245 lines
28 KiB
XML
245 lines
28 KiB
XML
<document id="E78A290CC3FE8BA93AC89469A77F57A3" ID-DOI="10.11646/zootaxa.5026.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5300736" ID-ZooBank="44594360-3408-4C4C-A4E6-1025F171A49F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630039285316" checkinUser="plazi" docAuthor="Michie, Laura A., Barnes, R. S. K., Clark, Paul F., Bennett, Wayne A. & Cragg, Simon M." docDate="2021" docId="947287DFFFA9FF857EAA17CFFC8EFCEC" docLanguage="en" docName="zootaxa.5026.4.2.pdf" docOrigin="Zootaxa 5026 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Austruca triangularis" docType="treatment" docVersion="4" lastPageNumber="491" masterDocId="684BFFA7FFA1FF8E7E3D124CFF94FFB1" masterDocTitle="Distinguishing ten sympatric species of fiddler crab (Decapoda: Ocypodidae) using a suite of phenotypic characteristics" masterLastPageNumber="506" masterPageNumber="480" pageNumber="488" updateTime="1699139032236" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="2372C200D531AF22506F6DE3CBCB9C75">Distinguishing ten sympatric species of fiddler crab (Decapoda: Ocypodidae) using a suite of phenotypic characteristics</mods:title>
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<mods:name id="9A54DCB44F82D5FCB13BD5BACC9070CF" type="personal">
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<mods:roleTerm id="46117D35D1932B54AC28AC3EE9AF1B9A">Author</mods:roleTerm>
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<mods:namePart id="2BF37D034563E2F12631D2328C06D477">Michie, Laura A.</mods:namePart>
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<mods:roleTerm id="C7D118CA2758C11E194421C8ECD8450F">Author</mods:roleTerm>
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<mods:namePart id="D88DB2550563472FAD0E182D93B4823B">Barnes, R. S. K.</mods:namePart>
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<mods:affiliation id="E2E8B5230080B45F89945F0E7124753B">Department of Zoology and Conservation Research Institute, University of Cambridge, CB 2 3 EJ, UK.</mods:affiliation>
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<mods:namePart id="9A8796D4D67D759B2FC14431E7D69C8A">Clark, Paul F.</mods:namePart>
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<mods:affiliation id="FDC8A3910555DC08BF2EAAF351B2EE47">Department of Life Sciences, The Natural History Museum, Cromwell Rad, London, SW 7 5 BD, UK.</mods:affiliation>
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<mods:roleTerm id="8882A638877A27F6D64B29CC21213205">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4D486D6ECD86481B90FDDBF221729AF5">Bennett, Wayne A.</mods:namePart>
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<mods:affiliation id="40EA3B4444822185FF5F56EC9CABDD66">University of West Florida, 11000 University Parkway, Pensacola, US.</mods:affiliation>
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<mods:namePart id="B6CBC7B4CB4CAEB32BD2A6ED159A592B">Cragg, Simon M.</mods:namePart>
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<mods:nameIdentifier id="3C92544AC18CBC22F8F16FF0876D8A65" type="ORCID">0000-0003-1082-7653</mods:nameIdentifier>
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<mods:affiliation id="A89B70ADD01725927A3E35C60293E297">simon. cragg @ port. ac. uk; https: // orcid. org / 0000 - 0003 - 1082 - 7653</mods:affiliation>
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<mods:nameIdentifier id="143D45B9CE98680AF8B507D053E5B565" type="email">simon.cragg@port.ac.uk</mods:nameIdentifier>
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<mods:title id="67B5F0BA1124ED18237616D7BF3A3B9D">Zootaxa</mods:title>
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<mods:part id="06480D2E8AC95DF19F77C15DE770BD69">
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<mods:date id="F6A88A5596B23D8336F9FD8DA7632A77">2021</mods:date>
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<mods:number id="33A56BF9740C867B6BEEEE472E93B7C1">2021-08-27</mods:number>
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<mods:number id="6305BCF5E87AED12081AEE6FA053DFA8">5026</mods:number>
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<mods:classification id="9AC1BA2F645CBA1BADAFE1157B66A7C3">journal article</mods:classification>
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<mods:identifier id="C416F0E01D1AC4CE6FC5BB0DCD24C01C" type="DOI">10.11646/zootaxa.5026.4.2</mods:identifier>
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<mods:identifier id="27C71E04FDE9BD7D213D8E77A5D65FB6" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="80388136280A6018ED69DCD5CB608E0B" type="Zenodo-Dep">5300736</mods:identifier>
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<treatment id="947287DFFFA9FF857EAA17CFFC8EFCEC" ID-DOI="http://doi.org/10.5281/zenodo.5307760" ID-Zenodo-Dep="5307760" LSID="urn:lsid:plazi:treatment:947287DFFFA9FF857EAA17CFFC8EFCEC" httpUri="http://treatment.plazi.org/id/947287DFFFA9FF857EAA17CFFC8EFCEC" lastPageId="11" lastPageNumber="491" pageId="8" pageNumber="488">
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<subSubSection id="54C16542FFA9FF867EAA17CFFD7CFA2C" box="[151,744,1411,1438]" pageId="8" pageNumber="488" type="nomenclature">
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<paragraph id="1C6436C9FFA9FF867EAA17CFFD7CFA2C" blockId="8.[151,744,1411,1474]" box="[151,744,1411,1438]" pageId="8" pageNumber="488">
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<heading id="472C81A5FFA9FF867EAA17CFFD7CFA2C" bold="true" box="[151,744,1411,1438]" fontSize="11" level="1" pageId="8" pageNumber="488" reason="1">
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<taxonomicName id="DBDB4D4AFFA9FF867EAA17CFFD7CFA2C" ID-CoL="JYP2" authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[151,744,1411,1438]" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="488" phylum="Arthropoda" rank="species" species="triangularis">
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<emphasis id="2EAFEADBFFA9FF867EAA17CFFD7CFA2C" bold="true" box="[151,744,1411,1438]" pageId="8" pageNumber="488">
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<emphasis id="2EAFEADBFFA9FF867EAA17CFFE08FA2F" bold="true" box="[151,412,1411,1438]" italics="true" pageId="8" pageNumber="488">Austruca triangularis</emphasis>
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(A. Milne-Edwards, 1873)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="54C16542FFA9FF867EAA17EBFEBEFA73" box="[151,298,1447,1474]" pageId="8" pageNumber="488" type="description">
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<paragraph id="1C6436C9FFA9FF867EAA17EBFEBEFA73" blockId="8.[151,744,1411,1474]" box="[151,298,1447,1474]" pageId="8" pageNumber="488">
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(
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<figureCitation id="84E02A4CFFA9FF867EA217EBFEB6FA73" box="[159,290,1447,1474]" captionStart="FIGURE 6" captionStartId="10.[151,250,1893,1917]" captionTargetBox="[204,1384,181,1865]" captionTargetId="figure-17@10.[204,1384,181,1869]" captionTargetPageId="10" captionText="FIGURE 6. Austruca triangularis (A. Milne-Edwards, 1873): a–d Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view.Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of left G1; e, lateral view; f, mesial view; CLSM images of apical part of G1; g, lateral view; h, mesial view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300748" httpUri="https://zenodo.org/record/5300748/files/figure.png" pageId="8" pageNumber="488">Figure 6a–j</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="54C16542FFA9FF867EAA17A3FBCDF9E0" pageId="8" pageNumber="488" type="materials_examined">
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<paragraph id="1C6436C9FFA9FF867EAA17A3FBCDF9E0" blockId="8.[151,1437,1519,2014]" pageId="8" pageNumber="488">
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<emphasis id="2EAFEADBFFA9FF867EAA17A3FE14F9B8" bold="true" box="[151,384,1519,1545]" pageId="8" pageNumber="488">Material examined.</emphasis>
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<materialsCitation id="ACB33C94FFA9FF867FBB17A3FD75F99C" collectingDate="2014-08-26" collectorName="L. Michie" country="Indonesia" latitude="-5.4975" location="Kaledupa" longLatPrecision="21" longitude="123.75584" pageId="8" pageNumber="488" specimenCount="20" specimenCount-female="10" specimenCount-male="10">
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<specimenCount id="0ADDFD40FFA9FF867FBB17A3FE29F9BB" box="[390,445,1519,1546]" pageId="8" pageNumber="488" type="female">10 ♀</specimenCount>
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,
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<specimenCount id="0ADDFD40FFA9FF867FF617A3FD90F9BB" box="[459,516,1519,1546]" pageId="8" pageNumber="488" type="male">10 ♂</specimenCount>
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,
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<location id="19046012FFA9FF867C3217A3FDE9F9B8" LSID="urn:lsid:plazi:treatment:947287DFFFA9FF857EAA17CFFC8EFCEC:19046012FFA9FF867C3217A3FDE9F9B8" box="[527,637,1519,1545]" country="Indonesia" latitude="-5.4975" longLatPrecision="21" longitude="123.75584" name="Kaledupa" pageId="8" pageNumber="488">Kaledupa</location>
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,
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<collectingCountry id="64CC7659FFA9FF867CB517A3FD6CF9B8" box="[648,760,1519,1545]" name="Indonesia" pageId="8" pageNumber="488">Indonesia</collectingCountry>
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,
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<geoCoordinate id="79EF500EFFA9FF867D3F17A3FC13F9BB" box="[770,903,1519,1546]" degrees="5" direction="south" minutes="29" orientation="latitude" pageId="8" pageNumber="488" precision="15" seconds="51" value="-5.4975">5º 29’ 51’’S</geoCoordinate>
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,
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<geoCoordinate id="79EF500EFFA9FF867DAF17A3FBA1F9B8" box="[914,1077,1519,1546]" degrees="123" direction="east" minutes="45" orientation="longitude" pageId="8" pageNumber="488" precision="15" seconds="21" value="123.75584">123º 45’ 21’’E</geoCoordinate>
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, mud flat in front of mangroves, under houses to the east, coll.
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<collectorName id="B12E531FFFA9FF867FD91458FDC0F99C" box="[484,596,1555,1582]" pageId="8" pageNumber="488">L. Michie</collectorName>
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,
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<date id="68651009FFA9FF867C62145FFD75F99C" box="[607,737,1555,1581]" pageId="8" pageNumber="488" value="2014-08-26">
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<collectingDate id="7821E9E1FFA9FF867C62145FFD75F99C" box="[607,737,1555,1581]" pageId="8" pageNumber="488" value="2014-08-26">26.08.2014</collectingDate>
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</date>
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</materialsCitation>
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.
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<materialsCitation id="ACB33C94FFA9FF867CD7145FFBC1F9E0" collectingDate="2014-08-28" collectorName="L. Michie" country="Indonesia" latitude="-5.4969444" location="Kaledupa" longLatPrecision="21" longitude="123.755554" pageId="8" pageNumber="488" specimenCount="20" specimenCount-female="10" specimenCount-male="10">
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<specimenCount id="0ADDFD40FFA9FF867CD7145FFCB5F99F" box="[746,801,1555,1582]" pageId="8" pageNumber="488" type="female">10 ♀</specimenCount>
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,
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<specimenCount id="0ADDFD40FFA9FF867D13145FFCFCF99F" box="[814,872,1555,1582]" pageId="8" pageNumber="488" type="male">10 ♂</specimenCount>
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,
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<location id="19046012FFA9FF867D4E145FFC75F99C" LSID="urn:lsid:plazi:treatment:947287DFFFA9FF857EAA17CFFC8EFCEC:19046012FFA9FF867D4E145FFC75F99C" box="[883,993,1555,1581]" country="Indonesia" latitude="-5.4969444" longLatPrecision="21" longitude="123.755554" name="Kaledupa" pageId="8" pageNumber="488">Kaledupa</location>
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,
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<collectingCountry id="64CC7659FFA9FF867DD6145FFBCFF99C" box="[1003,1115,1555,1581]" name="Indonesia" pageId="8" pageNumber="488">Indonesia</collectingCountry>
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,
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<geoCoordinate id="79EF500EFFA9FF867A58145FFB7EF99C" box="[1125,1258,1555,1582]" degrees="5" direction="south" minutes="29" orientation="latitude" pageId="8" pageNumber="488" precision="15" seconds="49" value="-5.4969444">5º 29’ 49’’S</geoCoordinate>
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,
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<geoCoordinate id="79EF500EFFA9FF867AC8145FFA0CF99C" box="[1269,1432,1555,1582]" degrees="123" direction="east" minutes="45" orientation="longitude" pageId="8" pageNumber="488" precision="15" seconds="20" value="123.755554">123º 45’ 20’’E</geoCoordinate>
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, mud flat in front of mangroves, under houses to the west, coll.
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<collectorName id="B12E531FFFA9FF867D6A1474FC5CF9E0" box="[855,968,1591,1618]" pageId="8" pageNumber="488">L. Michie</collectorName>
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,
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<date id="68651009FFA9FF867DEE147BFBC1F9E0" box="[979,1109,1591,1617]" pageId="8" pageNumber="488" value="2014-08-28">
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<collectingDate id="7821E9E1FFA9FF867DEE147BFBC1F9E0" box="[979,1109,1591,1617]" pageId="8" pageNumber="488" value="2014-08-28">28.08.2014</collectingDate>
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</date>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="54C16542FFA9FF857EFA1417FEE6FDA8" lastPageId="11" lastPageNumber="491" pageId="8" pageNumber="488" type="diagnosis">
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<paragraph id="1C6436C9FFA9FF867EFA1417FD71F950" blockId="8.[151,1437,1519,2014]" pageId="8" pageNumber="488">
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<emphasis id="2EAFEADBFFA9FF867EFA1417FED6F9C4" bold="true" box="[199,322,1627,1653]" pageId="8" pageNumber="488">Diagnosis.</emphasis>
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Carapace: broader than long; smooth; front broad; anterolateral angles strongly acute and produced; anterolateral margins absent; dorsolateral margins proceeding directly from anterolateral angle strongly converging. Major cheliped: dactyl and pollex relatively wide; pollex arched, dactyl also arched, but less so than pollex; pollex and manus have teeth in proximal and distal portion.
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</paragraph>
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<paragraph id="1C6436C9FFA9FF867EFA14A7FBB5F8C3" blockId="8.[151,1437,1519,2014]" pageId="8" pageNumber="488">
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<emphasis id="2EAFEADBFFA9FF867EFA14A7FECCF8B4" bold="true" box="[199,344,1771,1797]" pageId="8" pageNumber="488">Colouration</emphasis>
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(
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<figureCitation id="84E02A4CFFA9FF867F5414A7FE60F8B7" box="[361,500,1771,1798]" captionStart="FIGURE 6" captionStartId="10.[151,250,1893,1917]" captionTargetBox="[204,1384,181,1865]" captionTargetId="figure-17@10.[204,1384,181,1869]" captionTargetPageId="10" captionText="FIGURE 6. Austruca triangularis (A. Milne-Edwards, 1873): a–d Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view.Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of left G1; e, lateral view; f, mesial view; CLSM images of apical part of G1; g, lateral view; h, mesial view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300748" httpUri="https://zenodo.org/record/5300748/files/figure.png" pageId="8" pageNumber="488">Figure 6a–b</figureCitation>
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). Adult male. Carapace: mainly black, with white transverse markings varying in size and number, anterior ventral surface of carapace and merus of third maxilliped, black. Major cheliped: dactyl and pollex white/off-white with darker hue extending to manus and carpus, manus, carpus, merus and ischium with dark brown speckled pattern. Minor cheliped and ambulatory legs: black or grey.
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</paragraph>
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<paragraph id="1C6436C9FFA9FF867EFA1537FC92F86F" blockId="8.[151,1437,1519,2014]" pageId="8" pageNumber="488">Adult female. Carapace: similar to males, black with white transverse markings, with greater degree of variability than males, sometimes white and occasionally posteriorly black, frequently with turquoise transverse markings across posterior region. Ambulatory legs: black or grey.</paragraph>
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<caption id="48A46641FFA8FF877EAA1511FB0EF850" ID-DOI="http://doi.org/10.5281/zenodo.5300746" ID-Zenodo-Dep="5300746" httpUri="https://zenodo.org/record/5300746/files/figure.png" pageId="9" pageNumber="489" startId="9.[151,250,1885,1909]" targetBox="[196,1391,181,1859]" targetPageId="9">
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<paragraph id="1C6436C9FFA8FF877EAA1511FB0EF850" blockId="9.[151,1436,1885,2017]" pageId="9" pageNumber="489">
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<emphasis id="2EAFEADBFFA8FF877EAA1511FE80F8C4" bold="true" box="[151,276,1885,1909]" pageId="9" pageNumber="489">FIGURE 5.</emphasis>
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<taxonomicName id="DBDB4D4AFFA8FF877F211511FD74F8C4" authority="H. Milne Edwards, 1852" authorityName="H. Milne Edwards" authorityYear="1852" box="[284,736,1885,1909]" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="489" phylum="Arthropoda" rank="species" species="perplexa">
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<emphasis id="2EAFEADBFFA8FF877F211511FE4CF8C4" box="[284,472,1885,1909]" italics="true" pageId="9" pageNumber="489">Austruca perplexa</emphasis>
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H. Milne Edwards, 1852
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</taxonomicName>
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: a–d Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of right G1; e, mesial view; f, lateral view; CLSM images of apical part of G1; g, mesial view; h, lateral view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view.
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</paragraph>
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</caption>
|
||
<caption id="48A46641FFABFF847EAA1529FB0EF858" ID-DOI="http://doi.org/10.5281/zenodo.5300748" ID-Zenodo-Dep="5300748" httpUri="https://zenodo.org/record/5300748/files/figure.png" pageId="10" pageNumber="490" startId="10.[151,250,1893,1917]" targetBox="[204,1384,181,1865]" targetPageId="10">
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||
<paragraph id="1C6436C9FFABFF847EAA1529FB0EF858" blockId="10.[151,1437,1893,2025]" pageId="10" pageNumber="490">
|
||
<emphasis id="2EAFEADBFFABFF847EAA1529FE81F8CC" bold="true" box="[151,277,1893,1917]" pageId="10" pageNumber="490">FIGURE 6.</emphasis>
|
||
<taxonomicName id="DBDB4D4AFFABFF847F201529FC82F8CC" authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[285,790,1893,1917]" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="490" phylum="Arthropoda" rank="species" species="triangularis">
|
||
<emphasis id="2EAFEADBFFABFF847F201529FE6EF8CC" box="[285,506,1893,1917]" italics="true" pageId="10" pageNumber="490">Austruca triangularis</emphasis>
|
||
(A. Milne-Edwards, 1873)
|
||
</taxonomicName>
|
||
: a–d Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of left G1; e, lateral view; f, mesial view; CLSM images of apical part of G1; g, lateral view; h, mesial view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1C6436C9FFAAFF857EFA12DBFBF2FF48" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="491">Juvenile male and female. Carapace: usually white, sometimes light brown/russet with pattern and darker colours developing with maturity. Major cheliped: normally light brown at juvenile stage but becomes darker and develops patterning with maturation. Minor cheliped & ambulatory legs: black or grey.</paragraph>
|
||
<paragraph id="1C6436C9FFAAFF857EFA134EFDA7FE38" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="491">
|
||
<emphasis id="2EAFEADBFFAAFF857EFA134EFE44FEAC" bold="true" box="[199,464,258,285]" pageId="11" pageNumber="491">Gonopod morphology</emphasis>
|
||
(
|
||
<figureCitation id="84E02A4CFFAAFF857FDD134FFDF8FEAF" box="[480,620,259,286]" captionStart="FIGURE 6" captionStartId="10.[151,250,1893,1917]" captionTargetBox="[204,1384,181,1865]" captionTargetId="figure-17@10.[204,1384,181,1869]" captionTargetPageId="10" captionText="FIGURE 6. Austruca triangularis (A. Milne-Edwards, 1873): a–d Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view.Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of left G1; e, lateral view; f, mesial view; CLSM images of apical part of G1; g, lateral view; h, mesial view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300748" httpUri="https://zenodo.org/record/5300748/files/figure.png" pageId="11" pageNumber="491">Figure 6e–h</figureCitation>
|
||
). G1 curved in dorso-lateral direction; distal end long and narrow with flanges extend throughout length of distal part of shaft; distal edge of flanges rounded; pore of sperm channel large, extending beyond overlapping flanges; protuberance well developed, elongated, ending above base of flanges; distal end curved in lateral direction.
|
||
</paragraph>
|
||
<paragraph id="1C6436C9FFAAFF857EFA13DEFEE6FDA8" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="491">
|
||
<emphasis id="2EAFEADBFFAAFF857EFA13DEFEC2FE1D" bold="true" box="[199,342,402,428]" pageId="11" pageNumber="491">Gastric mill</emphasis>
|
||
(
|
||
<figureCitation id="84E02A4CFFAAFF857F5813DFFE72FE1F" box="[357,486,403,430]" captionStart="FIGURE 6" captionStartId="10.[151,250,1893,1917]" captionTargetBox="[204,1384,181,1865]" captionTargetId="figure-17@10.[204,1384,181,1869]" captionTargetPageId="10" captionText="FIGURE 6. Austruca triangularis (A. Milne-Edwards, 1873): a–d Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view.Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of left G1; e, lateral view; f, mesial view; CLSM images of apical part of G1; g, lateral view; h, mesial view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300748" httpUri="https://zenodo.org/record/5300748/files/figure.png" pageId="11" pageNumber="491">Figure 6i, j</figureCitation>
|
||
). Urocardiac ossicles complex, with 5 transverse ridges of median teeth, different in shape, 1–4 decreasing in size distally, 5 much larger being twice as broad as other ridges; stem region has 5 pairs of cusps, decreasing in size distally, fifth is much shorter and attached to the plate dorsally; lateral tooth plate with 20 comb-shaped teeth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="54C16542FFAAFF857EFA106EFF7FFD34" pageId="11" pageNumber="491" type="biology_ecology">
|
||
<paragraph id="1C6436C9FFAAFF857EFA106EFF7FFD34" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="491">
|
||
<emphasis id="2EAFEADBFFAAFF857EFA106EFEB5FD8D" bold="true" box="[199,289,546,572]" pageId="11" pageNumber="491">Biology</emphasis>
|
||
.
|
||
<taxonomicName id="DBDB4D4AFFAAFF857F13106FFD8AFD8C" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[302,542,547,573]" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="491" phylum="Arthropoda" rank="species" species="triangularis">
|
||
<emphasis id="2EAFEADBFFAAFF857F13106FFD8AFD8C" box="[302,542,547,573]" italics="true" pageId="11" pageNumber="491">Austruca triangularis</emphasis>
|
||
</taxonomicName>
|
||
is a small-size species (largest male CL =
|
||
<quantity id="DB239B2CFFAAFF857DCD106FFBD2FD8C" box="[1008,1094,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="491" unit="mm" value="8.2">8.2 mm</quantity>
|
||
, CB =
|
||
<quantity id="DB239B2CFFAAFF857AAC106FFB62FD8C" box="[1169,1270,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="11" pageNumber="491" unit="mm" value="12.1">12.1 mm</quantity>
|
||
, largest female CL =
|
||
<quantity id="DB239B2CFFAAFF857EE5100BFEA5FDD0" box="[216,305,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="491" unit="mm" value="8.2">8.2 mm</quantity>
|
||
, CB =
|
||
<quantity id="DB239B2CFFAAFF857FBD100BFE7DFDD0" box="[384,489,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="11" pageNumber="491" unit="mm" value="12.8">12.8 mm</quantity>
|
||
); one of least abundant species on shore, present in high intertidal zone under stilted houses.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="54C16542FFAAFF857EFA10C2FB72FD18" box="[199,1254,654,681]" pageId="11" pageNumber="491" type="distribution">
|
||
<paragraph id="1C6436C9FFAAFF857EFA10C2FB72FD18" blockId="11.[151,1437,151,861]" box="[199,1254,654,681]" pageId="11" pageNumber="491">
|
||
<emphasis id="2EAFEADBFFAAFF857EFA10C2FEF5FD19" bold="true" box="[199,353,654,680]" pageId="11" pageNumber="491">Distribution.</emphasis>
|
||
<collectingCountry id="64CC7659FFAAFF857F5B10C3FE44FD18" box="[358,464,655,681]" name="Australia" pageId="11" pageNumber="491">Australia</collectingCountry>
|
||
,
|
||
<collectingCountry id="64CC7659FFAAFF857FE610C3FDDFFD18" box="[475,587,655,681]" name="Indonesia" pageId="11" pageNumber="491">Indonesia</collectingCountry>
|
||
,
|
||
<collectingCountry id="64CC7659FFAAFF857C6810C3FD2AFD18" box="[597,702,655,681]" name="Malaysia" pageId="11" pageNumber="491">Malaysia</collectingCountry>
|
||
,
|
||
<collectingCountry id="64CC7659FFAAFF857CF710C3FCDFFD18" box="[714,843,655,681]" name="Philippines" pageId="11" pageNumber="491">Philippines</collectingCountry>
|
||
,
|
||
<collectingCountry id="64CC7659FFAAFF857D6B10C3FC3FFD18" box="[854,939,655,681]" name="Taiwan" pageId="11" pageNumber="491">Taiwan</collectingCountry>
|
||
,
|
||
<collectingCountry id="64CC7659FFAAFF857D8B10C3FC68FD18" box="[950,1020,655,681]" name="China" pageId="11" pageNumber="491">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="64CC7659FFAAFF857A3B10C3FB77FD18" box="[1030,1251,655,681]" name="Papua New Guinea" pageId="11" pageNumber="491">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="54C16542FFAAFF857EFA10FFFC8EFCEC" pageId="11" pageNumber="491" type="discussion">
|
||
<paragraph id="1C6436C9FFAAFF857EFA10FFFC8EFCEC" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="491">
|
||
<emphasis id="2EAFEADBFFAAFF857EFA10FFFEAFFD7C" bold="true" box="[199,315,691,717]" pageId="11" pageNumber="491">Remarks.</emphasis>
|
||
The specimens examined comply with the description of
|
||
<bibRefCitation id="784A4B38FFAAFF857DE010FFFB3EFD7C" author="Shih, H. T. & Ng, P. K. L. & Ravichandran, S. & Prema, M." box="[989,1194,691,717]" pageId="11" pageNumber="491" pagination="12" refId="ref14880" refString="Shih, H. T., Ng, P. K. L., Ravichandran, S. & Prema, M. (2019) Resurrection of Gelasimus variegatus Heller, 1862, A Fiddler Crab Closely Related to Austruca bengali (Crane, 1975) and A. triangularis (A. Milne-Edwards, 1873) (Decapoda, Brachyura, Ocypodidae), from the Bay of Bengal, Indian Ocean. Zoological Studies, 58, e 12. https: // doi. org / 10.6620 / ZS. 2019.58 - 12" type="journal article" year="2019">
|
||
Shih
|
||
<emphasis id="2EAFEADBFFAAFF857A2710FFFBDAFD7C" box="[1050,1102,691,717]" italics="true" pageId="11" pageNumber="491">et al</emphasis>
|
||
. (2019)
|
||
</bibRefCitation>
|
||
specifically with the elongated protuberance of the G1 and the complex morphology of the gastric mill, with the highest number of transverse ridges and cusps of all
|
||
<taxonomicName id="DBDB4D4AFFAAFF857C2D10B7FD59FCA4" box="[528,717,763,789]" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="491" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="2EAFEADBFFAAFF857C2D10B7FDE1FCA4" box="[528,629,763,789]" italics="true" pageId="11" pageNumber="491">Austruca</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
at Ambeua.
|
||
<taxonomicName id="DBDB4D4AFFAAFF857D6410B7FBDFFCA4" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[857,1099,763,789]" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="491" phylum="Arthropoda" rank="species" species="triangularis">
|
||
<emphasis id="2EAFEADBFFAAFF857D6410B7FBDFFCA4" box="[857,1099,763,789]" italics="true" pageId="11" pageNumber="491">Austruca triangularis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from the other
|
||
<taxonomicName id="DBDB4D4AFFAAFF857EE51153FE00FC88" box="[216,404,799,825]" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="491" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="2EAFEADBFFAAFF857EE51153FEA9FC88" box="[216,317,799,825]" italics="true" pageId="11" pageNumber="491">Austruca</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by the strongly acute anterolateral angles, the elongated protuberance of the G1, the distinct speckled pattern of the major cheliped and the small size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |