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<document id="2B772C5B49259C26A03C98E9F4A94AA3" ID="10.11646/zootaxa.4258.3.4SLASH10913" ID-CLB-Dataset="33143" ID-DOI="10.11646/zootaxa.4258.3.4" ID-GBIF-Dataset="78768fda-3413-47cf-96fa-d43cc3cf9f08" ID-ISSN="1175-5326" ID-Zenodo-Dep="569765" ID-ZooBank="24932262-6253-42D3-B6FB-02E5394564F2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493366146382" checkinUser="plazi" docAuthor="Viraktamath, C. A. &amp; Meshram, Naresh M." docDate="2017" docId="777B87B6FFAA7A03FF48F8FFFAF8A801" docLanguage="en" docName="zootaxa.4258.3.4SLASH10913.pdf" docOrigin="Zootaxa 4258 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mahellus ungulatus Viraktamath &amp; Meshram, 2017, sp. nov." docType="treatment" docVersion="9" lastPageNumber="280" masterDocId="8B42FFCEFFA27A0AFFDFFFB3FFF9AB14" masterDocTitle="A review of the coelidiine leafhopper genus Mahellus (Hemiptera: Cicadellidae: Coelidiinae) with description of two new species from the Oriental region" masterLastPageNumber="280" masterPageNumber="271" pageNumber="279" updateTime="1698439128449" updateUser="ExternalLinkService">
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<mods:title id="A822AB53D7EE88A24C3E3F2A4716547E">A review of the coelidiine leafhopper genus Mahellus (Hemiptera: Cicadellidae: Coelidiinae) with description of two new species from the Oriental region</mods:title>
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<mods:name id="2887024EDDFC25C6DE22AD34F92CC73D" type="personal">
<mods:role id="948AE1C4F03639F51ED546F25436EB7B">
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<mods:namePart id="017FC521D8AA74F1723148318B80E518">Viraktamath, C. A.</mods:namePart>
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<mods:namePart id="69A2ECE8B24F3720B2E537944EFDB155">Meshram, Naresh M.</mods:namePart>
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<mods:title id="2303F8DA68BB1F20521A43ADAEABEFE5">Zootaxa</mods:title>
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<mods:date id="B98365D24379A2D9CFCBDB35E5D8F11D">2017</mods:date>
<mods:detail id="9FDD5A93E581A99303B1FA97669CF5B4" type="volume">
<mods:number id="F7598F7890A082C8D634B49FF577896C">4258</mods:number>
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<mods:classification id="B74B83072B5519628DAA4C0CF37A1DAA">journal article</mods:classification>
<mods:identifier id="B605E91714DDEE641594088ABADF1B56" type="CLB-Dataset">33143</mods:identifier>
<mods:identifier id="6BC45C84215416A7723D1A8B5B85FC2C" type="DOI">10.11646/zootaxa.4258.3.4</mods:identifier>
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<mods:identifier id="5AFBBD8152BAF10AF832A291790673DB" type="ISSN">1175-5326</mods:identifier>
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<treatment id="777B87B6FFAA7A03FF48F8FFFAF8A801" ID-DOI="http://doi.org/10.5281/zenodo.6018226" ID-GBIF-Taxon="128810444" ID-Zenodo-Dep="6018226" LSID="urn:lsid:plazi:treatment:777B87B6FFAA7A03FF48F8FFFAF8A801" httpUri="http://treatment.plazi.org/id/777B87B6FFAA7A03FF48F8FFFAF8A801" lastPageId="9" lastPageNumber="280" pageId="8" pageNumber="279">
<subSubSection id="B7C8652BFFAA7A02FF48F8FFFE60AC93" pageId="8" pageNumber="279" type="nomenclature">
<paragraph id="FF6D36A0FFAA7A02FF48F8FFFE16AC72" blockId="8.[151,495,1868,1927]" box="[151,495,1868,1894]" pageId="8" pageNumber="279">
<heading id="A42581CCFFAA7A02FF48F8FFFE16AC72" bold="true" box="[151,495,1868,1894]" fontSize="11" level="1" pageId="8" pageNumber="279" reason="1">
<emphasis id="CDA6EAB2FFAA7A02FF48F8FFFE16AC72" bold="true" box="[151,495,1868,1894]" pageId="8" pageNumber="279">
<taxonomicName id="38D24D23FFAA7A02FF48F8FFFE73AC72" ID-CoL="8JWDT" box="[151,394,1868,1894]" class="Insecta" family="Cicadellidae" genus="Mahellus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="279" phylum="Arthropoda" rank="species" species="ungulatus" status="sp. nov.">
<emphasis id="CDA6EAB2FFAA7A02FF48F8FFFE73AC72" bold="true" box="[151,394,1868,1894]" italics="true" pageId="8" pageNumber="279">Mahellus ungulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D69557C9FFAA7A02FE4DF8FFFE16AC72" box="[402,495,1868,1894]" pageId="8" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FF6D36A0FFAA7A02FF48F8DDFE60AC93" blockId="8.[151,495,1868,1927]" box="[151,409,1902,1927]" pageId="8" pageNumber="279">Figs 89, 17, 40–47, 52.</paragraph>
</subSubSection>
<subSubSection id="B7C8652BFFAA7A03FF48F805FEF2A94B" lastPageId="9" lastPageNumber="280" pageId="8" pageNumber="279" type="description">
<paragraph id="FF6D36A0FFAA7A02FF48F805FA80ACE7" blockId="8.[151,1436,1974,2035]" pageId="8" pageNumber="279">Ochraceous brown, mesonotum and scutellum with few dark brown spots, forewing veins brownish with ochraceous spots. Apices of hind femora, hind tarsi, bases of setae on hind tibiae, brown and claws dark brown.</paragraph>
<paragraph id="FF6D36A0FFAB7A03FF7FFF24FD07A903" blockId="9.[151,1436,151,789]" pageId="9" pageNumber="280">
Male pygofer in lateral view more or less triangular, broad anteriorly and narrowed posteriorly with sclerotized process along dorsal margin, curved mesally and acuminate (
<figureCitation id="67E92A25FFAB7A03FCBBFF0FFC46ABC0" box="[868,959,188,213]" captionStart="FIGURES 40 – 47" captionStartId="7.[151,264,1953,1976]" captionTargetBox="[213,1386,201,1926]" captionTargetId="figure@7.[179,1408,193,1932]" captionTargetPageId="7" captionText="FIGURES 40 – 47. Mahellus ungulatus sp. nov., male genitalia. 40. Pygofer, lateral view; 41. Pygofer caudoventral process, lateral view; 42. Connective, dorsal view; 43. Style, dorsal view; 44. Subgenital plate, ventral view; 45. Aedeagus and dorsal connective, lateral view; 46. Aedeagus and dorsal connective, dorsal view; 47. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569770/files/figure.png" pageId="9" pageNumber="280">Figs 40</figureCitation>
,
<figureCitation id="67E92A25FFAB7A03FC10FF0FFC16ABC1" box="[975,1007,188,213]" captionStart="FIGURES 48 – 52" captionStartId="8.[151,264,1716,1739]" captionTargetBox="[168,1421,432,1697]" captionTargetId="figure@8.[151,1436,419,1713]" captionTargetPageId="8" captionText="FIGURES 48 – 52. Speceis of Mahellus. 48. Chaetotaxy of forefemur of M. distanti; 49. Chaetotaxy of metabasitarsus and second tarsomete of M. distanti, ventral view; 50 – 52. Posterior portion of male pygofer of Mahellus, dorsal view showing dorsal marginal sclerotized process: 50. M. cardoni sp. nov.; 51. M. distanti; 52. M. ungulatus sp. nov." httpUri="https://zenodo.org/record/569771/files/figure.png" pageId="9" pageNumber="280">52</figureCitation>
); with ventral blade-like process not exceeding apex of pygofer, broadened distally with 5–7 teeth-like projections (
<figureCitation id="67E92A25FFAB7A03FBC7FF6CFB9CABEC" box="[1048,1125,223,248]" captionStart="FIGURES 40 – 47" captionStartId="7.[151,264,1953,1976]" captionTargetBox="[213,1386,201,1926]" captionTargetId="figure@7.[179,1408,193,1932]" captionTargetPageId="7" captionText="FIGURES 40 – 47. Mahellus ungulatus sp. nov., male genitalia. 40. Pygofer, lateral view; 41. Pygofer caudoventral process, lateral view; 42. Connective, dorsal view; 43. Style, dorsal view; 44. Subgenital plate, ventral view; 45. Aedeagus and dorsal connective, lateral view; 46. Aedeagus and dorsal connective, dorsal view; 47. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569770/files/figure.png" pageId="9" pageNumber="280">Fig.41</figureCitation>
). Subgenital plate long and narrow with posterior spine-like projection; ventral surface with a few hair-like setae, posterior area with row of short stout setae (
<figureCitation id="67E92A25FFAB7A03FEBFFE94FE4FAA54" box="[352,438,295,320]" captionStart="FIGURES 40 – 47" captionStartId="7.[151,264,1953,1976]" captionTargetBox="[213,1386,201,1926]" captionTargetId="figure@7.[179,1408,193,1932]" captionTargetPageId="7" captionText="FIGURES 40 – 47. Mahellus ungulatus sp. nov., male genitalia. 40. Pygofer, lateral view; 41. Pygofer caudoventral process, lateral view; 42. Connective, dorsal view; 43. Style, dorsal view; 44. Subgenital plate, ventral view; 45. Aedeagus and dorsal connective, lateral view; 46. Aedeagus and dorsal connective, dorsal view; 47. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569770/files/figure.png" pageId="9" pageNumber="280">Fig. 44</figureCitation>
). Connective Y-shaped with median longitudinal apodeme, stem short (
<figureCitation id="67E92A25FFAB7A03FB39FE94FAC5AA54" box="[1254,1340,295,320]" captionStart="FIGURES 40 – 47" captionStartId="7.[151,264,1953,1976]" captionTargetBox="[213,1386,201,1926]" captionTargetId="figure@7.[179,1408,193,1932]" captionTargetPageId="7" captionText="FIGURES 40 – 47. Mahellus ungulatus sp. nov., male genitalia. 40. Pygofer, lateral view; 41. Pygofer caudoventral process, lateral view; 42. Connective, dorsal view; 43. Style, dorsal view; 44. Subgenital plate, ventral view; 45. Aedeagus and dorsal connective, lateral view; 46. Aedeagus and dorsal connective, dorsal view; 47. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569770/files/figure.png" pageId="9" pageNumber="280">Fig. 42</figureCitation>
). Dorsal connective membranous, unpigmented band-like. Style uniformly broad basally, tapered apically (
<figureCitation id="67E92A25FFAB7A03FAECFEFFFA69AA70" box="[1331,1424,332,357]" captionStart="FIGURES 40 – 47" captionStartId="7.[151,264,1953,1976]" captionTargetBox="[213,1386,201,1926]" captionTargetId="figure@7.[179,1408,193,1932]" captionTargetPageId="7" captionText="FIGURES 40 – 47. Mahellus ungulatus sp. nov., male genitalia. 40. Pygofer, lateral view; 41. Pygofer caudoventral process, lateral view; 42. Connective, dorsal view; 43. Style, dorsal view; 44. Subgenital plate, ventral view; 45. Aedeagus and dorsal connective, lateral view; 46. Aedeagus and dorsal connective, dorsal view; 47. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569770/files/figure.png" pageId="9" pageNumber="280">Fig. 43</figureCitation>
). Aedeagus asymmetrical, long and slender compared to that of
<taxonomicName id="38D24D23FFAB7A03FC8AFEC2FC37AA9C" box="[853,974,368,392]" class="Insecta" family="Cicadellidae" genus="Mahellus" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="280" phylum="Arthropoda" rank="species" species="distanti">
<emphasis id="CDA6EAB2FFAB7A03FC8AFEC2FC37AA9C" box="[853,974,368,392]" italics="true" pageId="9" pageNumber="280">M. distanti</emphasis>
</taxonomicName>
, in lateral aspect broad in basal half then narrowed and of uniform width to apex, with pair of subapical processes serrated distally, apical process arising on dorsal margin, other subapically on lateral margin, gonopore large on dorsal surface of shaft almost at midlength (
<figureCitation id="67E92A25FFAB7A03FF40FE6FFEDBAAE1" box="[159,290,476,501]" captionStart="FIGURES 40 – 47" captionStartId="7.[151,264,1953,1976]" captionTargetBox="[213,1386,201,1926]" captionTargetId="figure@7.[179,1408,193,1932]" captionTargetPageId="7" captionText="FIGURES 40 – 47. Mahellus ungulatus sp. nov., male genitalia. 40. Pygofer, lateral view; 41. Pygofer caudoventral process, lateral view; 42. Connective, dorsal view; 43. Style, dorsal view; 44. Subgenital plate, ventral view; 45. Aedeagus and dorsal connective, lateral view; 46. Aedeagus and dorsal connective, dorsal view; 47. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569770/files/figure.png" pageId="9" pageNumber="280">Figs 45–47</figureCitation>
). Segment X in lateral aspect longer than broad, with flat ventral surface, in dorsal aspect broad at base and slightly narrowed distally, without ventral process.
</paragraph>
<paragraph id="FF6D36A0FFAB7A03FF18FD97FEF2A94B" blockId="9.[151,1436,151,789]" pageId="9" pageNumber="280">
<emphasis id="CDA6EAB2FFAB7A03FF18FD97FE84A929" bold="true" box="[199,381,548,573]" pageId="9" pageNumber="280">Measurements.</emphasis>
Male 7.0 mm long, 1.9 mm wide across eyes and 2.1 mm wide across posterolateral angles of pronotum.
</paragraph>
</subSubSection>
<subSubSection id="B7C8652BFFAB7A03FF18FDDFFA62A990" box="[199,1435,620,645]" pageId="9" pageNumber="280" type="materials_examined">
<paragraph id="FF6D36A0FFAB7A03FF18FDDFFA62A990" blockId="9.[151,1436,151,789]" box="[199,1435,620,645]" pageId="9" pageNumber="280">
<materialsCitation id="4FBA3CFDFFAB7A03FF18FDDFFA62A990" ID-GBIF-Occurrence="1499315742" box="[199,1435,620,645]" collectingDate="1981-11-10" collectionCode="USAB" collectorName="S. Viraktamath Coll." country="India" location="Sukna" pageId="9" pageNumber="280" specimenCount="1" specimenCount-male="1" stateProvince="West Bengal" typeStatus="holotype">
<emphasis id="CDA6EAB2FFAB7A03FF18FDDFFE49A991" bold="true" box="[199,432,620,645]" pageId="9" pageNumber="280">Material examined.</emphasis>
<typeStatus id="20698802FFAB7A03FE68FDDFFDB2A990" box="[439,587,620,644]" pageId="9" pageNumber="280" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="E9D4FD29FFAB7A03FD8DFDDEFD90A990" box="[594,617,621,644]" pageId="9" pageNumber="280" type="male">♂</specimenCount>
,
<collectingCountry id="87C57630FFAB7A03FDAAFDDFFD30A990" box="[629,713,620,644]" name="India" pageId="9" pageNumber="280">INDIA</collectingCountry>
:
<collectingRegion id="3D16F842FFAB7A03FD0CFDDEFC9CA991" box="[723,869,620,645]" country="India" name="West Bengal" pageId="9" pageNumber="280">West Bengal</collectingRegion>
:
<location id="FA0D607BFFAB7A03FCB0FDDFFC41A991" LSID="urn:lsid:plazi:treatment:777B87B6FFAA7A03FF48F8FFFAF8A801:FA0D607BFFAB7A03FCB0FDDFFC41A991" box="[879,952,620,645]" country="India" name="Sukna" pageId="9" pageNumber="280" stateProvince="West Bengal">Sukna</location>
,
<collectingDate id="9B28E988FFAB7A03FC1CFDDFFBC4A991" box="[963,1085,620,645]" pageId="9" pageNumber="280" value="1981-11-10">
<date id="8B6C1060FFAB7A03FC1CFDDFFBC4A991" box="[963,1085,620,645]" pageId="9" pageNumber="280" value="1981-11-10">10.xi.1981</date>
</collectingDate>
,
<collectorName id="52275376FFAB7A03FB96FDDFFAC9A991" box="[1097,1328,620,645]" pageId="9" pageNumber="280">S. Viraktamath Coll.</collectorName>
(
<collectionCode id="99C3AE65FFAB7A03FAE1FDDFFA76A990" box="[1342,1423,620,644]" pageId="9" pageNumber="280">USAB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B7C8652BFFAB7A03FF18FD3CFB40A9B3" box="[199,1209,655,680]" pageId="9" pageNumber="280" type="etymology">
<paragraph id="FF6D36A0FFAB7A03FF18FD3CFB40A9B3" blockId="9.[151,1436,151,789]" box="[199,1209,655,680]" pageId="9" pageNumber="280">
<emphasis id="CDA6EAB2FFAB7A03FF18FD3CFEB7A9BC" bold="true" box="[199,334,655,680]" pageId="9" pageNumber="280">Etymology.</emphasis>
The species name alludes to the finger-like extensions of the pygofer process.
</paragraph>
</subSubSection>
<subSubSection id="B7C8652BFFAB7A03FF18FD07FAF8A801" pageId="9" pageNumber="280" type="discussion">
<paragraph id="FF6D36A0FFAB7A03FF18FD07FAF8A801" blockId="9.[151,1436,151,789]" pageId="9" pageNumber="280">
<emphasis id="CDA6EAB2FFAB7A03FF18FD07FEC2A9D9" bold="true" box="[199,315,692,717]" pageId="9" pageNumber="280">Remarks.</emphasis>
This species resembles
<taxonomicName id="38D24D23FFAB7A03FD87FD06FC7BA9D9" authority="Distant" authorityName="Distant" box="[600,898,692,717]" class="Insecta" family="Cicadellidae" genus="Jassus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="280" phylum="Arthropoda" rank="species" species="determinatus">
<emphasis id="CDA6EAB2FFAB7A03FD87FD06FCEAA9D8" box="[600,787,692,716]" italics="true" pageId="9" pageNumber="280">M. determinatus</emphasis>
(Distant)
</taxonomicName>
in aedeagal characters but differs in having a much narrower apophysis of the style. Both
<taxonomicName id="38D24D23FFAB7A03FD5CFD6AFD04A9E4" box="[643,765,728,752]" class="Insecta" family="Cicadellidae" genus="Mahellus" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="280" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis id="CDA6EAB2FFAB7A03FD5CFD6AFD04A9E4" box="[643,765,728,752]" italics="true" pageId="9" pageNumber="280">M. cardoni</emphasis>
</taxonomicName>
and
<taxonomicName id="38D24D23FFAB7A03FCECFD6AFC3CA9E4" box="[819,965,728,752]" class="Insecta" family="Cicadellidae" genus="Mahellus" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="280" phylum="Arthropoda" rank="species" species="ungulatus">
<emphasis id="CDA6EAB2FFAB7A03FCECFD6AFC3CA9E4" box="[819,965,728,752]" italics="true" pageId="9" pageNumber="280">M. ungulatus</emphasis>
</taxonomicName>
have a similar dorsal marginal sclerotized process which is mesally directed and acuminate distally, but differing in curvature (see
<figureCitation id="67E92A25FFAB7A03FBACFD4FFB0DA801" box="[1139,1268,764,789]" captionStart="FIGURES 48 – 52" captionStartId="8.[151,264,1716,1739]" captionTargetBox="[168,1421,432,1697]" captionTargetId="figure@8.[151,1436,419,1713]" captionTargetPageId="8" captionText="FIGURES 48 – 52. Speceis of Mahellus. 48. Chaetotaxy of forefemur of M. distanti; 49. Chaetotaxy of metabasitarsus and second tarsomete of M. distanti, ventral view; 50 – 52. Posterior portion of male pygofer of Mahellus, dorsal view showing dorsal marginal sclerotized process: 50. M. cardoni sp. nov.; 51. M. distanti; 52. M. ungulatus sp. nov." httpUri="https://zenodo.org/record/569771/files/figure.png" pageId="9" pageNumber="280">Figs 50, 52</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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