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<document ID-DOI="http://dx.doi.org/10.3897/jhr.24.1811" ID-GBIF-Taxon="182248524" ID-Pensoft-Pub="1314-2607-24-1" ID-Pensoft-UUID="6276A617FFD1156A810CFFEBB116B333" ID-Zenodo-Dep="574769" ID-ZooBank="487A7CCAB5E74D88B2BE24B7D4CE66F5" ModsDocID="1314-2607-24-1" checkinTime="1621552494608" checkinUser="pensoft" docAuthor="Smith, David R. &amp; Tripotin, Pierre" docDate="2012" docId="40B5DDCDB40E7AB2F15DC62B4D03BA52" docLanguage="en" docOrigin="Journal of Hymenoptera Research 24" docPubDate="2012-01-10" docSource="http://dx.doi.org/10.3897/jhr.24.1811" docTitle="Orthogonalys gigantea Benoit 1951" docType="treatment" docVersion="2" id="6276A617FFD1156A810CFFEBB116B333" lastPageId="7" lastPageNumber="8" masterDocId="6276A617FFD1156A810CFFEBB116B333" masterDocTitle="Trigonalidae (Hymenoptera) of Madagascar" masterLastPageNumber="25" masterPageNumber="1" pageId="4" pageNumber="5" updateTime="1643545408548" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Trigonalidae (Hymenoptera) of Madagascar</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, David R.</mods:namePart>
<mods:affiliation>Systematic Entomology Laboratory, PSI, Agricultural Research Service, c / o National Museum of Natural History, Smithsonian Institution, P. O. Box 37012, MRC 168, Washington, DC 20013 - 7012, USA</mods:affiliation>
<mods:nameIdentifier type="email">sawfly2@aol.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tripotin, Pierre</mods:namePart>
<mods:affiliation>10, rue de Thorigny, 76130 Mont Saint-Aignan, France</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-01-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>24</mods:number>
</mods:detail>
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<mods:start>1</mods:start>
<mods:end>25</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.24.1811</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.24.1811</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-24-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">6276A617FFD1156A810CFFEBB116B333</mods:identifier>
<mods:identifier type="Zenodo-Dep">574769</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182248524" LSID="urn:lsid:plazi:treatment:40B5DDCDB40E7AB2F15DC62B4D03BA52" httpUri="http://treatment.plazi.org/id/40B5DDCDB40E7AB2F15DC62B4D03BA52" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5">
<subSubSection pageId="4" pageNumber="5" type="nomenclature">
<paragraph pageId="4" pageNumber="5">
<taxonomicName LSID="http://species-id.net/wiki/Orthogonalys_gigantea" authority="Benoit" authorityName="Benoit" authorityYear="1951" class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Orthogonalys gigantea" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gigantea">Orthogonalys gigantea Benoit</taxonomicName>
<figureCitation captionStart="Figures 8–9" captionStartId="F3" captionText="Figures 8 - 9. Orthogonalys gigantea. 8 Female holotype, dorsolateral view. 9 Male, dorsolateral view." httpUri="https://binary.pensoft.net/fig/11359" pageId="4" pageNumber="5">Figs 8</figureCitation>
<figureCitation captionStart="Figures 10–13" captionStartId="F4" captionText="Figures 10 - 13. Orthogonalys gigantea. 10 Mesosoma, dorsal view, male. 11 Head front view, female holotype. 12 Head, dorsal view, female holotype. 13 Male paramere, lateral view (appearance is due to hairs matted against the surface)." httpUri="https://binary.pensoft.net/fig/11360" pageId="4" pageNumber="5">13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="reference_group">
<paragraph pageId="4" pageNumber="5">
<taxonomicName class="Insecta" family="Trigonalyidae" genus="Orthogonalos" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gigantea">Orthogonalos gigantea</taxonomicName>
Benoit, 1951: 143.-
<bibRefCitation author="Paulian, R" journalOrPublisher="American Entomologist" pageId="16" pageNumber="17" refId="B10" refString="Paulian, R, 1961. La zoogeographie de Madagascar et des iles voisines. Faune de Madagascar 13, Publications de l'Institut de Recherche Scientifique Tananarive - Tsimbazaza, 484 pp." title="La zoogeographie de Madagascar et des iles voisines. Faune de Madagascar 13, Publications de l'Institut de Recherche Scientifique Tananarive - Tsimbazaza, 484 pp." year="1961">Paulian 1961</bibRefCitation>
: 206, fig. 79.
</paragraph>
<paragraph pageId="4" pageNumber="5">
<taxonomicName authorityName="Benoit" authorityYear="1951" class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gigantea">Orthogonalys gigantea</taxonomicName>
:
<bibRefCitation author="Weinstein, P" journalOrPublisher="Journal of Natural History" pageId="16" pageNumber="17" pagination="399 - 433" publicationUrl="doi: 10.1080/00222939100770281" refId="B13" refString="Weinstein, P, Austin, AD, 1991. The host relationships of trigonalid wasps (Hymenoptera: Trigonalyidae), with a review of their biology and catalogue to world species. Journal of Natural History 25 (2): 399 - 433, doi: 10.1080/00222939100770281" title="The host relationships of trigonalid wasps (Hymenoptera: Trigonalyidae), with a review of their biology and catalogue to world species." url="doi: 10.1080/00222939100770281" volume="25" year="1991">Weinstein and Austin 1991</bibRefCitation>
: 422.-
<bibRefCitation author="Carmean, D" journalOrPublisher="Systematic Entomology" pageId="16" pageNumber="17" pagination="35 - 76" publicationUrl="doi: 10.1046/j.1365-3113.1998.00042.x" refId="B5" refString="Carmean, D, Kimsey, L, 1998. Phylogenetic revision of the parasitoid wasp family Trigonalidae (Hymenoptera). Systematic Entomology 23 (1): 35 - 76, doi: 10.1046/j.1365-3113.1998.00042.x" title="Phylogenetic revision of the parasitoid wasp family Trigonalidae (Hymenoptera)." url="doi: 10.1046/j.1365-3113.1998.00042.x" volume="23" year="1998">Carmean and Kimsey 1998</bibRefCitation>
: 53, 54.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="type material">
<paragraph pageId="4" pageNumber="5">Type material.</paragraph>
<paragraph pageId="4" pageNumber="5">
Holotype female at MNHN, labeled
<normalizedToken originalValue="“Muséum">&quot;Museum</normalizedToken>
Paris,&quot; &quot;Madagascar, Mgn
<normalizedToken originalValue="l’Ambre,”">l'Ambre,&quot;</normalizedToken>
<normalizedToken originalValue="“Muséum">&quot;Museum</normalizedToken>
Paris 1.34 A. Seyrig,&quot; &quot;TYPE [red],&quot; &quot;Orthogonalys gigantea sp. n. holotype ♀, det. P. L. G.
<bibRefCitation author="Benoit, PLG" journalOrPublisher="Revue de Zoologie et de Botanique Africaines" pageId="16" pageNumber="17" pagination="141 - 147" refId="B1" refString="Benoit, PLG, 1951. La systematique des Trigonaloidae (Hym.) ethiopiens. Revue de Zoologie et de Botanique Africaines 44 (2): 141 - 147" title="La systematique des Trigonaloidae (Hym.) ethiopiens." volume="44" year="1951">Benoit 1951</bibRefCitation>
.&quot;
<bibRefCitation author="Benoit, PLG" journalOrPublisher="Revue de Zoologie et de Botanique Africaines" pageId="16" pageNumber="17" pagination="141 - 147" refId="B1" refString="Benoit, PLG, 1951. La systematique des Trigonaloidae (Hym.) ethiopiens. Revue de Zoologie et de Botanique Africaines 44 (2): 141 - 147" title="La systematique des Trigonaloidae (Hym.) ethiopiens." volume="44" year="1951">Benoit (1951)</bibRefCitation>
described the species from &quot;Madagascar: Montagne
<normalizedToken originalValue="d’Ambre">d'Ambre</normalizedToken>
I.1934 (A. Seyrig ♀ holotype, au
<normalizedToken originalValue="Muséum">Museum</normalizedToken>
National
<normalizedToken originalValue="d’Histoire">d'Histoire</normalizedToken>
Naturelle de Paris.&quot; The antennae are missing except first three antennomeres of left and first antennomere of right and parts are glued onto a piece of cardboard on the pin.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="additional specimen examined">
<paragraph pageId="4" pageNumber="5">Additional specimen examined.</paragraph>
<paragraph pageId="4" pageNumber="5">
PROVINCE ANTSIRANANA: &quot;Madagascar Nord, dct
<normalizedToken originalValue="Diégo-Suarez">Diego-Suarez</normalizedToken>
, Analamerana, 80 m, 50 km SE
<normalizedToken originalValue="Diégo">Diego</normalizedToken>
, I-59, Andria R.&quot; (1 ♂, MNHN).
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph pageId="4" pageNumber="5">Description.</paragraph>
<paragraph pageId="4" pageNumber="5">
Female (
<figureCitation captionStart="Figures 8–9" captionStartId="F3" captionText="Figures 8 - 9. Orthogonalys gigantea. 8 Female holotype, dorsolateral view. 9 Male, dorsolateral view." httpUri="https://binary.pensoft.net/fig/11359" pageId="4" pageNumber="5">Fig. 8</figureCitation>
). Length, 14 mm. Head black; inner orbits broadly from top of eye through malar area and gena, clypeus except for median V-shaped stripe, and mandible except apex white (
<figureCitation captionStart="Figures 10–13" captionStartId="F4" captionText="Figures 10 - 13. Orthogonalys gigantea. 10 Mesosoma, dorsal view, male. 11 Head front view, female holotype. 12 Head, dorsal view, female holotype. 13 Male paramere, lateral view (appearance is due to hairs matted against the surface)." httpUri="https://binary.pensoft.net/fig/11360" pageId="4" pageNumber="5">Figs 11, 12</figureCitation>
). Mesosoma orange with hind corners of pronotum, lower anterior corner of pronotum, and metascutellum yellow. Legs dark orange; hind tibia and tarsus darker, nearly black. Metasoma dark orange with black spot at center of 1st tergite, small faint black spot at center of 2nd tergite, lateral and posterior bands on 3rd tergite, and most of tergite 4 except for white posterior margin; tergite 5 white; laterally and ventrally mostly yellow with black spot on 6th sternite and sheath black.
</paragraph>
<paragraph lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Antenna missing (see male). Head from above nearly straight behind, behind eyes slightly expanded, distance behind eyes about equal to eye length (
<figureCitation captionStart="Figures 10–13" captionStartId="F4" captionText="Figures 10 - 13. Orthogonalys gigantea. 10 Mesosoma, dorsal view, male. 11 Head front view, female holotype. 12 Head, dorsal view, female holotype. 13 Male paramere, lateral view (appearance is due to hairs matted against the surface)." httpUri="https://binary.pensoft.net/fig/11360" pageId="4" pageNumber="5">Fig. 12</figureCitation>
). Anterior
<pageBreakToken pageId="5" pageNumber="6" start="start">width</pageBreakToken>
of mesoscutellum 0.8
<normalizedToken originalValue="×">x</normalizedToken>
medial length. Metascutellum 2
<normalizedToken originalValue="×">x</normalizedToken>
broader than long (
<figureCitation captionStart="Figures 10–13" captionStartId="F4" captionText="Figures 10 - 13. Orthogonalys gigantea. 10 Mesosoma, dorsal view, male. 11 Head front view, female holotype. 12 Head, dorsal view, female holotype. 13 Male paramere, lateral view (appearance is due to hairs matted against the surface)." httpUri="https://binary.pensoft.net/fig/11360" pageId="5" pageNumber="6">Fig. 10</figureCitation>
). Mesonotum shiny with closely set punctures, mostly separated by shiny interspaces 1-2
<normalizedToken originalValue="×">x</normalizedToken>
puncture diameters. Propodeum with almost straight transverse carinae (
<figureCitation captionStart="Figures 10–13" captionStartId="F4" captionText="Figures 10 - 13. Orthogonalys gigantea. 10 Mesosoma, dorsal view, male. 11 Head front view, female holotype. 12 Head, dorsal view, female holotype. 13 Male paramere, lateral view (appearance is due to hairs matted against the surface)." httpUri="https://binary.pensoft.net/fig/11360" pageId="5" pageNumber="6">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="male">
<paragraph pageId="5" pageNumber="6">Male.</paragraph>
<paragraph pageId="5" pageNumber="6">
(
<figureCitation captionStart="Figures 8–9" captionStartId="F3" captionText="Figures 8 - 9. Orthogonalys gigantea. 8 Female holotype, dorsolateral view. 9 Male, dorsolateral view." httpUri="https://binary.pensoft.net/fig/11359" pageId="5" pageNumber="6">Fig. 9</figureCitation>
). Length, 15 mm. Antenna black; antennomeres 11-16 and basal half of 17 white. Similar to female except apex of mesoscutellum more yellowish, metasoma mostly black above with reddish brown at apex of tergite 1 and on most of tergites 2 and 3, tergites 1-3 laterally and ventrally orange yellow and tergites 4-6 with white spots. Paramere dark orange.
</paragraph>
<paragraph pageId="6" pageNumber="7">
<pageBreakToken pageId="6" pageNumber="7" start="start">Antenna</pageBreakToken>
with 28 antennomeres, otherwise similar to female. Paramere oval, about 1.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, broadly rounded at apex (
<figureCitation captionStart="Figures 10–13" captionStartId="F4" captionText="Figures 10 - 13. Orthogonalys gigantea. 10 Mesosoma, dorsal view, male. 11 Head front view, female holotype. 12 Head, dorsal view, female holotype. 13 Male paramere, lateral view (appearance is due to hairs matted against the surface)." httpUri="https://binary.pensoft.net/fig/11360" pageId="6" pageNumber="7">Fig. 13</figureCitation>
).
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/11359" pageId="6" pageNumber="7" start="Figures 8–9" startId="F3">
<paragraph pageId="6" pageNumber="7">
<emphasis bold="true" pageId="6" pageNumber="7">Figures 8-9.</emphasis>
<taxonomicName authorityName="Benoit" authorityYear="1951" class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis italics="true" pageId="6" pageNumber="7">Orthogonalys gigantea</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="6" pageNumber="7">8</emphasis>
Female holotype, dorsolateral view.
<emphasis bold="true" pageId="6" pageNumber="7">9</emphasis>
Male, dorsolateral view.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/11360" pageId="6" pageNumber="7" start="Figures 10–13" startId="F4">
<paragraph pageId="6" pageNumber="7">
<emphasis bold="true" pageId="6" pageNumber="7">Figures 10-13.</emphasis>
<taxonomicName authorityName="Benoit" authorityYear="1951" class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis italics="true" pageId="6" pageNumber="7">Orthogonalys gigantea</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="6" pageNumber="7">10</emphasis>
Mesosoma, dorsal view, male.
<emphasis bold="true" pageId="6" pageNumber="7">11</emphasis>
Head front view, female holotype.
<emphasis bold="true" pageId="6" pageNumber="7">12</emphasis>
Head, dorsal view, female holotype.
<emphasis bold="true" pageId="6" pageNumber="7">13</emphasis>
Male paramere, lateral view (appearance is due to hairs matted against the surface).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="distribution">
<paragraph pageId="6" pageNumber="7">Distribution.</paragraph>
<paragraph pageId="6" pageNumber="7">Madagascar: Antsiranana.</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="remarks">
<paragraph pageId="6" pageNumber="7">Remarks.</paragraph>
<paragraph lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
The status of this species has been questionable.
<bibRefCitation author="Carmean, D" journalOrPublisher="Systematic Entomology" pageId="16" pageNumber="17" pagination="35 - 76" publicationUrl="doi: 10.1046/j.1365-3113.1998.00042.x" refId="B5" refString="Carmean, D, Kimsey, L, 1998. Phylogenetic revision of the parasitoid wasp family Trigonalidae (Hymenoptera). Systematic Entomology 23 (1): 35 - 76, doi: 10.1046/j.1365-3113.1998.00042.x" title="Phylogenetic revision of the parasitoid wasp family Trigonalidae (Hymenoptera)." url="doi: 10.1046/j.1365-3113.1998.00042.x" volume="23" year="1998">Carmean and Kimsey (1998)</bibRefCitation>
thought that it might be only a large form of
<taxonomicName class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hova">
<emphasis italics="true" pageId="6" pageNumber="7">Orthogonalys hova</emphasis>
</taxonomicName>
.
<bibRefCitation author="Benoit, PLG" journalOrPublisher="Revue de Zoologie et de Botanique Africaines" pageId="16" pageNumber="17" pagination="141 - 147" refId="B1" refString="Benoit, PLG, 1951. La systematique des Trigonaloidae (Hym.) ethiopiens. Revue de Zoologie et de Botanique Africaines 44 (2): 141 - 147" title="La systematique des Trigonaloidae (Hym.) ethiopiens." volume="44" year="1951">Benoit (1951)</bibRefCitation>
described the species from a single female. He separated
<taxonomicName authorityName="Benoit" authorityYear="1951" class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis italics="true" pageId="6" pageNumber="7">Orthogonalys gigantea</emphasis>
</taxonomicName>
from
<taxonomicName class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hova">
<emphasis italics="true" pageId="6" pageNumber="7">Orthogonalys hova</emphasis>
</taxonomicName>
by its large size (14 mm) and by the following: teeth of mandibles broad but not very long, the inferior tooth directed downwards; clypeus very thickened; antennal sockets closer together; punctation of scutellum sparser but stronger; propodeum broader than long, with transverse carinae complete and regular (not tangled and less strong); color identical except the white
<normalizedToken originalValue="“atténué”">&quot;attenue&quot;</normalizedToken>
on the head and sides of abdomen. We have not seen additional females but have
<pageBreakToken pageId="7" pageNumber="8" start="start">examined</pageBreakToken>
one male that compares in color and structure with the holotype female and was collected not far from the type locality, and we consider them to be the same species.
</paragraph>
<paragraph pageId="7" pageNumber="8">
After study of both sexes, we note a number of characters that justify treating
<taxonomicName authorityName="Benoit" authorityYear="1951" class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis italics="true" pageId="7" pageNumber="8">Orthogonalys gigantea</emphasis>
</taxonomicName>
as a valid species. Size cannot be regarded as a valid character in trigonalids. However, in comparison to
<taxonomicName class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hova">
<emphasis italics="true" pageId="7" pageNumber="8">Orthogonalys hova</emphasis>
</taxonomicName>
, the specimens examined are consistently larger. The female holotype of
<taxonomicName authorityName="Benoit" authorityYear="1951" class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis italics="true" pageId="7" pageNumber="8">Orthogonalys gigantea</emphasis>
</taxonomicName>
is 14 mm and the male is 15 mm, whereas the largest
<taxonomicName class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hova">
<emphasis italics="true" pageId="7" pageNumber="8">Orthogonalys hova</emphasis>
</taxonomicName>
female is 9 mm and males 5.0-9.5 mm, with most in the 7 mm range. We are not sure about the clypeus, mandible, punctation of the scutellum, and closeness of the antennal sockets as Benoit described; these do not seem significant. However, we emphasize the following characters: the sculpture of the propodeum of
<taxonomicName authorityName="Benoit" authorityYear="1951" class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis italics="true" pageId="7" pageNumber="8">Orthogonalys gigantea</emphasis>
</taxonomicName>
is more transverse (
<figureCitation captionStart="Figures 10–13" captionStartId="F4" captionText="Figures 10 - 13. Orthogonalys gigantea. 10 Mesosoma, dorsal view, male. 11 Head front view, female holotype. 12 Head, dorsal view, female holotype. 13 Male paramere, lateral view (appearance is due to hairs matted against the surface)." httpUri="https://binary.pensoft.net/fig/11360" pageId="7" pageNumber="8">Fig. 10</figureCitation>
) than the more reticulate sculpture of
<taxonomicName class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hova">
<emphasis italics="true" pageId="7" pageNumber="8">Orthogonalys hova</emphasis>
</taxonomicName>
(
<figureCitation captionStart="Figures 16–20" captionStartId="F6" captionText="Figures 16 - 20. Orthogonalys hova, male. 16 Mesosoma, dorsal view. 17 Head, front view. 18, Head, dorsal view. 19 Genitalia, dorsal view. 20 Apex of abdomen and genitalia, lateral view." httpUri="https://binary.pensoft.net/fig/11362" pageId="7" pageNumber="8">Fig. 16</figureCitation>
) and other
<taxonomicName authorityName="Schulz" authorityYear="1905" class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="7" pageNumber="8">Orthogonalys</emphasis>
</taxonomicName>
species; the color of both are similar, except that
<taxonomicName authorityName="Benoit" authorityYear="1951" class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis italics="true" pageId="7" pageNumber="8">Orthogonalys gigantea</emphasis>
</taxonomicName>
has the metascutellum yellow and the male has the metasomal tergites mostly black whereas the metascutellum of
<taxonomicName class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hova">
<emphasis italics="true" pageId="7" pageNumber="8">Orthogonalys hova</emphasis>
</taxonomicName>
is orange and the metasomal tergites of the male are considerably orange; and the parameres of
<taxonomicName authorityName="Benoit" authorityYear="1951" class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis italics="true" pageId="7" pageNumber="8">Orthogonalys gigantea</emphasis>
</taxonomicName>
are dark orange and rounded at the apex (
<figureCitation captionStart="Figures 10–13" captionStartId="F4" captionText="Figures 10 - 13. Orthogonalys gigantea. 10 Mesosoma, dorsal view, male. 11 Head front view, female holotype. 12 Head, dorsal view, female holotype. 13 Male paramere, lateral view (appearance is due to hairs matted against the surface)." httpUri="https://binary.pensoft.net/fig/11360" pageId="7" pageNumber="8">Fig. 13</figureCitation>
), whereas those of
<taxonomicName class="Insecta" family="Trigonalidae" genus="Orthogonalys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hova">
<emphasis italics="true" pageId="7" pageNumber="8">Orthogonalys hova</emphasis>
</taxonomicName>
are white and more acute at the apex (
<figureCitation captionStart="Figures 16–20" captionStartId="F6" captionText="Figures 16 - 20. Orthogonalys hova, male. 16 Mesosoma, dorsal view. 17 Head, front view. 18, Head, dorsal view. 19 Genitalia, dorsal view. 20 Apex of abdomen and genitalia, lateral view." httpUri="https://binary.pensoft.net/fig/11362" pageId="7" pageNumber="8">Fig. 20</figureCitation>
).
</paragraph>
<paragraph pageId="7" pageNumber="8">This species is known only from northern Madagascar. The holotype and associated male were collected in the same area, around Diego Suarez, the two localities only about 50 km apart.</paragraph>
</subSubSection>
</treatment>
</document>
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