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<document id="DF424AD5E59903E6F0F53F1E9C633A0E" ID-DOI="10.11646/zootaxa.3888.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10086415" ID-ZooBank="1B437AE1-CF28-4C1B-95B6-C31A295905A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623730057475" checkinUser="felipe" docAuthor="Schwarzhans, Werner" docDate="2014" docId="463A8F36FF89FFC2129799D8E3BEFBE8" docLanguage="en" docName="zt03888p073.pdf.imf" docOrigin="Zootaxa 3888 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hymenocephalus aeger Gilbert &amp; Hubbs 1920" docType="treatment" docVersion="2" lastPageNumber="59" masterDocId="BA03F74EFFB0FFF812009814E16BFFF8" masterDocTitle="Head and otolith morphology of the genera Hymenocephalus, Hymenogadus and Spicomacrurus (Macrouridae), with the description of three new species" masterLastPageNumber="73" masterPageNumber="1" pageNumber="58" updateTime="1699488604623" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods id="D6E4703C08B02849A79115D1DE5B9A9F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="423D2DD32DADF9FFF92B5848AD4C9773">Head and otolith morphology of the genera Hymenocephalus, Hymenogadus and Spicomacrurus (Macrouridae), with the description of three new species</mods:title>
</mods:titleInfo>
<mods:name id="0042723A2C2331E8E29988B9882EA9C6" type="personal">
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<mods:namePart id="932653202363B9F53A97E70202615456">Schwarzhans, Werner</mods:namePart>
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<mods:title id="F4B036C792EC6F396BEE174DF14E745A">Zootaxa</mods:title>
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<mods:part id="C9BB8BE1971B52D4DFB345FFA3A005E6">
<mods:date id="493A0015B49D5B92A929EBAFDD60A7A0">2014</mods:date>
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<mods:number id="8A913F3500688B1AB3159B3AD58BE4D1">2014-11-28</mods:number>
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<mods:detail id="AD031D86B6ED72CA4047EDF7B1EF2AE1" type="volume">
<mods:number id="F194429F749503DF21A2C742111C5448">3888</mods:number>
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<mods:number id="984FE8BCABC249F6F16B50EC41E9FF3A">1</mods:number>
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<mods:start id="7B90FFBA21E170608F6FC96F1B2B5E7F">1</mods:start>
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<mods:classification id="C51B1FC82B036946271D14A2265F61DC">journal article</mods:classification>
<mods:identifier id="F85D1AE623226D8F867A286CB5B0AB82" type="DOI">10.11646/zootaxa.3888.1.1</mods:identifier>
<mods:identifier id="833359C1DE8CE99AB87D38317C8102A9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4E8307FFAA950AAE2AE8B9F278404F07" type="Zenodo-Dep">10086415</mods:identifier>
<mods:identifier id="3539E4D9236A300DD03C664537276837" type="ZooBank">1B437AE1-CF28-4C1B-95B6-C31A295905A0</mods:identifier>
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<treatment id="463A8F36FF89FFC2129799D8E3BEFBE8" LSID="urn:lsid:plazi:treatment:463A8F36FF89FFC2129799D8E3BEFBE8" httpUri="http://treatment.plazi.org/id/463A8F36FF89FFC2129799D8E3BEFBE8" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
<subSubSection id="86896DABFF89FFC1129799D8E3B2FE1E" box="[151,729,460,486]" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="CE2C3E20FF89FFC1129799D8E3B2FE1E" blockId="57.[151,729,460,519]" box="[151,729,460,486]" pageId="57" pageNumber="58">
<heading id="9564894CFF89FFC1129799D8E3B2FE1E" bold="true" box="[151,729,460,486]" fontSize="11" level="2" pageId="57" pageNumber="58" reason="4">
<taxonomicName id="099345A3FF89FFC1129799D8E3B2FE1E" ID-CoL="3NJ8K" authority="Gilbert &amp; Hubbs, 1920" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[151,729,460,486]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="aeger">
<emphasis id="FCE7E232FF89FFC1129799D8E3B2FE1E" bold="true" box="[151,729,460,486]" pageId="57" pageNumber="58">
<emphasis id="FCE7E232FF89FFC1129799D8E0D8FE1E" bold="true" box="[151,435,460,486]" italics="true" pageId="57" pageNumber="58">Hymenocephalus aeger</emphasis>
<bibRefCitation id="AA0243D1FF89FFC113BA99D8E3B2FE1E" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[442,729,460,486]" pageId="57" pageNumber="58" pagination="369 - 588" refId="ref50763" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1920) The macrouroid fishes of the Philippine Islands and the East Indies. United States National Museum Bulletin, 100 (1), 369 - 588. http: // dx. doi. org / 10.5962 / bhl. title. 13637" type="journal article" year="1920">Gilbert &amp; Hubbs, 1920</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF89FFC1129799FAE025FDFF" box="[151,334,494,519]" pageId="57" pageNumber="58" type="description">
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<figureCitation id="56A822A5FF89FFC1129799FAE043FDFF" box="[151,296,494,519]" captionStart="FIGURE 30" captionStartId="57.[151,250,1804,1826]" captionTargetBox="[181,1407,1109,1774]" captionTargetId="figure-361@57.[175,1412,1103,1779]" captionTargetPageId="57" captionText="FIGURE 30. Hymenocephalus aeger: A—ZMUC P375844, lateral view of head. B–H—Otoliths, B—Anterior view, C, E–H—Inner faces, D—Ventral View, B–D—ZMUC P375844, E—ZMUC unregistered, 05°46’S, 132°52’E, F—MNHN 2006- 0218, G—ZMUC P375857, H—ZMUC P375856." figureDoi="http://doi.org/10.5281/zenodo.10086479" httpUri="https://zenodo.org/record/10086479/files/figure.png" pageId="57" pageNumber="58">Figs. 30A–G</figureCitation>
,
<figureCitation id="56A822A5FF89FFC1133299FAE025FDFF" box="[306,334,494,519]" captionStart="FIGURE 37" captionStartId="66.[151,250,976,998]" captionTargetBox="[158,1430,272,949]" captionTargetId="figure-236@66.[152,1435,265,954]" captionTargetPageId="66" captionText="FIGURE 37. Geographical distribution of Hymenocephalus aeger, H. megalops, H. striatissimus, H. torvus, H. grimaldii, H. neglectissimus, H. semipellucidus, H. sp. 1 and H." figureDoi="http://doi.org/10.5281/zenodo.10086493" httpUri="https://zenodo.org/record/10086493/files/figure.png" pageId="57" pageNumber="58">37</figureCitation>
</paragraph>
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<paragraph id="CE2C3E20FF89FFC112979A22E5F6FDB4" blockId="57.[151,1181,566,619]" box="[151,1181,566,588]" pageId="57" pageNumber="58">
<treatmentCitationGroup id="EE83190EFF89FFC112979A22E5F6FDB4" box="[151,1181,566,588]" pageId="57" pageNumber="58">
<treatmentCitation id="4F321831FF89FFC112979A22E245FDB4" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[151,814,566,588]" page="531" pageId="57" pageNumber="58" year="1920">
<taxonomicName id="099345A3FF89FFC112979A22E245FDB4" ID-CoL="3NJ8K" authority="Gilbert &amp; Hubbs, 1920: 531" authorityName="Gilbert &amp; Hubbs" authorityPageNumber="531" authorityYear="1920" box="[151,814,566,588]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="57" pageNumber="58" phylum="Chordata" rank="subSpecies" species="striatissimus" subSpecies="aeger">
<emphasis id="FCE7E232FF89FFC112979A22E367FDB4" box="[151,524,566,588]" italics="true" pageId="57" pageNumber="58">Hymenocephalus striatissimus aeger</emphasis>
<bibRefCitation id="AA0243D1FF89FFC110119A22E245FDB4" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[529,814,566,588]" pageId="57" pageNumber="58" pagination="369 - 588" refId="ref50763" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1920) The macrouroid fishes of the Philippine Islands and the East Indies. United States National Museum Bulletin, 100 (1), 369 - 588. http: // dx. doi. org / 10.5962 / bhl. title. 13637" type="journal article" year="1920">Gilbert &amp; Hubbs, 1920: 531</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
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locality:
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,
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).
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</paragraph>
<paragraph id="CE2C3E20FF89FFC112979A41E203FD93" blockId="57.[151,1181,566,619]" box="[151,872,597,619]" pageId="57" pageNumber="58">
<treatmentCitationGroup id="EE83190EFF89FFC112979A41E203FD93" box="[151,872,597,619]" pageId="57" pageNumber="58">
<treatmentCitation id="4F321831FF89FFC112979A41E20FFD93" author="Iwamoto, T. &amp; Williams, A." box="[151,868,597,619]" page="178" pageId="57" pageNumber="58" year="1999">
<taxonomicName id="099345A3FF89FFC112979A41E20FFD93" ID-CoL="3NJ8K" authority=": Iwamoto &amp; Williams, 1999: 178" authorityName=": Iwamoto &amp; Williams" authorityPageNumber="178" authorityYear="1999" box="[151,868,597,619]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="57" pageNumber="58" phylum="Chordata" rank="subSpecies" species="striatissimus" subSpecies="aeger">
<emphasis id="FCE7E232FF89FFC112979A41E367FD93" box="[151,524,597,619]" italics="true" pageId="57" pageNumber="58">Hymenocephalus striatissimus aeger</emphasis>
:
<bibRefCitation id="AA0243D1FF89FFC110189A42E20FFD93" author="Iwamoto, T. &amp; Williams, A." box="[536,868,597,619]" pageId="57" pageNumber="58" pagination="105 - 243" refId="ref51219" refString="Iwamoto, T. &amp; Williams, A. (1999) Grenadiers (Pisces, Gadiformes) from the continental slope of Western and northwestern Australia. Proceedings of the California Academy of Sciences, 51 (3), 105 - 243." type="journal article" year="1999">Iwamoto &amp; Williams, 1999: 178</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="86896DABFF89FFC112979A83E01FFC9D" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="CE2C3E20FF89FFC112979A83E01FFC9D" blockId="57.[151,1437,663,1048]" pageId="57" pageNumber="58">
<emphasis id="FCE7E232FF89FFC112979A83E0E9FD48" bold="true" box="[151,386,663,688]" pageId="57" pageNumber="58">Material examined.</emphasis>
<specimenCount id="D895F5A9FF89FFC1138A9A83E34DFD48" box="[394,550,663,688]" pageId="57" pageNumber="58" type="generic">11 specimens</specimenCount>
;
<materialsCitation id="7EFB347DFF89FFC110339A83E465FD57" box="[563,1294,663,688]" collectionCode="MNHN" elevation="523" latitude="-7.7333336" longLatPrecision="1303" longitude="156.48334" pageId="57" pageNumber="58" specimenCode="MNHN 2006-0159" specimenCount="1">
<specimenCount id="D895F5A9FF89FFC110339A83E3DFFD48" box="[563,692,663,688]" pageId="57" pageNumber="58" type="generic">1 specimen</specimenCount>
<specimenCode id="9E35965BFF89FFC110BC9A8CE2FDFD48" box="[700,918,663,688]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="57" pageNumber="58">MNHN 2006-0159</specimenCode>
,
<geoCoordinate id="ABA758E7FF89FFC111A39A83E56AFD48" box="[931,1025,663,688]" degrees="07" direction="south" minutes="44" orientation="latitude" pageId="57" pageNumber="58" precision="925" value="-7.7333336">07°44’S</geoCoordinate>
,
<geoCoordinate id="ABA758E7FF89FFC1160F9A83E516FD48" box="[1039,1149,663,688]" degrees="156" direction="east" minutes="29" orientation="longitude" pageId="57" pageNumber="58" precision="925" value="156.48334">156°29’E</geoCoordinate>
,
<quantity id="096B93C5FF89FFC1168A9A83E465FD57" box="[1162,1294,663,688]" metricMagnitude="2" metricUnit="m" metricValue="5.225" metricValueMax="5.27" metricValueMin="5.18" pageId="57" pageNumber="58" unit="m" value="522.5" valueMax="527.0" valueMin="518.0">
<elevation id="45BED913FF89FFC1168A9A83E465FD57" box="[1162,1294,663,688]" metricMagnitude="2" metricUnit="m" metricValue="5.225" metricValueMax="5.27" metricValueMin="5.18" pageId="57" pageNumber="58" unit="m" value="522.5" valueMax="527.0" valueMin="518.0">518–527 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="7EFB347DFF89FFC1171B9A83E38EFD2B" collectionCode="MNHN" elevation="474" latitude="-8.15" longLatPrecision="1302" longitude="157.01666" pageId="57" pageNumber="58" specimenCode="MNHN 2006-0218" specimenCount="1">
<specimenCount id="D895F5A9FF89FFC1171B9A83E4F7FD48" box="[1307,1436,663,688]" pageId="57" pageNumber="58" type="generic">1 specimen</specimenCount>
<specimenCode id="9E35965BFF89FFC112979AA8E01BFD2D" box="[151,368,700,725]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="57" pageNumber="58">MNHN 2006-0218</specimenCode>
,
<geoCoordinate id="ABA758E7FF89FFC1137C9AA8E0B0FD2D" box="[380,475,700,725]" degrees="08" direction="south" minutes="09" orientation="latitude" pageId="57" pageNumber="58" precision="925" value="-8.15">08°09’S</geoCoordinate>
,
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,
<quantity id="096B93C5FF89FFC110629AA8E38EFD2B" box="[610,741,700,725]" metricMagnitude="2" metricUnit="m" metricValue="4.735" metricValueMax="4.87" metricValueMin="4.6" pageId="57" pageNumber="58" unit="m" value="473.5" valueMax="487.0" valueMin="460.0">
<elevation id="45BED913FF89FFC110629AA8E38EFD2B" box="[610,741,700,725]" metricMagnitude="2" metricUnit="m" metricValue="4.735" metricValueMax="4.87" metricValueMin="4.6" pageId="57" pageNumber="58" unit="m" value="473.5" valueMax="487.0" valueMin="460.0">460–487 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="7EFB347DFF89FFC110F19AA8E037FD0F" collectionCode="MNHN" elevation="430" latitude="-8.266666" longLatPrecision="1301" longitude="160.06667" pageId="57" pageNumber="58" specimenCode="MNHN 2006-0556, 91" specimenCount="3">
<specimenCount id="D895F5A9FF89FFC110F19AA8E217FD2D" box="[753,892,700,725]" pageId="57" pageNumber="58" type="generic">3 specimens</specimenCount>
<specimenCode id="9E35965BFF89FFC111839AA8E536FD2D" box="[899,1117,700,725]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="57" pageNumber="58">MNHN 2006-0556</specimenCode>
,
<specimenCode id="9E35965BFF89FFC116699AA8E5EEFD2D" box="[1129,1157,700,725]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="57" pageNumber="58">91</specimenCode>
+ –
<quantity id="096B93C5FF89FFC116AC9AA8E463FD2B" box="[1196,1288,700,725]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="57" pageNumber="58" unit="mm" value="140.0">140 mm</quantity>
TL,
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,
<geoCoordinate id="ABA758E7FF89FFC112979ACBE06FFD00" box="[151,260,735,760]" degrees="160" direction="east" minutes="04" orientation="longitude" pageId="57" pageNumber="58" precision="925" value="160.06667">160°04’E</geoCoordinate>
,
<quantity id="096B93C5FF89FFC113119ACBE037FD0F" box="[273,348,735,760]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="57" pageNumber="58" unit="m" value="430.0">
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</quantity>
</materialsCitation>
;
<materialsCitation id="7EFB347DFF89FFC113689ACBE52EFD00" box="[360,1093,735,760]" collectionCode="WAM" latitude="-17.95" longLatPrecision="1277" longitude="118.28333" pageId="57" pageNumber="58" specimenCount="1">
<specimenCount id="D895F5A9FF89FFC113689ACBE083FD00" box="[360,488,735,760]" pageId="57" pageNumber="58" type="generic">1 specimen</specimenCount>
<collectionCode id="A882A6E5FF89FFC113EF9ACBE358FD00" box="[495,563,735,760]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="57" pageNumber="58" type="Museum">WAM</collectionCode>
P.30578-002,
<quantity id="096B93C5FF89FFC110D79ACBE25EFD0F" box="[727,821,735,760]" metricMagnitude="-1" metricUnit="m" metricValue="1.21" pageId="57" pageNumber="58" unit="mm" value="121.0">121 mm</quantity>
TL,
<geoCoordinate id="ABA758E7FF89FFC1116D9ACBE2A0FD00" box="[877,971,735,760]" degrees="17" direction="south" minutes="57" orientation="latitude" pageId="57" pageNumber="58" precision="925" value="-17.95">17°57’S</geoCoordinate>
,
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</materialsCitation>
;
<materialsCitation id="7EFB347DFF89FFC116529ACBE0C6FCE5" collectionCode="ZMUC" location="Kai Islands" pageId="57" pageNumber="58" specimenCode="P375844, 84" specimenCount="1">
<specimenCount id="D895F5A9FF89FFC116529ACBE5B9FD00" box="[1106,1234,735,760]" pageId="57" pageNumber="58" type="generic">1 specimen</specimenCount>
<collectionCode id="A882A6E5FF89FFC116D99ACBE441FD00" box="[1241,1322,735,760]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="57" pageNumber="58">ZMUC</collectionCode>
<specimenCode id="9E35965BFF89FFC117319ACBE4F3FD00" box="[1329,1432,735,760]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="57" pageNumber="58">P375844</specimenCode>
,
<specimenCode id="9E35965BFF89FFC112979B10E1D8FCE5" box="[151,179,772,797]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="57" pageNumber="58">84</specimenCode>
+ mm TL,
<location id="CB4C68FBFF89FFC1132B9B10E0C6FCE5" LSID="urn:lsid:plazi:treatment:463A8F36FF89FFC2129799D8E3BEFBE8:CB4C68FBFF89FFC1132B9B10E0C6FCE5" box="[299,429,772,797]" name="Kai Islands" pageId="57" pageNumber="58">Kai Islands</location>
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;
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<collectionCode id="A882A6E5FF89FFC1103C9B10E3E6FCE5" box="[572,653,772,797]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="57" pageNumber="58">ZMUC</collectionCode>
P
<quantity id="096B93C5FF89FFC110A39B10E20AFCE3" box="[675,865,772,797]" metricMagnitude="5" metricUnit="m" metricValue="3.75854127" pageId="57" pageNumber="58" unit="mm" value="3.75854127E8">375854, 127 mm</quantity>
TL,
<location id="CB4C68FBFF89FFC111979B10E572FCE5" LSID="urn:lsid:plazi:treatment:463A8F36FF89FFC2129799D8E3BEFBE8:CB4C68FBFF89FFC111979B10E572FCE5" box="[919,1049,772,797]" name="Kai Islands" pageId="57" pageNumber="58">Kai Islands</location>
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;
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<specimenCount id="D895F5A9FF89FFC116239B10E5C9FCE5" box="[1059,1186,772,797]" pageId="57" pageNumber="58" type="generic">1 specimen</specimenCount>
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<specimenCode id="9E35965BFF89FFC116FF9B10E40DFCE5" box="[1279,1382,772,797]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="57" pageNumber="58">P375856</specimenCode>
,
<specimenCode id="9E35965BFF89FFC117709B10E4E7FCE5" box="[1392,1420,772,797]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="57" pageNumber="58">91</specimenCode>
+ mm TL,
<location id="CB4C68FBFF89FFC112FF9B33E0EFFCB8" LSID="urn:lsid:plazi:treatment:463A8F36FF89FFC2129799D8E3BEFBE8:CB4C68FBFF89FFC112FF9B33E0EFFCB8" box="[255,388,807,832]" name="Kai Islands" pageId="57" pageNumber="58">Kai Islands</location>
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;
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<collectionCode id="A882A6E5FF89FFC1101D9B33E305FCB8" box="[541,622,807,832]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="57" pageNumber="58">ZMUC</collectionCode>
P
<quantity id="096B93C5FF89FFC110919B33E23DFCC7" box="[657,854,807,832]" metricMagnitude="5" metricUnit="m" metricValue="3.75857126" pageId="57" pageNumber="58" unit="mm" value="3.75857126E8">375857, 126 mm</quantity>
TL,
<location id="CB4C68FBFF89FFC111929B33E57CFCB8" LSID="urn:lsid:plazi:treatment:463A8F36FF89FFC2129799D8E3BEFBE8:CB4C68FBFF89FFC111929B33E57CFCB8" box="[914,1047,807,832]" name="Kai Islands" pageId="57" pageNumber="58">Kai Islands</location>
</materialsCitation>
;
<materialsCitation id="7EFB347DFF89FFC116259B33E01BFC9D" collectionCode="ZMUC" latitude="-5.7666664" longLatPrecision="1305" longitude="132.86667" pageId="57" pageNumber="58" specimenCount="1">
<specimenCount id="D895F5A9FF89FFC116259B33E5CCFCB8" box="[1061,1191,807,832]" pageId="57" pageNumber="58" type="generic">1 specimen</specimenCount>
<collectionCode id="A882A6E5FF89FFC116B09B33E46AFCB8" box="[1200,1281,807,832]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="57" pageNumber="58">ZMUC</collectionCode>
unregistered,
<geoCoordinate id="ABA758E7FF89FFC112979B58E19EFC9D" box="[151,245,844,869]" degrees="05" direction="south" minutes="46" orientation="latitude" pageId="57" pageNumber="58" precision="925" value="-5.7666664">05°46’S</geoCoordinate>
,
<geoCoordinate id="ABA758E7FF89FFC113019B58E01BFC9D" box="[257,368,844,869]" degrees="132" direction="east" minutes="52" orientation="longitude" pageId="57" pageNumber="58" precision="925" value="132.86667">132°52’E</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF89FFC212C79B7BE5AFFF2D" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="diagnosis">
<paragraph id="CE2C3E20FF89FFC112C79B7BE065FBE0" blockId="57.[151,1437,663,1048]" pageId="57" pageNumber="58">
<emphasis id="FCE7E232FF89FFC112C79B7BE02AFC70" bold="true" box="[199,321,879,904]" pageId="57" pageNumber="58">Diagnosis.</emphasis>
Pelvic fin rays 8; pectoral fin rays 13–15; gill rakers 19–23. Barbel long, 20–25% HL, reaching beyond a vertical through anterior rim of orbit. Orbit 35–40% HL. Snout obtuse, slightly protruding, 15–18% HL. Ventral striae extending to about ⅔ the distance from pelvic fin bases to periproct. Infraorbital supporter long, 70–90% OD. Otolith compressed, OL:OH = 0.9–1.0; colliculi fused; pseudocolliculum long, TCL:PCL = 1.25–1.45.
</paragraph>
<caption id="9AEC6EA8FF89FFC112979F18E3FFF8A7" ID-DOI="http://doi.org/10.5281/zenodo.10086479" ID-Zenodo-Dep="10086479" httpUri="https://zenodo.org/record/10086479/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,250,1804,1826]" targetBox="[181,1407,1109,1774]" targetPageId="57">
<paragraph id="CE2C3E20FF89FFC112979F18E3FFF8A7" blockId="57.[151,1436,1804,1887]" pageId="57" pageNumber="58">
<emphasis id="FCE7E232FF89FFC112979F18E04FF8D9" bold="true" box="[151,292,1804,1826]" pageId="57" pageNumber="58">FIGURE 30.</emphasis>
<taxonomicName id="099345A3FF89FFC1132F9F18E348F8DA" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[303,547,1804,1826]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="aeger">
<emphasis id="FCE7E232FF89FFC1132F9F18E348F8DA" box="[303,547,1804,1826]" italics="true" pageId="57" pageNumber="58">Hymenocephalus aeger</emphasis>
</taxonomicName>
: A—ZMUC P375844, lateral view of head. B–H—Otoliths, B—Anterior view, C, E–H—Inner faces, D—Ventral View, B–D—ZMUC P375844, E—ZMUC unregistered, 05°46’S, 132°52’E, F—MNHN 2006- 0218, G—ZMUC P375857, H—ZMUC P375856.
</paragraph>
</caption>
<paragraph id="CE2C3E20FF89FFC212C79F98E5AFFF2D" blockId="57.[151,1436,1932,2029]" lastBlockId="58.[151,1436,151,1040]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
<emphasis id="FCE7E232FF89FFC112C79F98E009F85D" bold="true" box="[199,354,1932,1957]" pageId="57" pageNumber="58">Comparison.</emphasis>
<taxonomicName id="099345A3FF89FFC1136E9F98E310F85C" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[366,635,1932,1957]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="aeger">
<emphasis id="FCE7E232FF89FFC1136E9F98E310F85C" box="[366,635,1932,1957]" italics="true" pageId="57" pageNumber="58">Hymenocephalus aeger</emphasis>
</taxonomicName>
was established by
<bibRefCitation id="AA0243D1FF89FFC111719F98E5E4F85D" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[881,1167,1932,1957]" pageId="57" pageNumber="58" pagination="369 - 588" refId="ref50763" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1920) The macrouroid fishes of the Philippine Islands and the East Indies. United States National Museum Bulletin, 100 (1), 369 - 588. http: // dx. doi. org / 10.5962 / bhl. title. 13637" type="journal article" year="1920">Gilbert &amp; Hubbs (1920)</bibRefCitation>
as a subspecies of
<taxonomicName id="099345A3FF89FFC117819F99E04EF830" authorityName=": Weber" authorityYear="1913" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="striatissimus">
<emphasis id="FCE7E232FF89FFC117819F99E04EF830" italics="true" pageId="57" pageNumber="58">H. striatissimus</emphasis>
</taxonomicName>
on the basis of a longer barbel and a smaller eye. The difference in the eye (orbit) size was not verified from the specimens examined, but the longer barbel is quite distinct. In addition, the otoliths are slightly less compressed than those of the other species of the
<taxonomicName id="099345A3FF8AFFC210C8988CE23DFF57" authorityName=": Weber" authorityYear="1913" box="[712,854,152,175]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="striatissimus">
<emphasis id="FCE7E232FF8AFFC210C8988CE23DFF57" box="[712,854,152,175]" italics="true" pageId="58" pageNumber="59">striatissimus</emphasis>
</taxonomicName>
Group (OL:OH = 0.9–1.0 vs 0.75–0.9). This subtle difference in otolith proportions further supports the recognition of a separate taxonomic entity.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF8AFFC212C798CBE01CFD35" pageId="58" pageNumber="59" type="description">
<paragraph id="CE2C3E20FF8AFFC212C798CBE59DFE28" blockId="58.[151,1436,151,1040]" pageId="58" pageNumber="59">
<emphasis id="FCE7E232FF8AFFC212C798CBE033FF00" bold="true" box="[199,344,223,248]" pageId="58" pageNumber="59">Description.</emphasis>
Head morphology (n = 2) (
<figureCitation id="56A822A5FF8AFFC210A098CBE266FF00" box="[672,781,223,248]" captionStart="FIGURE 30" captionStartId="57.[151,250,1804,1826]" captionTargetBox="[181,1407,1109,1774]" captionTargetId="figure-361@57.[175,1412,1103,1779]" captionTargetPageId="57" captionText="FIGURE 30. Hymenocephalus aeger: A—ZMUC P375844, lateral view of head. B–H—Otoliths, B—Anterior view, C, E–H—Inner faces, D—Ventral View, B–D—ZMUC P375844, E—ZMUC unregistered, 05°46’S, 132°52’E, F—MNHN 2006- 0218, G—ZMUC P375857, H—ZMUC P375856." figureDoi="http://doi.org/10.5281/zenodo.10086479" httpUri="https://zenodo.org/record/10086479/files/figure.png" pageId="58" pageNumber="59">Fig. 30A</figureCitation>
): Snout rather short and obtuse, only slightly projecting, 15–18% HL, orbit diameter large, 35–40% HL, interorbital width 60–70% HW. Barbel long, 20–25% HL, reaching slightly beyond a vertical through anterior rim of orbit. Head canals well developed, infraorbital width 11–14% HL, supraorbital canal with 4 to 5 segments, width 10–15% HL, supratemporal canal rarely identifiable, above segment 3 or 4 of supraorbital canal, preopercular canal width 12–16% HL, postorbital-preopercular interspace 4–7% HL. Infranasal supporter moderately large; infraorbital supporter long, extended below almost entire length of orbit, 70–90% OD; preopercular supporter moderately long, 5–8% HL, with straight rear margin.
</paragraph>
<paragraph id="CE2C3E20FF8AFFC212C799C8E01CFD35" blockId="58.[151,1436,151,1040]" pageId="58" pageNumber="59">
Otolith morphology (n = 10) (
<figureCitation id="56A822A5FF8AFFC2102099C8E3C6FE0D" box="[544,685,476,501]" captionStart="FIGURE 30" captionStartId="57.[151,250,1804,1826]" captionTargetBox="[181,1407,1109,1774]" captionTargetId="figure-361@57.[175,1412,1103,1779]" captionTargetPageId="57" captionText="FIGURE 30. Hymenocephalus aeger: A—ZMUC P375844, lateral view of head. B–H—Otoliths, B—Anterior view, C, E–H—Inner faces, D—Ventral View, B–D—ZMUC P375844, E—ZMUC unregistered, 05°46’S, 132°52’E, F—MNHN 2006- 0218, G—ZMUC P375857, H—ZMUC P375856." figureDoi="http://doi.org/10.5281/zenodo.10086479" httpUri="https://zenodo.org/record/10086479/files/figure.png" pageId="58" pageNumber="59">Fig. 30B–H</figureCitation>
): Otolith large; OL:OH = 0.9–1.0; OH:OT = 3.5. Dorsal rim with very large, moderately broad, smooth predorsal lobe, distally marked by small concavity, inclining to posterior tip; posterior tip slightly expanded, positioned at about level of sulcus termination; ventral rim deep, regularly curved, smooth, deepest anterior of the middle; anterior rim high, subvertical to nearly vertical. Inner face slightly convex, with median sulcus. Colliculi completely fused, terminating far from anterior and posterior tips of otolith; pseudocolliculum very long. TCL:PCL = 1.25–1.45. Dorsal depression small, indistinct; ventral furrow distinct, close to ventral rim.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF8AFFC212C79AC3E427FC30" pageId="58" pageNumber="59" type="discussion">
<paragraph id="CE2C3E20FF8AFFC212C79AC3E427FC30" blockId="58.[151,1436,151,1040]" pageId="58" pageNumber="59">
<emphasis id="FCE7E232FF8AFFC212C79AC3E02EFD08" bold="true" box="[199,325,727,752]" pageId="58" pageNumber="59">Discussion</emphasis>
.
<bibRefCitation id="AA0243D1FF8AFFC213549AC3E30DFD08" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[340,614,727,752]" pageId="58" pageNumber="59" pagination="369 - 588" refId="ref50763" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1920) The macrouroid fishes of the Philippine Islands and the East Indies. United States National Museum Bulletin, 100 (1), 369 - 588. http: // dx. doi. org / 10.5962 / bhl. title. 13637" type="journal article" year="1920">Gilbert &amp; Hubbs (1920)</bibRefCitation>
studied a large number of specimens of the three nominal subspecies of
<taxonomicName id="099345A3FF8AFFC212979AE9E02DFCEC" authorityName=": Weber" authorityYear="1913" box="[151,326,765,788]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="striatissimus">
<emphasis id="FCE7E232FF8AFFC212979AE9E02DFCEC" box="[151,326,765,788]" italics="true" pageId="58" pageNumber="59">H. striatissimus</emphasis>
</taxonomicName>
, and recorded that they found intergrades between
<taxonomicName id="099345A3FF8AFFC211829AE9E513FCEC" authorityName=": Iwamoto &amp; Williams" authorityYear="1999" box="[898,1144,765,788]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="58" pageNumber="59" phylum="Chordata" rank="subSpecies" species="striatissimus" subSpecies="aeger">
<emphasis id="FCE7E232FF8AFFC211829AE9E513FCEC" box="[898,1144,765,788]" italics="true" pageId="58" pageNumber="59">H. striatissimus aeger</emphasis>
</taxonomicName>
and
<emphasis id="FCE7E232FF8AFFC216AD9AE9E445FCEC" box="[1197,1326,765,788]" italics="true" pageId="58" pageNumber="59">
H. s.
<taxonomicName id="099345A3FF8AFFC216E89AE9E445FCEC" authorityName="Smith &amp; Radcliffe" authorityYear="1912" box="[1256,1326,765,788]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="torvus">torvus</taxonomicName>
</emphasis>
along the Tawi Tawi Archipelago of the
<collectingCountry id="B6847EB0FF8AFFC213EA9B0BE300FCC0" box="[490,619,799,824]" name="Philippines" pageId="58" pageNumber="59">Philippines</collectingCountry>
, which led them to establish this form as a subspecies of
<taxonomicName id="099345A3FF8AFFC216E59B35E4FFFCCF" authorityName=": Weber" authorityYear="1913" box="[1253,1428,800,824]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="striatissimus">
<emphasis id="FCE7E232FF8AFFC216E59B35E4FFFCCF" box="[1253,1428,800,824]" italics="true" pageId="58" pageNumber="59">H. striatissimus</emphasis>
</taxonomicName>
. I was unable to verify their observation of intergrades, but found a specimen from off the nearby Jolo Island in the collection of ZMUC (P375860), which I consider as
<taxonomicName id="099345A3FF8AFFC210E49B7DE2F8FC87" authorityName=": Weber" authorityYear="1913" box="[740,915,872,896]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="striatissimus">
<emphasis id="FCE7E232FF8AFFC210E49B7DE2F8FC87" box="[740,915,872,896]" italics="true" pageId="58" pageNumber="59">H. striatissimus</emphasis>
</taxonomicName>
(
<figureCitation id="56A822A5FF8AFFC211A29B73E542FC78" box="[930,1065,871,896]" captionStart="FIGURE 32" captionStartId="60.[151,250,813,835]" captionTargetBox="[206,1385,199,785]" captionTargetId="figure-420@60.[198,1389,195,789]" captionTargetPageId="60" captionText="FIGURE 32. Hymenocephalus striatissimus: A—ZMUC P15-5-1914, lateral view of head. B–C—ZMUC P375860, B—Lateral view of head, C—Dorsal view of head. D–F—Otoliths, E–F—Inner faces, D—Ventral view, D–E—BSKU 110412, F—BSKU 110102." figureDoi="http://doi.org/10.5281/zenodo.10086483" httpUri="https://zenodo.org/record/10086483/files/figure.png" pageId="58" pageNumber="59">Fig. 32B–C</figureCitation>
). Since otoliths seem to present a further character helpful to identify
<emphasis id="FCE7E232FF8AFFC210329B99E3DBFC5C" box="[562,688,909,932]" italics="true" pageId="58" pageNumber="59">
H. s.
<taxonomicName id="099345A3FF8AFFC210719B99E3DBFC5C" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[625,688,909,932]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="aeger">aeger</taxonomicName>
</emphasis>
, I now tentatively propose full species rank for
<taxonomicName id="099345A3FF8AFFC216DA9B99E457FC5C" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[1242,1340,909,932]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="aeger">
<emphasis id="FCE7E232FF8AFFC216DA9B99E457FC5C" box="[1242,1340,909,932]" italics="true" pageId="58" pageNumber="59">H. aeger</emphasis>
</taxonomicName>
, subject however to verification of a more detailed and possibly statistical assessment of this difficult species group.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF8AFFC212C79BC0E3BEFBE8" pageId="58" pageNumber="59" type="distribution">
<paragraph id="CE2C3E20FF8AFFC212C79BC0E3BEFBE8" blockId="58.[151,1436,151,1040]" pageId="58" pageNumber="59">
<emphasis id="FCE7E232FF8AFFC212C79BC0E032FC15" bold="true" box="[199,345,980,1005]" pageId="58" pageNumber="59">Distribution</emphasis>
(
<figureCitation id="56A822A5FF8AFFC2136A9BC0E0ABFC15" box="[362,448,980,1005]" captionStart="FIGURE 37" captionStartId="66.[151,250,976,998]" captionTargetBox="[158,1430,272,949]" captionTargetId="figure-236@66.[152,1435,265,954]" captionTargetPageId="66" captionText="FIGURE 37. Geographical distribution of Hymenocephalus aeger, H. megalops, H. striatissimus, H. torvus, H. grimaldii, H. neglectissimus, H. semipellucidus, H. sp. 1 and H." figureDoi="http://doi.org/10.5281/zenodo.10086493" httpUri="https://zenodo.org/record/10086493/files/figure.png" pageId="58" pageNumber="59">Fig. 37</figureCitation>
). Specimens studied and considered valid for
<taxonomicName id="099345A3FF8AFFC211CF9BC1E558FC14" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[975,1075,981,1004]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="aeger">
<emphasis id="FCE7E232FF8AFFC211CF9BC1E558FC14" box="[975,1075,981,1004]" italics="true" pageId="58" pageNumber="59">H. aeger</emphasis>
</taxonomicName>
are from off NW-Australia, off southern New
<collectingCountry id="B6847EB0FF8AFFC2133A9BE3E0E0FBE8" box="[314,395,1015,1040]" name="Guinea-Bissau" pageId="58" pageNumber="59">Guinea</collectingCountry>
and off the
<collectingCountry id="B6847EB0FF8AFFC210129BE3E3BAFBE8" box="[530,721,1015,1040]" name="Solomon Islands" pageId="58" pageNumber="59">Solomon Islands</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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