huila, Colombia, habitat of A. guaitipan sp. nov. (~ 2000 m); c) Andean Oak forest (Quercus humboldtii) at Quinini Natural Reserve, Cundinamarca, Colombia, habitat of A. quinini sp. nov. (~ 1700 m); d) Secondary lowland semi wet forest on the outskirts of the city of Villavicencio, Meta, Colombia, habitat of A. sikuani sp. nov. (~ 500 m)." httpUri="https://zenodo.org/record/828258/files/figure.png" pageId="8" pageNumber="9">Fig. 26b</figureCitation>
|
||
), beating low shrubs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C335458457209078FF71FAD8FB29FAD8" box="[199,1192,1303,1328]" pageId="8" pageNumber="9" type="distribution">
|
||
<paragraph id="8B90160F57209078FF71FAD8FB29FAD8" blockId="8.[151,1437,151,1328]" box="[199,1192,1303,1328]" pageId="8" pageNumber="9">
|
||
<emphasis id="B95BCA1D57209078FF71FAD8FEE1FAD8" bold="true" box="[199,352,1303,1328]" pageId="8" pageNumber="9">Distribution.</emphasis>
|
||
<collectingCountry id="F338569F57209078FED1FAD8FE54FAD8" box="[359,469,1303,1328]" name="Colombia" pageId="8" pageNumber="9">Colombia</collectingCountry>
|
||
(
|
||
<collectingRegion id="49EBD8ED57209078FE52FAD8FDA4FAD8" box="[484,549,1303,1328]" country="Colombia" name="Huila" pageId="8" pageNumber="9">Huila</collectingRegion>
|
||
) (
|
||
<figureCitation id="13140A8A57209078FD8BFAD8FD10FAD8" box="[573,657,1303,1328]" captionStart="FIGURE 27" captionStartId="38.[151,250,1317,1339]" captionTargetBox="[138,1418,182,1273]" captionTargetId="figure@38.[137,1422,179,1282]" captionTargetPageId="38" captionText="FIGURE 27. Distribution records of the species of the genus Amphidraus Simon from Colombia treated here. Symbols with a dot inside represent species that have distributions in the Cundiboyacense high-Andean plateau, in the Boyacá and Cundinamarca departments." httpUri="https://zenodo.org/record/828260/files/figure.png" pageId="8" pageNumber="9">Fig. 27</figureCitation>
|
||
). Known altitudinal distribution:
|
||
<quantity id="4CD7BBEA57209078FBB0FAD8FB22FAD8" box="[1030,1187,1303,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.66" metricValueMax="2.17" metricValueMin="1.15" pageId="8" pageNumber="9" unit="m" value="1660.0" valueMax="2170.0" valueMin="1150.0">1150–2170 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |