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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1041.64460" ID-GBIF-Dataset="5dba2b2e-dcd9-43f5-92e5-ac318c319307" ID-PMC="PMC8184736" ID-Pensoft-Pub="1313-2970-1041-27" ID-Pensoft-UUID="5AE03537388755CFAF1E06C3CC9EFA72" ID-PubMed="34140825" ID-ZooBank="EEE8490BB41D4A6CA963234C256C99BF" ModsDocID="1313-2970-1041-27" checkinTime="1622492499165" checkinUser="pensoft" docAuthor="Brunke, Adam J., Pentinsaari, Mikko &amp; Klimaszewski, Jan" docDate="2021" docId="0A48DBD450F356F5813B2D277A51D955" docLanguage="en" docName="ZooKeys 1041: 27-99" docOrigin="ZooKeys 1041" docPubDate="2021-05-31" docSource="http://dx.doi.org/10.3897/zookeys.1041.64460" docTitle="Isoglossa triangularis Klimaszewski, Brunke &amp; Pentinsaari 2021, sp. nov." docType="treatment" docUuid="A8A0402E-2950-4394-B9CF-25E3DD629804" docUuidSource="ZooBank" docVersion="4" id="5AE03537388755CFAF1E06C3CC9EFA72" lastPageNumber="27" masterDocId="5AE03537388755CFAF1E06C3CC9EFA72" masterDocTitle="Integrative taxonomy of Nearctic and Palaearctic Aleocharinae: new species, synonymies, and records (Coleoptera, Staphylinidae)" masterLastPageNumber="99" masterPageNumber="27" pageNumber="27" updateTime="1668150356826" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Integrative taxonomy of Nearctic and Palaearctic Aleocharinae: new species, synonymies, and records (Coleoptera, Staphylinidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brunke, Adam J.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1158-936X</mods:nameIdentifier>
<mods:affiliation>Agriculture and Agri-Food Canada, Canadian National Collection of Insects, Arachnids and Nematodes, 960 Carling Avenue, Ottawa, Ontario, K 1 A 0 C 6, Canada</mods:affiliation>
<mods:nameIdentifier type="email">adam.j.brunke@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pentinsaari, Mikko</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7241-3873</mods:nameIdentifier>
<mods:affiliation>Centre for Biodiversity Genomics, 50 Stone Road East, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Klimaszewski, Jan</mods:namePart>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, 1055 du PEPS, PO Box 10380, Stn. Sainte-Foy, Quebec, QC, G 1 V 4 C 7, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-05-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1041</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>27</mods:start>
<mods:end>99</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1041.64460</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1041.64460</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1041-27</mods:identifier>
<mods:identifier type="ZooBank">EEE8490BB41D4A6CA963234C256C99BF</mods:identifier>
<mods:identifier type="Pensoft-UUID">5AE03537388755CFAF1E06C3CC9EFA72</mods:identifier>
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<treatment ID-GBIF-Taxon="192735627" LSID="urn:lsid:zoobank.org:act:A8A0402E-2950-4394-B9CF-25E3DD629804" httpUri="http://treatment.plazi.org/id/0A48DBD450F356F5813B2D277A51D955" lastPageNumber="27" pageId="0" pageNumber="27">
<subSubSection pageId="0" pageNumber="27" type="nomenclature">
<paragraph pageId="0" pageNumber="27">
<taxonomicName LSID="http://zoobank.org/A8A0402E-2950-4394-B9CF-25E3DD629804" authority="Klimaszewski, Brunke &amp; Pentinsaari" authorityName="Klimaszewski, Brunke &amp; Pentinsaari" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Isoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoglossa triangularis" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="triangularis" status="sp. nov.">Isoglossa triangularis Klimaszewski, Brunke &amp; Pentinsaari</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="27">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Isoglossa triangularis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure5" httpUri="https://binary.pensoft.net/fig/550270" pageId="0" pageNumber="27">Fig. 5A-H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="type material">
<paragraph pageId="0" pageNumber="27">Type material.</paragraph>
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">
<emphasis italics="true" pageId="0" pageNumber="27">Holotype</emphasis>
.
</emphasis>
(male): Canada, British Columbia, Prince George, Nukko Lake Elementary EQP-CLL-574,
<geoCoordinate degrees="54.0831" direction="north" orientation="latitude" precision="5" value="54.0831">54.0831°N</geoCoordinate>
,
<geoCoordinate degrees="122.988" direction="west" orientation="longitude" precision="55" value="-122.988">122.988°W</geoCoordinate>
, 764 m asl, Holly Sapun 04/20/2015 to 05/08/2015, Barcode of life, DNA voucher specimen, Sample ID: BIOUG22036-B02, Process ID: SMTPM2682-15 (CNC).
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Paratypes</emphasis>
(3, CBG): Canada, British Columbia, Prince George, Nukko Lake Elementary EQP-CLL-574,
<geoCoordinate degrees="54.0831" direction="north" orientation="latitude" precision="5" value="54.0831">54.0831°N</geoCoordinate>
,
<geoCoordinate degrees="122.988" direction="west" orientation="longitude" precision="55" value="-122.988">122.988°W</geoCoordinate>
, 764 m asl, Holly Sapun 04/20/2015 to 05/08/2015, Barcode of life, DNA voucher specimen, Sample ID: BIOUG22036-B07, Process ID: SMTPM2682-15 (1 male, CBG); same label data except: Sample ID: BIOUG22035-H08, Process ID: SMTPM2665-15 (1 female, CBG); Sample ID: BIOUG22036-A04, Process ID: SMTPM2672-15 (1 female, CBG).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="etymology">
<paragraph pageId="0" pageNumber="27">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="27">
The species epithet refers to the remarkably separated triangular apex of the median lobe of the aedeagus, distinguishing it from all other members of the
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ocalea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ocalea" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Ocalea</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="distribution">
<paragraph pageId="0" pageNumber="27">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Origin</emphasis>
: Nearctic.
<emphasis bold="true" pageId="0" pageNumber="27">Canada</emphasis>
: BC.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="diagnosis">
<paragraph pageId="0" pageNumber="27">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="27">
<taxonomicName authorityName="Klimaszewski, Brunke &amp; Pentinsaari" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Isoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoglossa triangularis" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="27">Isoglossa triangularis</emphasis>
</taxonomicName>
can be easily distinguished from all Nearctic species of the
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ocalea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ocalea" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Ocalea</emphasis>
</taxonomicName>
group of genera by a combination of the strongly transverse and sparsely punctate pronotum, transverse antennomere 4, distinct triangular apex of the median lobe in lateral view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Isoglossa triangularis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure5" httpUri="https://binary.pensoft.net/fig/550270" pageId="0" pageNumber="27">5B</figureCitation>
), and distinct and simple 'walking
<normalizedToken originalValue="cane">cane'</normalizedToken>
shape of the spermatheca (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Isoglossa triangularis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure5" httpUri="https://binary.pensoft.net/fig/550270" pageId="0" pageNumber="27">5D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1041.64460.figure5" httpUri="https://binary.pensoft.net/fig/550270" pageId="0" pageNumber="27" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Figure 5.</emphasis>
<taxonomicName authorityName="Klimaszewski, Brunke &amp; Pentinsaari" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Isoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoglossa triangularis" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="27">Isoglossa triangularis</emphasis>
</taxonomicName>
Klimaszewski, Brunke &amp; Pentinsaari, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="27">A</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="27">B</emphasis>
median lobe of aedeagus in lateral view
<emphasis bold="true" pageId="0" pageNumber="27">C</emphasis>
median lobe of aedeagus in dorsal view
<emphasis bold="true" pageId="0" pageNumber="27">D</emphasis>
spermatheca
<emphasis bold="true" pageId="0" pageNumber="27">E</emphasis>
male tergite VIII
<emphasis bold="true" pageId="0" pageNumber="27">F</emphasis>
male sternite VIII
<emphasis bold="true" pageId="0" pageNumber="27">G</emphasis>
female tergite VIII
<emphasis bold="true" pageId="0" pageNumber="27">H</emphasis>
female sternite VIII. Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="27">A</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="27">B-H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="description">
<paragraph pageId="0" pageNumber="27">Description.</paragraph>
<paragraph pageId="0" pageNumber="27">
Body length 3.0-3.3 mm, dark brown with elytra, antennomeres 1-2 or 1-3, legs and apical part of abdomen yellow-brown, forebody moderately glossy and abdomen strongly so (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Isoglossa triangularis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure5" httpUri="https://binary.pensoft.net/fig/550270" pageId="0" pageNumber="27">5A</figureCitation>
); antenna moderately stout, antennomere 4 slightly transverse, antennomeres 5-10 strongly transverse, terminal antennomere ca. as long as two preceding ones combined; pronotum transverse (width/length ratio = 1.6), impressed medially at base, lateral edges evenly arcuate, length ratio of base to apex 1.2
<normalizedToken originalValue="×">x</normalizedToken>
, punctures fine and sparse, distance between punctures ~ 3
<normalizedToken originalValue="×">x</normalizedToken>
diameter of a puncture, space between punctures with faint isodiametric microsculpture, pubescence directed laterad from midline of disc forming arcuate lines on both sides; elytra transverse (width/length ratio = 1.3), 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as pronotum; abdomen arcuate laterally and gradually narrowing toward apex. MALE. Tergite VIII broadly arcuate apically (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Isoglossa triangularis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure5" httpUri="https://binary.pensoft.net/fig/550270" pageId="0" pageNumber="27">5E</figureCitation>
); sternite VIII with apical part broadly triangularly produced (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Isoglossa triangularis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure5" httpUri="https://binary.pensoft.net/fig/550270" pageId="0" pageNumber="27">5F</figureCitation>
); median lobe of aedeagus in lateral view with narrowly elongate crista apicalis at base of bulbus, tubus moderately long, strongly produced ventrally, apex narrowly triangular constricted baso-dorsally in lateral view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Isoglossa triangularis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure5" httpUri="https://binary.pensoft.net/fig/550270" pageId="0" pageNumber="27">5B</figureCitation>
), internal sac structures not pronounced (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Isoglossa triangularis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure5" httpUri="https://binary.pensoft.net/fig/550270" pageId="0" pageNumber="27">5B, C</figureCitation>
). FEMALE. Tergite VIII truncate apico-medially (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Isoglossa triangularis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure5" httpUri="https://binary.pensoft.net/fig/550270" pageId="0" pageNumber="27">5G</figureCitation>
); sternite VIII arcuate apically (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Isoglossa triangularis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure5" httpUri="https://binary.pensoft.net/fig/550270" pageId="0" pageNumber="27">5H</figureCitation>
); spermatheca with capsule approximately spherical with short neck, stem narrow, long and straight (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Isoglossa triangularis Klimaszewski, Brunke &amp; Pentinsaari, sp. nov. A habitus B median lobe of aedeagus in lateral view C median lobe of aedeagus in dorsal view D spermatheca E male tergite VIII F male sternite VIII G female tergite VIII H female sternite VIII. Scale bars: 1 mm (A); 0.2 mm (B-H)." figureDoi="10.3897/zookeys.1041.64460.figure5" httpUri="https://binary.pensoft.net/fig/550270" pageId="0" pageNumber="27">5D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="bionomics">
<paragraph pageId="0" pageNumber="27">Bionomics.</paragraph>
<paragraph pageId="0" pageNumber="27">The specimens were collected in a Malaise trap on an open field surrounded by mixed forest.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="comments">
<paragraph pageId="0" pageNumber="27">Comments.</paragraph>
<paragraph pageId="0" pageNumber="27">
Based on a combination of small size (&lt;4.5 mm), superficial, meshed microsculpture, sparse pronotal punctation, with punctures separated by more than two puncture diameters, pronotum transverse, shorter and narrower than elytra, and the transverse antennomeres 5-10,
<taxonomicName lsidName="I. triangularis" pageId="0" pageNumber="27" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="27">I. triangularis</emphasis>
</taxonomicName>
keys to genus
<taxonomicName authorityName="Casey" authorityYear="1893" class="Insecta" family="Staphylinidae" genus="Isoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoglossa" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Isoglossa</emphasis>
</taxonomicName>
Casey in
<bibRefCitation DOI="https://doi.org/10.4039/n03-069" author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="0" pageNumber="27" pagination="443 - 500" refId="B31" refString="Klimaszewski, J, Pelletier, G, 2004. Review of the Ocalea group of genera (Coleoptera, Staphylinidae, Aleocharinae) in Canada and Alaska: new taxa, bionomics and distribution. The Canadian Entomologist 136: 443 - 500, DOI: https://doi.org/10.4039/n03-069" title="Review of the Ocalea group of genera (Coleoptera, Staphylinidae, Aleocharinae) in Canada and Alaska: new taxa, bionomics and distribution." url="https://doi.org/10.4039/n03-069" volume="136" year="2004">Klimaszewski and Pelletier (2004)</bibRefCitation>
. However, barcode sequences of this species do not cluster with
<taxonomicName baseAuthorityName="Casey" baseAuthorityYear="1911" class="Insecta" family="Staphylinidae" genus="Isoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoglossa agnita" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="agnita">
<emphasis italics="true" pageId="0" pageNumber="27">Isoglossa agnita</emphasis>
</taxonomicName>
but rather form a cluster with
<taxonomicName authorityName="Casey" authorityYear="1906" class="Insecta" family="Staphylinidae" genus="Gennadota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Gennadota canadensis" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="canadensis">
<emphasis italics="true" pageId="0" pageNumber="27">Gennadota canadensis</emphasis>
</taxonomicName>
and the species of
<taxonomicName authorityName="Klimaszewski" authorityYear="2004" class="Insecta" family="Staphylinidae" genus="Neothetalia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Neothetalia" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Neothetalia</emphasis>
</taxonomicName>
which bear a spermatheca with broad, circular loops, similar to those of
<taxonomicName authorityName="Casey" authorityYear="1906" class="Insecta" family="Staphylinidae" genus="Gennadota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Gennadota" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Gennadota</emphasis>
</taxonomicName>
.
<taxonomicName authorityName="Klimaszewski, Brunke &amp; Pentinsaari" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Isoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoglossa triangularis" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="27">Isoglossa triangularis</emphasis>
</taxonomicName>
has a simple spermatheca with a long straight stem and is not externally similar to these taxa (see above), and the barcode divergence between these species and
<taxonomicName lsidName="I. triangularis" pageId="0" pageNumber="27" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="27">I. triangularis</emphasis>
</taxonomicName>
is 11-12%. It is likely that
<taxonomicName lsidName="I. triangularis" pageId="0" pageNumber="27" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="27">I. triangularis</emphasis>
</taxonomicName>
belongs in a separate genus, but this is outside of the scope of this study. We here place
<taxonomicName lsidName="I. triangularis" pageId="0" pageNumber="27" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="27">I. triangularis</emphasis>
</taxonomicName>
tentatively in
<taxonomicName authorityName="Casey" authorityYear="1893" class="Insecta" family="Staphylinidae" genus="Isoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoglossa" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Isoglossa</emphasis>
</taxonomicName>
as not to disturb the existing morphological diagnoses of the genera and identification keys (e.g.,
<bibRefCitation DOI="https://doi.org/10.4039/n03-069" author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="0" pageNumber="27" pagination="443 - 500" refId="B31" refString="Klimaszewski, J, Pelletier, G, 2004. Review of the Ocalea group of genera (Coleoptera, Staphylinidae, Aleocharinae) in Canada and Alaska: new taxa, bionomics and distribution. The Canadian Entomologist 136: 443 - 500, DOI: https://doi.org/10.4039/n03-069" title="Review of the Ocalea group of genera (Coleoptera, Staphylinidae, Aleocharinae) in Canada and Alaska: new taxa, bionomics and distribution." url="https://doi.org/10.4039/n03-069" volume="136" year="2004">Klimaszewski and Pelletier 2004</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1007/978-3-030-36174-7" author="Klimaszewski, J" journalOrPublisher="Springer, Cham" pageId="0" pageNumber="27" refId="B41" refString="Klimaszewski, J, Hoebeke, ER, Godin, B, Davies, A, Perry, KI, Bourdon, C, Winchester, N, 2020. Aleocharine rove beetles of British Columbia: a hotspot of Canadian biodiversity (Coleoptera, Staphylinidae). Springer, Cham, DOI: https://doi.org/10.1007/978-3-030-36174-7" title="Aleocharine rove beetles of British Columbia: a hotspot of Canadian biodiversity (Coleoptera, Staphylinidae)." url="https://doi.org/10.1007/978-3-030-36174-7" year="2020">Klimaszewski et al. 2020</bibRefCitation>
), pending generic revision of the
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ocalea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ocalea" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Ocalea</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
</treatment>
</document>