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<document id="BBACE97543574A39C48FDDEBC7982DA5" ID-CLB-Dataset="27597" ID-DOI="10.11646/zootaxa.4551.3.4" ID-GBIF-Dataset="eddea689-dc54-4802-a738-abcb61384604" ID-ISSN="1175-5326" ID-Zenodo-Dep="2622947" ID-ZooBank="946C9A0F-D5C8-4EB6-8939-48BC7DE400A9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554270235618" checkinUser="plazi" docAuthor="Grisales, Diana &amp; De Carvalho, Claudio J. B." docDate="2019" docId="6909C857F20BFF805390BD14FE7D3A12" docLanguage="en" docName="zootaxa.4551.3.4.pdf" docOrigin="Zootaxa 4551 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fannia palta Grisales &amp; De Carvalho 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="345" masterDocId="9530B02FF205FF8F5307B92BFFB83E02" masterDocTitle="Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America" masterLastPageNumber="360" masterPageNumber="330" pageNumber="344" updateTime="1698716131619" updateUser="ExternalLinkService">
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<mods:title id="C8046F2FE609A25D8B26F07A3C0F608C">Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America</mods:title>
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<mods:namePart id="EAF4062054B67E13F6985C0618F5D9A6">Grisales, Diana</mods:namePart>
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<mods:namePart id="08839FC81DE773DB8C37850838A2131A">De Carvalho, Claudio J. B.</mods:namePart>
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<treatment id="6909C857F20BFF805390BD14FE7D3A12" ID-DOI="http://doi.org/10.5281/zenodo.5931480" ID-GBIF-Taxon="154533669" ID-Zenodo-Dep="5931480" LSID="urn:lsid:plazi:treatment:6909C857F20BFF805390BD14FE7D3A12" httpUri="http://treatment.plazi.org/id/6909C857F20BFF805390BD14FE7D3A12" lastPageId="15" lastPageNumber="345" pageId="14" pageNumber="344">
<subSubSection id="A9BA2ACAF20BFF815390BD14FE213A58" box="[151,409,1087,1114]" pageId="14" pageNumber="344" type="nomenclature">
<paragraph id="E11F7941F20BFF815390BD14FE213A58" blockId="14.[151,480,1087,1147]" box="[151,409,1087,1114]" pageId="14" pageNumber="344">
<heading id="BA57CE2DF20BFF815390BD14FE213A58" bold="true" box="[151,409,1087,1114]" fontSize="11" level="1" pageId="14" pageNumber="344" reason="1">
<emphasis id="D3D4A553F20BFF815390BD14FE213A58" bold="true" box="[151,409,1087,1114]" pageId="14" pageNumber="344">
<taxonomicName id="26A002C2F20BFF815390BD14FE8C3A5B" ID-CoL="9J258" authority="Grisales &amp; De Carvalho, 2019" authorityName="Grisales &amp; De Carvalho" authorityYear="2019" box="[151,308,1087,1113]" class="Insecta" family="Fanniidae" genus="Fannia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="344" phylum="Arthropoda" rank="species" species="palta" status="sp. nov.">
<emphasis id="D3D4A553F20BFF815390BD14FE8C3A5B" bold="true" box="[151,308,1087,1113]" italics="true" pageId="14" pageNumber="344">Fannia palta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8E71828F20BFF81523CBD6BFE213A58" box="[315,409,1088,1114]" pageId="14" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="A9BA2ACAF20BFF815390BD49FE583A79" box="[151,480,1122,1147]" pageId="14" pageNumber="344" type="description">
<paragraph id="E11F7941F20BFF815390BD49FE583A79" blockId="14.[151,480,1087,1147]" box="[151,480,1122,1147]" pageId="14" pageNumber="344">
(
<figureCitation id="799B65C4F20BFF815398BD49FEBB3A79" box="[159,259,1122,1147]" captionStart="FIGURE 1" captionStartId="7.[151,250,1880,1902]" captionTargetBox="[151,1435,193,1859]" captionTargetId="figure@7.[151,1435,193,1859]" captionTargetPageId="7" captionText="FIGURE 1. Male holotype, complete lateral view: (A) Fannia awa sp. nov.; (B) Fannia bari sp. nov., (C) Fannia boruca sp. nov.; (D) Fannia bribri sp. nov.; (E) Fannia huetare sp. nov.; (F) Fannia humahuaca sp. nov.; (G) Fannia kabekwa sp. nov.; (H) Fannia palta sp. nov.; (I) Fannia polleti sp. nov.; (J) Fannia quillacingas sp. nov.; (K) Fannia teribi sp. nov.; (L) Fannia terraba sp. nov.; (M) Fannia tsachilas sp. nov.; (N) Fannia yukpa sp. nov. Scale: 1 mm." httpUri="https://zenodo.org/record/2622949/files/figure.png" pageId="14" pageNumber="344">Figs. 1H</figureCitation>
,
<figureCitation id="799B65C4F20BFF815209BD48FEA23A79" box="[270,282,1123,1147]" captionStart="FIGURE 3" captionStartId="12.[151,250,1857,1879]" captionTargetBox="[186,1392,195,1831]" captionTargetId="figure@12.[186,1392,195,1831]" captionTargetPageId="12" captionText="FIGURE 3. Fannia huetare sp. nov., male holotype: (A) Head, anterior view, vibrissal and subvibrisal setae; (B) Fore coxa, lateral view, spine on posterior surface; (C) Fore tarsomeres; (D) Hind femur, anterior view; (E) Hind femur, posterior view. Fannia humahuaca sp. nov., male holotype: (F) Hind femur, anterior view. Fannia kabekwa sp. nov., male holotype: (G) Fore tibia, anterior view; (H) Fore tarsi; (I) Hind femur, anterior view. Fannia palta sp. nov., male holotype: (J) First fore tarsomere, leaflike spine; (K) Fore tarsomeres; (L) Mid tibia, posterior view; (M) Hind femur, anterior view; (N) Hind femur, posterior view. Fannia polleti sp. nov., male holotype: (O) Fore tarsi; (P) Hind femur, anterior view." httpUri="https://zenodo.org/record/2622953/files/figure.png" pageId="14" pageNumber="344">3</figureCitation>
JL, 5H, 6H, 8H)
</paragraph>
</subSubSection>
<subSubSection id="A9BA2ACAF20BFF815390BD81FC5E3AE5" pageId="14" pageNumber="344" type="diagnosis">
<paragraph id="E11F7941F20BFF815390BD81FC5E3AE5" blockId="14.[151,1437,1194,2011]" pageId="14" pageNumber="344">
<emphasis id="D3D4A553F20BFF815390BD81FEB23AC1" bold="true" box="[151,266,1194,1219]" pageId="14" pageNumber="344">Diagnosis</emphasis>
. Fore tarsomere 1 with 1 broad leaflike spine at the tip of posterior surface and another tiny leaflike spine; tarsomeres 35 flattened; tarsomere 1 and 2 yellowish on apical half.
</paragraph>
</subSubSection>
<subSubSection id="A9BA2ACAF20BFF8053C0BDD9FE253FAF" lastPageId="15" lastPageNumber="345" pageId="14" pageNumber="344" type="description">
<paragraph id="E11F7941F20BFF8153C0BDD9FD3B3B09" blockId="14.[151,1437,1194,2011]" box="[199,643,1266,1291]" pageId="14" pageNumber="344">
<emphasis id="D3D4A553F20BFF8153C0BDD9FEE93B09" bold="true" box="[199,337,1266,1291]" pageId="14" pageNumber="344">Description</emphasis>
.
<emphasis id="D3D4A553F20BFF815258BDD9FDB43B09" bold="true" box="[351,524,1266,1291]" pageId="14" pageNumber="344">
<typeStatus id="3E1BC7E3F20BFF815258BDD9FE743B09" box="[351,460,1266,1291]" pageId="14" pageNumber="344" type="holotype">Holotype</typeStatus>
male
</emphasis>
(
<figureCitation id="799B65C4F20BFF81511BBDD9FDCD3B09" box="[540,629,1266,1291]" captionStart="FIGURE 1" captionStartId="7.[151,250,1880,1902]" captionTargetBox="[151,1435,193,1859]" captionTargetId="figure@7.[151,1435,193,1859]" captionTargetPageId="7" captionText="FIGURE 1. Male holotype, complete lateral view: (A) Fannia awa sp. nov.; (B) Fannia bari sp. nov., (C) Fannia boruca sp. nov.; (D) Fannia bribri sp. nov.; (E) Fannia huetare sp. nov.; (F) Fannia humahuaca sp. nov.; (G) Fannia kabekwa sp. nov.; (H) Fannia palta sp. nov.; (I) Fannia polleti sp. nov.; (J) Fannia quillacingas sp. nov.; (K) Fannia teribi sp. nov.; (L) Fannia terraba sp. nov.; (M) Fannia tsachilas sp. nov.; (N) Fannia yukpa sp. nov. Scale: 1 mm." httpUri="https://zenodo.org/record/2622949/files/figure.png" pageId="14" pageNumber="344">Fig. 1H</figureCitation>
).
</paragraph>
<paragraph id="E11F7941F20BFF8153C0BC3DFCD83BE1" blockId="14.[151,1437,1194,2011]" pageId="14" pageNumber="344">
Head: eye densely setulose. Frontal vitta velvety black, pollinose. 15
<emphasis id="D3D4A553F20BFF8150E6BC3CFC4A3B2D" box="[993,1010,1303,1327]" italics="true" pageId="14" pageNumber="344">fr</emphasis>
. Ocellar triangle black;
<emphasis id="D3D4A553F20BFF81560EBC3CFA9C3B2C" box="[1289,1316,1303,1326]" italics="true" pageId="14" pageNumber="344">oc</emphasis>
proclinate and developed, 5 less developed setulae on ocellar area. Frontoorbital plate black and pollinose. Parafacial bare, narrowing ventrally, golden pollinose. Frontogenal suture and gena golden pollinose, gena with long setae. Scape black, 3 setae reaching pedicel. Pedicel black pollinose, dorsal setae developed, one much longer than the others. Postpedicel black, golden pilose, 2 times the length of pedicel. Arista black, with short pubescence, rays shorter than width of base of arista. Palpus black and slightly spatulate.
</paragraph>
<paragraph id="E11F7941F20BFF8153C0BCC5FCC33829" blockId="14.[151,1437,1194,2011]" pageId="14" pageNumber="344">
Thorax: black. Scutum without vitae.
<emphasis id="D3D4A553F20BFF81516BBCC4FD2D3804" box="[620,661,1519,1542]" italics="true" pageId="14" pageNumber="344">Acr</emphasis>
2:34, not arranged in rows. Dc 2:3, long, posteriormost postsutural pair developed.
<emphasis id="D3D4A553F20BFF81524DBF38FE3B3828" box="[330,387,1555,1578]" italics="true" pageId="14" pageNumber="344">Pprn</emphasis>
2, developed.
<emphasis id="D3D4A553F20BFF815121BF38FDE93828" box="[550,593,1555,1578]" italics="true" pageId="14" pageNumber="344">Pra</emphasis>
2. Proepimeron setulose.
</paragraph>
<paragraph id="E11F7941F20BFF8153C0BF1DFC113871" blockId="14.[151,1437,1194,2011]" pageId="14" pageNumber="344">
Wing: brownish, with upper third between vein C and half of vein R 2+3, and cells
<emphasis id="D3D4A553F20BFF815790BF1DFB17384C" box="[1175,1199,1590,1614]" italics="true" pageId="14" pageNumber="344">br</emphasis>
,
<emphasis id="D3D4A553F20BFF8157B8BF1DFB59384C" box="[1215,1249,1590,1614]" italics="true" pageId="14" pageNumber="344">bm</emphasis>
and
<emphasis id="D3D4A553F20BFF81561ABF1CFAFE384C" box="[1309,1350,1591,1614]" italics="true" pageId="14" pageNumber="344">cup</emphasis>
darker. Calypters whitish. Haltere brown with stem and base lighter in colour.
</paragraph>
<paragraph id="E11F7941F20BFF8053C0BF54FF033ED6" blockId="14.[151,1437,1194,2011]" lastBlockId="15.[151,1436,151,1040]" lastPageId="15" lastPageNumber="345" pageId="14" pageNumber="344">
Legs: black, pulvilli brownish and tarsi black, except for fore tarsomere 1 and 2 with apical half yellowish. Fore femur setulose on posterior surface with 1 row of long
<emphasis id="D3D4A553F20BFF81503CBF88FCEE38B8" box="[827,854,1699,1722]" italics="true" pageId="14" pageNumber="344">pv</emphasis>
and
<emphasis id="D3D4A553F20BFF81508BBF88FC2238B9" box="[908,922,1699,1723]" italics="true" pageId="14" pageNumber="344">d</emphasis>
; ventral surface bare. Fore tibia with 1 strong preapical
<emphasis id="D3D4A553F20BFF815215BFEDFE9838DC" box="[274,288,1734,1758]" italics="true" pageId="14" pageNumber="344">d</emphasis>
; 1 apical
<emphasis id="D3D4A553F20BFF81528BBFECFE1E38DC" box="[396,422,1735,1758]" italics="true" pageId="14" pageNumber="344">pv</emphasis>
; 1 apical
<emphasis id="D3D4A553F20BFF815114BFECFDA638DC" box="[531,542,1735,1758]" italics="true" pageId="14" pageNumber="344">v</emphasis>
. Fore tarsomere 1 with 1 broad leaflike spine at the tip of posterior surface and another tiny leaflike spine (
<figureCitation id="799B65C4F20BFF8152DABFC1FD963901" box="[477,558,1770,1795]" captionStart="FIGURE 3" captionStartId="12.[151,250,1857,1879]" captionTargetBox="[186,1392,195,1831]" captionTargetId="figure@12.[186,1392,195,1831]" captionTargetPageId="12" captionText="FIGURE 3. Fannia huetare sp. nov., male holotype: (A) Head, anterior view, vibrissal and subvibrisal setae; (B) Fore coxa, lateral view, spine on posterior surface; (C) Fore tarsomeres; (D) Hind femur, anterior view; (E) Hind femur, posterior view. Fannia humahuaca sp. nov., male holotype: (F) Hind femur, anterior view. Fannia kabekwa sp. nov., male holotype: (G) Fore tibia, anterior view; (H) Fore tarsi; (I) Hind femur, anterior view. Fannia palta sp. nov., male holotype: (J) First fore tarsomere, leaflike spine; (K) Fore tarsomeres; (L) Mid tibia, posterior view; (M) Hind femur, anterior view; (N) Hind femur, posterior view. Fannia polleti sp. nov., male holotype: (O) Fore tarsi; (P) Hind femur, anterior view." httpUri="https://zenodo.org/record/2622953/files/figure.png" pageId="14" pageNumber="344">Fig. 3J</figureCitation>
); tarsomeres 35 flattened (
<figureCitation id="799B65C4F20BFF81506CBFC1FC7E3901" box="[875,966,1770,1795]" captionStart="FIGURE 3" captionStartId="12.[151,250,1857,1879]" captionTargetBox="[186,1392,195,1831]" captionTargetId="figure@12.[186,1392,195,1831]" captionTargetPageId="12" captionText="FIGURE 3. Fannia huetare sp. nov., male holotype: (A) Head, anterior view, vibrissal and subvibrisal setae; (B) Fore coxa, lateral view, spine on posterior surface; (C) Fore tarsomeres; (D) Hind femur, anterior view; (E) Hind femur, posterior view. Fannia humahuaca sp. nov., male holotype: (F) Hind femur, anterior view. Fannia kabekwa sp. nov., male holotype: (G) Fore tibia, anterior view; (H) Fore tarsi; (I) Hind femur, anterior view. Fannia palta sp. nov., male holotype: (J) First fore tarsomere, leaflike spine; (K) Fore tarsomeres; (L) Mid tibia, posterior view; (M) Hind femur, anterior view; (N) Hind femur, posterior view. Fannia polleti sp. nov., male holotype: (O) Fore tarsi; (P) Hind femur, anterior view." httpUri="https://zenodo.org/record/2622953/files/figure.png" pageId="14" pageNumber="344">Fig. 3K</figureCitation>
). Mid femur with 1 row of decreasing
<emphasis id="D3D4A553F20BFF815686BFC0FA243900" box="[1409,1436,1771,1794]" italics="true" pageId="14" pageNumber="344">av</emphasis>
from base to apex; 2 rows of long
<emphasis id="D3D4A553F20BFF815122BE24FDF83924" box="[549,576,1807,1830]" italics="true" pageId="14" pageNumber="344">pv</emphasis>
with small tuft of setae with hooked apex on apical third. Mid tibia with 2 pronounced constrictions on ventral surface on basal half area, densely setulose on entire surface,
<date id="951E5F81F20BFF8157E4BE18FABC3948" box="[1251,1284,1843,1867]" pageId="14" pageNumber="344">
1
<emphasis id="D3D4A553F20BFF8157FFBE18FABC3948" box="[1272,1284,1843,1866]" italics="true" pageId="14" pageNumber="344">v</emphasis>
</date>
; 1 developed preapical
<emphasis id="D3D4A553F20BFF815214BE7DFE97396C" box="[275,303,1878,1902]" italics="true" pageId="14" pageNumber="344">ad</emphasis>
; 1 median
<emphasis id="D3D4A553F20BFF8152AABE7CFE03396C" box="[429,443,1879,1902]" italics="true" pageId="14" pageNumber="344">a</emphasis>
and 1 apical; 1 apical
<emphasis id="D3D4A553F20BFF8151BEBE7CFD6C396C" box="[697,724,1879,1902]" italics="true" pageId="14" pageNumber="344">av</emphasis>
; 1 apical modified
<emphasis id="D3D4A553F20BFF8150A9BE7CFC70396C" box="[942,968,1879,1902]" italics="true" pageId="14" pageNumber="344">pv</emphasis>
; 2
<emphasis id="D3D4A553F20BFF8150EABE7DFBB1396C" box="[1005,1033,1878,1902]" italics="true" pageId="14" pageNumber="344">pd</emphasis>
; (
<figureCitation id="799B65C4F20BFF815726BE7DFBC0396D" box="[1057,1144,1878,1903]" captionStart="FIGURE 3" captionStartId="12.[151,250,1857,1879]" captionTargetBox="[186,1392,195,1831]" captionTargetId="figure@12.[186,1392,195,1831]" captionTargetPageId="12" captionText="FIGURE 3. Fannia huetare sp. nov., male holotype: (A) Head, anterior view, vibrissal and subvibrisal setae; (B) Fore coxa, lateral view, spine on posterior surface; (C) Fore tarsomeres; (D) Hind femur, anterior view; (E) Hind femur, posterior view. Fannia humahuaca sp. nov., male holotype: (F) Hind femur, anterior view. Fannia kabekwa sp. nov., male holotype: (G) Fore tibia, anterior view; (H) Fore tarsi; (I) Hind femur, anterior view. Fannia palta sp. nov., male holotype: (J) First fore tarsomere, leaflike spine; (K) Fore tarsomeres; (L) Mid tibia, posterior view; (M) Hind femur, anterior view; (N) Hind femur, posterior view. Fannia polleti sp. nov., male holotype: (O) Fore tarsi; (P) Hind femur, anterior view." httpUri="https://zenodo.org/record/2622953/files/figure.png" pageId="14" pageNumber="344">Fig. 3L</figureCitation>
). Hind coxa with setae on posterior margin. Hind femur on ventral surface with strong preapical protuberance and tuft of long setae on the protuberance; 1 row of
<emphasis id="D3D4A553F20BFF8152BBBEB5FE6039B4" box="[444,472,1950,1974]" italics="true" pageId="14" pageNumber="344">ad</emphasis>
, 4 stronger setae running towards dorsal surface; posterior surface with strong protuberance on median third (
<figureCitation id="799B65C4F20BFF8152FDBEE9FD8539D9" box="[506,573,1986,2011]" captionStart="FIGURE 3" captionStartId="12.[151,250,1857,1879]" captionTargetBox="[186,1392,195,1831]" captionTargetId="figure@12.[186,1392,195,1831]" captionTargetPageId="12" captionText="FIGURE 3. Fannia huetare sp. nov., male holotype: (A) Head, anterior view, vibrissal and subvibrisal setae; (B) Fore coxa, lateral view, spine on posterior surface; (C) Fore tarsomeres; (D) Hind femur, anterior view; (E) Hind femur, posterior view. Fannia humahuaca sp. nov., male holotype: (F) Hind femur, anterior view. Fannia kabekwa sp. nov., male holotype: (G) Fore tibia, anterior view; (H) Fore tarsi; (I) Hind femur, anterior view. Fannia palta sp. nov., male holotype: (J) First fore tarsomere, leaflike spine; (K) Fore tarsomeres; (L) Mid tibia, posterior view; (M) Hind femur, anterior view; (N) Hind femur, posterior view. Fannia polleti sp. nov., male holotype: (O) Fore tarsi; (P) Hind femur, anterior view." httpUri="https://zenodo.org/record/2622953/files/figure.png" pageId="14" pageNumber="344">Fig. 3</figureCitation>
MN). Hind tibia with 1 median
<emphasis id="D3D4A553F20BFF8150B9BEE8FC7439D9" box="[958,972,1987,2011]" italics="true" pageId="14" pageNumber="344">d</emphasis>
and 1 preapical; 1 median
<emphasis id="D3D4A553F20BFF81561FBEE8FA8C39D9" box="[1304,1332,1987,2011]" italics="true" pageId="14" pageNumber="344">ad</emphasis>
; 3
<emphasis id="D3D4A553F20BFF81565BBEE8FACE39D8" box="[1372,1398,1987,2010]" italics="true" pageId="14" pageNumber="344">av</emphasis>
on apical half and 1 apical; anterior surface with short setae and 1 short apical; 2 strong median
<emphasis id="D3D4A553F20AFF8057A2B9B2FB063EB2" box="[1189,1214,153,176]" italics="true" pageId="15" pageNumber="345">av</emphasis>
, 1 apical (
<figureCitation id="799B65C4F20AFF805634B9BCFACB3EB2" box="[1331,1395,151,176]" captionStart="FIGURE 3" captionStartId="12.[151,250,1857,1879]" captionTargetBox="[186,1392,195,1831]" captionTargetId="figure@12.[186,1392,195,1831]" captionTargetPageId="12" captionText="FIGURE 3. Fannia huetare sp. nov., male holotype: (A) Head, anterior view, vibrissal and subvibrisal setae; (B) Fore coxa, lateral view, spine on posterior surface; (C) Fore tarsomeres; (D) Hind femur, anterior view; (E) Hind femur, posterior view. Fannia humahuaca sp. nov., male holotype: (F) Hind femur, anterior view. Fannia kabekwa sp. nov., male holotype: (G) Fore tibia, anterior view; (H) Fore tarsi; (I) Hind femur, anterior view. Fannia palta sp. nov., male holotype: (J) First fore tarsomere, leaflike spine; (K) Fore tarsomeres; (L) Mid tibia, posterior view; (M) Hind femur, anterior view; (N) Hind femur, posterior view. Fannia polleti sp. nov., male holotype: (O) Fore tarsi; (P) Hind femur, anterior view." httpUri="https://zenodo.org/record/2622953/files/figure.png" pageId="15" pageNumber="345">Fig. 3</figureCitation>
M N).
</paragraph>
<paragraph id="E11F7941F20AFF8053C0B9F4FE013F8A" blockId="15.[151,1436,151,1040]" pageId="15" pageNumber="345">
Abdomen: elongate, black, weakly greyish pollinose with developed setae. Sternite 1 densely setulose. Sternite 5 (
<figureCitation id="799B65C4F20AFF8053BFB82FFEAE3F1F" box="[184,278,260,285]" captionStart="FIGURE 5" captionStartId="23.[151,250,1877,1899]" captionText="FIGURE 5. Fannia, male, sternite 5: (A) F. awa sp. nov.; (B) F. bari sp. nov.; (C) F. boruca sp. nov.; (D) F. bribri sp. nov.; (E) F. huetare sp. nov.; (F) F. humahuaca sp. nov.; (G) F. kabekwa sp. nov.; (H) F. palta sp. nov.; (I) F. polleti sp. nov.; (J) F. quillacingas sp. nov.; (K) F. teribi sp. nov.; (L) F. terraba sp. nov.; (M) F. tsachilas sp. nov.; (N) F. yukpa sp. nov. Scale: 1 mm." pageId="15" pageNumber="345">Fig. 5H</figureCitation>
). Terminalia (
<figureCitation id="799B65C4F20AFF8052BEB82FFDA03F1F" box="[441,536,260,285]" captionStart="FIGURE 6" captionStartId="24.[151,250,1904,1926]" captionTargetBox="[206,1368,187,1878]" captionTargetId="figure@24.[171,1417,187,1890]" captionTargetPageId="24" captionText="FIGURE 6. Fannia, male, epandrium, cercal plate and surstyli, dorsal view: (A) F. awa sp. nov.; (B) F. bari sp. nov.; (C) F. boruca sp. nov.; (D) F. bribri sp. nov.; (E) F. huetare sp. nov.; (F) F. humahuaca sp. nov.; (G) F. kabekwa sp. nov.; (H) F. palta sp. nov. Scale: 1 mm." httpUri="https://zenodo.org/record/2622957/files/figure.png" pageId="15" pageNumber="345">Fig. 6H</figureCitation>
): epandrium wider than long, strong setae on basal half, short apical internal expansions; cercal plate elongate, globose, apically wide, short weak setae on the plate; surstylus articulates with epandrium, curved apically, shaped as a comma in lateral view, basal half with internal basal expansion; bacilliform process spiral and simple.
</paragraph>
<paragraph id="E11F7941F20AFF8053C0B8BFFE253FAF" blockId="15.[151,1436,151,1040]" box="[199,413,404,429]" pageId="15" pageNumber="345">
<emphasis id="D3D4A553F20AFF8053C0B8BFFEA63FAF" bold="true" box="[199,286,404,429]" pageId="15" pageNumber="345">Female</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="A9BA2ACAF20AFF8053C0B89CFAF83FF7" pageId="15" pageNumber="345" type="biology_ecology">
<paragraph id="E11F7941F20AFF8053C0B89CFAF83FF7" blockId="15.[151,1436,151,1040]" pageId="15" pageNumber="345">
<emphasis id="D3D4A553F20AFF8053C0B89CFEA73FD2" bold="true" box="[199,287,439,464]" pageId="15" pageNumber="345">Biology</emphasis>
. Label information indicates that this species occurs in the Andean highlands, above
<quantity id="2658D4A4F20AFF8057FAB893FAEF3FCD" box="[1277,1367,440,464]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="345" unit="m" value="3000.0">3000 m</quantity>
, from southern
<collectingCountry id="99B739D1F20AFF8053F9B8F7FEE33FF7" box="[254,347,476,501]" name="Ecuador" pageId="15" pageNumber="345">Ecuador</collectingCountry>
to the Central Cordillera in
<collectingCountry id="99B739D1F20AFF805190B8F7FCB13FF7" box="[663,777,476,501]" name="Colombia" pageId="15" pageNumber="345">Colombia</collectingCountry>
. Nothing else is known about its habits or biology.
</paragraph>
</subSubSection>
<subSubSection id="A9BA2ACAF20AFF8053C0B8D4FD443C87" pageId="15" pageNumber="345" type="discussion">
<paragraph id="E11F7941F20AFF8053C0B8D4FD443C87" blockId="15.[151,1436,151,1040]" pageId="15" pageNumber="345">
<emphasis id="D3D4A553F20AFF8053C0B8D4FEF03C1A" bold="true" box="[199,328,511,536]" pageId="15" pageNumber="345">Comments</emphasis>
.
<taxonomicName id="26A002C2F20AFF805251BB2BFE503C1A" authority="Grisales &amp; De Carvalho, 2019" authorityName="Grisales &amp; De Carvalho" authorityYear="2019" box="[342,488,512,536]" class="Insecta" family="Fanniidae" genus="Fannia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="345" phylum="Arthropoda" rank="species" species="palta" status="sp. nov.">
<emphasis id="D3D4A553F20AFF805251BB2BFE503C1A" box="[342,488,512,536]" italics="true" pageId="15" pageNumber="345">Fannia palta</emphasis>
</taxonomicName>
<emphasis id="D3D4A553F20AFF8052E8B8D4FDF13C1A" bold="true" box="[495,585,511,536]" pageId="15" pageNumber="345">
<taxonomicNameLabel id="C8E71828F20AFF8052E8B8D4FDF13C1A" box="[495,585,511,536]" pageId="15" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically similar to
<taxonomicName id="26A002C2F20AFF80509ABB2BFB5F3C1A" authority="Stein, 1911" authorityName="Stein" authorityYear="1911" box="[925,1255,511,536]" class="Insecta" family="Fanniidae" genus="Fannia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="345" phylum="Arthropoda" rank="species" species="albitarsis">
<emphasis id="D3D4A553F20AFF80509ABB2BFBD83C1A" box="[925,1120,512,536]" italics="true" pageId="15" pageNumber="345">Fannia albitarsis</emphasis>
<bibRefCitation id="853104B0F20AFF805760B8D4FB5F3C1A" author="Stein, P." box="[1127,1255,511,536]" pageId="15" pageNumber="345" pagination="61 - 189" refId="ref26397" refString="Stein, P. (1911) Die von Schnuse in Sudamerika gefangenen Anthomyiden. Archiv fur Naturgeschichte, 77, 61 - 189." type="journal article" year="1911">Stein, 1911</bibRefCitation>
</taxonomicName>
, in the first fore tarsomere having 1 broad leaflike spine at the tip of posterior surface but differ in the fore tarsi that are not strong, flattened and yellowish.
<taxonomicName id="26A002C2F20AFF8052B1BB63FDF43C62" authority="Grisales &amp; De Carvalho, 2019" authorityName="Grisales &amp; De Carvalho" authorityYear="2019" box="[438,588,584,608]" class="Insecta" family="Fanniidae" genus="Fannia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="345" phylum="Arthropoda" rank="species" species="palta" status="sp. nov.">
<emphasis id="D3D4A553F20AFF8052B1BB63FDF43C62" box="[438,588,584,608]" italics="true" pageId="15" pageNumber="345">Fannia palta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8E71828F20AFF805151BB63FD093C5D" box="[598,689,584,607]" pageId="15" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
has fore tarsomeres 1 and 2 with apical half yellowish and the remainder brownish, tarsomeres 35 slightly flattened.
</paragraph>
</subSubSection>
<subSubSection id="A9BA2ACAF20AFF8053C0BBA4FDDD3CCE" pageId="15" pageNumber="345" type="etymology">
<paragraph id="E11F7941F20AFF8053C0BBA4FDDD3CCE" blockId="15.[151,1436,151,1040]" pageId="15" pageNumber="345">
<emphasis id="D3D4A553F20AFF8053C0BBA4FEFE3CAA" bold="true" box="[199,326,655,680]" pageId="15" pageNumber="345">Etymology</emphasis>
. Name in apposition. The species epithet refers to the Palta indigenous people that once inhabited the Province of
<collectingRegion id="2364B7A3F20AFF80524EBB9FFEC43CCF" box="[329,380,692,717]" country="Ecuador" name="Loja" pageId="15" pageNumber="345">Loja</collectingRegion>
in the pre-Incan era.
</paragraph>
</subSubSection>
<subSubSection id="A9BA2ACAF20AFF8053C0BBFCFBB63DCA" pageId="15" pageNumber="345" type="materials_examined">
<paragraph id="E11F7941F20AFF8053C0BBFCFF493D82" blockId="15.[151,1436,151,1040]" pageId="15" pageNumber="345">
<emphasis id="D3D4A553F20AFF8053C0BBFCFEC93CF2" bold="true" box="[199,369,727,752]" pageId="15" pageNumber="345">Type material</emphasis>
.
<materialsCitation id="51C8731CF20AFF805285BBFCFC2C3D3A" ID-GBIF-Occurrence="2234220113" collectingDate="2009" collectingDateMax="2037" collectingDateMin="2009" collectionCode="UTPL, EC, YPT, MP" collectorName="de Anteojos &amp; Marc Pollet" country="Ecuador" county="Cajanuma" elevation="3000" location="Oso" municipality="Podocarpus National Park" pageId="15" pageNumber="345" specimenCount="1" stateProvince="Loja" typeStatus="holotype">
<emphasis id="D3D4A553F20AFF805285BBFCFE573CF2" bold="true" box="[386,495,727,752]" pageId="15" pageNumber="345">
<typeStatus id="3E1BC7E3F20AFF805285BBFCFE573CF2" box="[386,495,727,752]" pageId="15" pageNumber="345" type="holotype">Holotype</typeStatus>
</emphasis>
male (
<collectionCode id="87B1E184F20AFF805145BBF3FD363CF2" box="[578,654,728,752]" pageId="15" pageNumber="345">UTPL</collectionCode>
).
<collectingCountry id="99B739D1F20AFF8051A3BBF3FC883CF2" box="[676,816,728,752]" name="Ecuador" pageId="15" pageNumber="345">ECUADOR</collectingCountry>
: 13155 (
<collectionCode id="87B1E184F20AFF8050AABBF3FC6B3CF2" box="[941,979,728,752]" pageId="15" pageNumber="345">EC</collectionCode>
)
<collectingRegion id="2364B7A3F20AFF8050E1BBFCFBA63CF2" box="[998,1054,727,752]" country="Ecuador" name="Loja" pageId="15" pageNumber="345">Loja</collectingRegion>
:
<collectingCounty id="087E01CDF20AFF80572CBBFCFB243CF2" box="[1067,1180,727,752]" pageId="15" pageNumber="345">Cajanuma</collectingCounty>
/
<collectingMunicipality id="017BE33BF20AFF8057A8BBFCFF753D17" pageId="15" pageNumber="345">Podocarpus National Park</collectingMunicipality>
, trail
<location id="E47F2F9AF20AFF805214BBD7FE873D16" LSID="urn:lsid:plazi:treatment:6909C857F20BFF805390BD14FE7D3A12:E47F2F9AF20AFF805214BBD7FE873D16" box="[275,319,764,788]" country="Ecuador" county="Cajanuma" municipality="Podocarpus National Park" name="Oso" pageId="15" pageNumber="345" stateProvince="Loja">Oso</location>
/
<collectorName id="4C551C97F20AFF805255BBD7FE673D17" box="[338,479,764,789]" pageId="15" pageNumber="345">de Anteojos</collectorName>
,
<quantity id="2658D4A4F20AFF8052E9BBD7FD873D16" box="[494,575,764,788]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="345" unit="m" value="3000.0">
<elevation id="6A8D9E72F20AFF8052E9BBD7FD873D16" box="[494,575,764,788]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="345" unit="m" value="3000.0">3000m</elevation>
</quantity>
,
<date id="951E5F81F20AFF80514ABBD7FD743D16" box="[589,716,764,788]" pageId="15" pageNumber="345" value="2009-02-27">
<collectingDate id="855AA669F20AFF80514ABBD7FD743D16" box="[589,716,764,788]" pageId="15" pageNumber="345" value="2009-02-27">27/02/2009</collectingDate>
</date>
<date id="951E5F81F20AFF8051E8BBD7FCD93D17" box="[751,865,764,789]" pageId="15" pageNumber="345" value="2009-03-05">
<collectingDate id="855AA669F20AFF8051E8BBD7FCD93D17" box="[751,865,764,789]" pageId="15" pageNumber="345" value="2009-03-05">5/03/2009</collectingDate>
</date>
(
<collectionCode id="87B1E184F20AFF805074BBD7FC153D16" box="[883,941,764,788]" pageId="15" pageNumber="345">YPT</collectionCode>
), leg.
<collectorName id="4C551C97F20AFF8050FFBBD6FBC63D17" box="[1016,1150,764,789]" pageId="15" pageNumber="345">Marc Pollet</collectorName>
&amp;/
<location id="E47F2F9AF20AFF8057B6BBD7FA203D17" LSID="urn:lsid:plazi:treatment:6909C857F20BFF805390BD14FE7D3A12:E47F2F9AF20AFF8057B6BBD7FA203D17" box="[1201,1432,764,789]" country="Ecuador" county="Cajanuma" municipality="Podocarpus National Park" name="Anja De Braekeleer" pageId="15" pageNumber="345" stateProvince="Loja">Anja De Braekeleer</location>
, sorted by
<collectorName id="4C551C97F20AFF805202BA0BFE303D3A" box="[261,392,799,824]" pageId="15" pageNumber="345">Marc Pollet</collectorName>
sample code: EC/
<collectingDate id="855AA669F20AFF80516DBA0BFD613D3A" box="[618,729,800,824]" pageId="15" pageNumber="345" value="2009" valueMax="2037">
<date id="951E5F81F20AFF80516DBA0BFD613D3A" box="[618,729,800,824]" pageId="15" pageNumber="345" value="2009" valueMax="2037">2009 37</date>
</collectingDate>
/
<collectionCode id="87B1E184F20AFF8051E6BA0BFCB33D3A" box="[737,779,800,824]" country="0" name="Mohonk Preserve, Inc." pageId="15" pageNumber="345">MP</collectionCode>
&amp;ACB011
</materialsCitation>
.
<materialsCitation id="51C8731CF20AFF805099BA34FF553D82" ID-GBIF-Occurrence="2234220147" collectionCode="CEUA" collectorName="El Congo. &amp; L. Rios" country="Colombia" elevation="3000" location="San Jose" pageId="15" pageNumber="345" specimenCount="1" specimenCount-male="1" stateProvince="Antioquia" typeStatus="paratype">
<emphasis id="D3D4A553F20AFF805099BA34FBB33D3A" bold="true" box="[926,1035,799,824]" pageId="15" pageNumber="345">
<typeStatus id="3E1BC7E3F20AFF805099BA34FBB33D3A" box="[926,1035,799,824]" pageId="15" pageNumber="345" type="paratype">Paratype</typeStatus>
</emphasis>
.
<collectingCountry id="99B739D1F20AFF80571EBA0BFB0A3D3A" box="[1049,1202,800,824]" name="Colombia" pageId="15" pageNumber="345">COLOMBIA</collectingCountry>
.
<collectingRegion id="2364B7A3F20AFF8057BBBA34FA883D3A" box="[1212,1328,799,824]" country="Colombia" name="Antioquia" pageId="15" pageNumber="345">Antioquia</collectingRegion>
:
<location id="E47F2F9AF20AFF80563CBA0BFA243D3A" LSID="urn:lsid:plazi:treatment:6909C857F20BFF805390BD14FE7D3A12:E47F2F9AF20AFF80563CBA0BFA243D3A" box="[1339,1436,800,824]" country="Colombia" name="San Jose" pageId="15" pageNumber="345" stateProvince="Antioquia">San José</location>
de la montaña/
<collectorName id="4C551C97F20AFF80524EBA6FFE793D5E" box="[329,449,836,861]" pageId="15" pageNumber="345">El Congo.</collectorName>
1015\set\2011/
<quantity id="2658D4A4F20AFF805186BA6FFD6A3D5F" box="[641,722,836,861]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="345" unit="m" value="3000.0">
<elevation id="6A8D9E72F20AFF805186BA6FFD6A3D5F" box="[641,722,836,861]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="345" unit="m" value="3000.0">3000m</elevation>
</quantity>
.
<collectorName id="4C551C97F20AFF8051E4BA6FFC943D5F" box="[739,812,836,861]" pageId="15" pageNumber="345">L.Rios</collectorName>
/VSR pez trampa 5 (
<specimenCount id="F7A6B2C8F20AFF805728BA6FFB3A3D5F" box="[1071,1154,836,861]" pageId="15" pageNumber="345" type="male">1 male</specimenCount>
,
<collectionCode id="87B1E184F20AFF805793BA6FFB5D3D5E" box="[1172,1253,836,860]" pageId="15" pageNumber="345">CEUA</collectionCode>
). Voucher code DG013
</materialsCitation>
.
</paragraph>
<paragraph id="E11F7941F20AFF8053C0BAA7FBB63DCA" blockId="15.[151,1436,151,1040]" pageId="15" pageNumber="345">
The abdomen of
<typeStatus id="3E1BC7E3F20AFF805280BAA7FE513DA7" box="[391,489,908,933]" pageId="15" pageNumber="345" type="holotype">holotype</typeStatus>
was taken to dissect the terminalia and it was fixed with the
<typeStatus id="3E1BC7E3F20AFF80579FBAA7FB463DA7" box="[1176,1278,908,933]" pageId="15" pageNumber="345" type="holotype">holotype</typeStatus>
. The
<typeStatus id="3E1BC7E3F20AFF80563BBAA6FA243DA7" box="[1340,1436,909,933]" pageId="15" pageNumber="345" type="paratype">paratype</typeStatus>
was used as voucher in a molecular study of
<taxonomicName id="26A002C2F20AFF80518BBA84FD423DCA" box="[652,762,943,968]" class="Insecta" family="Fanniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="345" phylum="Arthropoda" rank="family">Fanniidae</taxonomicName>
(Voucher code DG013).
</paragraph>
</subSubSection>
<subSubSection id="A9BA2ACAF20AFF8053C0BAFFFE7D3A12" pageId="15" pageNumber="345" type="distribution">
<paragraph id="E11F7941F20AFF8053C0BAFFFE7D3A12" blockId="15.[151,1436,151,1040]" pageId="15" pageNumber="345">
<emphasis id="D3D4A553F20AFF8053C0BAFFFEE13DEF" bold="true" box="[199,345,980,1005]" pageId="15" pageNumber="345">Distribution</emphasis>
.
<collectingCountry id="99B739D1F20AFF80526EBAFFFE643DEF" box="[361,476,980,1005]" name="Colombia" pageId="15" pageNumber="345">Colombia</collectingCountry>
:
<collectingRegion id="2364B7A3F20AFF8052EFBAFFFD673DEF" box="[488,735,980,1005]" country="Colombia" name="Antioquia" pageId="15" pageNumber="345">Antioquia department</collectingRegion>
(Central Cordillera, Andean region);
<collectingCountry id="99B739D1F20AFF805797BAFFFB4A3DEF" box="[1168,1266,980,1005]" name="Ecuador" pageId="15" pageNumber="345">Ecuador</collectingCountry>
:
<collectingRegion id="2364B7A3F20AFF8057F9BAFFFA243DEF" box="[1278,1436,980,1005]" country="Ecuador" name="Loja" pageId="15" pageNumber="345">Loja province</collectingRegion>
(Andean region) (
<figureCitation id="799B65C4F20AFF805259BADCFE003A12" box="[350,440,1015,1040]" captionStart="FIGURE 8" captionStartId="26.[151,250,1774,1796]" captionTargetBox="[150,1436,193,1753]" captionTargetId="figure@26.[150,1436,193,1753]" captionTargetPageId="26" captionText="FIGURE 8. Fannia, maps of distribution: (A) F. awa sp. nov.; (B) F. bari sp. nov.; (C) F. boruca sp. nov.; (D) F. bribri sp. nov.; (E) F. huetare sp. nov.; (F) F. humahuaca sp. nov.; (G) F. kabekwa sp. nov.; (H) F. palta sp. nov.; (I) F. polleti sp. nov.; (J) F. quillacingas sp. nov.; (K) F. teribi sp. nov.; (L) F. terraba sp. nov.; (M) F. tsachilas sp. nov.; (N) F. yukpa sp. nov." httpUri="https://zenodo.org/record/2622962/files/figure.png" pageId="15" pageNumber="345">Fig. 8H</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>